378 lines
46 KiB
XML
378 lines
46 KiB
XML
<document id="3F81DE0CECD702E95A6EA1AEEC237B06" ID-CLB-Dataset="28412" ID-DOI="10.11646/zootaxa.4565.1.1" ID-GBIF-Dataset="e9ffcfc7-9f7c-4212-ae6e-b842b31de6e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="2589489" ID-ZooBank="B9919693-2478-4089-B82E-C25EDFDB2A96" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552246080949" checkinUser="plazi" docAuthor="Chen, Hao, Zhang, Hui, Chen, Yongxia & Freyhof, Jörg" docDate="2019" docId="5212F95FFFC9AA7FB5CAFAE2FA1B3350" docLanguage="en" docName="zootaxa.4565.1.1.pdf" docOrigin="Zootaxa 4565 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Barbatula liaoyangensis Chen & Zhang & Chen & Freyhof 2019, new species" docType="treatment" docVersion="8" lastPageNumber="25" masterDocId="AE2B8127FFDCAA67B55DFFF6FF9E324D" masterDocTitle="A review of the Barbatula loaches (Teleostei: Nemacheilidae) from north-eastern China, with the description of four new species" masterLastPageNumber="36" masterPageNumber="1" pageNumber="22" updateTime="1698701226448" updateUser="ExternalLinkService">
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<mods:mods id="27D919F0091875445F0206A814213E2E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="74ED55FD9927D23D9D1816AA295A553A">
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<mods:title id="A99D12ADA2A908213EABE7EAB2EB7FC6">A review of the Barbatula loaches (Teleostei: Nemacheilidae) from north-eastern China, with the description of four new species</mods:title>
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</mods:titleInfo>
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<mods:name id="5C02A8D60380CBA32ED5C3C1F4AE85A0" type="personal">
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<mods:roleTerm id="3E114E9569A307BA794853C677D3E6F3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C313A5CB6AECB5B841592F87D1485471">Chen, Hao</mods:namePart>
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</mods:name>
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<mods:name id="117694C6908E89DFCA36BE4C66E5AF14" type="personal">
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<mods:roleTerm id="66153F70D2A798DCC4C76E77355B7AFE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6FDE131C9E7B2605694E844FADF7758B">Zhang, Hui</mods:namePart>
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</mods:name>
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<mods:name id="B4D283E54BFDDE17F107FDEFB653857E" type="personal">
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<mods:roleTerm id="B6F99E83D194F4495E90D4FAA52F41B5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="384CFE9C10273C0C58DC1EFEDF347FAF">Chen, Yongxia</mods:namePart>
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</mods:name>
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<mods:name id="AA9E8443A0D04EB88F6C40B6D38CD25B" type="personal">
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<mods:roleTerm id="A8EB7140F3EDA1F2202ED07BF98716C0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="02D3CBD26CE94FE14A8E63DC3E25BE57">Freyhof, Jörg</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="73655519E88C3713E23B0B5E6554469E">text</mods:typeOfResource>
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<mods:title id="C24581B04F1F7A5A37F5E2B8B3B340A1">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="351524EFEBAC1752ED0EE79A96AB97C3">
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<mods:date id="591C3382FB2F26678AE90CEE424DD395">2019</mods:date>
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<mods:number id="80EE477776C2A1C42D747FED7A69BF2E">2019-03-07</mods:number>
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<mods:detail id="417408BA2721C568BD46EB4FF9F48DDF" type="volume">
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<mods:number id="36E82FBB932356B26654D98DC038E49C">4565</mods:number>
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<mods:detail id="6B647F4871390E915353BCB6574550C9" type="issue">
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<mods:number id="48CAFA99A6A4C7904B0246685547D2EB">1</mods:number>
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<mods:classification id="68A45D99A6A9170AF20062A33BA1FCE6">journal article</mods:classification>
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<mods:identifier id="6DE3B9FFC4EFDC7C419EA84E8A4AD1B6" type="CLB-Dataset">28412</mods:identifier>
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<mods:identifier id="2754F6552847ECD8A8D26F91F2F52EAA" type="DOI">10.11646/zootaxa.4565.1.1</mods:identifier>
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<mods:identifier id="964A5FEEEAFF35089FF6C6ABF9F9FF20" type="ISSN">1175-5326</mods:identifier>
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<treatment id="5212F95FFFC9AA7FB5CAFAE2FA1B3350" ID-DOI="http://doi.org/10.5281/zenodo.5923833" ID-GBIF-Taxon="154366724" ID-Zenodo-Dep="5923833" LSID="urn:lsid:plazi:treatment:5212F95FFFC9AA7FB5CAFAE2FA1B3350" httpUri="http://treatment.plazi.org/id/5212F95FFFC9AA7FB5CAFAE2FA1B3350" lastPageId="24" lastPageNumber="25" pageId="21" pageNumber="22">
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<subSubSection id="92A11BC2FFC9AA72B5CAFAE2FDC53763" box="[151,603,1300,1326]" pageId="21" pageNumber="22" type="nomenclature">
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<paragraph id="DA044849FFC9AA72B5CAFAE2FDC53763" blockId="21.[151,603,1300,1359]" box="[151,603,1300,1326]" pageId="21" pageNumber="22">
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<heading id="814CFF25FFC9AA72B5CAFAE2FDC53763" bold="true" box="[151,603,1300,1326]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
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||
<emphasis id="E8CF945BFFC9AA72B5CAFAE2FDC53763" bold="true" box="[151,603,1300,1326]" pageId="21" pageNumber="22">
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<taxonomicName id="1DBB33CAFFC9AA72B5CAFAE2FE223763" authority="Chen & Zhang & Chen & Freyhof, 2019" authorityName="Chen & Zhang & Chen & Freyhof" authorityYear="2019" box="[151,444,1300,1326]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="liaoyangensis" status="sp. nov.">
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<emphasis id="E8CF945BFFC9AA72B5CAFAE2FE223763" bold="true" box="[151,444,1300,1326]" italics="true" pageId="21" pageNumber="22">Barbatula liaoyangensis</emphasis>
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||
</taxonomicName>
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||
,
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||
<taxonomicNameLabel id="F3FC2920FFC9AA72B496FAE2FDC53763" box="[459,603,1300,1326]" pageId="21" pageNumber="22" rank="species">new species</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="92A11BC2FFC9AA72B5CAFAC0FEBD3702" box="[151,291,1334,1359]" pageId="21" pageNumber="22" type="description">
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||
<paragraph id="DA044849FFC9AA72B5CAFAC0FEBD3702" blockId="21.[151,603,1300,1359]" box="[151,291,1334,1359]" pageId="21" pageNumber="22">
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(
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<figureCitation id="428054CCFFC9AA72B5C2FAC0FE853702" box="[159,283,1334,1359]" captionStart-0="FIGURE 23" captionStart-1="FIGURE 24" captionStart-2="FIGURE 25" captionStartId-0="22.[151,250,1258,1280]" captionStartId-1="23.[151,250,1491,1513]" captionStartId-2="23.[151,250,1966,1988]" captionTargetBox-0="[220,1414,412,1232]" captionTargetBox-1="[247,1338,719,1389]" captionTargetBox-2="[151,1400,1593,1944]" captionTargetId-0="figure@22.[151,1436,337,1237]" captionTargetId-1="figure@23.[151,1436,651,1479]" captionTargetId-2="figure@23.[151,1436,1593,1944]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionTargetPageId-2="23" captionText-0="FIGURE 23. Barbatula liaoyangensis, from the top: HU 1608150, holotype, 75.9 mm SL; HU1608320, 75.9 mm SL; IHB88v0157, 74.2 mm SL; China: Liaoning prov.: Liao River drainage." captionText-1="FIGURE 24. Barbatula liaoyangensis, from the top: HU1608279, paratype, 82.3 mm SL; HU1608320, 75.9 mm SL; IHB88v0157, 74.2 mm SL; China: Liaoning prov.: Liao River drainage." captionText-2="FIGURE 25. Barbatula liaoyangensis, HU1608150, holotype, 75.9 mm SL; China: Liaoning prov.: Liao River drainage. Scale bars: a–b = 1 mm, c = 0.1 mm." httpUri-0="https://zenodo.org/record/2589535/files/figure.png" httpUri-1="https://zenodo.org/record/2589537/files/figure.png" httpUri-2="https://zenodo.org/record/2589539/files/figure.png" pageId="21" pageNumber="22">Fig. 23–25</figureCitation>
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||
)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="92A11BC2FFC9AA72B5CAFA88FB6D34D9" pageId="21" pageNumber="22" type="materials_examined">
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||
<paragraph id="DA044849FFC9AA72B5CAFA88FC9237F1" blockId="21.[151,1437,1406,2007]" pageId="21" pageNumber="22">
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<materialsCitation id="6AD34214FFC9AA72B5CAFA88FC9237F1" ID-GBIF-Occurrence="2012822322" collectingDate="2016-10" country="China" county="Taizi River" latitude="41.6" location="Liao River" municipality="Xiaobeihezhen" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Liaoning" typeStatus="holotype">
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||
<emphasis id="E8CF945BFFC9AA72B5CAFA88FE9537DA" bold="true" box="[151,267,1406,1431]" pageId="21" pageNumber="22">
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||
<typeStatus id="0500F6EBFFC9AA72B5CAFA88FE9837DA" box="[151,262,1406,1431]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
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||
.
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||
</emphasis>
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||
HU 1608150,
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||
<quantity id="1D43E5ACFFC9AA72B49DFA88FDB737DA" box="[448,553,1406,1431]" metricMagnitude="-2" metricUnit="m" metricValue="7.59" pageId="21" pageNumber="22" unit="mm" value="75.9">75.9 mm</quantity>
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SL;
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||
<collectingCountry id="A2AC08D9FFC9AA72B735FA88FD3137DA" box="[616,687,1406,1431]" name="China" pageId="21" pageNumber="22">China</collectingCountry>
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||
:
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||
<collectingRegion id="187F86ABFFC9AA72B7E2FA88FCBA37DA" box="[703,804,1406,1431]" country="China" name="Liaoning" pageId="21" pageNumber="22">Liaoning</collectingRegion>
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||
prov.:
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||
<collectingCounty id="336530C5FFC9AA72B621FA88FC6337DA" box="[892,1021,1406,1431]" pageId="21" pageNumber="22">Taizi River</collectingCounty>
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||
east of
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||
<collectingMunicipality id="3A60D233FFC9AA72B139FA88FA9137DA" box="[1124,1295,1406,1431]" pageId="21" pageNumber="22">Xiaobeihezhen</collectingMunicipality>
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||
,
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||
<location id="DF641E92FFC9AA72B042FA88FA0237DA" LSID="urn:lsid:plazi:treatment:5212F95FFFC9AA7FB5CAFAE2FA1B3350:DF641E92FFC9AA72B042FA88FA0237DA" box="[1311,1436,1406,1431]" country="China" county="Taizi River" latitude="41.6" municipality="Xiaobeihezhen" name="Liao River" pageId="21" pageNumber="22" stateProvince="Liaoning">Liao River</location>
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||
drainage,
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||
<geoCoordinate id="BF8F2E8EFFC9AA72B458FA55FEFD37F1" box="[261,355,1443,1468]" direction="north" orientation="latitude" pageId="21" pageNumber="22" precision="925" value="41.6">41°36′N</geoCoordinate>
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||
122°85′E (12);
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||
<location id="DF641E92FFC9AA72B74BFA52FD1437F1" LSID="urn:lsid:plazi:treatment:5212F95FFFC9AA7FB5CAFAE2FA1B3350:DF641E92FFC9AA72B74BFA52FD1437F1" box="[534,650,1443,1468]" country="China" county="Taizi River" latitude="41.6" municipality="Xiaobeihezhen" name="Hao Chen" pageId="21" pageNumber="22" stateProvince="Liaoning">Hao Chen</location>
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||
,
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||
<date id="AE056E89FFC9AA72B7CBFA55FC9637F1" box="[662,776,1443,1468]" pageId="21" pageNumber="22" value="2016-10">
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||
<collectingDate id="BE419761FFC9AA72B7CBFA55FC9637F1" box="[662,776,1443,1468]" pageId="21" pageNumber="22" value="2016-10">Oct. 2016</collectingDate>
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||
</date>
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||
.
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</materialsCitation>
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||
</paragraph>
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||
<paragraph id="DA044849FFC9AA72B59AFA30FE0B3449" blockId="21.[151,1437,1406,2007]" pageId="21" pageNumber="22">
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||
<materialsCitation id="6AD34214FFC9AA72B59AFA30FA063792" ID-GBIF-Occurrence="2012822318" box="[199,1432,1478,1503]" collectionCode="FSJF" pageId="21" pageNumber="22" specimenCode="FSJF 4047, 2, 69.2, 71.8" specimenCount="1" typeStatus="paratype">
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||
<emphasis id="E8CF945BFFC9AA72B59AFA30FED83792" bold="true" box="[199,326,1478,1503]" pageId="21" pageNumber="22">
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||
<typeStatus id="0500F6EBFFC9AA72B59AFA30FEDF3792" box="[199,321,1478,1503]" pageId="21" pageNumber="22" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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HU 1608320, 1608152, 1608851–59, 11,
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<quantity id="1D43E5ACFFC9AA72B669FA30FC443792" box="[820,986,1478,1503]" metricMagnitude="-2" metricUnit="m" metricValue="7.595" metricValueMax="8.98" metricValueMin="6.21" pageId="21" pageNumber="22" unit="mm" value="75.95" valueMax="89.8" valueMin="62.1">62.1–89.8 mm</quantity>
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SL,
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||
<specimenCode id="8A1DE032FFC9AA72B14BFA31FB0B3792" box="[1046,1173,1479,1503]" collectionCode="FSJF" pageId="21" pageNumber="22">FSJF 4047</specimenCode>
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||
,
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||
<specimenCode id="8A1DE032FFC9AA72B1F9FA31FB2A3792" box="[1188,1204,1479,1503]" collectionCode="FSJF" pageId="21" pageNumber="22">2</specimenCode>
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||
,
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||
<quantity id="1D43E5ACFFC9AA72B19EFA30FAF43792" box="[1219,1386,1478,1503]" metricMagnitude="-2" metricUnit="m" metricValue="7.05" metricValueMax="7.18" metricValueMin="6.92" pageId="21" pageNumber="22" unit="mm" value="70.5" valueMax="71.8" valueMin="69.2">
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||
<specimenCode id="8A1DE032FFC9AA72B19EFA30FB6A3792" box="[1219,1268,1478,1503]" collectionCode="FSJF" pageId="21" pageNumber="22">69.2</specimenCode>
|
||
–
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||
<specimenCode id="8A1DE032FFC9AA72B05EFA30FAAA3792" box="[1283,1332,1478,1503]" collectionCode="FSJF" pageId="21" pageNumber="22">71.8</specimenCode>
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||
mm
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||
</quantity>
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||
SL
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="6AD34214FFC9AA72B5CAFA1AFE0F3449" ID-GBIF-Occurrence="2012822337" box="[151,401,1515,1540]" collectingDate="2016-10" country="China" county="Taizi River" latitude="41.6" location="Liao River" municipality="Xiaobeihezhen" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Liaoning" typeStatus="paratype">same data as holotype</materialsCitation>
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||
.
|
||
</paragraph>
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||
<paragraph id="DA044849FFC9AA72B59AF9F8FC513406" blockId="21.[151,1437,1406,2007]" pageId="21" pageNumber="22">
|
||
<emphasis id="E8CF945BFFC9AA72B59AF9F8FDA7346A" bold="true" box="[199,569,1550,1575]" pageId="21" pageNumber="22">Additional material examined.</emphasis>
|
||
<materialsCitation id="6AD34214FFC9AA72B71FF9E6FC553406" ID-GBIF-Occurrence="2012822336" collectingDate="1988-05" collectionCode="IHB" country="China" county="Taizi River" location="Liao River" municipality="Liaoyang" pageId="21" pageNumber="22" specimenCode="IHB 88" specimenCount="1" stateProvince="Liaoning">
|
||
<specimenCode id="8A1DE032FFC9AA72B71FF9E6FD08346A" box="[578,662,1551,1575]" collectionCode="IHB" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35160" lsid="urn:lsid:biocol.org:col:35160" name="Institute of Hydrobiology, Chinese Academy of Sciences" pageId="21" pageNumber="22">
|
||
IHB
|
||
<date id="AE056E89FFC9AA72B726F9F9FD08346A" box="[635,662,1551,1575]" pageId="21" pageNumber="22" value="1988-05">88</date>
|
||
</specimenCode>
|
||
v0139,
|
||
<date id="AE056E89FFC9AA72B7B3F9F9FC89346A" box="[750,791,1551,1575]" pageId="21" pageNumber="22" value="1988-05">
|
||
<collectingDate id="BE419761FFC9AA72B7B3F9F9FC89346A" box="[750,791,1551,1575]" pageId="21" pageNumber="22" value="1988-05">88v</collectingDate>
|
||
</date>
|
||
0143,
|
||
<date id="AE056E89FFC9AA72B63DF9F8FC17346A" box="[864,905,1550,1575]" pageId="21" pageNumber="22" value="1988-05">
|
||
<collectingDate id="BE419761FFC9AA72B63DF9F8FC17346A" box="[864,905,1550,1575]" pageId="21" pageNumber="22" value="1988-05">88v</collectingDate>
|
||
</date>
|
||
0152–53,
|
||
<date id="AE056E89FFC9AA72B6A0F9F8FBB8346A" box="[1021,1062,1550,1575]" pageId="21" pageNumber="22" value="1988-05">
|
||
<collectingDate id="BE419761FFC9AA72B6A0F9F8FBB8346A" box="[1021,1062,1550,1575]" pageId="21" pageNumber="22" value="1988-05">88v</collectingDate>
|
||
</date>
|
||
0155–57. 7,
|
||
<quantity id="1D43E5ACFFC9AA72B1EAF9F8FAF5346A" box="[1207,1387,1550,1575]" metricMagnitude="-2" metricUnit="m" metricValue="8.59" metricValueMax="10.59" metricValueMin="6.59" pageId="21" pageNumber="22" unit="mm" value="85.9" valueMax="105.9" valueMin="65.9">65.9–105.9 mm</quantity>
|
||
SL;
|
||
<collectingCountry id="A2AC08D9FFC9AA72B5CAF9C5FF413401" box="[151,223,1587,1612]" name="China" pageId="21" pageNumber="22">China</collectingCountry>
|
||
:
|
||
<collectingRegion id="187F86ABFFC9AA72B5B5F9C5FED33401" box="[232,333,1587,1612]" country="China" name="Liaoning" pageId="21" pageNumber="22">Liaoning</collectingRegion>
|
||
prov.:
|
||
<collectingCounty id="336530C5FFC9AA72B4C1F9C5FD863401" box="[412,536,1587,1612]" pageId="21" pageNumber="22">Taizi River</collectingCounty>
|
||
at
|
||
<collectingMunicipality id="3A60D233FFC9AA72B766F9C5FD363401" box="[571,680,1587,1612]" pageId="21" pageNumber="22">Liaoyang</collectingMunicipality>
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||
,
|
||
<location id="DF641E92FFC9AA72B7EEF9C5FCB53401" LSID="urn:lsid:plazi:treatment:5212F95FFFC9AA7FB5CAFAE2FA1B3350:DF641E92FFC9AA72B7EEF9C5FCB53401" box="[691,811,1587,1612]" country="China" county="Taizi River" municipality="Liaoyang" name="Liao River" pageId="21" pageNumber="22" stateProvince="Liaoning">Liao River</location>
|
||
drainage (12)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="DA044849FFC9AA72B59AF9A0FB6D34D9" blockId="21.[151,1437,1406,2007]" pageId="21" pageNumber="22">
|
||
<emphasis id="E8CF945BFFC9AA72B59AF9A0FC933422" bold="true" box="[199,781,1622,1647]" pageId="21" pageNumber="22">Material used in the molecular genetic analysis.</emphasis>
|
||
HU 1608278–79, 1608320;
|
||
<collectingCountry id="A2AC08D9FFC9AA72B105F9A0FB013422" box="[1112,1183,1622,1647]" name="China" pageId="21" pageNumber="22">China</collectingCountry>
|
||
:
|
||
<collectingRegion id="187F86ABFFC9AA72B1F1F9A0FA8F3422" box="[1196,1297,1622,1647]" country="China" name="Liaoning" pageId="21" pageNumber="22">Liaoning</collectingRegion>
|
||
prov.: Taizi River, Liao River drainage,
|
||
<geoCoordinate id="BF8F2E8EFFC9AA72B48DF98DFDB034D9" box="[464,558,1659,1684]" direction="north" orientation="latitude" pageId="21" pageNumber="22" precision="925" value="41.6">41°36′N</geoCoordinate>
|
||
122°85′E (12). (GenBank accession numbers:
|
||
<accessionNumber id="C5E8D5AAFFC9AA72B161F98DFB2134D9" box="[1084,1215,1659,1684]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KY451929" pageId="21" pageNumber="22">KY451929</accessionNumber>
|
||
–31).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92A11BC2FFC9AA71B59AF968FA0E3729" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="diagnosis">
|
||
<paragraph id="DA044849FFC9AA72B59AF968FEF234B2" blockId="21.[151,1437,1406,2007]" pageId="21" pageNumber="22">
|
||
<emphasis id="E8CF945BFFC9AA72B59AF968FEDF34FA" bold="true" box="[199,321,1694,1719]" pageId="21" pageNumber="22">Diagnosis.</emphasis>
|
||
<taxonomicName id="1DBB33CAFFC9AA72B41AF968FDC434FA" authorityName="Chen & Zhang & Chen & Freyhof" authorityYear="2019" box="[327,602,1694,1719]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="liaoyangensis">
|
||
<emphasis id="E8CF945BFFC9AA72B41AF968FDC434FA" box="[327,602,1694,1719]" italics="true" pageId="21" pageNumber="22">Barbatula liaoyangensis</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from all other species of
|
||
<taxonomicName id="1DBB33CAFFC9AA72B177F968FB0234FA" box="[1066,1180,1694,1719]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="genus">
|
||
<emphasis id="E8CF945BFFC9AA72B177F968FB0234FA" box="[1066,1180,1694,1719]" italics="true" pageId="21" pageNumber="22">Barbatula</emphasis>
|
||
</taxonomicName>
|
||
in north-eastern
|
||
<collectingCountry id="A2AC08D9FFC9AA72B004F968FA0234FA" box="[1369,1436,1694,1719]" name="China" pageId="21" pageNumber="22">China</collectingCountry>
|
||
by having a pointed snout, head width at posterior margin of operculum 1.5–1.8 times in its width at anterior nostril (vs. blunt, 13–1.4).
|
||
</paragraph>
|
||
<paragraph id="DA044849FFC9AA71B59AF8FDFE7E3351" blockId="21.[151,1437,1406,2007]" lastBlockId="22.[151,1436,151,285]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
|
||
<taxonomicName id="1DBB33CAFFC9AA72B59AF8FDFE473569" authorityName="Chen & Zhang & Chen & Freyhof" authorityYear="2019" box="[199,473,1803,1828]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="liaoyangensis">
|
||
<emphasis id="E8CF945BFFC9AA72B59AF8FDFE473569" box="[199,473,1803,1828]" italics="true" pageId="21" pageNumber="22">Barbatula liaoyangensis</emphasis>
|
||
</taxonomicName>
|
||
is further distinguished from the other species of
|
||
<taxonomicName id="1DBB33CAFFC9AA72B15BF8FDFBE63569" box="[1030,1144,1803,1828]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="genus">
|
||
<emphasis id="E8CF945BFFC9AA72B15BF8FDFBE63569" box="[1030,1144,1803,1828]" italics="true" pageId="21" pageNumber="22">Barbatula</emphasis>
|
||
</taxonomicName>
|
||
in north-eastern
|
||
<collectingCountry id="A2AC08D9FFC9AA72B06BF8FDFAE73569" box="[1334,1401,1803,1828]" name="China" pageId="21" pageNumber="22">China</collectingCountry>
|
||
by a combination of characters, none of them unique. It is distinguished by having the nostrils widely spaced (vs. closely set in
|
||
<taxonomicName id="1DBB33CAFFC9AA72B467F8A2FE0B3521" baseAuthorityName="Bleeker" baseAuthorityYear="1865" box="[314,405,1875,1900]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="nuda">
|
||
<emphasis id="E8CF945BFFC9AA72B467F8A2FE0B3521" box="[314,405,1875,1900]" italics="true" pageId="21" pageNumber="22">B. nuda</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="1DBB33CAFFC9AA72B48FF8A2FDC53526" authorityName="Chen & Zhang & Chen & Freyhof" authorityYear="2019" box="[466,603,1876,1899]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="emuensis">
|
||
<emphasis id="E8CF945BFFC9AA72B48FF8A2FDC53526" box="[466,603,1876,1899]" italics="true" pageId="21" pageNumber="22">B. emuensis</emphasis>
|
||
</taxonomicName>
|
||
), the snout being shorter than (vs. equal to in
|
||
<taxonomicName id="1DBB33CAFFC9AA72B1DDF8A2FB453521" baseAuthorityName="Bleeker" baseAuthorityYear="1865" box="[1152,1243,1875,1900]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="nuda">
|
||
<emphasis id="E8CF945BFFC9AA72B1DDF8A2FB453521" box="[1152,1243,1875,1900]" italics="true" pageId="21" pageNumber="22">B. nuda</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="1DBB33CAFFC9AA72B045F8A2FAF83526" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[1304,1382,1876,1899]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="toni">
|
||
<emphasis id="E8CF945BFFC9AA72B045F8A2FAF83526" box="[1304,1382,1876,1899]" italics="true" pageId="21" pageNumber="22">B. toni</emphasis>
|
||
</taxonomicName>
|
||
) the postorbital head length, a shallow median incision in the upper lip, its depth 10–30% of the width of the upper-lip (vs. deep, 40–70% in
|
||
<taxonomicName id="1DBB33CAFFC9AA72B4D0F86AFE7B35F9" baseAuthorityName="Bleeker" baseAuthorityYear="1865" box="[397,485,1947,1972]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="nuda">
|
||
<emphasis id="E8CF945BFFC9AA72B4D0F86AFE7B35F9" box="[397,485,1947,1972]" italics="true" pageId="21" pageNumber="22">B. nuda</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="1DBB33CAFFC9AA72B4A9F86AFDCC35F9" authorityName="Cao, Causse & Zhang" authorityYear="2012" box="[500,594,1947,1972]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="gibba">
|
||
<emphasis id="E8CF945BFFC9AA72B4A9F86AFDCC35F9" box="[500,594,1947,1972]" italics="true" pageId="21" pageNumber="22">B. gibba</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="1DBB33CAFFC9AA72B7D7F86AFC8F35FE" authorityName="Chen & Zhang & Chen & Freyhof" authorityYear="2019" box="[650,785,1948,1971]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="emuensis">
|
||
<emphasis id="E8CF945BFFC9AA72B7D7F86AFC8F35FE" box="[650,785,1948,1971]" italics="true" pageId="21" pageNumber="22">B. emuensis</emphasis>
|
||
</taxonomicName>
|
||
), a blade-shaped lateral expansion of the mental lobe (vs. without expansion in
|
||
<taxonomicName id="1DBB33CAFFC9AA72B4D6F836FE7D359A" baseAuthorityName="Bleeker" baseAuthorityYear="1865" box="[395,483,1982,2007]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="nuda">
|
||
<emphasis id="E8CF945BFFC9AA72B4D6F836FE7D359A" box="[395,483,1982,2007]" italics="true" pageId="21" pageNumber="22">B. nuda</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="1DBB33CAFFC9AA72B4AFF836FDCF359A" authorityName="Cao, Causse & Zhang" authorityYear="2012" box="[498,593,1982,2007]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="gibba">
|
||
<emphasis id="E8CF945BFFC9AA72B4AFF836FDCF359A" box="[498,593,1982,2007]" italics="true" pageId="21" pageNumber="22">B. gibba</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="1DBB33CAFFC9AA72B73DF836FD32359A" baseAuthorityName="Dybowski" baseAuthorityYear="1869" box="[608,684,1984,2007]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="toni">
|
||
<emphasis id="E8CF945BFFC9AA72B73DF836FD32359A" box="[608,684,1984,2007]" italics="true" pageId="21" pageNumber="22">B. toni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="1DBB33CAFFC9AA72B7E7F836FCD9359A" authorityName="Tchang" authorityYear="1932" box="[698,839,1982,2007]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="kirinensis">
|
||
<emphasis id="E8CF945BFFC9AA72B7E7F836FCD9359A" box="[698,839,1982,2007]" italics="true" pageId="21" pageNumber="22">B. kirinensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="1DBB33CAFFC9AA72B622F836FB98359A" authorityName="Chen & Zhang & Chen & Freyhof" authorityYear="2019" box="[895,1030,1984,2007]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="emuensis">
|
||
<emphasis id="E8CF945BFFC9AA72B622F836FB98359A" box="[895,1030,1984,2007]" italics="true" pageId="21" pageNumber="22">B. emuensis</emphasis>
|
||
</taxonomicName>
|
||
), the upper jaw completely covered by the upper lip (vs. partly covered in
|
||
<taxonomicName id="1DBB33CAFFCAAA71B71BFF6FFD0332FD" baseAuthorityName="Bleeker" baseAuthorityYear="1865" box="[582,669,151,176]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="nuda">
|
||
<emphasis id="E8CF945BFFCAAA71B71BFF6FFD0332FD" box="[582,669,151,176]" italics="true" pageId="22" pageNumber="23">B. nuda</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="1DBB33CAFFCAAA71B789FF6FFCFF32FD" authorityName="Tchang" authorityYear="1932" box="[724,865,151,176]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="kirinensis">
|
||
<emphasis id="E8CF945BFFCAAA71B789FF6FFCFF32FD" box="[724,865,151,176]" italics="true" pageId="22" pageNumber="23">B. kirinensis</emphasis>
|
||
</taxonomicName>
|
||
), a slightly convex dorsal profile (vs. convex in
|
||
<taxonomicName id="1DBB33CAFFCAAA71B0D9FF6FFF5D3299" baseAuthorityName="Dybowski" baseAuthorityYear="1869" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="toni">
|
||
<emphasis id="E8CF945BFFCAAA71B0D9FF6FFF5D3299" italics="true" pageId="22" pageNumber="23">B. toni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="1DBB33CAFFCAAA71B5A0FF4BFE3C3298" baseAuthorityName="Fowler" baseAuthorityYear="1899" box="[253,418,188,213]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="pechiliensis">
|
||
<emphasis id="E8CF945BFFCAAA71B5A0FF4BFE3C3298" box="[253,418,188,213]" italics="true" pageId="22" pageNumber="23">B. pechiliensis</emphasis>
|
||
</taxonomicName>
|
||
), 7½ branched dorsal-fin rays (vs. 6½ in
|
||
<taxonomicName id="1DBB33CAFFCAAA71B6D9FF4BFBA93299" authorityName="Prokofiev" authorityYear="2007" box="[900,1079,189,212]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="potaninorum">
|
||
<emphasis id="E8CF945BFFCAAA71B6D9FF4BFBA93299" box="[900,1079,189,212]" italics="true" pageId="22" pageNumber="23">B. potaninorum</emphasis>
|
||
</taxonomicName>
|
||
) and sparsely set scales on the flank anterior to the vertical through the dorsal-fin origin (vs. without scales anterior to dorsal-fin origin in
|
||
<taxonomicName id="1DBB33CAFFCAAA71B063FF17FA0A32B5" baseAuthorityName="Bleeker" baseAuthorityYear="1865" box="[1342,1428,223,248]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="nuda">
|
||
<emphasis id="E8CF945BFFCAAA71B063FF17FA0A32B5" box="[1342,1428,223,248]" italics="true" pageId="22" pageNumber="23">B. nuda</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="1DBB33CAFFCAAA71B5CAFEF3FF683350" authorityName="Cao, Causse & Zhang" authorityYear="2012" box="[151,246,260,285]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="gibba">
|
||
<emphasis id="E8CF945BFFCAAA71B5CAFEF3FF683350" box="[151,246,260,285]" italics="true" pageId="22" pageNumber="23">B. gibba</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="1DBB33CAFFCAAA71B471FEF3FE513350" baseAuthorityName="Fowler" baseAuthorityYear="1899" box="[300,463,260,285]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="pechiliensis">
|
||
<emphasis id="E8CF945BFFCAAA71B471FEF3FE513350" box="[300,463,260,285]" italics="true" pageId="22" pageNumber="23">B. pechiliensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<caption id="8EC418C1FFCAAA71B5CAFB1CFCEE3752" httpUri="https://zenodo.org/record/2589535/files/figure.png" pageId="22" pageNumber="23" startId="22.[151,250,1258,1280]" targetBox="[220,1414,412,1232]" targetPageId="22">
|
||
<paragraph id="DA044849FFCAAA71B5CAFB1CFCEE3752" blockId="22.[151,1436,1258,1311]" pageId="22" pageNumber="23">
|
||
<emphasis id="E8CF945BFFCAAA71B5CAFB1CFEBB36B2" bold="true" box="[151,293,1258,1280]" pageId="22" pageNumber="23">FIGURE 23.</emphasis>
|
||
<taxonomicName id="1DBB33CAFFCAAA71B46DFB1CFDB0374D" authorityName="Chen & Zhang & Chen & Freyhof" authorityYear="2019" box="[304,558,1258,1280]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="liaoyangensis">
|
||
<emphasis id="E8CF945BFFCAAA71B46DFB1CFDB0374D" box="[304,558,1258,1280]" italics="true" pageId="22" pageNumber="23">Barbatula liaoyangensis</emphasis>
|
||
</taxonomicName>
|
||
, from the top: HU 1608150, holotype, 75.9 mm SL; HU1608320, 75.9 mm SL; IHB88v0157, 74.2 mm SL; China: Liaoning prov.: Liao River drainage.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DA044849FFCAAA71B59AFABAFA0E3729" blockId="22.[151,1437,1355,2029]" box="[199,1424,1355,1380]" pageId="22" pageNumber="23">
|
||
See
|
||
<tableCitation id="97397DF2FFCAAA71B5ABFABDFED73729" box="[246,329,1355,1380]" captionStart="TABLE 3" captionStartId="31.[147,216,807,825]" captionText="TABLE 3. (Continued)" pageId="22" pageNumber="23">Table 3</tableCitation>
|
||
for the character states shown by the different species of
|
||
<taxonomicName id="1DBB33CAFFCAAA71B690FABDFBA13729" box="[973,1087,1355,1380]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="22" pageNumber="23" phylum="Chordata" rank="genus">
|
||
<emphasis id="E8CF945BFFCAAA71B690FABDFBA13729" box="[973,1087,1355,1380]" italics="true" pageId="22" pageNumber="23">Barbatula</emphasis>
|
||
</taxonomicName>
|
||
found in north-eastern
|
||
<collectingCountry id="A2AC08D9FFCAAA71B018FABDFA123729" box="[1349,1420,1355,1380]" name="China" pageId="22" pageNumber="23">China</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92A11BC2FFCAAA7FB59AFA99FA953298" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23" type="description">
|
||
<paragraph id="DA044849FFCAAA71B59AFA99FF4E3559" blockId="22.[151,1437,1355,2029]" pageId="22" pageNumber="23">
|
||
<emphasis id="E8CF945BFFCAAA71B59AFA99FEC737C5" bold="true" box="[199,345,1391,1416]" pageId="22" pageNumber="23">Description.</emphasis>
|
||
See
|
||
<figureCitation id="428054CCFFCAAA71B4C8FA99FDB037C5" box="[405,558,1391,1416]" captionStart-0="FIGURE 23" captionStart-1="FIGURE 24" captionStartId-0="22.[151,250,1258,1280]" captionStartId-1="23.[151,250,1491,1513]" captionTargetBox-0="[220,1414,412,1232]" captionTargetBox-1="[247,1338,719,1389]" captionTargetId-0="figure@22.[151,1436,337,1237]" captionTargetId-1="figure@23.[151,1436,651,1479]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionText-0="FIGURE 23. Barbatula liaoyangensis, from the top: HU 1608150, holotype, 75.9 mm SL; HU1608320, 75.9 mm SL; IHB88v0157, 74.2 mm SL; China: Liaoning prov.: Liao River drainage." captionText-1="FIGURE 24. Barbatula liaoyangensis, from the top: HU1608279, paratype, 82.3 mm SL; HU1608320, 75.9 mm SL; IHB88v0157, 74.2 mm SL; China: Liaoning prov.: Liao River drainage." httpUri-0="https://zenodo.org/record/2589535/files/figure.png" httpUri-1="https://zenodo.org/record/2589537/files/figure.png" pageId="22" pageNumber="23">Figure 23–24</figureCitation>
|
||
for general appearance and
|
||
<tableCitation id="97397DF2FFCAAA71B626FA99FC4F37C5" box="[891,977,1391,1416]" captionStart="TABLE 2" captionStartId="28.[147,222,518,538]" captionText="TABLE 2. Morphometric characters of eight species of Barbatula. Ho = holotype; SD = standard deviation." pageId="22" pageNumber="23">Table 2</tableCitation>
|
||
for morphometric data. Body elongate, roundish; caudal peduncle compressed. Dorsal profile slightly convex, parallel with ventral profile; caudalpeduncle depth 59–68% of body depth at dorsal-fin origin. Ventral profile between anal-fin base and caudal-fin origin concave. Head wider than deep, head depth 75–94% in its width at posterior margin of operculum. Mouth inferior and arched (
|
||
<figureCitation id="428054CCFFCAAA71B426FA09FE423455" box="[379,476,1535,1560]" captionStart="FIGURE 25" captionStartId="23.[151,250,1966,1988]" captionTargetBox="[151,1400,1593,1944]" captionTargetId="figure@23.[151,1436,1593,1944]" captionTargetPageId="23" captionText="FIGURE 25. Barbatula liaoyangensis, HU1608150, holotype, 75.9 mm SL; China: Liaoning prov.: Liao River drainage. Scale bars: a–b = 1 mm, c = 0.1 mm." httpUri="https://zenodo.org/record/2589539/files/figure.png" pageId="22" pageNumber="23">Fig. 25a</figureCitation>
|
||
). Snout rounded, short and pointed; head width at posterior margin of operculum 1.5– 1.8 times in its width at anterior nostril. Snout shorter than postorbital head length. Eye small, set close to dorsal profile. Anterior and posterior nostril widely spaced, gap between nostrils equal or shorter than width of base of posterior nostril (
|
||
<figureCitation id="428054CCFFCAAA71B403F99DFE5A34C9" box="[350,452,1643,1668]" captionStart="FIGURE 25" captionStartId="23.[151,250,1966,1988]" captionTargetBox="[151,1400,1593,1944]" captionTargetId="figure@23.[151,1436,1593,1944]" captionTargetPageId="23" captionText="FIGURE 25. Barbatula liaoyangensis, HU1608150, holotype, 75.9 mm SL; China: Liaoning prov.: Liao River drainage. Scale bars: a–b = 1 mm, c = 0.1 mm." httpUri="https://zenodo.org/record/2589539/files/figure.png" pageId="22" pageNumber="23">Fig. 25b</figureCitation>
|
||
). Caudal fin truncate. Caudal-peduncle length 75–97% in HL, caudal-peduncle width 45–65% its depth. Distance between anus and anal-fin origin shorter than or equal to eye diameter. Dorsal-fin origin situated behind middle between tip of snout and caudal-fin base. Pelvic-fin origin situated below vertical through dorsal-fin origin or in front. Anal-fin origin anterior to midpoint between pelvic-fin origin and caudal-fin base.
|
||
</paragraph>
|
||
<paragraph id="DA044849FFCAAA70B59AF8E9FA033350" blockId="22.[151,1437,1355,2029]" lastBlockId="23.[151,1437,151,608]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
|
||
Lateral line complete, reaching to caudal-fin base. Scales sparsely set on flank anterior to dorsal-fin origin, absent on back and belly anterior to dorsal-fin origin, sparsely set on back and flank behind dorsal-fin origin, densely set on caudal peduncle. Scales on caudal peduncle with small focal zone and 24–26 radial grooves (
|
||
<figureCitation id="428054CCFFCAAA71B032F891FF5D35E9" captionStart="FIGURE 25" captionStartId="23.[151,250,1966,1988]" captionTargetBox="[151,1400,1593,1944]" captionTargetId="figure@23.[151,1436,1593,1944]" captionTargetPageId="23" captionText="FIGURE 25. Barbatula liaoyangensis, HU1608150, holotype, 75.9 mm SL; China: Liaoning prov.: Liao River drainage. Scale bars: a–b = 1 mm, c = 0.1 mm." httpUri="https://zenodo.org/record/2589539/files/figure.png" pageId="22" pageNumber="23">Fig. 25c</figureCitation>
|
||
). Anterior nostril formed by short tube, posterior tube shorter or without tube. Width of anterior nostril equal to width of posterior nostril (
|
||
<figureCitation id="428054CCFFCAAA71B49FF859FDBB3585" box="[450,549,1967,1992]" captionStart="FIGURE 25" captionStartId="23.[151,250,1966,1988]" captionTargetBox="[151,1400,1593,1944]" captionTargetId="figure@23.[151,1436,1593,1944]" captionTargetPageId="23" captionText="FIGURE 25. Barbatula liaoyangensis, HU1608150, holotype, 75.9 mm SL; China: Liaoning prov.: Liao River drainage. Scale bars: a–b = 1 mm, c = 0.1 mm." httpUri="https://zenodo.org/record/2589539/files/figure.png" pageId="22" pageNumber="23">Fig. 25b</figureCitation>
|
||
). Upper jaw unexposed, completely covered by upper lip. Upper lip with very shallow median incision, its depth 10–30% of width of upper lip. Lower lip widely separated in middle. Mental lobe with blade-shaped lateral expansion. Interspace between mental lobes narrow, exposing small part of lower jaw. Lower lip with short lateral expansion, its length 30–40% of width of upper-lip (
|
||
<figureCitation id="428054CCFFCBAA70B135FF4AFB543298" box="[1128,1226,188,213]" captionStart="FIGURE 25" captionStartId="23.[151,250,1966,1988]" captionTargetBox="[151,1400,1593,1944]" captionTargetId="figure@23.[151,1436,1593,1944]" captionTargetPageId="23" captionText="FIGURE 25. Barbatula liaoyangensis, HU1608150, holotype, 75.9 mm SL; China: Liaoning prov.: Liao River drainage. Scale bars: a–b = 1 mm, c = 0.1 mm." httpUri="https://zenodo.org/record/2589539/files/figure.png" pageId="23" pageNumber="24">Fig. 25a</figureCitation>
|
||
). Maxillary barbel usually reaching to anterior half or middle of eye, inner rostral barbel usually reaching to posterior margin of anterior nostril, outer rostral barbel usually reaching to middle of snout. In few individuals, barbels slightly shorter.
|
||
</paragraph>
|
||
<paragraph id="DA044849FFCBAA70B59AFED1FA08339D" blockId="23.[151,1437,151,608]" pageId="23" pageNumber="24">Dorsal fin with 4 unbranched and 7½ branched rays. Anal fin with 2 unbranched and 5½ branched rays. Caudal fin with 7–8+8 branched rays. Pectoral fin with 1 unbranched and 10–11 branched rays. Pelvic fin with 1 unbranched and 6–7 branched rays. Infraorbital canal confluent with occipital canal, not confluent with supraorbital canal. Infraorbital canal with 12–14, supraorbital with 8–9, occipital with 3, mandibular and preopecular with 13–14 pores. Intestine Z-shaped, anterior margin of zigzag loop not touching U-shaped stomach.</paragraph>
|
||
<paragraph id="DA044849FFCBAA70B59AFE2AFB0F3012" blockId="23.[151,1437,151,608]" pageId="23" pageNumber="24">
|
||
<emphasis id="E8CF945BFFCBAA70B59AFE2AFE7F33B8" bold="true" box="[199,481,476,501]" pageId="23" pageNumber="24">Colouration in alcohol.</emphasis>
|
||
Background colour pale yellow with dark-brown pattern. Back with 11–13 large, dark-brown bars. Interspaces between the bars equal to width of them. Flank with large cloud-like mottling. Caudal fin hyaline with 5–6 dark-brown bands formed by spots. Dorsal fin hyaline with 3–5 dark-brown bands formed by spots. Pectoral and pelvic fins hyaline without pigmentation or with few dark-brown spots.
|
||
</paragraph>
|
||
<caption id="8EC418C1FFCBAA70B5CAFA25FCEE344A" httpUri="https://zenodo.org/record/2589537/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,250,1491,1513]" targetBox="[247,1338,719,1389]" targetPageId="23">
|
||
<paragraph id="DA044849FFCBAA70B5CAFA25FCEE344A" blockId="23.[151,1436,1491,1543]" pageId="23" pageNumber="24">
|
||
<emphasis id="E8CF945BFFCBAA70B5CAFA25FEB837A5" bold="true" box="[151,294,1491,1513]" pageId="23" pageNumber="24">FIGURE 24.</emphasis>
|
||
<taxonomicName id="1DBB33CAFFCBAA70B46FFA25FDAE37A4" authorityName="Chen & Zhang & Chen & Freyhof" authorityYear="2019" box="[306,560,1491,1513]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="liaoyangensis">
|
||
<emphasis id="E8CF945BFFCBAA70B46FFA25FDAE37A4" box="[306,560,1491,1513]" italics="true" pageId="23" pageNumber="24">Barbatula liaoyangensis</emphasis>
|
||
</taxonomicName>
|
||
, from the top: HU1608279, paratype, 82.3 mm SL; HU1608320, 75.9 mm SL; IHB88v0157, 74.2 mm SL; China: Liaoning prov.: Liao River drainage.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="8EC418C1FFCBAA70B5CAF858FE5135AF" httpUri="https://zenodo.org/record/2589539/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,250,1966,1988]" targetBox="[151,1400,1593,1944]" targetPageId="23">
|
||
<paragraph id="DA044849FFCBAA70B5CAF858FE5135AF" blockId="23.[151,1436,1966,2018]" pageId="23" pageNumber="24">
|
||
<emphasis id="E8CF945BFFCBAA70B5CAF858FE81358E" bold="true" box="[151,287,1966,1988]" pageId="23" pageNumber="24">FIGURE 25.</emphasis>
|
||
<taxonomicName id="1DBB33CAFFCBAA70B478F858FD833589" authorityName="Chen & Zhang & Chen & Freyhof" authorityYear="2019" box="[293,541,1966,1988]" class="Actinopterygii" family="Nemacheilidae" genus="Barbatula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="liaoyangensis">
|
||
<emphasis id="E8CF945BFFCBAA70B478F858FD833589" box="[293,541,1966,1988]" italics="true" pageId="23" pageNumber="24">Barbatula liaoyangensis</emphasis>
|
||
</taxonomicName>
|
||
, HU1608150, holotype, 75.9 mm SL; China: Liaoning prov.: Liao River drainage. Scale bars: a–b = 1 mm, c = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DA044849FFC4AA7FB59AFF61FA953298" blockId="24.[151,1436,151,285]" pageId="24" pageNumber="25">
|
||
<emphasis id="E8CF945BFFC4AA7FB59AFF61FE2F32FD" bold="true" box="[199,433,151,176]" pageId="24" pageNumber="25">Sexual dimorphism</emphasis>
|
||
. Males have a longer pectoral-fin than females (pectoral-fin length 15–16% SL vs. 12– 15% in female) and the 2nd–7rd branched pectoral-fin rays are thickened in males (vs. not in female).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92A11BC2FFC4AA7FB59AFF29FA9B32B5" box="[199,1285,223,248]" pageId="24" pageNumber="25" type="distribution">
|
||
<paragraph id="DA044849FFC4AA7FB59AFF29FA9B32B5" blockId="24.[151,1436,151,285]" box="[199,1285,223,248]" pageId="24" pageNumber="25">
|
||
<emphasis id="E8CF945BFFC4AA7FB59AFF29FEFE32B5" bold="true" box="[199,352,223,248]" pageId="24" pageNumber="25">Distribution.</emphasis>
|
||
Known from the Taizi River, a tributary of the Liao River. See
|
||
<figureCitation id="428054CCFFC4AA7FB17AFF29FB1832B5" box="[1063,1158,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1882,1904]" captionTargetBox="[173,1389,888,1818]" captionTargetId="figure@3.[151,1436,876,1861]" captionTargetPageId="3" captionText="FIGURE 2. Map showing the collection sites. Details for each number are listed in material examined." httpUri="https://zenodo.org/record/2589493/files/figure.png" pageId="24" pageNumber="25">Figure 2</figureCitation>
|
||
for details.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92A11BC2FFC4AA7FB59AFEF2FA1B3350" box="[199,1413,260,285]" pageId="24" pageNumber="25" type="etymology">
|
||
<paragraph id="DA044849FFC4AA7FB59AFEF2FA1B3350" blockId="24.[151,1436,151,285]" box="[199,1413,260,285]" pageId="24" pageNumber="25">
|
||
<emphasis id="E8CF945BFFC4AA7FB59AFEF2FED03350" bold="true" box="[199,334,260,285]" pageId="24" pageNumber="25">Etymology.</emphasis>
|
||
The name of the species is derived from its
|
||
<typeStatus id="0500F6EBFFC4AA7FB660FEF3FCF33350" box="[829,877,261,285]" pageId="24" pageNumber="25">type</typeStatus>
|
||
locality in the Chinese Liaoyang. An adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |