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<document ID-DOI="10.11646/zootaxa.5278.2.6" ID-GBIF-Dataset="869b1ddb-d171-4dc3-9dd7-22084bd360e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="7906289" ID-ZooBank="E84EA49C-11B0-4022-8C66-87A54CEA71A9" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683529750663" checkinUser="plazi" docAuthor="Smolis, Adrian, Skarżyński, Dariusz &amp; Gwiazdowicz, Dariusz J." docDate="2023" docId="5211B4092819FFFFFF30515BFAF1FD49" docLanguage="en" docName="zootaxa.5278.2.6.pdf" docOrigin="Zootaxa 5278 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Friesea schulzi Smolis &amp; Skarżyński &amp; Gwiazdowicz 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="336" masterDocId="AE28CC712818FFFCFFA7523CFF92FF84" masterDocTitle="New species of Neanuridae (Collembola) living on the endemic tree Zelkova abelicea in Crete" masterLastPageNumber="350" masterPageNumber="333" pageNumber="334" updateTime="1683530458092" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New species of Neanuridae (Collembola) living on the endemic tree Zelkova abelicea in Crete</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smolis, Adrian</mods:namePart>
<mods:affiliation>University of Wrocław, Faculty of Biological Sciences, Department of Invertebrate Biology, Evolution and Conservation, Przybyszewskiego 65, 51 - 148 Wrocław, Poland.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Skarżyński, Dariusz</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-1767-5747</mods:nameIdentifier>
<mods:affiliation>University of Wrocław, Faculty of Biological Sciences, Department of Invertebrate Biology, Evolution and Conservation, Przybyszewskiego 65, 51 - 148 Wrocław, Poland. &amp; dariusz. skarzynski @ uwr. edu. pl; https: // orcid. org / 0000 - 0002 - 1767 - 5747</mods:affiliation>
<mods:nameIdentifier type="email">dariusz.skarzynski@uwr.edu.pl</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gwiazdowicz, Dariusz J.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-0064-2316</mods:nameIdentifier>
<mods:affiliation>Poznań University of Life Sciences, Faculty of Forestry and Wood Technology, Department of Forest Entomology and Pathology, Wojska Polskiego 71 c, 60 - 625, Poznań, Poland. dariusz. gwiazdowicz @ up. poznan. pl; https: // orcid. org / 0000 - 0002 - 0064 - 2316</mods:affiliation>
<mods:nameIdentifier type="email">dariusz.gwiazdowicz@up.poznan.pl</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-05-05</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5278</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>333</mods:start>
<mods:end>350</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5278.2.6</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5278.2.6</mods:identifier>
<mods:identifier type="GBIF-Dataset">869b1ddb-d171-4dc3-9dd7-22084bd360e9</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7906289</mods:identifier>
<mods:identifier type="ZooBank">E84EA49C-11B0-4022-8C66-87A54CEA71A9</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7906291" ID-Zenodo-Dep="7906291" LSID="urn:lsid:plazi:treatment:5211B4092819FFFFFF30515BFAF1FD49" httpUri="http://treatment.plazi.org/id/5211B4092819FFFFFF30515BFAF1FD49" lastPageId="3" lastPageNumber="336" pageId="1" pageNumber="334">
<subSubSection box="[151,435,870,897]" pageId="1" pageNumber="334" type="nomenclature">
<paragraph blockId="1.[151,435,870,933]" box="[151,435,870,897]" pageId="1" pageNumber="334">
<heading bold="true" box="[151,435,870,897]" fontSize="11" level="1" pageId="1" pageNumber="334" reason="1">
<emphasis bold="true" box="[151,435,870,897]" pageId="1" pageNumber="334">
<taxonomicName authority="Smolis &amp; Skarżyński &amp; Gwiazdowicz, 2023" authorityName="Smolis &amp; Skarżyński &amp; Gwiazdowicz" authorityYear="2023" box="[151,335,870,897]" class="Collembola" family="Neanuridae" genus="Friesea" kingdom="Animalia" order="Poduromorpha" pageId="1" pageNumber="334" phylum="Arthropoda" rank="species" species="schulzi" status="sp. nov.">
<emphasis bold="true" box="[151,335,870,897]" italics="true" pageId="1" pageNumber="334">Friesea schulzi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[342,435,871,897]" pageId="1" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,248,907,933]" pageId="1" pageNumber="334" type="description">
<paragraph blockId="1.[151,435,870,933]" box="[151,248,907,933]" pageId="1" pageNumber="334">
<figureCitation box="[151,248,907,933]" captionStart="FIGURES 16" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[168,1418,185,1910]" captionTargetId="figure-18@2.[165,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 16. Friesea schulzi sp. nov.: 1, dorsal chaetotaxy of head, Th. and Abd. I (holotype); 2, tibiotarsus and claw of second pair of legs, ventrolateral view; 3, tibiotarsus and claw of second pair of legs, dorsolateral view; 4, ventral chaetotaxy of Abd. IVV; 5, ventral chaetotaxy of Abd. VI; 6, dorsal chaetotaxy of Abd. IVVI (holotype)." figureDoi="http://doi.org/10.5281/zenodo.7906293" httpUri="https://zenodo.org/record/7906293/files/figure.png" pageId="1" pageNumber="334">Figs 16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="334" type="materials_examined">
<paragraph blockId="1.[151,1437,978,1941]" pageId="1" pageNumber="334">
<emphasis bold="true" box="[151,330,978,1005]" pageId="1" pageNumber="334">Type material.</emphasis>
<materialsCitation collectingDate="2018-10-11" collectorName="D. J. Gwiazdowicz." country="Greece" elevation="1255" latitude="35.194828" location="Mt. Kedros" longLatPrecision="1" longitude="24.606712" municipality="Gerakari village" pageId="1" pageNumber="334" specimenCount="1" stateProvince="Kriti" typeStatus="holotype">
<typeStatus box="[342,451,979,1005]" pageId="1" pageNumber="334">Holotype</typeStatus>
: female, bark of
<taxonomicName box="[659,854,979,1005]" class="Magnoliopsida" family="Ulmaceae" genus="Zelkova" kingdom="Plantae" order="Rosales" pageId="1" pageNumber="334" phylum="Tracheophyta" rank="species" species="abelicea">
<emphasis box="[659,854,979,1005]" italics="true" pageId="1" pageNumber="334">Zelkova abelicea</emphasis>
</taxonomicName>
trees, slope of
<location LSID="urn:lsid:plazi:treatment:5211B4092819FFFFFF30515BFAF1FD49:DF6753C42819FFFDFBB251EFFB08FC69" box="[1045,1178,979,1005]" country="Greece" latitude="35.194828" longLatPrecision="1" longitude="24.606712" municipality="Gerakari village" name="Mt. Kedros" pageId="1" pageNumber="334" stateProvince="Kriti">Mt. Kedros</location>
near
<collectingMunicipality box="[1250,1436,979,1005]" pageId="1" pageNumber="334">Gerakari village</collectingMunicipality>
(
<geoCoordinate box="[159,308,1014,1041]" degrees="35.194829" direction="north" orientation="latitude" pageId="1" pageNumber="334" precision="1" value="35.194828">35.194829 N</geoCoordinate>
,
<geoCoordinate box="[319,464,1015,1041]" degrees="24.606713" direction="east" orientation="longitude" pageId="1" pageNumber="334" precision="1" value="24.606712">24.606713 E</geoCoordinate>
),
<elevation box="[483,637,1014,1041]" metricMagnitude="3" metricUnit="m" metricValue="1.255" pageId="1" pageNumber="334" unit="m" value="1255.0">
<quantity box="[483,567,1014,1041]" metricMagnitude="3" metricUnit="m" metricValue="1.255" pageId="1" pageNumber="334" unit="m" value="1255.0">1255 m</quantity>
a. s. l.
</elevation>
,
<collectingRegion box="[649,712,1015,1041]" country="Greece" name="Kriti" pageId="1" pageNumber="334">Crete</collectingRegion>
,
<collectingCountry box="[723,805,1015,1041]" name="Greece" pageId="1" pageNumber="334">Greece</collectingCountry>
,
<date box="[815,934,1015,1041]" pageId="1" pageNumber="334" value="2018-10-11">
<collectingDate box="[815,934,1015,1041]" pageId="1" pageNumber="334" value="2018-10-11">11.X.2018</collectingDate>
</date>
, leg.
<collectorName box="[993,1205,1015,1041]" pageId="1" pageNumber="334">D.J. Gwiazdowicz.</collectorName>
</materialsCitation>
<materialsCitation collectingDate="2018-10-11" collectorName="D. J. Gwiazdowicz." country="Greece" elevation="1255" latitude="35.194828" location="Mt. Kedros" longLatPrecision="1" longitude="24.606712" municipality="Gerakari village" pageId="1" pageNumber="334" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Kriti" typeStatus="paratype">
<typeStatus box="[1210,1323,1015,1041]" pageId="1" pageNumber="334">Paratypes</typeStatus>
:
<specimenCount box="[1333,1432,1015,1041]" pageId="1" pageNumber="334" type="female">1 female</specimenCount>
,
<specimenCount box="[151,229,1051,1077]" pageId="1" pageNumber="334" type="male">1 male</specimenCount>
, same data as above
</materialsCitation>
,
<materialsCitation box="[470,985,1051,1077]" collectingDate="2021-09-29" collectorName="D. J. Gwiazdowicz." country="Greece" elevation="1255" latitude="35.194828" location="Mt. Kedros" longLatPrecision="1" longitude="24.606712" municipality="Gerakari village" pageId="1" pageNumber="334" specimenCount="1" specimenCount-female="1" stateProvince="Kriti" typeStatus="paratype">
<specimenCount box="[470,570,1051,1077]" pageId="1" pageNumber="334" type="female">1 female</specimenCount>
, same data as above, but
<date box="[853,985,1051,1077]" pageId="1" pageNumber="334" value="2021-09-29">
<collectingDate box="[853,985,1051,1077]" pageId="1" pageNumber="334" value="2021-09-29">29.IX.2021</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="334" type="description">
<paragraph blockId="1.[151,1437,978,1941]" pageId="1" pageNumber="334">
<emphasis bold="true" box="[199,345,1086,1112]" pageId="1" pageNumber="334">Description.</emphasis>
Habitus typical of the genus. Body length 0.550.68 mm. Colour of the body white, ocelli with black pigment.
</paragraph>
<paragraph blockId="1.[151,1437,978,1941]" pageId="1" pageNumber="334">Antennae shorter than head. Ant. I with 7 chaetae, Ant. II with 12 chaetae. Ant. III and IV fused dorsally. AOIII with two small internal curved sensilla, two cylindrical guard sensilla and ventral microsensillum. Ant. IV with simple apical bulb, small subapical organite, microsensillum, chaeta i and 6 cylindrical, curved sensilla (S1 and S3 clearly thinner than others).</paragraph>
<paragraph blockId="1.[151,1437,978,1941]" box="[199,737,1303,1329]" pageId="1" pageNumber="334">
Ocelli 3 + 3 (
<figureCitation box="[348,413,1303,1329]" captionStart="FIGURES 16" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[168,1418,185,1910]" captionTargetId="figure-18@2.[165,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 16. Friesea schulzi sp. nov.: 1, dorsal chaetotaxy of head, Th. and Abd. I (holotype); 2, tibiotarsus and claw of second pair of legs, ventrolateral view; 3, tibiotarsus and claw of second pair of legs, dorsolateral view; 4, ventral chaetotaxy of Abd. IVV; 5, ventral chaetotaxy of Abd. VI; 6, dorsal chaetotaxy of Abd. IVVI (holotype)." figureDoi="http://doi.org/10.5281/zenodo.7906293" httpUri="https://zenodo.org/record/7906293/files/figure.png" pageId="1" pageNumber="334">Fig. 1</figureCitation>
). Postantennal organ absent.
</paragraph>
<paragraph blockId="1.[151,1437,978,1941]" pageId="1" pageNumber="334">Labrum with 5, 3, 4 chaetae. Prelabral chetae 2. Labium with complete chaetotaxy including papillated chaeta L. Mandible with 8 teeth. Head of maxilla with two dentate lamellae.</paragraph>
<paragraph blockId="1.[151,1437,978,1941]" pageId="1" pageNumber="334">
<materialsCitation collectionCode="II, III, IV, V, VI" collectorName="Th. I &amp; Abd. I" location="Microsensillum on Th." pageId="1" pageNumber="334" specimenCount="1">
Dorsal chaetotaxy as in
<figureCitation box="[471,572,1411,1437]" captionStart="FIGURES 16" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[168,1418,185,1910]" captionTargetId="figure-18@2.[165,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 16. Friesea schulzi sp. nov.: 1, dorsal chaetotaxy of head, Th. and Abd. I (holotype); 2, tibiotarsus and claw of second pair of legs, ventrolateral view; 3, tibiotarsus and claw of second pair of legs, dorsolateral view; 4, ventral chaetotaxy of Abd. IVV; 5, ventral chaetotaxy of Abd. VI; 6, dorsal chaetotaxy of Abd. IVVI (holotype)." figureDoi="http://doi.org/10.5281/zenodo.7906293" httpUri="https://zenodo.org/record/7906293/files/figure.png" pageId="1" pageNumber="334">Figs 1, 6</figureCitation>
. Chaetae short and smooth, body sensilla fine and smooth. Sensillar formula per half tergum: 022/11111. Lateral sensilla on Th.
<collectionCode box="[721,739,1447,1473]" pageId="1" pageNumber="334">II</collectionCode>
<collectionCode box="[754,782,1447,1473]" pageId="1" pageNumber="334">III</collectionCode>
distinctly longer than nearest ordinary chaetae, other body sensilla only slightly longer.
<location LSID="urn:lsid:plazi:treatment:5211B4092819FFFFFF30515BFAF1FD49:DF6753C42819FFFDFE7B57F7FD4FFA61" box="[476,733,1483,1509]" name="Microsensillum on Th." pageId="1" pageNumber="334">Microsensillum on Th.</location>
<collectionCode box="[740,759,1483,1509]" pageId="1" pageNumber="334">II</collectionCode>
present. Chaetae a
<subScript attach="right" box="[972,980,1501,1515]" fontSize="6" pageId="1" pageNumber="334">0</subScript>
, d
<subScript attach="left" box="[1008,1016,1501,1515]" fontSize="6" pageId="1" pageNumber="334">1</subScript>
and p
<subScript attach="left" box="[1083,1091,1501,1515]" fontSize="6" pageId="1" pageNumber="334">1</subScript>
on head present (
<figureCitation box="[1291,1357,1483,1509]" captionStart="FIGURES 16" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[168,1418,185,1910]" captionTargetId="figure-18@2.[165,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 16. Friesea schulzi sp. nov.: 1, dorsal chaetotaxy of head, Th. and Abd. I (holotype); 2, tibiotarsus and claw of second pair of legs, ventrolateral view; 3, tibiotarsus and claw of second pair of legs, dorsolateral view; 4, ventral chaetotaxy of Abd. IVV; 5, ventral chaetotaxy of Abd. VI; 6, dorsal chaetotaxy of Abd. IVVI (holotype)." figureDoi="http://doi.org/10.5281/zenodo.7906293" httpUri="https://zenodo.org/record/7906293/files/figure.png" pageId="1" pageNumber="334">Fig. 1</figureCitation>
).
<collectorName box="[1381,1436,1483,1509]" pageId="1" pageNumber="334">Th. I</collectorName>
with 4 + 4 chaetae. Chaetae a
<subScript attach="left" box="[479,487,1537,1551]" fontSize="6" pageId="1" pageNumber="334">2</subScript>
on Th.
<collectionCode box="[574,593,1519,1545]" pageId="1" pageNumber="334">II</collectionCode>
present, m
<subScript attach="left" box="[716,724,1537,1551]" fontSize="6" pageId="1" pageNumber="334">4</subScript>
absent.
<collectorName box="[812,883,1519,1545]" pageId="1" pageNumber="334">Abd. I</collectorName>
<collectionCode box="[897,925,1519,1545]" pageId="1" pageNumber="334">III</collectionCode>
with chaetae m
<subScript attach="left" box="[1101,1109,1537,1551]" fontSize="6" pageId="1" pageNumber="334">3</subScript>
and m
<subScript attach="left" box="[1185,1193,1537,1551]" fontSize="6" pageId="1" pageNumber="334">4</subScript>
usually present. Abd.
<collectionCode box="[151,180,1555,1581]" pageId="1" pageNumber="334">IV</collectionCode>
with chaetae a
<subScript attach="left" box="[351,359,1573,1587]" fontSize="6" pageId="1" pageNumber="334">3</subScript>
usually present and chaetae m
<subScript attach="left" box="[708,716,1573,1587]" fontSize="6" pageId="1" pageNumber="334">4</subScript>
and m
<subScript attach="left" box="[794,802,1573,1587]" fontSize="6" pageId="1" pageNumber="334">5</subScript>
rarely present. Abd.
<collectionCode box="[1042,1062,1555,1581]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="334" type="Museum">V</collectionCode>
without chaetae p
<subScript attach="left" box="[1269,1277,1573,1587]" fontSize="6" pageId="1" pageNumber="334">2</subScript>
(
<figureCitation box="[1291,1360,1555,1581]" captionStart="FIGURES 16" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[168,1418,185,1910]" captionTargetId="figure-18@2.[165,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 16. Friesea schulzi sp. nov.: 1, dorsal chaetotaxy of head, Th. and Abd. I (holotype); 2, tibiotarsus and claw of second pair of legs, ventrolateral view; 3, tibiotarsus and claw of second pair of legs, dorsolateral view; 4, ventral chaetotaxy of Abd. IVV; 5, ventral chaetotaxy of Abd. VI; 6, dorsal chaetotaxy of Abd. IVVI (holotype)." figureDoi="http://doi.org/10.5281/zenodo.7906293" httpUri="https://zenodo.org/record/7906293/files/figure.png" pageId="1" pageNumber="334">Fig. 6</figureCitation>
). Abd.
<collectionCode box="[151,181,1591,1617]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="1" pageNumber="334">VI</collectionCode>
with spines embedded directly on the tegument in positions a
<subScript attach="left" box="[888,896,1609,1623]" fontSize="6" pageId="1" pageNumber="334">1</subScript>
and m
<subScript attach="left" box="[976,984,1609,1623]" fontSize="6" pageId="1" pageNumber="334">1</subScript>
, and with thickened, smooth chaetae in positions a
<subScript attach="left" box="[272,280,1645,1659]" fontSize="6" pageId="1" pageNumber="334">2</subScript>
and p
<subScript attach="left" box="[348,356,1645,1659]" fontSize="6" pageId="1" pageNumber="334">2</subScript>
(in
<specimenCount box="[402,555,1627,1653]" pageId="1" pageNumber="334" type="generic">one specimen</specimenCount>
unpaired p
<subScript attach="left" box="[681,689,1645,1659]" fontSize="6" pageId="1" pageNumber="334">2</subScript>
as small spine)
</materialsCitation>
.
</paragraph>
<paragraph blockId="1.[151,1437,978,1941]" pageId="1" pageNumber="334">
Thoracic sterna without chaetae.Ventral abdominal chaetotaxy as in
<figureCitation box="[983,1087,1662,1689]" captionStart="FIGURES 16" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[168,1418,185,1910]" captionTargetId="figure-18@2.[165,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 16. Friesea schulzi sp. nov.: 1, dorsal chaetotaxy of head, Th. and Abd. I (holotype); 2, tibiotarsus and claw of second pair of legs, ventrolateral view; 3, tibiotarsus and claw of second pair of legs, dorsolateral view; 4, ventral chaetotaxy of Abd. IVV; 5, ventral chaetotaxy of Abd. VI; 6, dorsal chaetotaxy of Abd. IVVI (holotype)." figureDoi="http://doi.org/10.5281/zenodo.7906293" httpUri="https://zenodo.org/record/7906293/files/figure.png" pageId="1" pageNumber="334">Figs 4, 5</figureCitation>
. Each anal valve with 3 small chaetae hr.
</paragraph>
<paragraph blockId="1.[151,1437,978,1941]" pageId="1" pageNumber="334">
Tibiotarsi I, II, III with 17, 17, 16 chaetae respectively, clavate tenant hairs absent (
<figureCitation box="[1135,1237,1735,1761]" captionStart="FIGURES 16" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[168,1418,185,1910]" captionTargetId="figure-18@2.[165,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 16. Friesea schulzi sp. nov.: 1, dorsal chaetotaxy of head, Th. and Abd. I (holotype); 2, tibiotarsus and claw of second pair of legs, ventrolateral view; 3, tibiotarsus and claw of second pair of legs, dorsolateral view; 4, ventral chaetotaxy of Abd. IVV; 5, ventral chaetotaxy of Abd. VI; 6, dorsal chaetotaxy of Abd. IVVI (holotype)." figureDoi="http://doi.org/10.5281/zenodo.7906293" httpUri="https://zenodo.org/record/7906293/files/figure.png" pageId="1" pageNumber="334">Figs 2, 3</figureCitation>
). Femora I, II, III with 12, 11, 10 chaetae respectively, trochanters with 5 chaetae each, coxae I, II, III with 3, 78, 7 chaetae, subcoxae 2 of legs I, II and III with 0, 2 and 23 chaetae, subcoxae 1 of legs I, II and III with 1, 2 and 2 chaetae. Claws with small inner tooth (
<figureCitation box="[356,457,1843,1869]" captionStart="FIGURES 16" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[168,1418,185,1910]" captionTargetId="figure-18@2.[165,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 16. Friesea schulzi sp. nov.: 1, dorsal chaetotaxy of head, Th. and Abd. I (holotype); 2, tibiotarsus and claw of second pair of legs, ventrolateral view; 3, tibiotarsus and claw of second pair of legs, dorsolateral view; 4, ventral chaetotaxy of Abd. IVV; 5, ventral chaetotaxy of Abd. VI; 6, dorsal chaetotaxy of Abd. IVVI (holotype)." figureDoi="http://doi.org/10.5281/zenodo.7906293" httpUri="https://zenodo.org/record/7906293/files/figure.png" pageId="1" pageNumber="334">Figs 2, 3</figureCitation>
). Empodial appendage absent.
</paragraph>
<paragraph blockId="1.[151,1437,978,1941]" pageId="1" pageNumber="334">
Ventral tube with 4 + 4 chaetae. Furca and retinaculum absent (stage 5 according to
<bibRefCitation author="Cassagnau, P." box="[1140,1327,1878,1905]" pageId="1" pageNumber="334" pagination="17 - 29" refId="ref8661" refString="Cassagnau, P. (1958) Les especes europeennes du genre Friesea (Collemboles Poduromorphes). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 93, 17 - 29." type="journal article" year="1958">Cassagnau 1958</bibRefCitation>
), Abd. IV ventrally with 56 microchaetae in two rows instead of furca (
<figureCitation box="[848,918,1915,1941]" captionStart="FIGURES 16" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[168,1418,185,1910]" captionTargetId="figure-18@2.[165,1422,181,1919]" captionTargetPageId="2" captionText="FIGURES 16. Friesea schulzi sp. nov.: 1, dorsal chaetotaxy of head, Th. and Abd. I (holotype); 2, tibiotarsus and claw of second pair of legs, ventrolateral view; 3, tibiotarsus and claw of second pair of legs, dorsolateral view; 4, ventral chaetotaxy of Abd. IVV; 5, ventral chaetotaxy of Abd. VI; 6, dorsal chaetotaxy of Abd. IVVI (holotype)." figureDoi="http://doi.org/10.5281/zenodo.7906293" httpUri="https://zenodo.org/record/7906293/files/figure.png" pageId="1" pageNumber="334">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7906293" ID-Zenodo-Dep="7906293" httpUri="https://zenodo.org/record/7906293/files/figure.png" pageId="2" pageNumber="335" startId="2.[151,264,1943,1967]" targetBox="[168,1418,185,1910]" targetPageId="2" targetType="figure">
<paragraph blockId="2.[151,1436,1943,2040]" pageId="2" pageNumber="335">
<emphasis bold="true" box="[151,316,1943,1967]" pageId="2" pageNumber="335">FIGURES 16.</emphasis>
<taxonomicName authority="Smolis &amp; Skarżyński &amp; Gwiazdowicz, 2023" authorityName="Smolis &amp; Skarżyński &amp; Gwiazdowicz" authorityYear="2023" box="[324,480,1944,1968]" class="Collembola" family="Neanuridae" genus="Friesea" kingdom="Animalia" order="Poduromorpha" pageId="2" pageNumber="335" phylum="Arthropoda" rank="species" species="schulzi" status="sp. nov.">
<emphasis box="[324,480,1944,1968]" italics="true" pageId="2" pageNumber="335">Friesea schulzi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[488,571,1944,1968]" pageId="2" pageNumber="335">
<taxonomicNameLabel box="[488,571,1944,1968]" pageId="2" pageNumber="335" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 1, dorsal chaetotaxy of head, Th. and Abd. I (holotype); 2, tibiotarsus and claw of second pair of legs, ventrolateral view; 3, tibiotarsus and claw of second pair of legs, dorsolateral view; 4, ventral chaetotaxy of Abd. IVV; 5, ventral chaetotaxy of Abd. VI; 6, dorsal chaetotaxy of Abd. IVVI (holotype).
</paragraph>
</caption>
<subSubSection pageId="3" pageNumber="336" type="etymology">
<paragraph blockId="3.[151,1437,151,717]" pageId="3" pageNumber="336">
<emphasis bold="true" box="[199,334,151,177]" pageId="3" pageNumber="336">Etymology.</emphasis>
Dedicated to Dr. Hans-J̧rgen Schulz, who made a significant contribution to the knowledge of the fauna of
<collectingRegion box="[251,315,187,213]" country="Greece" name="Kriti" pageId="3" pageNumber="336">Crete</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="336" type="discussion">
<paragraph blockId="3.[151,1437,151,717]" pageId="3" pageNumber="336">
<emphasis bold="true" box="[199,315,223,249]" pageId="3" pageNumber="336">Remarks.</emphasis>
Based on the key features proposed by
<bibRefCitation author="Cassagnau, P." box="[786,991,222,249]" pageId="3" pageNumber="336" pagination="17 - 29" refId="ref8661" refString="Cassagnau, P. (1958) Les especes europeennes du genre Friesea (Collemboles Poduromorphes). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 93, 17 - 29." type="journal article" year="1958">Cassagnau (1958)</bibRefCitation>
<taxonomicName authority="Smolis &amp; Skarżyński &amp; Gwiazdowicz, 2023" authorityName="Smolis &amp; Skarżyński &amp; Gwiazdowicz" authorityYear="2023" box="[1001,1175,223,249]" class="Collembola" family="Neanuridae" genus="Friesea" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="schulzi" status="sp. nov.">
<emphasis box="[1001,1175,223,249]" italics="true" pageId="3" pageNumber="336">Friesea schulzi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1185,1277,223,249]" pageId="3" pageNumber="336">
<taxonomicNameLabel box="[1185,1277,223,249]" pageId="3" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is close to
<taxonomicName authority="Tamura &amp; Yue, 1998" authorityName="Tamura &amp; Yue" authorityYear="1998" class="Collembola" family="Neanuridae" genus="Friesea" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="daliensis">
<emphasis italics="true" pageId="3" pageNumber="336">F. daliensis</emphasis>
<bibRefCitation author="Tamura, H. &amp; Yue, Q." box="[258,495,259,285]" pageId="3" pageNumber="336" pagination="623 - 626" refId="ref10087" refString="Tamura, H. &amp; Yue, Q. (1998) Two new species of the genus Friesea from northwest region of Yunnan Province, China (Collembola, Neanuridae). Entomological Science, 1 (4), 623 - 626." type="journal article" year="1998">Tamura &amp; Yue, 1998</bibRefCitation>
</taxonomicName>
from
<collectingCountry box="[565,636,259,285]" name="China" pageId="3" pageNumber="336">China</collectingCountry>
. Both species share such features as white body, 3 + 3 ocelli and furca and retinaculum absent (stage 5). However, they differ clearly in the number of anal spines (
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<taxonomicName authorityName="Tamura &amp; Yue" authorityYear="1998" box="[1250,1378,295,321]" class="Collembola" family="Neanuridae" genus="Friesea" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="daliensis">
<emphasis box="[1250,1378,295,321]" italics="true" pageId="3" pageNumber="336">F. daliensis</emphasis>
</taxonomicName>
,
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positions a
<subScript attach="left" box="[272,280,349,363]" fontSize="6" pageId="3" pageNumber="336">1</subScript>
and p
<quantity box="[350,385,331,363]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="336" unit="in" value="1.0">
<subScript attach="left" box="[350,358,349,363]" fontSize="6" pageId="3" pageNumber="336">1</subScript>
in
</quantity>
<taxonomicName authority="Smolis &amp; Skarżyński &amp; Gwiazdowicz, 2023" authorityName="Smolis &amp; Skarżyński &amp; Gwiazdowicz" authorityYear="2023" box="[392,499,331,357]" class="Collembola" family="Neanuridae" genus="Friesea" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="schulzi" status="sp. nov.">
<emphasis box="[392,499,331,357]" italics="true" pageId="3" pageNumber="336">F. schulzi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[506,596,331,357]" pageId="3" pageNumber="336">
<taxonomicNameLabel box="[506,596,331,357]" pageId="3" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, which has also thickened, smooth chaetae in positions a
<subScript attach="left" box="[1241,1249,349,363]" fontSize="6" pageId="3" pageNumber="336">2</subScript>
and p
<subScript attach="left" box="[1319,1327,349,363]" fontSize="6" pageId="3" pageNumber="336">2</subScript>
), number of chaetae on tibiotarsi I, II, III (16, 16,
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<taxonomicName authorityName="Tamura &amp; Yue" authorityYear="1998" box="[660,788,367,393]" class="Collembola" family="Neanuridae" genus="Friesea" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="daliensis">
<emphasis box="[660,788,367,393]" italics="true" pageId="3" pageNumber="336">F. daliensis</emphasis>
</taxonomicName>
, 17, 17, 16 respectively in
<taxonomicName authority="Smolis &amp; Skarżyński &amp; Gwiazdowicz, 2023" authorityName="Smolis &amp; Skarżyński &amp; Gwiazdowicz" authorityYear="2023" box="[1090,1196,367,393]" class="Collembola" family="Neanuridae" genus="Friesea" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="schulzi" status="sp. nov.">
<emphasis box="[1090,1196,367,393]" italics="true" pageId="3" pageNumber="336">F. schulzi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1202,1292,367,393]" pageId="3" pageNumber="336">
<taxonomicNameLabel box="[1202,1292,367,393]" pageId="3" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), chaetotaxy of labrum (3, 3, 4 chaetae in
<taxonomicName authorityName="Tamura &amp; Yue" authorityYear="1998" box="[486,616,403,429]" class="Collembola" family="Neanuridae" genus="Friesea" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="daliensis">
<emphasis box="[486,616,403,429]" italics="true" pageId="3" pageNumber="336">F. daliensis</emphasis>
</taxonomicName>
, 5, 3,
<quantity box="[691,735,403,429]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="3" pageNumber="336" unit="in" value="4.0">4 in</quantity>
<taxonomicName authority="Smolis &amp; Skarżyński &amp; Gwiazdowicz, 2023" authorityName="Smolis &amp; Skarżyński &amp; Gwiazdowicz" authorityYear="2023" box="[744,851,403,429]" class="Collembola" family="Neanuridae" genus="Friesea" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="schulzi" status="sp. nov.">
<emphasis box="[744,851,403,429]" italics="true" pageId="3" pageNumber="336">F. schulzi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[860,951,403,429]" pageId="3" pageNumber="336">
<taxonomicNameLabel box="[860,951,403,429]" pageId="3" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and chaetotaxy of labium (in
<taxonomicName authorityName="Tamura &amp; Yue" authorityYear="1998" box="[1307,1437,403,429]" class="Collembola" family="Neanuridae" genus="Friesea" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="daliensis">
<emphasis box="[1307,1437,403,429]" italics="true" pageId="3" pageNumber="336">F. daliensis</emphasis>
</taxonomicName>
chaetae B absent, in
<taxonomicName authority="Smolis &amp; Skarżyński &amp; Gwiazdowicz, 2023" authorityName="Smolis &amp; Skarżyński &amp; Gwiazdowicz" authorityYear="2023" box="[383,489,439,465]" class="Collembola" family="Neanuridae" genus="Friesea" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="schulzi" status="sp. nov.">
<emphasis box="[383,489,439,465]" italics="true" pageId="3" pageNumber="336">F. schulzi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[496,586,439,465]" pageId="3" pageNumber="336">
<taxonomicNameLabel box="[496,586,439,465]" pageId="3" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
present). The same number of ocelli and missing furca and retinaculum are characteristic for three European species
<taxonomicName authority="Denis, 1932" authorityName="Denis" authorityYear="1932" box="[615,842,475,501]" class="Collembola" family="Neanuridae" genus="Friesea" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="fagei">
<emphasis box="[615,699,475,501]" italics="true" pageId="3" pageNumber="336">F. fagei</emphasis>
<bibRefCitation author="Denis, J. R." box="[706,842,475,501]" pageId="3" pageNumber="336" pagination="357 - 383" refId="ref8936" refString="Denis, J. R. (1932) Sur la faune francaise des Apterygotes, XII. Archives de Zoologie Experimentale et Generale, Paris, 74, 357 - 383." type="journal article" year="1932">Denis, 1932</bibRefCitation>
</taxonomicName>
(
<collectingCountry box="[857,938,475,501]" name="France" pageId="3" pageNumber="336">France</collectingCountry>
,
<collectingCountry box="[950,1002,475,501]" name="Italy" pageId="3" pageNumber="336">Italy</collectingCountry>
,
<collectingCountry box="[1014,1082,475,501]" name="Spain" pageId="3" pageNumber="336">Spain</collectingCountry>
),
<taxonomicName authority="Simon-Benito, 2005" authorityName="Simon-Benito" authorityYear="2005" class="Collembola" family="Neanuridae" genus="Friesea" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="oteruelensis">
<emphasis box="[1101,1264,475,501]" italics="true" pageId="3" pageNumber="336">F. oteruelensis</emphasis>
<bibRefCitation author="Simon-Benito, J. C." pageId="3" pageNumber="336" pagination="327 - 330" refId="ref9429" refString="Simon-Benito, J. C. (2005) Una nueva especie de Friesea de la peninsula Iberica y consideraciones sobre F. decipiens Steiner, 1959 (Collembola, Frieseinae). Bulletin de la Societe Entomologique de France, 110 (3), 327 - 330. https: // doi. org / 10.3406 / bsef. 2005.16239" type="journal article" year="2005">Simón-Benito, 2005</bibRefCitation>
</taxonomicName>
(
<collectingCountry box="[223,290,511,537]" name="Spain" pageId="3" pageNumber="336">Spain</collectingCountry>
) and
<taxonomicName authority="Giuga &amp; Jordana, 2013" authorityName="Giuga &amp; Jordana" authorityYear="2013" box="[353,750,511,537]" class="Collembola" family="Neanuridae" genus="Friesea" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="guarinoi">
<emphasis box="[353,478,511,536]" italics="true" pageId="3" pageNumber="336">F. guarinoi</emphasis>
<bibRefCitation author="Giuga, L. &amp; Jordana, R." box="[486,750,511,537]" pageId="3" pageNumber="336" pagination="283 - 300" refId="ref8996" refString="Giuga, L. &amp; Jordana, R. (2013) Three new species of Collembola from soils of Mediterranean cork-oak forests of Sicily (Italy). Zootaxa, 3664 (2), 283 - 300. https: // doi. org / 10.11646 / zootaxa. 3664.2.9" type="journal article" year="2013">Giuga &amp; Jordana, 2013</bibRefCitation>
</taxonomicName>
(
<collectingCountry box="[766,823,511,537]" name="Italy" pageId="3" pageNumber="336">Italy</collectingCountry>
). However, all of them possess exclusively chaetae on Abd. VI and not spines. Similar in respect of degree of furca reduction are also other European species
<taxonomicName authority="Cassagnau, 1958" authorityName="Cassagnau" authorityYear="1958" class="Collembola" family="Neanuridae" genus="Friesea" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="troglophila">
<emphasis box="[1285,1436,547,573]" italics="true" pageId="3" pageNumber="336">F. troglophila</emphasis>
<bibRefCitation author="Cassagnau, P." box="[151,342,582,609]" pageId="3" pageNumber="336" pagination="17 - 29" refId="ref8661" refString="Cassagnau, P. (1958) Les especes europeennes du genre Friesea (Collemboles Poduromorphes). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 93, 17 - 29." type="journal article" year="1958">Cassagnau, 1958</bibRefCitation>
</taxonomicName>
(
<collectingCountry box="[357,438,583,609]" name="France" pageId="3" pageNumber="336">France</collectingCountry>
,
<collectingCountry box="[450,517,583,609]" name="Spain" pageId="3" pageNumber="336">Spain</collectingCountry>
,
<collectingCountry box="[529,628,583,609]" name="Portugal" pageId="3" pageNumber="336">Portugal</collectingCountry>
),
<taxonomicName authority="Jordana &amp; Asiain, 1981" authorityName="Jordana &amp; Asiain" authorityYear="1981" box="[648,1057,583,609]" class="Collembola" family="Neanuridae" genus="Friesea" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="bioculata">
<emphasis box="[648,782,583,609]" italics="true" pageId="3" pageNumber="336">F. bioculata</emphasis>
<bibRefCitation author="Jordana, R. &amp; Asiain, A." box="[790,1057,583,609]" pageId="3" pageNumber="336" pagination="47 - 59" refId="ref9135" refString="Jordana, R. &amp; Asiain, A. (1981) Estudio de Friesea en Navarra, con description de una nueva forma dentro del grupo, Friesea subterranea. Publicaciones de Biologia de la Universidad de Navarra, Serie Zoologica, 6, 47 - 59." type="journal article" year="1981">Jordana &amp; Asiain, 1981</bibRefCitation>
</taxonomicName>
(
<collectingCountry box="[1073,1141,583,609]" name="Spain" pageId="3" pageNumber="336">Spain</collectingCountry>
,
<collectingCountry box="[1153,1252,583,609]" name="Portugal" pageId="3" pageNumber="336">Portugal</collectingCountry>
),
<taxonomicName authority="Cassagnau, 1958" authorityName="Cassagnau" authorityYear="1958" class="Collembola" family="Neanuridae" genus="Friesea" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="subterranea">
<emphasis box="[1272,1436,583,609]" italics="true" pageId="3" pageNumber="336">F. subterranea</emphasis>
<bibRefCitation author="Cassagnau, P." box="[151,340,618,645]" pageId="3" pageNumber="336" pagination="17 - 29" refId="ref8661" refString="Cassagnau, P. (1958) Les especes europeennes du genre Friesea (Collemboles Poduromorphes). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 93, 17 - 29." type="journal article" year="1958">Cassagnau, 1958</bibRefCitation>
</taxonomicName>
(
<collectingCountry box="[354,435,619,645]" name="France" pageId="3" pageNumber="336">France</collectingCountry>
,
<collectingCountry box="[446,513,619,645]" name="Spain" pageId="3" pageNumber="336">Spain</collectingCountry>
,
<collectingCountry box="[523,620,619,645]" name="Portugal" pageId="3" pageNumber="336">Portugal</collectingCountry>
) and
<taxonomicName authority="Cassagnau, 1958" authorityName="Cassagnau" authorityYear="1958" box="[680,999,618,645]" class="Collembola" family="Neanuridae" genus="Friesea" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="tolosana">
<emphasis box="[680,803,619,645]" italics="true" pageId="3" pageNumber="336">F. tolosana</emphasis>
<bibRefCitation author="Cassagnau, P." box="[810,999,618,645]" pageId="3" pageNumber="336" pagination="17 - 29" refId="ref8661" refString="Cassagnau, P. (1958) Les especes europeennes du genre Friesea (Collemboles Poduromorphes). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 93, 17 - 29." type="journal article" year="1958">Cassagnau, 1958</bibRefCitation>
</taxonomicName>
(
<collectingCountry box="[1013,1094,619,645]" name="France" pageId="3" pageNumber="336">France</collectingCountry>
,
<collectingCountry box="[1104,1171,619,645]" name="Spain" pageId="3" pageNumber="336">Spain</collectingCountry>
,
<collectingCountry box="[1181,1280,619,645]" name="Portugal" pageId="3" pageNumber="336">Portugal</collectingCountry>
), in which the number of the anal spines varies from 4 to 6. Nevertheless, they differ from
<taxonomicName authority="Smolis &amp; Skarżyński &amp; Gwiazdowicz, 2023" authorityName="Smolis &amp; Skarżyński &amp; Gwiazdowicz" authorityYear="2023" box="[1001,1107,655,681]" class="Collembola" family="Neanuridae" genus="Friesea" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="336" phylum="Arthropoda" rank="species" species="schulzi" status="sp. nov.">
<emphasis box="[1001,1107,655,681]" italics="true" pageId="3" pageNumber="336">F. schulzi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1113,1203,655,681]" pageId="3" pageNumber="336">
<taxonomicNameLabel box="[1113,1203,655,681]" pageId="3" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in the number of the eyes. The first two species mentioned above have 2 + 2 ocelli, the penultimate 1 + 1, and the last one is blind.
</paragraph>
</subSubSection>
</treatment>
</document>