387 lines
46 KiB
XML
387 lines
46 KiB
XML
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<mods:title id="DBFCD60194B22E8604711FD9C918FE21">Further review of Orthochirus Karsch, 1892 (Scorpiones: Buthidae) from Asia: taxonomic position of O. melanurus, O. persa, O. scrobiculosus, and description of six new species</mods:title>
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<mods:namePart id="7C54700483B7E4FD17B39DA1989373DA">Kovařík, FrantIšek</mods:namePart>
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<mods:namePart id="BE2D13ECF65CD7FD85C039BE7F5A7ECD">Fet, Victor</mods:namePart>
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<mods:namePart id="D10177971075C53ADF7C81E905B85AA6">Yağmur, Ersen Aydın</mods:namePart>
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<mods:date id="0EAA47BC14991A0DA4549EEED40BD932">2020</mods:date>
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(
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<figureCitation id="429FAA98E836FFABFEC0FBA7FE6EFBBB" box="[299,442,1107,1132]" captionStart-0="Figures 3–4" captionStart-1="Figures 5–20" captionStart-2="Figures 21–22" captionStart-3="Figures 23–28" captionStartId-0="6.[115,192,2005,2027]" captionStartId-1="7.[116,193,1907,1929]" captionStartId-2="8.[115,192,1979,2001]" captionStartId-3="9.[115,192,1970,1992]" captionTargetBox-0="[129,1464,334,1892]" captionTargetBox-1="[116,1472,247,1897]" captionTargetBox-2="[115,1476,263,1877]" captionTargetBox-3="[123,1474,267,1924]" captionTargetId-0="figure-39@6.[113,1474,306,1942]" captionTargetId-1="figure-106@7.[116,1477,234,1897]" captionTargetId-2="figure-41@8.[115,1476,253,1877]" captionTargetId-3="figure-47@9.[113,1474,257,1945]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="Figures 3–4. Orthochirus birulai sp. n., holotype female, dorsal (3) and ventral (4) views. Scale bar: 10 mm." captionText-1="Figures 5–20: Orthochirus birulai sp. n., holotype female. Figures 5–14. Segments of pedipalps. Pedipalp chela, dorsal (5), external (6), and ventral (7) views. Pedipalp patella, dorsal (8), external (9), and ventral (10) views. Pedipalp femur and trochanter, dorsal (11), internal (12), and ventral (13) views. Pedipalp chela, movable finger (14) dentate margins. The trichobothrial pattern is indicated in Figures 6–9, 11–12 (white circles). Figures 15–16. Carapace and tergites I–IV (15), and sternopectinal region and sternites (16). Figures 17–20. Right legs I–IV, retrolateral aspect." captionText-2="Figures 21–22. Orthochirus birulai sp. n., holotype female, carapace and tergites (21), and sternopectinal region and sternites (22) under UV light." captionText-3="Figures 23–28. Orthochirus birulai sp. n., holotype female, metasoma and telson, lateral (23, 26), dorsal (24, 27) and ventral (25, 28), under white light (23–25) and UV light (26–28). Scale bar: 10 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5741622" figureDoi-1="http://doi.org/10.5281/zenodo.5741624" figureDoi-2="http://doi.org/10.5281/zenodo.5741626" figureDoi-3="http://doi.org/10.5281/zenodo.5741628" httpUri-0="https://zenodo.org/record/5741622/files/figure.png" httpUri-1="https://zenodo.org/record/5741624/files/figure.png" httpUri-2="https://zenodo.org/record/5741626/files/figure.png" httpUri-3="https://zenodo.org/record/5741628/files/figure.png" pageId="5" pageNumber="4">Figures 3–28</figureCitation>
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,
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<figureCitation id="429FAA98E836FFABFE2DFBA0FE24FBBB" box="[454,496,1108,1132]" captionStart="Figure 343" captionStartId="66.[115,186,1950,1972]" captionTargetBox="[116,1475,234,1817]" captionTargetId="figure-15@66.[116,1477,234,1832]" captionTargetPageId="66" captionText="Figure 343. Geographic distribution of the Orthochirus spp. found from Turkey to India. An approximate area where the lectotype of O. melanurus was collected is marked by red ellipse." figureDoi="http://doi.org/10.5281/zenodo.5741722" httpUri="https://zenodo.org/record/5741722/files/figure.png" pageId="5" pageNumber="4">343</figureCitation>
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,
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<tableCitation id="972683A6E836FFABFE10FBA7FD9CFBBB" box="[507,584,1107,1132]" captionStart="Table 1" captionStartId="5.[113,169,985,1006]" captionText="Table 1. Comparative measurements of Orthochirus birulai sp. n., O. kryzhanovskyi sp. n., and O. formozovi sp. n. types from type locality. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="5" pageNumber="4">Table 1</tableCitation>
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)
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||
,
|
||
<collectionCode id="BCB52ED8E836FFABFF01FA60FEF9FA7B" box="[234,301,1428,1452]" pageId="5" pageNumber="4">FKCP</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92BEE596E836FFABFF99FA20FD93F99C" pageId="5" pageNumber="4" type="etymology">
|
||
<paragraph id="DA1BB61DE836FFABFF99FA20FD93F99C" blockId="5.[114,772,1491,1611]" pageId="5" pageNumber="4">ETYMOLOGY. The specific epithet is a patronym honoring Alexei Andreevich Byalynitskii-Birulya (or A. A. Birula) (1864–1937), a renowned Russian zoologist and arachnologist who specialized in scorpions and solpugids.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92BEE596E836FFABFF99F980FB2FFA9F" pageId="5" pageNumber="4" type="diagnosis">
|
||
<paragraph id="DA1BB61DE836FFABFF99F980FB2FFA9F" blockId="5.[114,773,1651,2027]" lastBlockId="5.[817,1476,1072,2024]" pageId="5" pageNumber="4">
|
||
DIAGNOSIS (
|
||
<specimenCount id="CCA27D94E836FFABFF11F980FED8F95B" box="[250,268,1652,1676]" pageId="5" pageNumber="4" type="female">♀</specimenCount>
|
||
). Total length
|
||
<quantity id="1D5C1BF8E836FFABFE53F987FDC8F95C" box="[440,540,1651,1675]" metricMagnitude="-2" metricUnit="m" metricValue="4.05" pageId="5" pageNumber="4" unit="mm" value="40.5">40.5 mm</quantity>
|
||
of female
|
||
<typeStatus id="051F08BFE836FFABFD4BF987FCD5F95C" box="[672,769,1651,1675]" pageId="5" pageNumber="4" type="holotype">holotype</typeStatus>
|
||
.
|
||
<materialsCitation id="6ACCBC40E836FFABFF99F967FAC3FAFF" collectionCode="V, III, IV" collectorName="Trichobothrium & Pectinal & Movable & Dorsal & Pedipalp & Metasoma & Metasoma I & Ventral & Tergites & Sternite & Moderate & Tarsomere I & Ratio" pageId="5" pageNumber="4" specimenCount="1" typeStatus="holotype">
|
||
<collectorName id="7751D3CBE836FFABFF99F967FECEF97C" box="[114,282,1683,1707]" pageId="5" pageNumber="4">Trichobothrium</collectorName>
|
||
<emphasis id="E8D06A0FE836FFABFEC0F967FEECF97C" box="[299,312,1683,1707]" italics="true" pageId="5" pageNumber="4">d</emphasis>
|
||
2 on dorsal surface of pedipalp femur present.
|
||
<collectorName id="7751D3CBE836FFABFF24F947FEF1F91C" box="[207,293,1715,1739]" pageId="5" pageNumber="4">Pectinal</collectorName>
|
||
teeth number
|
||
<quantity id="1D5C1BF8E836FFABFE2FF940FE29F91C" box="[452,509,1715,1740]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" pageId="5" pageNumber="4" unit="in" value="22.0">22 in</quantity>
|
||
female.
|
||
<collectorName id="7751D3CBE836FFABFDB5F947FD69F91C" box="[606,701,1715,1739]" pageId="5" pageNumber="4">Movable</collectorName>
|
||
finger of pedipalps with 9 rows of denticles, 10 ID and 9–10 OD.
|
||
<collectorName id="7751D3CBE836FFABFF99F907FF6DF8DC" box="[114,185,1779,1803]" pageId="5" pageNumber="4">Dorsal</collectorName>
|
||
carinae on pedipalp patella developed and smooth.
|
||
<collectorName id="7751D3CBE836FFABFF99F8E7FF1BF8FC" box="[114,207,1811,1835]" pageId="5" pageNumber="4">Pedipalp</collectorName>
|
||
femur dorsal granulated.
|
||
<collectorName id="7751D3CBE836FFABFE0AF8E0FD84F8FC" box="[481,592,1812,1835]" pageId="5" pageNumber="4">Metasoma</collectorName>
|
||
<collectionCode id="BCB52ED8E836FFABFDBDF8E0FDBDF8FC" box="[598,617,1812,1835]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="4" type="Museum">V</collectionCode>
|
||
dorsal surface mesially with several fine granules only.
|
||
<collectorName id="7751D3CBE836FFABFDC6F8C0FD78F89C" box="[557,684,1844,1867]" pageId="5" pageNumber="4">Metasoma I</collectorName>
|
||
–II with 10 carinae, metasoma
|
||
<collectionCode id="BCB52ED8E836FFABFE99F8A0FE59F8BC" box="[370,397,1876,1899]" pageId="5" pageNumber="4">III</collectionCode>
|
||
with 8 carinae, metasoma
|
||
<collectionCode id="BCB52ED8E836FFABFD2CF8A0FD37F8BC" box="[711,739,1876,1899]" pageId="5" pageNumber="4">IV</collectionCode>
|
||
–
|
||
<collectionCode id="BCB52ED8E836FFABFD1AF8A0FCD0F8BC" box="[753,772,1876,1899]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="4" type="Museum">V</collectionCode>
|
||
with 2 dorsolateral carinae; incomplete ventrolateral carinae present on metasoma
|
||
<collectionCode id="BCB52ED8E836FFABFE83F860FEAFF87C" box="[360,379,1940,1963]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="4" type="Museum">V</collectionCode>
|
||
.
|
||
<collectorName id="7751D3CBE836FFABFE62F867FE02F87C" box="[393,470,1939,1963]" pageId="5" pageNumber="4">Ventral</collectorName>
|
||
carinae of metasoma I–
|
||
<collectionCode id="BCB52ED8E836FFABFD02F860FCD0F87C" box="[745,772,1940,1963]" pageId="5" pageNumber="4">III</collectionCode>
|
||
consist of small granules irregularly in wide row.
|
||
<collectorName id="7751D3CBE836FFABFD7EF840FCD0F81C" box="[661,772,1972,1995]" pageId="5" pageNumber="4">Metasoma</collectorName>
|
||
<collectionCode id="BCB52ED8E836FFABFF99F820FF5AF83C" box="[114,142,2004,2027]" pageId="5" pageNumber="4">IV</collectionCode>
|
||
–
|
||
<collectionCode id="BCB52ED8E836FFABFF70F820FF7AF83C" box="[155,174,2004,2027]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="4" type="Museum">V</collectionCode>
|
||
ventrally and laterally with fine punctation developed, spaces among punctae smooth ventral and granulated lateral; metasoma I–
|
||
<collectionCode id="BCB52ED8E836FFABFC50FBA4FC02FBB0" box="[955,982,1104,1127]" pageId="5" pageNumber="4">III</collectionCode>
|
||
ventrally and laterally granulated and bumpy with punctation reduced.
|
||
<collectorName id="7751D3CBE836FFABFBD7FB84FB45FB5F" box="[1084,1169,1136,1160]" pageId="5" pageNumber="4">Tergites</collectorName>
|
||
roughly to finely granulated.
|
||
<collectorName id="7751D3CBE836FFABFCDAFB64FC50FB7F" box="[817,900,1168,1192]" pageId="5" pageNumber="4">Sternite</collectorName>
|
||
VII densely granulated, with four granulated carinae present.
|
||
<collectorName id="7751D3CBE836FFABFC64FB44FC3BFB1F" box="[911,1007,1200,1224]" pageId="5" pageNumber="4">Pedipalp</collectorName>
|
||
, metasoma and telson glabrous.
|
||
<collectorName id="7751D3CBE836FFABFAB7FB44FA16FB1F" box="[1372,1474,1200,1224]" pageId="5" pageNumber="4">Moderate</collectorName>
|
||
to strong tibial spurs present on legs
|
||
<collectionCode id="BCB52ED8E836FFABFB38FB24FB3AFB30" box="[1235,1262,1232,1255]" pageId="5" pageNumber="4">III</collectionCode>
|
||
and
|
||
<collectionCode id="BCB52ED8E836FFABFAC2FB24FA91FB30" box="[1321,1349,1232,1255]" pageId="5" pageNumber="4">IV</collectionCode>
|
||
.
|
||
<collectorName id="7751D3CBE836FFABFAB9FB24FCEEFAD0" pageId="5" pageNumber="4">Tarsomere I</collectorName>
|
||
of legs I–
|
||
<collectionCode id="BCB52ED8E836FFABFC46FB04FC1CFAD0" box="[941,968,1264,1287]" pageId="5" pageNumber="4">III</collectionCode>
|
||
with 8–10 long setae in female.
|
||
<collectorName id="7751D3CBE836FFABFADDFB04FAA4FADF" box="[1334,1392,1264,1288]" pageId="5" pageNumber="4">Ratio</collectorName>
|
||
length/ width of metasoma
|
||
<collectionCode id="BCB52ED8E836FFABFBEAFAE4FBC0FAF0" box="[1025,1044,1296,1319]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="4" type="Museum">V</collectionCode>
|
||
<quantity id="1D5C1BF8E836FFABFBF2FAE4FBB7FAFF" box="[1049,1123,1296,1320]" metricMagnitude="-2" metricUnit="m" metricValue="2.9718" pageId="5" pageNumber="4" unit="in" value="1.17">1.17 in</quantity>
|
||
female
|
||
<typeStatus id="051F08BFE836FFABFB5DFAE4FAC3FAFF" box="[1206,1303,1296,1320]" pageId="5" pageNumber="4" type="holotype">holotype</typeStatus>
|
||
</materialsCitation>
|
||
. Pedipalp femur length/width ratio
|
||
<quantity id="1D5C1BF8E836FFABFC1DFAC4FB94FA9F" box="[1014,1088,1328,1352]" metricMagnitude="-2" metricUnit="m" metricValue="7.5438" pageId="5" pageNumber="4" unit="in" value="2.97">2.97 in</quantity>
|
||
female
|
||
<typeStatus id="051F08BFE836FFABFB7DFAC4FB23FA9F" box="[1174,1271,1328,1352]" pageId="5" pageNumber="4" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92BEE596E836FFA4FCDAFAA4FE31FAB4" lastPageId="10" lastPageNumber="9" pageId="5" pageNumber="4" type="description">
|
||
<paragraph id="DA1BB61DE836FFABFCDAFAA4FB9EFA1F" blockId="5.[817,1476,1072,2024]" pageId="5" pageNumber="4">
|
||
DESCRIPTION. Total length of adult female
|
||
<quantity id="1D5C1BF8E836FFABFAA4FAA4FA6BFAB0" box="[1359,1471,1360,1384]" metricMagnitude="-2" metricUnit="m" metricValue="4.05" pageId="5" pageNumber="4" unit="mm" value="40.5">40.5 mm</quantity>
|
||
. Measurements of the carapace, telson, segments of the metasoma and segments of the pedipalps are given in
|
||
<tableCitation id="972683A6E836FFABFA86FA64FA69FA7F" box="[1389,1469,1424,1448]" captionStart="Table 1" captionStartId="5.[113,169,985,1006]" captionText="Table 1. Comparative measurements of Orthochirus birulai sp. n., O. kryzhanovskyi sp. n., and O. formozovi sp. n. types from type locality. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="5" pageNumber="4">Table 1</tableCitation>
|
||
. For habitus, see
|
||
<figureCitation id="429FAA98E836FFABFC0BFA44FB93FA1F" box="[992,1095,1456,1480]" captionStart="Figures 3–4" captionStartId="6.[115,192,2005,2027]" captionTargetBox="[129,1464,334,1892]" captionTargetId="figure-39@6.[113,1474,306,1942]" captionTargetPageId="6" captionText="Figures 3–4. Orthochirus birulai sp. n., holotype female, dorsal (3) and ventral (4) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5741622" httpUri="https://zenodo.org/record/5741622/files/figure.png" pageId="5" pageNumber="4">Figs. 3–4</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="DA1BB61DE836FFABFCDAFA24FAD7F97F" blockId="5.[817,1476,1072,2024]" pageId="5" pageNumber="4">
|
||
<emphasis id="E8D06A0FE836FFABFCDAFA24FC78FA3F" bold="true" box="[817,940,1488,1512]" pageId="5" pageNumber="4">Coloration</emphasis>
|
||
(
|
||
<figureCitation id="429FAA98E836FFABFC2AFA24FBE4FA3F" box="[961,1072,1488,1512]" captionStart="Figures 3–4" captionStartId="6.[115,192,2005,2027]" captionTargetBox="[129,1464,334,1892]" captionTargetId="figure-39@6.[113,1474,306,1942]" captionTargetPageId="6" captionText="Figures 3–4. Orthochirus birulai sp. n., holotype female, dorsal (3) and ventral (4) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5741622" httpUri="https://zenodo.org/record/5741622/files/figure.png" pageId="5" pageNumber="4">Figs. 3–4</figureCitation>
|
||
). Carapace, tergites, and metasoma black. Femur and patella of pedipalps reddish black to black, chela of pedipalps yellowish to reddish brown. Femur and patella of legs reddish black, other segments of legs yellowish brown. Sternite VII black, other sternites reddish black with yellow median area in posterior margin of sternites III–V, mainly on sternite V. Telson reddish brown.
|
||
</paragraph>
|
||
<paragraph id="DA1BB61DE836FFABFCDAF944FB2EF89F" blockId="5.[817,1476,1072,2024]" pageId="5" pageNumber="4">
|
||
<emphasis id="E8D06A0FE836FFABFCDAF944FBEBF91F" bold="true" box="[817,1087,1712,1736]" pageId="5" pageNumber="4">Mesosoma and carapace</emphasis>
|
||
(
|
||
<figureCitation id="429FAA98E836FFABFBA6F944FB1EF91F" box="[1101,1226,1712,1736]" captionStart="Figures 5–20" captionStartId="7.[116,193,1907,1929]" captionTargetBox="[116,1472,247,1897]" captionTargetId="figure-106@7.[116,1477,234,1897]" captionTargetPageId="7" captionText="Figures 5–20: Orthochirus birulai sp. n., holotype female. Figures 5–14. Segments of pedipalps. Pedipalp chela, dorsal (5), external (6), and ventral (7) views. Pedipalp patella, dorsal (8), external (9), and ventral (10) views. Pedipalp femur and trochanter, dorsal (11), internal (12), and ventral (13) views. Pedipalp chela, movable finger (14) dentate margins. The trichobothrial pattern is indicated in Figures 6–9, 11–12 (white circles). Figures 15–16. Carapace and tergites I–IV (15), and sternopectinal region and sternites (16). Figures 17–20. Right legs I–IV, retrolateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5741624" httpUri="https://zenodo.org/record/5741624/files/figure.png" pageId="5" pageNumber="4">Figs. 15–16</figureCitation>
|
||
,
|
||
<figureCitation id="429FAA98E836FFABFB3EF944FACDF91F" box="[1237,1305,1712,1736]" captionStart="Figures 21–22" captionStartId="8.[115,192,1979,2001]" captionTargetBox="[115,1476,263,1877]" captionTargetId="figure-41@8.[115,1476,253,1877]" captionTargetPageId="8" captionText="Figures 21–22. Orthochirus birulai sp. n., holotype female, carapace and tergites (21), and sternopectinal region and sternites (22) under UV light." figureDoi="http://doi.org/10.5281/zenodo.5741626" httpUri="https://zenodo.org/record/5741626/files/figure.png" pageId="5" pageNumber="4">21–22</figureCitation>
|
||
). Tergites with a median carina and is roughly to finely granulated.Seventh sternite densely granulated and with four granulated carinae, the other sternites partly granulated but smooth in middle and posteriorly. Pectinal teeth number
|
||
<quantity id="1D5C1BF8E836FFABFBFEF8C4FB9CF89F" box="[1045,1096,1840,1864]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" pageId="5" pageNumber="4" unit="in" value="22.0">22 in</quantity>
|
||
female
|
||
<typeStatus id="051F08BFE836FFABFB71F8C4FB23F89F" box="[1178,1271,1840,1864]" pageId="5" pageNumber="4" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="DA1BB61DE836FFA4FCDAF8A4FF26FDF4" blockId="5.[817,1476,1072,2024]" lastBlockId="10.[113,772,235,1059]" lastPageId="10" lastPageNumber="9" pageId="5" pageNumber="4">
|
||
<emphasis id="E8D06A0FE836FFABFCDAF8A4FBFAF8BF" bold="true" box="[817,1070,1872,1896]" pageId="5" pageNumber="4">Metasoma and telson</emphasis>
|
||
(
|
||
<figureCitation id="429FAA98E836FFABFBA8F8A4FB1FF8BF" box="[1091,1227,1872,1896]" captionStart="Figures 23–28" captionStartId="9.[115,192,1970,1992]" captionTargetBox="[123,1474,267,1924]" captionTargetId="figure-47@9.[113,1474,257,1945]" captionTargetPageId="9" captionText="Figures 23–28. Orthochirus birulai sp. n., holotype female, metasoma and telson, lateral (23, 26), dorsal (24, 27) and ventral (25, 28), under white light (23–25) and UV light (26–28). Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5741628" httpUri="https://zenodo.org/record/5741628/files/figure.png" pageId="5" pageNumber="4">Figs. 23–28</figureCitation>
|
||
). Metasoma I–II with 10 granulated carinae. Metasoma III–IV lacks lateral and ventromedian carinae. Ventrolateral carinae are developed on metasoma I–III and indicated on metasoma IV–V, dorsolateral carinae are present on all metasomal segments. Ventral carinae of metasoma I–III consist of small granules irregularly in wide row. Metasoma I–IV and partly metasoma V are granulated laterally, metasoma III–V laterally punctate; granulation absent on dorsal surfaces of all metasomal segments mesially except several fine granules on metasoma; several large granules are present on metasoma I–IV rather laterally, metasoma I with ca. 10 of these granules. Fine punctation on metasoma IV–V ventrally developed, spaces among punctae smooth. Entire metasoma and telson glabrous. Telson without punctation and granulation.
|
||
</paragraph>
|
||
<caption id="8EDBE695E835FFA8FF98F821FB5FF83C" ID-DOI="http://doi.org/10.5281/zenodo.5741622" ID-Zenodo-Dep="5741622" box="[115,1163,2005,2027]" httpUri="https://zenodo.org/record/5741622/files/figure.png" pageId="6" pageNumber="5" startId="6.[115,192,2005,2027]" targetBox="[129,1464,334,1892]" targetPageId="6">
|
||
<paragraph id="DA1BB61DE835FFA8FF98F821FB5FF83C" blockId="6.[115,1163,2005,2027]" box="[115,1163,2005,2027]" pageId="6" pageNumber="5">
|
||
<emphasis id="E8D06A0FE835FFA8FF98F821FF3EF83D" bold="true" box="[115,234,2005,2027]" pageId="6" pageNumber="5">Figures 3–4</emphasis>
|
||
.
|
||
<taxonomicName id="1DA4CD9EE835FFA8FF1DF821FE66F83D" authority="Kovařík & Fet & Yağmur, 2020" authorityName="Kovařík & Fet & Yağmur" authorityYear="2020" box="[246,434,2005,2026]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="birulai" status="sp. nov.">
|
||
<emphasis id="E8D06A0FE835FFA8FF1DF821FE66F83D" box="[246,434,2005,2026]" italics="true" pageId="6" pageNumber="5">Orthochirus birulai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F3E3D774E835FFA8FE53F821FE39F83C" box="[440,493,2005,2027]" pageId="6" pageNumber="5" rank="species">
|
||
<emphasis id="E8D06A0FE835FFA8FE53F821FE1BF83D" bold="true" box="[440,463,2005,2026]" pageId="6" pageNumber="5">sp</emphasis>
|
||
.
|
||
<emphasis id="E8D06A0FE835FFA8FE30F821FE3CF83D" bold="true" box="[475,488,2005,2026]" pageId="6" pageNumber="5">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, holotype female, dorsal (3) and ventral (4) views. Scale bar: 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="8EDBE695E834FFA9FF9FF887FEF2F82C" ID-DOI="http://doi.org/10.5281/zenodo.5741624" ID-Zenodo-Dep="5741624" httpUri="https://zenodo.org/record/5741624/files/figure.png" pageId="7" pageNumber="6" startId="7.[116,193,1907,1929]" targetBox="[116,1472,247,1897]" targetPageId="7">
|
||
<paragraph id="DA1BB61DE834FFA9FF9FF887FEF2F82C" blockId="7.[116,1475,1907,2044]" pageId="7" pageNumber="6">
|
||
<emphasis id="E8D06A0FE834FFA9FF9FF887FF23F85F" bold="true" box="[116,247,1907,1929]" pageId="7" pageNumber="6">Figures 5–20</emphasis>
|
||
:
|
||
<taxonomicName id="1DA4CD9EE834FFA9FEE8F887FE14F85F" authority="Kovařík & Fet & Yağmur, 2020" authorityName="Kovařík & Fet & Yağmur" authorityYear="2020" box="[259,448,1907,1928]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="birulai" status="sp. nov.">
|
||
<emphasis id="E8D06A0FE834FFA9FEE8F887FE14F85F" box="[259,448,1907,1928]" italics="true" pageId="7" pageNumber="6">Orthochirus birulai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F3E3D774E834FFA9FE2DF887FE2FF85F" box="[454,507,1907,1928]" pageId="7" pageNumber="6" rank="species">
|
||
<emphasis id="E8D06A0FE834FFA9FE2DF887FE09F85F" bold="true" box="[454,477,1907,1928]" pageId="7" pageNumber="6">sp</emphasis>
|
||
.
|
||
<emphasis id="E8D06A0FE834FFA9FE02F887FE22F85F" bold="true" box="[489,502,1907,1928]" pageId="7" pageNumber="6">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, holotype female.
|
||
<emphasis id="E8D06A0FE834FFA9FD44F887FCE6F85F" bold="true" box="[687,818,1907,1929]" pageId="7" pageNumber="6">Figures 5–14</emphasis>
|
||
. Segments of pedipalps. Pedipalp chela, dorsal (5), external (6), and ventral (7) views. Pedipalp patella, dorsal (8), external (9), and ventral (10) views. Pedipalp femur and trochanter, dorsal (11), internal (12), and ventral (13) views. Pedipalp chela, movable finger (14) dentate margins. The trichobothrial pattern is indicated in Figures 6–9, 11–12 (white circles).
|
||
<emphasis id="E8D06A0FE834FFA9FEE1F83DFE4DF809" bold="true" box="[266,409,1993,2015]" pageId="7" pageNumber="6">Figures 15–16</emphasis>
|
||
. Carapace and tergites I–IV (15), and sternopectinal region and sternites (16).
|
||
<emphasis id="E8D06A0FE834FFA9FB65F83DFAC9F809" bold="true" box="[1166,1309,1993,2015]" pageId="7" pageNumber="6">Figures 17–20</emphasis>
|
||
. Right legs I–IV, retrolateral aspect.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="8EDBE695E83BFFA6FF98F84FFF72F839" ID-DOI="http://doi.org/10.5281/zenodo.5741626" ID-Zenodo-Dep="5741626" httpUri="https://zenodo.org/record/5741626/files/figure.png" pageId="8" pageNumber="7" startId="8.[115,192,1979,2001]" targetBox="[115,1476,263,1877]" targetPageId="8">
|
||
<paragraph id="DA1BB61DE83BFFA6FF98F84FFF72F839" blockId="8.[115,1473,1979,2030]" pageId="8" pageNumber="7">
|
||
<emphasis id="E8D06A0FE83BFFA6FF98F84FFED6F807" bold="true" box="[115,258,1979,2001]" pageId="8" pageNumber="7">Figures 21–22</emphasis>
|
||
.
|
||
<taxonomicName id="1DA4CD9EE83BFFA6FEE5F84FFE1EF807" authority="Kovařík & Fet & Yağmur, 2020" authorityName="Kovařík & Fet & Yağmur" authorityYear="2020" box="[270,458,1979,2000]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="birulai" status="sp. nov.">
|
||
<emphasis id="E8D06A0FE83BFFA6FEE5F84FFE1EF807" box="[270,458,1979,2000]" italics="true" pageId="8" pageNumber="7">Orthochirus birulai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F3E3D774E83BFFA6FE3BF84FFDD1F807" box="[464,517,1979,2000]" pageId="8" pageNumber="7" rank="species">
|
||
<emphasis id="E8D06A0FE83BFFA6FE3BF84FFE33F807" bold="true" box="[464,487,1979,2000]" pageId="8" pageNumber="7">sp</emphasis>
|
||
.
|
||
<emphasis id="E8D06A0FE83BFFA6FE18F84FFDD4F807" bold="true" box="[499,512,1979,2000]" pageId="8" pageNumber="7">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, holotype female, carapace and tergites (21), and sternopectinal region and sternites (22) under UV light.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="8EDBE695E83AFFA7FF98F846FD69F832" ID-DOI="http://doi.org/10.5281/zenodo.5741628" ID-Zenodo-Dep="5741628" httpUri="https://zenodo.org/record/5741628/files/figure.png" pageId="9" pageNumber="8" startId="9.[115,192,1970,1992]" targetBox="[123,1474,267,1924]" targetPageId="9">
|
||
<paragraph id="DA1BB61DE83AFFA7FF98F846FD69F832" blockId="9.[115,1473,1970,2021]" pageId="9" pageNumber="8">
|
||
<emphasis id="E8D06A0FE83AFFA7FF98F846FED6F81F" bold="true" box="[115,258,1970,1992]" pageId="9" pageNumber="8">Figures 23–28</emphasis>
|
||
.
|
||
<taxonomicName id="1DA4CD9EE83AFFA7FEE5F847FE1FF81F" authority="Kovařík & Fet & Yağmur, 2020" authorityName="Kovařík & Fet & Yağmur" authorityYear="2020" box="[270,459,1971,1992]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="birulai" status="sp. nov.">
|
||
<emphasis id="E8D06A0FE83AFFA7FEE5F847FE1FF81F" box="[270,459,1971,1992]" italics="true" pageId="9" pageNumber="8">Orthochirus birulai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F3E3D774E83AFFA7FE3AF847FDD3F81F" box="[465,519,1971,1992]" pageId="9" pageNumber="8" rank="species">
|
||
<emphasis id="E8D06A0FE83AFFA7FE3AF847FE3CF81F" bold="true" box="[465,488,1971,1992]" pageId="9" pageNumber="8">sp</emphasis>
|
||
.
|
||
<emphasis id="E8D06A0FE83AFFA7FE1FF847FDD5F81F" bold="true" box="[500,513,1971,1992]" pageId="9" pageNumber="8">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, holotype female, metasoma and telson, lateral (23, 26), dorsal (24, 27) and ventral (25, 28), under white light (23–25) and UV light (26–28). Scale bar: 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DA1BB61DE839FFA4FF9AFDDEFDC4FCF4" blockId="10.[113,772,235,1059]" pageId="10" pageNumber="9">
|
||
<emphasis id="E8D06A0FE839FFA4FF9AFDDEFF34FD95" bold="true" box="[113,224,554,578]" pageId="10" pageNumber="9">Pedipalps</emphasis>
|
||
(
|
||
<figureCitation id="429FAA98E839FFA4FF1AFDDFFEBCFD95" box="[241,360,554,579]" captionStart="Figures 5–20" captionStartId="7.[116,193,1907,1929]" captionTargetBox="[116,1472,247,1897]" captionTargetId="figure-106@7.[116,1477,234,1897]" captionTargetPageId="7" captionText="Figures 5–20: Orthochirus birulai sp. n., holotype female. Figures 5–14. Segments of pedipalps. Pedipalp chela, dorsal (5), external (6), and ventral (7) views. Pedipalp patella, dorsal (8), external (9), and ventral (10) views. Pedipalp femur and trochanter, dorsal (11), internal (12), and ventral (13) views. Pedipalp chela, movable finger (14) dentate margins. The trichobothrial pattern is indicated in Figures 6–9, 11–12 (white circles). Figures 15–16. Carapace and tergites I–IV (15), and sternopectinal region and sternites (16). Figures 17–20. Right legs I–IV, retrolateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5741624" httpUri="https://zenodo.org/record/5741624/files/figure.png" pageId="10" pageNumber="9">Figs. 5–14</figureCitation>
|
||
). Trichobothrium
|
||
<emphasis id="E8D06A0FE839FFA4FDC7FDDFFDEDFD94" box="[556,569,555,579]" italics="true" pageId="10" pageNumber="9">d</emphasis>
|
||
<subScript id="4620B458E839FFA4FDD2FDCFFD95FD9E" attach="left" box="[569,577,571,585]" fontSize="6" pageId="10" pageNumber="9">2</subScript>
|
||
on dorsal surface of pedipalp femur present; trichobothrium
|
||
<emphasis id="E8D06A0FE839FFA4FDB8FDBFFD8BFDB5" box="[595,607,587,610]" italics="true" pageId="10" pageNumber="9">e</emphasis>
|
||
<subScript id="4620B458E839FFA4FDB4FDAFFDB3FDBE" attach="left" box="[607,615,603,617]" fontSize="6" pageId="10" pageNumber="9">1</subScript>
|
||
is situated at level with
|
||
<emphasis id="E8D06A0FE839FFA4FF09FD9FFF3BFD54" box="[226,239,619,643]" italics="true" pageId="10" pageNumber="9">d</emphasis>
|
||
<subScript id="4620B458E839FFA4FF04FD8FFF23FD5E" attach="left" box="[239,247,635,649]" fontSize="6" pageId="10" pageNumber="9">4</subScript>
|
||
. Femur of pedipalps with five granulated carinae and is dorsally granulated. Patella has seven smooth carinae, and the chela has smooth carinae which may be discernible throughout the length of the fixed finger. The entire pedipalps are glabrous, with several setae only. Movable fingers with 9 rows of denticles, 10 ID and 9–10 OD.
|
||
</paragraph>
|
||
<paragraph id="DA1BB61DE839FFA4FF9AFCDFFE79FBD5" blockId="10.[113,772,235,1059]" pageId="10" pageNumber="9">
|
||
<emphasis id="E8D06A0FE839FFA4FF9AFCDFFF72FC94" bold="true" box="[113,166,811,835]" pageId="10" pageNumber="9">Legs</emphasis>
|
||
(
|
||
<figureCitation id="429FAA98E839FFA4FF52FCDFFEEBFC94" box="[185,319,811,835]" captionStart="Figures 5–20" captionStartId="7.[116,193,1907,1929]" captionTargetBox="[116,1472,247,1897]" captionTargetId="figure-106@7.[116,1477,234,1897]" captionTargetPageId="7" captionText="Figures 5–20: Orthochirus birulai sp. n., holotype female. Figures 5–14. Segments of pedipalps. Pedipalp chela, dorsal (5), external (6), and ventral (7) views. Pedipalp patella, dorsal (8), external (9), and ventral (10) views. Pedipalp femur and trochanter, dorsal (11), internal (12), and ventral (13) views. Pedipalp chela, movable finger (14) dentate margins. The trichobothrial pattern is indicated in Figures 6–9, 11–12 (white circles). Figures 15–16. Carapace and tergites I–IV (15), and sternopectinal region and sternites (16). Figures 17–20. Right legs I–IV, retrolateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5741624" httpUri="https://zenodo.org/record/5741624/files/figure.png" pageId="10" pageNumber="9">Figs. 17–20</figureCitation>
|
||
). Moderate to strong tibial spurs present on legs III and IV. Femur with four partly granulated carinae; patella with five rather smooth carinae; tibia smooth. Patella with only a few setae. Tibia with setae on the outer side of legs I–II. Tarsomere I of legs I–III with 8–10 long setae, legs IV with 0–2 setae. Tarsomeres I–II of all legs internal with two rather irregular rows of setae.
|
||
</paragraph>
|
||
<paragraph id="DA1BB61DE839FFA4FF9AFBFFFE7CFBF4" blockId="10.[113,772,235,1059]" box="[113,424,1035,1059]" pageId="10" pageNumber="9">
|
||
<emphasis id="E8D06A0FE839FFA4FF9AFBFFFECCFBF4" bold="true" box="[113,280,1035,1059]" pageId="10" pageNumber="9">Measurements</emphasis>
|
||
. See
|
||
<tableCitation id="972683A6E839FFA4FEB9FBFFFE76FBF4" box="[338,418,1035,1059]" captionStart="Table 1" captionStartId="5.[113,169,985,1006]" captionText="Table 1. Comparative measurements of Orthochirus birulai sp. n., O. kryzhanovskyi sp. n., and O. formozovi sp. n. types from type locality. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="10" pageNumber="9">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="DA1BB61DE839FFA4FF9AFBBFFE94FAF5" blockId="10.[113,771,1099,1315]" pageId="10" pageNumber="9">
|
||
AFFINITIES. The combination of six characters (metasoma V dorsal mesially with several fine granules only; entire metasoma glabrous; punctation on the ventral surfaces of metasoma IV–V developed; lateral surfaces of metasoma I–V granulated; sternite VII densely granulated; and pedipalp movable finger with 9–10 ID and OD) is unique in the entire genus
|
||
<taxonomicName id="1DA4CD9EE839FFA4FF5DFAFFFEECFAF4" authorityName=": Fet & Lowe" authorityYear="2000" box="[182,312,1291,1315]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E8D06A0FE839FFA4FF5DFAFFFEECFAF4" box="[182,312,1291,1315]" italics="true" pageId="10" pageNumber="9">Orthochirus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="DA1BB61DE839FFA4FF9AFABFFE31FAB4" blockId="10.[113,485,1355,1379]" box="[113,485,1355,1379]" pageId="10" pageNumber="9">
|
||
DISTRIBUTION.
|
||
<collectingCountry id="A2B3F68DE839FFA4FEF9FABFFEB8FAB4" box="[274,364,1355,1379]" name="Pakistan" pageId="10" pageNumber="9">Pakistan</collectingCountry>
|
||
(
|
||
<figureCitation id="429FAA98E839FFA4FE90FABFFE0DFAB4" box="[379,473,1355,1379]" captionStart="Figure 343" captionStartId="66.[115,186,1950,1972]" captionTargetBox="[116,1475,234,1817]" captionTargetId="figure-15@66.[116,1477,234,1832]" captionTargetPageId="66" captionText="Figure 343. Geographic distribution of the Orthochirus spp. found from Turkey to India. An approximate area where the lectotype of O. melanurus was collected is marked by red ellipse." figureDoi="http://doi.org/10.5281/zenodo.5741722" httpUri="https://zenodo.org/record/5741722/files/figure.png" pageId="10" pageNumber="9">Fig. 343</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |