351 lines
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XML
351 lines
43 KiB
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<mods:title id="D333F743CC202ED9AA69D78B094C7F36">Further review of Orthochirus Karsch, 1892 (Scorpiones: Buthidae) from Asia: taxonomic position of O. melanurus, O. persa, O. scrobiculosus, and description of six new species</mods:title>
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<mods:namePart id="2E07444E45529A235AC7D8E163E9B6BA">Kovařík, FrantIšek</mods:namePart>
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<mods:namePart id="99561C89F175EAFE24A74DBD139D4DBD">Fet, Victor</mods:namePart>
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<mods:namePart id="6F085CDCB493BD0574104DD7BC5B67BE">Yağmur, Ersen Aydın</mods:namePart>
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(
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<figureCitation id="429FAA98E821FFBCFC37F9E5FB53F9FE" box="[988,1159,1553,1577]" captionStart-0="Figures 79–80" captionStart-1="Figures 81–97" captionStart-2="Figures 98–99" captionStart-3="Figures 100–105" captionStartId-0="19.[117,194,2006,2028]" captionStartId-1="20.[113,190,1901,1923]" captionStartId-2="21.[116,193,1720,1742]" captionStartId-3="22.[115,192,1983,2005]" captionTargetBox-0="[137,1456,307,1887]" captionTargetBox-1="[115,1470,234,1876]" captionTargetBox-2="[113,1474,232,1713]" captionTargetBox-3="[116,1474,238,1960]" captionTargetId-0="figure-31@19.[114,1475,285,1919]" captionTargetId-1="figure-115@20.[113,1474,234,1883]" captionTargetId-2="figure-185@21.[113,1474,231,1713]" captionTargetId-3="figure-15@22.[113,1474,234,1973]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionTargetPageId-3="22" captionText-0="Figures 79–80. Orthochirus grosseri sp. n., holotype female, dorsal (79) and ventral (80) views. Scale bar: 10 mm." captionText-1="Figures 81–97: Orthochirus grosseri sp. n., holotype female. Figures 81–82. Carapace and tergites I–IV (81), and sternopectinal region and sternites (82). Figures 83–86. Right legs I–IV, retrolateral aspect. Figures 87–97. Segments of pedipalps. Pedipalp chela, dorsal (87), external (88), and ventral (89) views. Pedipalp patella, dorsal (90), external (91), and ventral (92) views. Pedipalp femur and trochanter, internal (93), dorsal (94), and ventral (95) views. Pedipalp chela, movable (96) and fixed (97) fingers dentate margins. The trichobothrial pattern is indicated in Figures 87–91, 93–94 (white circles)." captionText-2="Figures 98–99. Orthochirus grosseri sp. n., holotype female, carapace and tergites (98), and sternopectinal region and sternites (99) under UV light." captionText-3="Figures 100–105. Orthochirus grosseri sp. n., holotype female, metasoma and telson, lateral (100, 103), dorsal (101, 104) and ventral (102, 105), under white light (100–102) and UV light (103–105). Scale bar: 10 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5741644" figureDoi-1="http://doi.org/10.5281/zenodo.5741646" figureDoi-2="http://doi.org/10.5281/zenodo.5741648" figureDoi-3="http://doi.org/10.5281/zenodo.5741650" httpUri-0="https://zenodo.org/record/5741644/files/figure.png" httpUri-1="https://zenodo.org/record/5741646/files/figure.png" httpUri-2="https://zenodo.org/record/5741648/files/figure.png" httpUri-3="https://zenodo.org/record/5741650/files/figure.png" pageId="18" pageNumber="17">Figures 79–105</figureCitation>
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,
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||
</date>
|
||
,
|
||
<specimenCount id="CCA27D94E821FFBCFACCF8A5FA9CF8BE" box="[1319,1352,1873,1897]" pageId="18" pageNumber="17" type="female">1♀</specimenCount>
|
||
(
|
||
<typeStatus id="051F08BFE821FFBCFAB1F8A5FA6FF8BE" box="[1370,1467,1873,1897]" pageId="18" pageNumber="17" type="holotype">holotype</typeStatus>
|
||
)
|
||
<specimenCount id="CCA27D94E821FFBCFCDBF885FC9DF85E" box="[816,841,1905,1929]" pageId="18" pageNumber="17" type="female">1♀</specimenCount>
|
||
<specimenCount id="CCA27D94E821FFBCFCA2F885FCBDF85E" box="[841,873,1905,1929]" pageId="18" pageNumber="17" type="male">1♂</specimenCount>
|
||
juv. (
|
||
<typeStatus id="051F08BFE821FFBCFC43F885FBC0F85E" box="[936,1044,1905,1929]" pageId="18" pageNumber="17" type="paratype">paratypes</typeStatus>
|
||
), leg.
|
||
<collectorName id="7751D3CBE821FFBCFBB8F886FB18F85E" box="[1107,1228,1905,1929]" pageId="18" pageNumber="17">W. Grosser</collectorName>
|
||
</materialsCitation>
|
||
,
|
||
<collectionCode id="BCB52ED8E821FFBCFB3CF885FACEF85E" box="[1239,1306,1905,1929]" pageId="18" pageNumber="17">FKCP</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92BEE596E821FFBCFCDBF845FC0BF7DE" pageId="18" pageNumber="17" type="etymology">
|
||
<paragraph id="DA1BB61DE821FFBCFCDBF845FC0BF7DE" blockId="18.[816,1474,1969,2057]" pageId="18" pageNumber="17">
|
||
ETYMOLOGY. The species epithet is a patronym honoring a Czech entomologist Walter Grosser, the collector of
|
||
<typeStatus id="051F08BFE821FFBCFA80F826FA77F83D" box="[1387,1443,2002,2026]" pageId="18" pageNumber="17">types</typeStatus>
|
||
of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="8EDBE695E820FFBDFF9EF822FB1EF83C" ID-DOI="http://doi.org/10.5281/zenodo.5741644" ID-Zenodo-Dep="5741644" box="[117,1226,2006,2028]" httpUri="https://zenodo.org/record/5741644/files/figure.png" pageId="19" pageNumber="18" startId="19.[117,194,2006,2028]" targetBox="[137,1456,307,1887]" targetPageId="19">
|
||
<paragraph id="DA1BB61DE820FFBDFF9EF822FB1EF83C" blockId="19.[117,1226,2006,2028]" box="[117,1226,2006,2028]" pageId="19" pageNumber="18">
|
||
<emphasis id="E8D06A0FE820FFBDFF9EF822FED7F83C" bold="true" box="[117,259,2006,2028]" pageId="19" pageNumber="18">Figures 79–80</emphasis>
|
||
.
|
||
<taxonomicName id="1DA4CD9EE820FFBDFEE4F822FE0DF83B" authority="Kovařík & Fet & Yağmur, 2020" authorityName="Kovařík & Fet & Yağmur" authorityYear="2020" box="[271,473,2006,2028]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="grosseri" status="sp. nov.">
|
||
<emphasis id="E8D06A0FE820FFBDFEE4F822FE0DF83B" box="[271,473,2006,2028]" italics="true" pageId="19" pageNumber="18">Orthochirus grosseri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F3E3D774E820FFBDFE34F822FDC0F83C" box="[479,532,2006,2027]" pageId="19" pageNumber="18" rank="species">
|
||
<emphasis id="E8D06A0FE820FFBDFE34F822FE22F83C" bold="true" box="[479,502,2006,2027]" pageId="19" pageNumber="18">sp</emphasis>
|
||
.
|
||
<emphasis id="E8D06A0FE820FFBDFDEAF822FDDAF83C" bold="true" box="[513,526,2006,2027]" pageId="19" pageNumber="18">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, holotype female, dorsal (79) and ventral (80) views. Scale bar: 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="8EDBE695E827FFBAFF9AF899FE27F820" ID-DOI="http://doi.org/10.5281/zenodo.5741646" ID-Zenodo-Dep="5741646" httpUri="https://zenodo.org/record/5741646/files/figure.png" pageId="20" pageNumber="19" startId="20.[113,190,1901,1923]" targetBox="[115,1470,234,1876]" targetPageId="20">
|
||
<paragraph id="DA1BB61DE827FFBAFF9AF899FE27F820" blockId="20.[113,1472,1901,2039]" pageId="20" pageNumber="19">
|
||
<emphasis id="E8D06A0FE827FFBAFF9AF899FED5F855" bold="true" box="[113,257,1901,1923]" pageId="20" pageNumber="19">Figures 81–97</emphasis>
|
||
:
|
||
<taxonomicName id="1DA4CD9EE827FFBAFEE5F899FE0CF854" authority="Kovařík & Fet & Yağmur, 2020" authorityName="Kovařík & Fet & Yağmur" authorityYear="2020" box="[270,472,1901,1923]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="grosseri" status="sp. nov.">
|
||
<emphasis id="E8D06A0FE827FFBAFEE5F899FE0CF854" box="[270,472,1901,1923]" italics="true" pageId="20" pageNumber="19">Orthochirus grosseri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F3E3D774E827FFBAFE35F89AFDC0F854" box="[478,532,1902,1923]" pageId="20" pageNumber="19" rank="species">
|
||
<emphasis id="E8D06A0FE827FFBAFE35F89AFE21F854" bold="true" box="[478,501,1902,1923]" pageId="20" pageNumber="19">sp</emphasis>
|
||
.
|
||
<emphasis id="E8D06A0FE827FFBAFDEAF89AFDDAF854" bold="true" box="[513,526,1902,1923]" pageId="20" pageNumber="19">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, holotype female.
|
||
<emphasis id="E8D06A0FE827FFBAFD22F899FC8DF855" bold="true" box="[713,857,1901,1923]" pageId="20" pageNumber="19">Figures 81–82</emphasis>
|
||
. Carapace and tergites I–IV (81), and sternopectinal region and sternites (82).
|
||
<emphasis id="E8D06A0FE827FFBAFF10F87EFE5DF848" bold="true" box="[251,393,1930,1952]" pageId="20" pageNumber="19">Figures 83–86</emphasis>
|
||
. Right legs I–IV, retrolateral aspect.
|
||
<emphasis id="E8D06A0FE827FFBAFD03F87EFCA3F848" bold="true" box="[744,887,1930,1952]" pageId="20" pageNumber="19">Figures 87–97</emphasis>
|
||
. Segments of pedipalps. Pedipalp chela, dorsal (87), external (88), and ventral (89) views. Pedipalp patella, dorsal (90), external (91), and ventral (92) views. Pedipalp femur and trochanter, internal (93), dorsal (94), and ventral (95) views. Pedipalp chela, movable (96) and fixed (97) fingers dentate margins. The trichobothrial pattern is indicated in Figures 87–91, 93–94 (white circles).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="8EDBE695E826FFBBFF9FF94CFF1BF93C" ID-DOI="http://doi.org/10.5281/zenodo.5741648" ID-Zenodo-Dep="5741648" httpUri="https://zenodo.org/record/5741648/files/figure.png" pageId="21" pageNumber="20" startId="21.[116,193,1720,1742]" targetBox="[113,1474,232,1713]" targetPageId="21">
|
||
<paragraph id="DA1BB61DE826FFBBFF9FF94CFF1BF93C" blockId="21.[113,1475,1720,2064]" pageId="21" pageNumber="20">
|
||
<emphasis id="E8D06A0FE826FFBBFF9FF94CFED0F91A" bold="true" box="[116,260,1720,1742]" pageId="21" pageNumber="20">Figures 98–99</emphasis>
|
||
.
|
||
<taxonomicName id="1DA4CD9EE826FFBBFEF9F94CFE09F919" authority="Kovařík & Fet & Yağmur, 2020" authorityName="Kovařík & Fet & Yağmur" authorityYear="2020" box="[274,477,1720,1742]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="grosseri" status="sp. nov.">
|
||
<emphasis id="E8D06A0FE826FFBBFEF9F94CFE09F919" box="[274,477,1720,1742]" italics="true" pageId="21" pageNumber="20">Orthochirus grosseri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F3E3D774E826FFBBFE0FF94CFDCFF91A" box="[484,539,1720,1741]" pageId="21" pageNumber="20" rank="species">
|
||
<emphasis id="E8D06A0FE826FFBBFE0FF94CFE2FF91A" bold="true" box="[484,507,1720,1741]" pageId="21" pageNumber="20">sp</emphasis>
|
||
.
|
||
<emphasis id="E8D06A0FE826FFBBFDE3F94CFDC1F91A" bold="true" box="[520,533,1720,1741]" pageId="21" pageNumber="20">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, holotype female, carapace and tergites (98), and sternopectinal region and sternites (99) under UV light.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="92BEE596E826FFBBFF9AF90CFBB3F7C7" pageId="21" pageNumber="20" type="diagnosis">
|
||
<paragraph id="DA1BB61DE826FFBBFF9AF90CFBB3F7C7" blockId="21.[113,1475,1720,2064]" pageId="21" pageNumber="20">
|
||
DIAGNOSIS (
|
||
<specimenCount id="CCA27D94E826FFBBFF12F90CFED3F8C7" box="[249,263,1784,1808]" pageId="21" pageNumber="20" type="female">♀</specimenCount>
|
||
).Total length of adult females
|
||
<quantity id="1D5C1BF8E826FFBBFD6AF90CFCD4F8C7" box="[641,768,1784,1808]" metricMagnitude="-2" metricUnit="m" metricValue="4.15" metricValueMax="4.3" metricValueMin="4.0" pageId="21" pageNumber="20" unit="mm" value="41.5" valueMax="43.0" valueMin="40.0">40–43 mm</quantity>
|
||
. laterally with fine punctation developed, spaces among punctae Trichobothrium
|
||
<emphasis id="E8D06A0FE826FFBBFEC8F8ECFEE4F8E7" box="[291,304,1816,1840]" italics="true" pageId="21" pageNumber="20">d</emphasis>
|
||
<subScript id="4620B458E826FFBBFEDAF8DCFEEDF8E1" attach="left" box="[305,313,1832,1846]" fontSize="6" pageId="21" pageNumber="20">2</subScript>
|
||
on pedipalp femur is present but located smooth; metasoma II–III ventrally and laterally bumpy with on internal surface. Pectinal teeth number
|
||
<quantity id="1D5C1BF8E826FFBBFDAAF8CCFD75F887" box="[577,673,1848,1872]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" metricValueMax="5.08" metricValueMin="4.572" pageId="21" pageNumber="20" unit="in" value="19.0" valueMax="20.0" valueMin="18.0">18–20 in</quantity>
|
||
females. punctation reduced. Metasoma V dorsal surface mesially Movable finger of pedipalps with 9 rows of denticles, 8 ID smooth; metasoma I dorsally with 11–14 large granules. and no OD. Dorsal carinae on pedipalp patella present and Tergites roughly granulated. Sternite VII densely granulated. smooth. Metasoma I–II with 10 carinae, metasoma III with Pedipalp, metasoma and telson glabrous. Moderate tibial 6–8 carinae, metasoma IV–V with 2 incomplete dorsolateral spurs present on legs III and IV. Tarsomere I of legs I–III with carinae; ventrolateral carinae indicated on metasoma IV bristlecombs composed of 9–13 setae. Ratio length/width of and present on metasoma V. Metasoma IV–V ventrally and metasoma V
|
||
<quantity id="1D5C1BF8E826FFBBFC57F80CFBD2F7C7" box="[956,1030,2040,2064]" metricMagnitude="-2" metricUnit="m" metricValue="2.6162" pageId="21" pageNumber="20" unit="in" value="1.03">1.03 in</quantity>
|
||
females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="8EDBE695E825FFB8FF98F84BFCB5F826" ID-DOI="http://doi.org/10.5281/zenodo.5741650" ID-Zenodo-Dep="5741650" httpUri="https://zenodo.org/record/5741650/files/figure.png" pageId="22" pageNumber="21" startId="22.[115,192,1983,2005]" targetBox="[116,1474,238,1960]" targetPageId="22">
|
||
<paragraph id="DA1BB61DE825FFB8FF98F84BFCB5F826" blockId="22.[115,1473,1983,2034]" pageId="22" pageNumber="21">
|
||
<emphasis id="E8D06A0FE825FFB8FF98F84BFECEF803" bold="true" box="[115,282,1983,2005]" pageId="22" pageNumber="21">Figures 100–105</emphasis>
|
||
.
|
||
<taxonomicName id="1DA4CD9EE825FFB8FECCF84BFE25F802" authority="Kovařík & Fet & Yağmur, 2020" authorityName="Kovařík & Fet & Yağmur" authorityYear="2020" box="[295,497,1983,2005]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="22" pageNumber="21" phylum="Arthropoda" rank="species" species="grosseri" status="sp. nov.">
|
||
<emphasis id="E8D06A0FE825FFB8FECCF84BFE25F802" box="[295,497,1983,2005]" italics="true" pageId="22" pageNumber="21">Orthochirus grosseri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F3E3D774E825FFB8FE13F84BFDF9F803" box="[504,557,1983,2004]" pageId="22" pageNumber="21" rank="species">
|
||
<emphasis id="E8D06A0FE825FFB8FE13F84BFDDBF803" bold="true" box="[504,527,1983,2004]" pageId="22" pageNumber="21">sp</emphasis>
|
||
.
|
||
<emphasis id="E8D06A0FE825FFB8FDF0F84BFDFCF803" bold="true" box="[539,552,1983,2004]" pageId="22" pageNumber="21">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, holotype female, metasoma and telson, lateral (100, 103), dorsal (101, 104) and ventral (102, 105), under white light (100–102) and UV light (103–105). Scale bar: 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="92BEE596E824FFB9FF9AFF06FF28F87D" pageId="23" pageNumber="22" type="description">
|
||
<paragraph id="DA1BB61DE824FFB9FF9AFF06FE72FEBD" blockId="23.[113,772,242,1770]" pageId="23" pageNumber="22">
|
||
DESCRIPTION. Total length
|
||
<quantity id="1D5C1BF8E824FFB9FE45FF06FDFFFEDD" box="[430,555,242,266]" metricMagnitude="-2" metricUnit="m" metricValue="4.15" metricValueMax="4.3" metricValueMin="4.0" pageId="23" pageNumber="22" unit="mm" value="41.5" valueMax="43.0" valueMin="40.0">40–43 mm</quantity>
|
||
of adult females. Measurements of the carapace, telson, segments of the metasoma and segments of the pedipalps are given in
|
||
<tableCitation id="972683A6E824FFB9FD45FEC6FD2BFE9D" box="[686,767,306,330]" captionStart="Table 2" captionStartId="28.[117,173,976,997]" captionText="Table 2. Comparative measurements of Orthochirus scrobiculosus, O. melanurus, and O. grosseri sp. n. specimens.Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="23" pageNumber="22">Table 2</tableCitation>
|
||
. For habitus, see
|
||
<figureCitation id="429FAA98E824FFB9FECBFEA6FE75FEBD" box="[288,417,338,362]" captionStart="Figures 79–80" captionStartId="19.[117,194,2006,2028]" captionTargetBox="[137,1456,307,1887]" captionTargetId="figure-31@19.[114,1475,285,1919]" captionTargetPageId="19" captionText="Figures 79–80. Orthochirus grosseri sp. n., holotype female, dorsal (79) and ventral (80) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5741644" httpUri="https://zenodo.org/record/5741644/files/figure.png" pageId="23" pageNumber="22">Figs. 79–80</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="DA1BB61DE824FFB9FF9AFE86FF1AFD9D" blockId="23.[113,772,242,1770]" pageId="23" pageNumber="22">
|
||
<emphasis id="E8D06A0FE824FFB9FF9AFE86FF38FE5D" bold="true" box="[113,236,370,394]" pageId="23" pageNumber="22">Coloration</emphasis>
|
||
(
|
||
<figureCitation id="429FAA98E824FFB9FEEEFE86FE40FE5D" box="[261,404,370,394]" captionStart="Figures 79–80" captionStartId="19.[117,194,2006,2028]" captionTargetBox="[137,1456,307,1887]" captionTargetId="figure-31@19.[114,1475,285,1919]" captionTargetPageId="19" captionText="Figures 79–80. Orthochirus grosseri sp. n., holotype female, dorsal (79) and ventral (80) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5741644" httpUri="https://zenodo.org/record/5741644/files/figure.png" pageId="23" pageNumber="22">Figs. 79–80</figureCitation>
|
||
). Carapace, tergites, metasoma, and femur and patella of pedipalps and legs black. Chela of pedipalps reddish brown with black spots, fingers lighter. Tarsomeres of legs yellowish brown. Sternites reddish black with yellow median area indicated in posterior margin of sternite IV and developed on sternite V. Telson reddish black to black.
|
||
</paragraph>
|
||
<paragraph id="DA1BB61DE824FFB9FF9AFDA6FE22FCDD" blockId="23.[113,772,242,1770]" pageId="23" pageNumber="22">
|
||
<emphasis id="E8D06A0FE824FFB9FF9AFDA6FE43FDBD" bold="true" box="[113,407,594,618]" pageId="23" pageNumber="22">Mesosoma and carapace</emphasis>
|
||
(
|
||
<figureCitation id="429FAA98E824FFB9FE45FDA6FDE3FDBD" box="[430,567,594,618]" captionStart="Figures 81–97" captionStartId="20.[113,190,1901,1923]" captionTargetBox="[115,1470,234,1876]" captionTargetId="figure-115@20.[113,1474,234,1883]" captionTargetPageId="20" captionText="Figures 81–97: Orthochirus grosseri sp. n., holotype female. Figures 81–82. Carapace and tergites I–IV (81), and sternopectinal region and sternites (82). Figures 83–86. Right legs I–IV, retrolateral aspect. Figures 87–97. Segments of pedipalps. Pedipalp chela, dorsal (87), external (88), and ventral (89) views. Pedipalp patella, dorsal (90), external (91), and ventral (92) views. Pedipalp femur and trochanter, internal (93), dorsal (94), and ventral (95) views. Pedipalp chela, movable (96) and fixed (97) fingers dentate margins. The trichobothrial pattern is indicated in Figures 87–91, 93–94 (white circles)." figureDoi="http://doi.org/10.5281/zenodo.5741646" httpUri="https://zenodo.org/record/5741646/files/figure.png" pageId="23" pageNumber="22">Figs. 81–82</figureCitation>
|
||
,
|
||
<figureCitation id="429FAA98E824FFB9FDA1FDA6FD45FDBD" box="[586,657,594,618]" captionStart="Figures 98–99" captionStartId="21.[116,193,1720,1742]" captionTargetBox="[113,1474,232,1713]" captionTargetId="figure-185@21.[113,1474,231,1713]" captionTargetPageId="21" captionText="Figures 98–99. Orthochirus grosseri sp. n., holotype female, carapace and tergites (98), and sternopectinal region and sternites (99) under UV light." figureDoi="http://doi.org/10.5281/zenodo.5741648" httpUri="https://zenodo.org/record/5741648/files/figure.png" pageId="23" pageNumber="22">98–99</figureCitation>
|
||
). Tergites with a median carina and is roughly granulated. Seventh sternite densely granulated with four granulated carinae, the other sternites partly granulated but smooth in middle and posteriorly. Pectine teeth number
|
||
<quantity id="1D5C1BF8E824FFB9FE32FD26FDC4FD3D" box="[473,528,722,746]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" pageId="23" pageNumber="22" unit="in" value="20.0">20 in</quantity>
|
||
juvenile male and 18– 20 (1 x 18, 2 x 19, 1x20) in females.
|
||
</paragraph>
|
||
<paragraph id="DA1BB61DE824FFB9FF9AFCE6FEFEFB1D" blockId="23.[113,772,242,1770]" pageId="23" pageNumber="22">
|
||
<emphasis id="E8D06A0FE824FFB9FF9AFCE6FEB3FCFD" bold="true" box="[113,359,786,810]" pageId="23" pageNumber="22">Metasoma and telson</emphasis>
|
||
(
|
||
<figureCitation id="429FAA98E824FFB9FE9CFCE6FDCCFCFD" box="[375,536,786,810]" captionStart="Figures 100–105" captionStartId="22.[115,192,1983,2005]" captionTargetBox="[116,1474,238,1960]" captionTargetId="figure-15@22.[113,1474,234,1973]" captionTargetPageId="22" captionText="Figures 100–105. Orthochirus grosseri sp. n., holotype female, metasoma and telson, lateral (100, 103), dorsal (101, 104) and ventral (102, 105), under white light (100–102) and UV light (103–105). Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5741650" httpUri="https://zenodo.org/record/5741650/files/figure.png" pageId="23" pageNumber="22">Figs. 100–105</figureCitation>
|
||
). Metasoma I–II with 10 granulated carinae. Metasoma III–V lacks lateral and metasoma IV–V ventromedian carinae. Ventrolateral carinae are developed on metasoma I–III and indicated on metasoma IV–V, more on metasoma V, dorsolateral carinae are present on metasoma I–IV. Ventral carinae of metasoma I–III consist of large granules in one (metasoma II–III) or two (usually on metasoma I) rows. Metasoma I granulated laterally, metasoma III–V laterally punctate and smooth; granulation absent on dorsal surfaces of metasoma II–V. Metasoma I dorsally with 11–14 large granules. Fine punctation on metasoma IV–V ventrally developed, spaces among punctae smooth. Entire metasoma and telson glabrous. Telson smooth with punctation present ventrally.
|
||
</paragraph>
|
||
<paragraph id="DA1BB61DE824FFB9FF9AFB26FF15FA3D" blockId="23.[113,772,242,1770]" pageId="23" pageNumber="22">
|
||
<emphasis id="E8D06A0FE824FFB9FF9AFB26FF34FB3D" bold="true" box="[113,224,1234,1258]" pageId="23" pageNumber="22">Pedipalps</emphasis>
|
||
(
|
||
<figureCitation id="429FAA98E824FFB9FF06FB26FEBAFB3D" box="[237,366,1234,1258]" captionStart="Figures 81–97" captionStartId="20.[113,190,1901,1923]" captionTargetBox="[115,1470,234,1876]" captionTargetId="figure-115@20.[113,1474,234,1883]" captionTargetPageId="20" captionText="Figures 81–97: Orthochirus grosseri sp. n., holotype female. Figures 81–82. Carapace and tergites I–IV (81), and sternopectinal region and sternites (82). Figures 83–86. Right legs I–IV, retrolateral aspect. Figures 87–97. Segments of pedipalps. Pedipalp chela, dorsal (87), external (88), and ventral (89) views. Pedipalp patella, dorsal (90), external (91), and ventral (92) views. Pedipalp femur and trochanter, internal (93), dorsal (94), and ventral (95) views. Pedipalp chela, movable (96) and fixed (97) fingers dentate margins. The trichobothrial pattern is indicated in Figures 87–91, 93–94 (white circles)." figureDoi="http://doi.org/10.5281/zenodo.5741646" httpUri="https://zenodo.org/record/5741646/files/figure.png" pageId="23" pageNumber="22">Figs. 87–97</figureCitation>
|
||
). Trichobothrium
|
||
<emphasis id="E8D06A0FE824FFB9FDC1FB26FDE3FB3D" box="[554,567,1234,1258]" italics="true" pageId="23" pageNumber="22">d</emphasis>
|
||
<subScript id="4620B458E824FFB9FDDCFB16FDEBFB27" attach="left" box="[567,575,1250,1264]" fontSize="6" pageId="23" pageNumber="22">2</subScript>
|
||
on pedipalp femur is present but located on internal surface; trichobothrium
|
||
<emphasis id="E8D06A0FE824FFB9FD3CFB07FD37FADD" box="[727,739,1267,1290]" italics="true" pageId="23" pageNumber="22">e</emphasis>
|
||
<subScript id="4620B458E824FFB9FD08FAF6FD3FFAC7" attach="left" box="[739,747,1282,1296]" fontSize="6" pageId="23" pageNumber="22">1</subScript>
|
||
is situated between trichobothria
|
||
<emphasis id="E8D06A0FE824FFB9FE2AFAE6FE1AFAFD" box="[449,462,1298,1322]" italics="true" pageId="23" pageNumber="22">d</emphasis>
|
||
<subScript id="4620B458E824FFB9FE24FAD6FE03FAE7" attach="left" box="[463,471,1314,1328]" fontSize="6" pageId="23" pageNumber="22">2</subScript>
|
||
and
|
||
<emphasis id="E8D06A0FE824FFB9FDE0FAE6FDCCFAFD" box="[523,536,1298,1322]" italics="true" pageId="23" pageNumber="22">d</emphasis>
|
||
<subScript id="4620B458E824FFB9FDF3FAD6FDF4FAE7" attach="left" box="[536,544,1314,1328]" fontSize="6" pageId="23" pageNumber="22">4</subScript>
|
||
. Femur of pedipalps with five granulated carinae and is dorsally rather smooth. Patella has seven smooth carinae, and the chela has smooth carinae which may be discernible throughout the length of the fixed finger. The entire pedipalps are glabrous with several setae only. Movable fingers with 9 rows of denticles, 8 ID and no OD.
|
||
</paragraph>
|
||
<paragraph id="DA1BB61DE824FFB9FF9AFA06FE0FF91D" blockId="23.[113,772,242,1770]" pageId="23" pageNumber="22">
|
||
<emphasis id="E8D06A0FE824FFB9FF9AFA06FF72F9DD" bold="true" box="[113,166,1522,1546]" pageId="23" pageNumber="22">Legs</emphasis>
|
||
(
|
||
<figureCitation id="429FAA98E824FFB9FF5CFA06FEEFF9DD" box="[183,315,1522,1546]" captionStart="Figures 81–97" captionStartId="20.[113,190,1901,1923]" captionTargetBox="[115,1470,234,1876]" captionTargetId="figure-115@20.[113,1474,234,1883]" captionTargetPageId="20" captionText="Figures 81–97: Orthochirus grosseri sp. n., holotype female. Figures 81–82. Carapace and tergites I–IV (81), and sternopectinal region and sternites (82). Figures 83–86. Right legs I–IV, retrolateral aspect. Figures 87–97. Segments of pedipalps. Pedipalp chela, dorsal (87), external (88), and ventral (89) views. Pedipalp patella, dorsal (90), external (91), and ventral (92) views. Pedipalp femur and trochanter, internal (93), dorsal (94), and ventral (95) views. Pedipalp chela, movable (96) and fixed (97) fingers dentate margins. The trichobothrial pattern is indicated in Figures 87–91, 93–94 (white circles)." figureDoi="http://doi.org/10.5281/zenodo.5741646" httpUri="https://zenodo.org/record/5741646/files/figure.png" pageId="23" pageNumber="22">Figs. 83–86</figureCitation>
|
||
). Moderate tibial spurs present on legs III and IV. Femur with four partly granulated carinae; patella with five rather smooth carinae; tibia smooth. Patella with only a few setae. Tibia with setae on the outer side of legs I–III. Tarsomere I of legs I–III with 9–13 long setae in females, legs IV with 2–4 setae. Tarsomeres I–II of all legs internally with two rather irregular rows of setae.
|
||
</paragraph>
|
||
<paragraph id="DA1BB61DE824FFB9FF9AF926FE7CF93D" blockId="23.[113,772,242,1770]" box="[113,424,1746,1770]" pageId="23" pageNumber="22">
|
||
<emphasis id="E8D06A0FE824FFB9FF9AF926FECCF93D" bold="true" box="[113,280,1746,1770]" pageId="23" pageNumber="22">Measurements</emphasis>
|
||
. See
|
||
<tableCitation id="972683A6E824FFB9FEB9F926FE70F93D" box="[338,420,1746,1770]" captionStart="Table 2" captionStartId="28.[117,173,976,997]" captionText="Table 2. Comparative measurements of Orthochirus scrobiculosus, O. melanurus, and O. grosseri sp. n. specimens.Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="23" pageNumber="22">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="DA1BB61DE824FFB9FF9AF8E6FF7AF89D" blockId="23.[113,771,1810,1866]" pageId="23" pageNumber="22">
|
||
AFFINITIES. See affinities under description of
|
||
<taxonomicName id="1DA4CD9EE824FFB9FD9FF8E7FCD7F8FD" authority="Kovařík & Fet & Yağmur, 2020" authorityName="Kovařík & Fet & Yağmur" authorityYear="2020" box="[628,771,1810,1834]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="23" pageNumber="22" phylum="Arthropoda" rank="species" species="formozovi" status="sp. nov.">
|
||
<emphasis id="E8D06A0FE824FFB9FD9FF8E7FCD7F8FD" box="[628,771,1810,1834]" italics="true" pageId="23" pageNumber="22">O. formozovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F3E3D774E824FFB9FF9AF8C6FF7AF89D" box="[113,174,1842,1866]" pageId="23" pageNumber="22" rank="species">
|
||
<emphasis id="E8D06A0FE824FFB9FF9AF8C6FF5EF89D" bold="true" box="[113,138,1842,1866]" pageId="23" pageNumber="22">sp</emphasis>
|
||
.
|
||
<emphasis id="E8D06A0FE824FFB9FF73F8C6FF73F89D" bold="true" box="[152,167,1842,1866]" pageId="23" pageNumber="22">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="DA1BB61DE824FFB9FF9AF886FF28F87D" blockId="23.[113,771,1906,1963]" pageId="23" pageNumber="22">
|
||
<materialsCitation id="6ACCBC40E824FFB9FF9AF886FF2CF87D" country="Uzbekistan" location="Known" pageId="23" pageNumber="22" specimenCount="1">
|
||
DISTRIBUTION.
|
||
<collectingCountry id="A2B3F68DE824FFB9FEFEF886FE59F85D" box="[277,397,1906,1930]" name="Uzbekistan" pageId="23" pageNumber="22">Uzbekistan</collectingCountry>
|
||
(
|
||
<figureCitation id="429FAA98E824FFB9FE75F886FE2BF85D" box="[414,511,1906,1930]" captionStart="Figure 343" captionStartId="66.[115,186,1950,1972]" captionTargetBox="[116,1475,234,1817]" captionTargetId="figure-15@66.[116,1477,234,1832]" captionTargetPageId="66" captionText="Figure 343. Geographic distribution of the Orthochirus spp. found from Turkey to India. An approximate area where the lectotype of O. melanurus was collected is marked by red ellipse." figureDoi="http://doi.org/10.5281/zenodo.5741722" httpUri="https://zenodo.org/record/5741722/files/figure.png" pageId="23" pageNumber="22">Fig. 343</figureCitation>
|
||
).
|
||
<location id="DF7BE0C6E824FFB9FDFFF887FDB7F85D" LSID="urn:lsid:plazi:treatment:520D070BE821FFB9FC3CFA1BFF28F87D:DF7BE0C6E824FFB9FDFFF887FDB7F85D" box="[532,611,1907,1930]" country="Uzbekistan" name="Known" pageId="23" pageNumber="22">Known</location>
|
||
only from the
|
||
<typeStatus id="051F08BFE824FFB9FF9AF867FF4BF87C" box="[113,159,1939,1963]" pageId="23" pageNumber="22">type</typeStatus>
|
||
locality
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |