185 lines
22 KiB
XML
185 lines
22 KiB
XML
<document id="CB5FEA99EA171AB770DC9DC3C7728E25" ID-CLB-Dataset="28787" ID-DOI="http://dx.doi.org/10.3897/zookeys.800.26292" ID-GBIF-Dataset="216546ea-85b8-4a1d-89c3-4c88818cc79d" ID-PMC="PMC6284011" ID-Pensoft-Pub="1313-2970-800-1" ID-PubMed="30532621" ID-ZooBank="EB5EC9C8D9804F5ABD9AE48DB4158D59" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-800-1" ModsDocOrigin="ZooKeys 800" ModsDocTitle="Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae)" checkinTime="1543542350729" checkinUser="pensoft" docAuthor="Huemer, Peter & Karsholt, Ole" docDate="2018" docId="518F2983F7A6F5D61163E9A7FEB28F79" docLanguage="en" docName="ZooKeys 800: 1-278" docOrigin="ZooKeys 800" docSource="http://dx.doi.org/10.3897/zookeys.800.26292" docTitle="Megacraspedus leuca Filipjev 1929" docType="treatment" docVersion="7" lastPageNumber="137" masterDocId="FF97FFAFDB35FFAAFFD1B44AFF8EBF02" masterDocTitle="Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae)" masterLastPageNumber="278" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1543542350729" originalUpdateUser="pensoft" pageNumber="135" updateTime="1701384233526" updateUser="ExternalLinkService">
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<mods:title id="44AF3BC642431D140460A50BDD4B7E3C">Revision of the genus Megacraspedus Zeller, 1839, a challenging taxonomic tightrope of species delimitation (Lepidoptera, Gelechiidae)</mods:title>
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<mods:namePart id="E607606F85976458FAF9956CE9A38BF3">Huemer, Peter</mods:namePart>
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<mods:namePart id="C5F44264AD62BA96F79B37EB5971FFD0">Karsholt, Ole</mods:namePart>
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<mods:date id="CBB5A11C692BDE9E56B7B239BCBBFE2A">2018</mods:date>
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<mods:identifier id="CB8A6C7EA03928F6370CCBC7E7AAECCF" type="DOI">http://dx.doi.org/10.3897/zookeys.800.26292</mods:identifier>
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<treatment id="518F2983F7A6F5D61163E9A7FEB28F79" ID-GBIF-Taxon="149858180" LSID="urn:lsid:plazi:treatment:518F2983F7A6F5D61163E9A7FEB28F79" httpUri="http://treatment.plazi.org/id/518F2983F7A6F5D61163E9A7FEB28F79" lastPageId="136" lastPageNumber="137" pageId="134" pageNumber="135">
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<subSubSection id="306022453D027E2E856FE9C1363A76F4" pageId="134" pageNumber="135" type="nomenclature">
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<paragraph id="1E2ADC1B4E7745F1ED6F236736BE4BDE" pageId="134" pageNumber="135">
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<taxonomicName id="54EC24654129904999AB83935CC56938" ID-CoL="93JG2" authority="Filipjev, 1929" authorityName="Filipjev" authorityYear="1929" class="Insecta" family="Gelechiidae" genus="Megacraspedus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megacraspedus leuca" order="Lepidoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="leuca">Megacraspedus leuca (Filipjev, 1929)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C82BD956E98A76AB4FD040D6685BBC04" pageId="134" pageNumber="135" type="reference_group">
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<paragraph id="3E7BE243DFA8240948F055C2F4D5FEED" pageId="134" pageNumber="135">
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<taxonomicName id="48363ECFF5C0310A0E0D3870EF2549D0" class="Insecta" family="Gelechiidae" genus="Nothris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothris leuca" order="Lepidoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="leuca">Nothris leuca</taxonomicName>
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Filipjev, 1929: 9, pl. 2a, fig. 3.
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</paragraph>
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<paragraph id="3049093F160C4369C64547DB3D975225" pageId="134" pageNumber="135">
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<taxonomicName id="B1462AA7D58E3837B398DA32DA642805" class="Insecta" family="Gelechiidae" genus="Megacraspedus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megacraspedus kaszabianus" order="Lepidoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="kaszabianus">Megacraspedus kaszabianus</taxonomicName>
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<normalizedToken id="035967E9FA9910FF5F3B67F3B2BDE526" originalValue="Povolný">Povolny</normalizedToken>
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, 1982: 193, figs 1-2, syn. n.
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</paragraph>
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</subSubSection>
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<subSubSection id="32FDC8BED1790638F1451B5EAD95DADD" lastPageId="135" lastPageNumber="136" pageId="134" pageNumber="135" type="materials_examined">
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<paragraph id="4C7FDFDEBFFDF412D1BFB2ACA1C46C87" pageId="134" pageNumber="135">Examined material.</paragraph>
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<paragraph id="E347B3C56961E7164181F55FF59D6384" lastPageId="135" lastPageNumber="136" pageId="134" pageNumber="135">
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Paratypes of
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<taxonomicName id="06DC46063424694D620FBBEFB43E4860" lsidName="M. kaszabianus" pageId="134" pageNumber="135" rank="species" species="kaszabianus">M. kaszabianus</taxonomicName>
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. Mongolia. 1 ♂, Gobi Altaj aimak, Chasat chajrchan ul mts, cca. 20 km S Somon Zargalan, 2400 m, 15.vii.1966, leg. Z. Kaszab; 1 ♂,
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<normalizedToken id="80D0DE01A9E20C888155C848FE6771DC" originalValue="Südgobi">Suedgobi</normalizedToken>
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aimak, Gurban Sajchan ul, 15 km S Dalanzadgad, 1750 m, 13.vi.1967, leg. Z. Kaszab; 1 ♂, 1 ♀,
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<normalizedToken id="D4F2C665427CFBB8129A88C9AC204250" originalValue="Südgobi">Suedgobi</normalizedToken>
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aimak, Somon Bulgan Talyn bulag, 1350 m, 5.vii.1967, genitalia slides GU 16/1446 ♂ Huemer, GU 16/1447 ♀ Huemer, leg. Z. Kaszab (HNHM). Non-type material. Russia. 1 ♂ Alai mts, Kuraisk hrebet, 2300 m, 13.vii.2001, leg. K. Nupponen, genitalia slide no. 3/21.X.2008 K. Nupponen (RCKN); 2 ♂, Altai rep., 15 km S Kosh-Agach, Thuja steppe, 1800 m, 10.viii.2000, leg. A. Bidzilya (ZMUC); 4 ♂, Altai rep., 6.5 km SW Kosh-Agach, Kurai env., 1550 m, 9-10.vii.2014, leg. J.
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<normalizedToken id="3E0413202473063CDAECC4CBC099F2D6" originalValue="Šumpich">Sumpich</normalizedToken>
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, genitalia slide GU 16/1408 ♂ Huemer (NMPC, TLMF); 19 ♂, Altai rep., 10 km NE Kosh-Agach, Kurai mts, Range valley of Tabozhok river, 2100 m, 2-4.
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<pageBreakToken id="54AFDC500431AF3B4C9DE641462576B3" pageId="135" pageNumber="136" start="start">viii</pageBreakToken>
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.2016, leg. P. Huemer & B. Wiesmair; 1 ♂, Altai rep., 17 km NNE Kokorya village, Chikhacheva mts range, Talduair Mt., valley of Sajlyugem river, 30.
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<normalizedToken id="2F4A92B62FF50E76E2550F41ACE12AA4" originalValue="vii–">vii-</normalizedToken>
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2.viii.2016, leg. P. Huemer & B. Wiesmair; 2 ♂, Altai rep., Aktasch env., 17.
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<normalizedToken id="19F524C53C574FFEA52F4AD761EAAB7F" originalValue="vi–">vi-</normalizedToken>
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2.vii.2009, leg. B. Schacht (all TLMF); 4 ♂, Tuva rep.,
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<geoCoordinate id="2172C6D7C1607F900A6055426F29BD87" direction="north" orientation="latitude" precision="925" value="52.066666">52°04'N</geoCoordinate>
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,
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<geoCoordinate id="4401EA016B989ECC9FC0E6946E07B7DF" direction="east" orientation="longitude" precision="925" value="94.36667">94°22'E</geoCoordinate>
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, Ust-Uljuk, steppe hills, 670 m, 3-5.vi.1995, leg. J. Jalava & J. Kullberg (MZH); 26 ♂, Tuva rep.,
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<geoCoordinate id="E0E8D9A8B464FB2634B011C2D7945AEC" direction="north" orientation="latitude" precision="925" value="50.016666">50°01'N</geoCoordinate>
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,
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<geoCoordinate id="E1486C40143D451A6CC97801260E020A" direction="east" orientation="longitude" precision="925" value="95.05">95°03'E</geoCoordinate>
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, Lake Tere Khol, sand dunes, 1150 m, 9-12.vi.1995, leg. J. Jalava & J. Kullberg (MZH, ZMUC); 8 ♂, 1 ♀, Tuva rep.,
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<geoCoordinate id="E945E928C3CF0A14C763C9D182D877BA" direction="north" orientation="latitude" precision="925" value="50.266666">50°16'N</geoCoordinate>
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,
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<geoCoordinate id="8D3AC99C68B1D69313638C6AB9FAB15A" direction="east" orientation="longitude" precision="925" value="94.9">94°54'E</geoCoordinate>
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, ca. 25 km W Erzin, steppe/stony slopes, 1250 m, 7-11.vi.1995, leg. J. Jalava & J. Kullberg (MZH); 18 ♂, Tuva rep.,
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<geoCoordinate id="766ADECAA5E56BB8BF1644C145DD26C2" direction="north" orientation="latitude" precision="925" value="50.733334">50°44'N</geoCoordinate>
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,
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<geoCoordinate id="603E6E14091CA92F0D0EB9350C5586C2" direction="east" orientation="longitude" precision="925" value="93.13333">93°08'E</geoCoordinate>
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, E Tannu-Ola mts, Irbitei r., stony steppe slopes, 1000 m, 13-16.vi.1995, leg. J. Jalava & J. Kullberg (MZH); 1 ♂, Tuva rep.,
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<geoCoordinate id="1701146C0DB90C7E76FA3AE8A5E5C984" direction="north" orientation="latitude" precision="925" value="50.75">50°45'N</geoCoordinate>
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,
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<geoCoordinate id="E12281D92E9DE6C1C79427AF484DE1D3" direction="east" orientation="longitude" precision="925" value="94.48333">94°29'E</geoCoordinate>
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, E Tannu-Ola mts, 5 km ENE Khol-Oozha, steppe slopes, 1000 m, 13-16.vi.1995, leg. J. Jalava & J. Kullberg (MZH).
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</paragraph>
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</subSubSection>
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<subSubSection id="6557E298784B6B57D3C383FAEC152810" lastPageId="136" lastPageNumber="137" pageId="135" pageNumber="136" type="redescription">
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<paragraph id="55BA60BCECF6BC5ECB4ECE11E0C6C709" pageId="135" pageNumber="136">Redescription.</paragraph>
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<paragraph id="3D0E2EAD19788CE06BFFA34F1E2A5836" pageId="135" pageNumber="136">Adult. Male (Figure 111). Wingspan 15-18 mm. Segment 2 of labial palpus with sub-rectangular scale brush longer than segment 3, blackish brown mottled with white, especially on upper surface; segment 3 about two-thirds length of segment 2, white mottled with black. Antennal scape with pecten of several hairs; flagellum light brown, indistinctly ringed with black. Head, thorax and tegula whitish mottled with light brown. Forewing cream whitish, mottled with brownish- or blackish-tipped scales; indistinct blackish blown spots in yellow-white fold and at 2/3 and 3/4 in middle of wing; fringes whitish grey with two grey fringe lines. Hindwing light grey with concolorous fringes.</paragraph>
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<paragraph id="1C3D5DFA028AB89EAB9EB389C30F78C1" pageId="135" pageNumber="136">Female (Figure 112). Wingspan 15 mm. Otherwise similar to male.</paragraph>
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<paragraph id="821DFDB31CEB3A742DF7DDAEA0BDCB75" pageId="135" pageNumber="136">Variation. The amount of brownish- or blackish-tipped scales of the forewing varies; they are normally scattered over the wing but can also form lines along the veins. The blackish brown dots on the forewing can be more or less distinct. One specimen (in good condition) is almost without dark scales.</paragraph>
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<paragraph id="89A47605CF596F631E34C052E8368DE1" pageId="135" pageNumber="136">Male genitalia (Figure 237). Uncus large, sub-square, apical corners rounded, apical edge straight to weakly emarginated; gnathos hook evenly slender, apically pointed, almost twice length of uncus, bent at right angles at about one-quarter; anterior margin of tegumen with broad and shallow U-shaped emargination, anteromedially small additional emargination; pedunculi small, suboval; valva stout, extending slightly beyond base of uncus, basally weakly inflated, digitate distal part weakly tapered to slightly pointed apex; saccular area densely covered with setae, longitudinal ridge present, without separated sacculus; posterior margin of vinculum shallow medial emargination, without distinct lateral humps, vincular sclerite elongated sub-triangular, with sclerotised posterior edge; saccus prominent, longer than valva, U-shaped, ratio maximum width to length approximately 0.7, posterior margin with broadly rounded shallow projections, separated by small incision, medial part with long sclerotised ridge from posterior margin to apex of saccus, lateral sclerites approximately 0.8 times length of maximum width of saccus, with distinctly bulged apex; phallus with weakly inflated coecum, about two times wider than distal part, distal part more than twice length of coecum, sclerotised dorsal ridge with one to five small teeth.</paragraph>
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<paragraph id="10A439F1B1556DFE4D3AC0DBE8B7EA4C" pageId="136" pageNumber="137">
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<pageBreakToken id="5006A0333FA9533ADE037A53B88707A2" pageId="136" pageNumber="137" start="start">Female</pageBreakToken>
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genitalia (Figure 292). Papilla analis weakly sclerotised, moderately small, apically rounded; apophysis posterior slender rod-like, short, slightly longer than apophysis anterior, posterior end with small transverse sclerotised zone; segment VIII very short, laterally strongly sclerotised, medially largely membranous; anterior edge of segment VIII laterally rod-like, connected with apophysis anterior, subgenital plate reduced, ostium bursae without bordering sclerotisations; apophysis anterior rod-like, almost five times length of segment VIII, posteriorly broadly connected with segment VIII by sclerotised band; colliculum short, ring-shaped; ductus bursae slender, weakly widening to hardly delimited, oblong corpus bursae; signum reduced.
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</paragraph>
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</subSubSection>
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<subSubSection id="3E8AF440FFB6A5A40905C4E1E0FF1834" pageId="136" pageNumber="137" type="diagnosis">
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<paragraph id="6713106F4A7F2FFABF9BC6DE66DCE869" pageId="136" pageNumber="137">Diagnosis.</paragraph>
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<paragraph id="1B6B1249387BF429386EC098C7D4AE46" pageId="136" pageNumber="137">
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<taxonomicName id="2C92F9F4673017ACADE583B1C66C1AE9" class="Insecta" family="Gelechiidae" genus="Megacraspedus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megacraspedus leuca" order="Lepidoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="leuca">Megacraspedus leuca</taxonomicName>
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is characterised by having the antennal scape with a pecten of several hairs, and by the having the cream whitish grey forewings mottled with brownish- or blackish-tipped scales. It resembles
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<taxonomicName id="E7D5CCDFB54CD93E15ECEE7E21AA7228" lsidName="M. multispinella" pageId="136" pageNumber="137" rank="species" species="multispinella">M. multispinella</taxonomicName>
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(Figs 103-104), which is smaller and has greyish white forewings with black scales along the apical margins. It is furthermore similar to
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<taxonomicName id="7D9F7A71576C0BE617869CCB1847962D" lsidName="M. imparellus" pageId="136" pageNumber="137" rank="species" species="imparellus">M. imparellus</taxonomicName>
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(p 130),
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<taxonomicName id="A240BEDED5EA19C8AC8DF3A66FF7AC32" lsidName="M. orenburgensis" pageId="136" pageNumber="137" rank="species" species="orenburgensis">M. orenburgensis</taxonomicName>
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(p 144), and
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<taxonomicName id="59BB3BADF651343ABBBC897457F70F99" lsidName="M. attritellus" pageId="136" pageNumber="137" rank="species" species="attritellus">M. attritellus</taxonomicName>
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(p 138). The male genitalia are unmistakable and in particular the prominent, long and slender U-shaped saccus is unique in
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<taxonomicName id="5E6378D315CA411CF893A0BACB13DE5C" class="Insecta" family="Gelechiidae" genus="Megacraspedus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megacraspedus" order="Lepidoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="genus">Megacraspedus</taxonomicName>
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. The female genitalia are characterised by several unique structures such as the short segment VIII with a membranous medial part, the comparatively very long apophysis anterior, and the absence of a signum.
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</paragraph>
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</subSubSection>
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<subSubSection id="9FC593F4B1580F9C03DD550C235D83F6" pageId="136" pageNumber="137" type="molecular data">
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<paragraph id="C8526B4B6E96A1F5BC95D4F39D5553C4" pageId="136" pageNumber="137">Molecular data.</paragraph>
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<paragraph id="F19FE882892B45984FD8A5E6ED2027C1" pageId="136" pageNumber="137">
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BIN BOLD:ACB3260 (n = 10). The intraspecific divergence of the barcode region is low with mean 0.03% and maximum divergence of 0.2%. The distance to the nearest neighbour
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<taxonomicName id="699555B03722C35C6982902B3D4B95A2" lsidName="M. skulei" pageId="136" pageNumber="137" rank="species" species="skulei">M. skulei</taxonomicName>
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is 6.6% (p-dist).
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</paragraph>
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</subSubSection>
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<subSubSection id="A14792C679E5D25BA3C5F47C3631C7A4" pageId="136" pageNumber="137" type="distribution">
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<paragraph id="0C894B8DEDFB870CBF8441EDC9390E38" pageId="136" pageNumber="137">Distribution.</paragraph>
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<paragraph id="4AF810FC509FDDB2EE5CB695D3518243" pageId="136" pageNumber="137">Asian part of Russia, Mongolia.</paragraph>
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</subSubSection>
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<subSubSection id="BF9B89C1B33284143131E3DB4F54BA76" pageId="136" pageNumber="137" type="biology_ecology">
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<paragraph id="57C7A921E4683D6CB25319F823307064" pageId="136" pageNumber="137">Biology.</paragraph>
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<paragraph id="E3DA7C5B6D54EBB8C25BDB4CA1285CE1" pageId="136" pageNumber="137">
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Host plant and early stages are unknown.
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<bibRefCitation id="5A7E502B1D2DCF720C1108552C68334C" author="Bidzilya, O" journalOrPublisher="The Kharkov entomological Society Gazette" pageId="185" pageNumber="186" pagination="64 - 72" title="On the distribution of gelechiid moths (Lepidoptera: Gelechiidae) in the southern Siberia (information 1)." volume="9" year="2002">Bidzilya (2002</bibRefCitation>
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: 64) collected adults in numbers at artificial light placed among tussocks of
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<taxonomicName id="AF6E7F735CDF5EA3C851354859CE3CB8" class="Liliopsida" family="Poaceae" genus="Achnatherum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Achnatherum splendens" order="Poales" pageId="136" pageNumber="137" phylum="Tracheophyta" rank="species" species="splendens">Achnatherum splendens</taxonomicName>
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(Trin.) Nevski (=
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<taxonomicName id="01F4AAC78E0DE914FE9DC4E875DFC6D7" class="Liliopsida" family="Poaceae" genus="Stipa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Stipa splendens" order="Poales" pageId="136" pageNumber="137" phylum="Tracheophyta" rank="species" species="splendens">Stipa splendens</taxonomicName>
|
||
Trin.).
|
||
<taxonomicName id="0331F7E51EDAC695AD14E21167EB7382" lsidName="M. leuca" pageId="136" pageNumber="137" rank="species" species="leuca">M. leuca</taxonomicName>
|
||
is by far the most common species of the genus in Central Asia, inhabiting a wide range of different steppe habitats. According to
|
||
<bibRefCitation id="EE1AC5466DEC3BE805588D1ADB61EB20" pageId="136" pageNumber="137">
|
||
<normalizedToken id="6DB41E1240CAAFBC3E4FFE0E61316C22" originalValue="Povolný">Povolny</normalizedToken>
|
||
(1982)
|
||
</bibRefCitation>
|
||
it was collected from June to August in grass-, sand- and salt steppes at altitudes of between 1000 and 2400 m. Adults are easily attracted to articificial light (PH, pers. obs.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="56D1BD86ED84008058BBF83E89F7208E" pageId="136" pageNumber="137" type="remarks">
|
||
<paragraph id="8819070B397771F010F8160AA3170D18" pageId="136" pageNumber="137">Remarks.</paragraph>
|
||
<paragraph id="9F5A01D95C4D89F8A329B6644D65729B" pageId="136" pageNumber="137">
|
||
<taxonomicName id="6E02B16F8A2F9BBFF5AC5A07E6FBA6E4" class="Insecta" family="Gelechiidae" genus="Nothris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothris leuca" order="Lepidoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="leuca">Nothris leuca</taxonomicName>
|
||
was described from one male collected at Munku-Sardyk (=
|
||
<normalizedToken id="D727B7ED96D8B189BF8134B9C04BD279" originalValue="Mönkh-Saridak">Moenkh-Saridak</normalizedToken>
|
||
) in the Sajan Mountains at the border between Russia and Mongolia (Filipjev 1929). The holotype in ZISP was studied and figured by
|
||
<bibRefCitation id="906EBE211B107487D76FCEFB789E622F" author="Junnilainen, J" journalOrPublisher="Zootaxa" pageId="188" pageNumber="189" pagination="1 - 34" title="The gelechiid fauna of the Southern Ural Mountains, part I: descriptions of seventeen new species (Lepidoptera: Gelechiidae)." volume="2366" year="2010">Junnilainen and Nupponen (2010</bibRefCitation>
|
||
: 11).
|
||
</paragraph>
|
||
<paragraph id="31EBF63020C1FEA4A9CDB7372F15766F" pageId="136" pageNumber="137">
|
||
<taxonomicName id="55FEF57854FC5CA36525894F74698C0F" class="Insecta" family="Gelechiidae" genus="Megacraspedus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megacraspedus kaszabianus" order="Lepidoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="kaszabianus">Megacraspedus kaszabianus</taxonomicName>
|
||
was described from several hundred males collected in Mongolia by Z. Kaszab (
|
||
<bibRefCitation id="053BCA60B526CFCF56617CABF85A8980" pageId="136" pageNumber="137">
|
||
<normalizedToken id="CE9C9602BA77DA879A645B17FDCB46D9" originalValue="Povolný">Povolny</normalizedToken>
|
||
1982
|
||
</bibRefCitation>
|
||
). We have been able to examine paratypes, which fully agree with
|
||
<taxonomicName id="54FED9E3063987A363364560E8AAF8D5" lsidName="M. leuca" pageId="136" pageNumber="137" rank="species" species="leuca">M. leuca</taxonomicName>
|
||
in every detail. We therefore formally synonymise
|
||
<taxonomicName id="300A9545F6999B8977087A5BD9F7E8EC" lsidName="M. kaszabianus" pageId="136" pageNumber="137" rank="species" species="kaszabianus">M. kaszabianus</taxonomicName>
|
||
with
|
||
<taxonomicName id="6DF947833B086E526C2946F4E123B056" lsidName="M. leuca" pageId="136" pageNumber="137" rank="species" species="leuca">M. leuca</taxonomicName>
|
||
(syn. n.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |