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<document id="21FBBB9FD7E712C085A686F6EAF19DDC" ID-DOI="10.5281/zenodo.174318" ID-GBIF-Dataset="b8813703-00e8-48ff-bb05-667752b9a7f2" ID-ISSN="1175-5326" ID-Zenodo-Dep="174318" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459795368271" checkinUser="plazi" docAuthor="Hogan, James E." docDate="2006" docId="517C8791FFCD6843FEA2132E2B2288F4" docLanguage="en" docName="zt01337p068.pdf" docOrigin="Zootaxa 1337" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Psammocoryza Hogan" docType="treatment" docVersion="10" lastPageNumber="64" masterDocId="AD45FFE9FFCC6840FFAA100A280A890A" masterDocTitle="Psammocoryza vanemdeni, a new genus and species of scaritine beetle from the Atlantic coast of Brazil (Coleoptera: Carabidae: Clivinini)" masterLastPageNumber="68" masterPageNumber="61" pageNumber="62" updateTime="1698219676329" updateUser="plazi">
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<mods:title id="29BE17A8C60F820BAB83D15AF1BA6F2C">Psammocoryza vanemdeni, a new genus and species of scaritine beetle from the Atlantic coast of Brazil (Coleoptera: Carabidae: Clivinini)</mods:title>
</mods:titleInfo>
<mods:name id="2B2C5249D5B96A5F421FC91EBE321B05" type="personal">
<mods:role id="A0DE5520E4DEBFB03F8D32F9E3C9850F">
<mods:roleTerm id="920A580B56D2561A198E572496548866">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="51E1F6681FB7E64D61E71EF55720E5EF">Hogan, James E.</mods:namePart>
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<mods:typeOfResource id="301432941B08E2BEE0A28FC0E69CDD39">text</mods:typeOfResource>
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<mods:date id="AF7B268D6691AA5FF90AEB1D8C2654D4">2006</mods:date>
<mods:detail id="16D702D73C6932637FE8988C5872F0CC" type="volume">
<mods:number id="2044E944CBF29C7B14C5C9DA2A1CE6CA">1337</mods:number>
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<mods:classification id="B0429A496583C594B908786912B827F3">journal article</mods:classification>
<mods:identifier id="BA063444A8D13CB93283C7C6B2F61786" type="DOI">10.5281/zenodo.174318</mods:identifier>
<mods:identifier id="66E9E469E814F1BC0FAAF8F5C9E0A9D4" type="GBIF-Dataset">b8813703-00e8-48ff-bb05-667752b9a7f2</mods:identifier>
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<mods:identifier id="570FF2432D9A6ED2A259A4E6D43C6856" type="Zenodo-Dep">174318</mods:identifier>
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<treatment id="517C8791FFCD6843FEA2132E2B2288F4" ID-DOI="http://doi.org/10.5281/zenodo.5670922" ID-GBIF-Taxon="119351021" ID-Zenodo-Dep="5670922" LSID="urn:lsid:plazi:treatment:517C8791FFCD6843FEA2132E2B2288F4" httpUri="http://treatment.plazi.org/id/517C8791FFCD6843FEA2132E2B2288F4" lastPageId="3" lastPageNumber="64" pageId="1" pageNumber="62">
<subSubSection id="91CF650CFFCD6841FEA2132E29528A6B" pageId="1" pageNumber="62" type="nomenclature">
<paragraph id="D96A3687FFCD6841FEA2132E2AA38A34" blockId="1.[264,681,804,865]" box="[264,681,804,830]" pageId="1" pageNumber="62">
<heading id="822281EBFFCD6841FEA2132E2AA38A34" bold="true" box="[264,681,804,830]" fontSize="11" level="1" pageId="1" pageNumber="62" reason="1">
<emphasis id="EBA1EA95FFCD6841FEA2132E2AA38A34" bold="true" box="[264,681,804,830]" pageId="1" pageNumber="62">
<taxonomicName id="1ED54D04FFCD6841FEA2132E2A118A34" ID-CoL="8KYKX" authority="Hogan" authorityName="Hogan" box="[264,539,804,830]" class="Insecta" family="Carabidae" genus="Psammocoryza" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FEA2132E29B48A34" bold="true" box="[264,446,804,830]" italics="true" pageId="1" pageNumber="62">Psammocoryza</emphasis>
Hogan
</taxonomicName>
, new genus
</emphasis>
</heading>
</paragraph>
<paragraph id="D96A3687FFCD6841FEA2134D29528A6B" blockId="1.[264,681,804,865]" box="[264,344,839,865]" pageId="1" pageNumber="62">
(
<figureCitation id="41EE2A02FFCD6841FEBB134D29458A6B" box="[273,335,839,865]" captionStart="FIGURES 1 – 4" captionStartId="2.[264,382,1571,1595]" captionTargetBox="[264,1323,414,1531]" captionTargetId="figure@2.[264,1323,406,1547]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Psammocoryza vanemdeni Hogan, new genus and species, holotype, female. 1, habitus, latero-dorsal aspect (total length of specimen 3.64 mm); 2, labium, ventral aspect, showing palpi and ligula; 3, left maxilla, ventral aspect; 4, ovipositor styli, dorsal aspect, Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/174319/files/figure.png" pageId="1" pageNumber="62">Fig.1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="91CF650CFFCD6841FEA2139C2CA78ABB" box="[264,1197,918,945]" pageId="1" pageNumber="62" type="materials_examined">
<paragraph id="D96A3687FFCD6841FEA2139C2CA78ABB" blockId="1.[264,1323,918,1905]" box="[264,1197,918,945]" pageId="1" pageNumber="62">
<emphasis id="EBA1EA95FFCD6841FEA2139C29A08ABA" bold="true" box="[264,426,918,944]" pageId="1" pageNumber="62">
<typeStatus id="066E8825FFCD6841FEA2139C294F8ABA" box="[264,325,918,944]" pageId="1" pageNumber="62">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="1ED54D04FFCD6841FE18139D2B498ABB" authority="Hogan" authorityName="Hogan" box="[434,835,919,945]" class="Insecta" family="Carabidae" genus="Psammocoryza" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="vanemdeni" status="sp. nov.">
<emphasis id="EBA1EA95FFCD6841FE18139D2AE18ABA" box="[434,747,919,944]" italics="true" pageId="1" pageNumber="62">Psammocoryza vanemdeni</emphasis>
Hogan
</taxonomicName>
,
<taxonomicNameLabel id="F09257EEFFCD6841FCE5139C2BD58ABA" box="[847,991,918,944]" pageId="1" pageNumber="62" rank="species">
<emphasis id="EBA1EA95FFCD6841FCE5139C2BD58ABA" bold="true" box="[847,991,918,944]" pageId="1" pageNumber="62">new species</emphasis>
</taxonomicNameLabel>
, here designated.
</paragraph>
</subSubSection>
<subSubSection id="91CF650CFFCD6841FE9213B42CBE8CEB" pageId="1" pageNumber="62" type="diagnosis">
<paragraph id="D96A3687FFCD6841FE9213B42CBE8CEB" blockId="1.[264,1323,918,1905]" pageId="1" pageNumber="62">
<emphasis id="EBA1EA95FFCD6841FE9213B429BD8AD2" bold="true" box="[312,439,958,984]" pageId="1" pageNumber="62">Diagnosis.</emphasis>
Immediately recognised by the multiple longitudinal carinae of the pronotum, a character not seen in any other genus of
<taxonomicName id="1ED54D04FFCD6841FCDF13ED2BEB8D0B" box="[885,993,999,1025]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="tribe" tribe="Clivinini">Clivinini</taxonomicName>
. Anterior margin of clypeus trituberculate as in
<taxonomicName id="1ED54D04FFCD6841FE5014052A898D22" box="[506,643,1039,1064]" class="Insecta" family="Carabidae" genus="Halocoryza" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FE5014052A898D22" box="[506,643,1039,1064]" italics="true" pageId="1" pageNumber="62">Halocoryza</emphasis>
</taxonomicName>
or
<taxonomicName id="1ED54D04FFCD6841FD1F14052B448D22" box="[693,846,1039,1064]" class="Insecta" family="Carabidae" genus="Schizogenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FD1F14052B448D22" box="[693,846,1039,1064]" italics="true" pageId="1" pageNumber="62">Schizogenius</emphasis>
</taxonomicName>
. Clypeal suture obsolete medially and partially engraved laterally, in contrast to
<taxonomicName id="1ED54D04FFCD6841FD5B143D2B808D5A" box="[753,906,1079,1104]" class="Insecta" family="Carabidae" genus="Schizogenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FD5B143D2B808D5A" box="[753,906,1079,1104]" italics="true" pageId="1" pageNumber="62">Schizogenius</emphasis>
</taxonomicName>
where the suture is fully impressed to a greater or lesser extent and in
<taxonomicName id="1ED54D04FFCD6841FD6514552B528D72" box="[719,856,1119,1144]" class="Insecta" family="Carabidae" genus="Halocoryza" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FD6514552B528D72" box="[719,856,1119,1144]" italics="true" pageId="1" pageNumber="62">Halocoryza</emphasis>
</taxonomicName>
where it is obsolete. Eyes large and prominent, not reduced as in in
<taxonomicName id="1ED54D04FFCD6841FD3E148D2B178DAA" box="[660,797,1159,1184]" class="Insecta" family="Carabidae" genus="Halocoryza" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FD3E148D2B178DAA" box="[660,797,1159,1184]" italics="true" pageId="1" pageNumber="62">Halocoryza</emphasis>
</taxonomicName>
(in
<taxonomicName id="1ED54D04FFCD6841FCF9148D2BE68DAA" box="[851,1004,1159,1184]" class="Insecta" family="Carabidae" genus="Schizogenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FCF9148D2BE68DAA" box="[851,1004,1159,1184]" italics="true" pageId="1" pageNumber="62">Schizogenius</emphasis>
</taxonomicName>
the eyes vary in size and prominence). Head bearing multiple longitudinal carinae and sulci as in
<taxonomicName id="1ED54D04FFCD6841FBD414A52D0D8DC2" box="[1150,1287,1199,1224]" class="Insecta" family="Carabidae" genus="Halocoryza" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FBD414A52D0D8DC2" box="[1150,1287,1199,1224]" italics="true" pageId="1" pageNumber="62">Halocoryza</emphasis>
</taxonomicName>
or
<taxonomicName id="1ED54D04FFCD6841FEA214DD29AB8DFA" box="[264,417,1239,1264]" class="Insecta" family="Carabidae" genus="Schizogenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FEA214DD29AB8DFA" box="[264,417,1239,1264]" italics="true" pageId="1" pageNumber="62">Schizogenius</emphasis>
</taxonomicName>
, but unlike these two genera the median sulcus is narrower than the paramedian sulci. Gula narrow as in
<taxonomicName id="1ED54D04FFCD6841FD1114F52B5E8C12" box="[699,852,1279,1304]" class="Insecta" family="Carabidae" genus="Schizogenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FD1114F52B5E8C12" box="[699,852,1279,1304]" italics="true" pageId="1" pageNumber="62">Schizogenius</emphasis>
</taxonomicName>
, at narrowest point 0.2 times maximum width of mentum. Base of profemora with row of round tubercles, in
<taxonomicName id="1ED54D04FFCD6841FBC6152D2CFE8C4A" box="[1132,1268,1319,1344]" class="Insecta" family="Carabidae" genus="Halocoryza" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FBC6152D2CFE8C4A" box="[1132,1268,1319,1344]" italics="true" pageId="1" pageNumber="62">Halocoryza</emphasis>
</taxonomicName>
and
<taxonomicName id="1ED54D04FFCD6841FEA2154529AB8C62" box="[264,417,1359,1384]" class="Insecta" family="Carabidae" genus="Schizogenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FEA2154529AB8C62" box="[264,417,1359,1384]" italics="true" pageId="1" pageNumber="62">Schizogenius</emphasis>
</taxonomicName>
these tubercles are lacking. Protibiae similar in form to
<taxonomicName id="1ED54D04FFCD6841FB9E15452CB68C62" box="[1076,1212,1359,1384]" class="Insecta" family="Carabidae" genus="Halocoryza" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FB9E15452CB68C62" box="[1076,1212,1359,1384]" italics="true" pageId="1" pageNumber="62">Halocoryza</emphasis>
</taxonomicName>
, with the three distal spines elongate and not widened basally. Lacinia of maxilla setose along inner and outer margin as in
<taxonomicName id="1ED54D04FFCD6841FDBF15952AA48CB2" box="[533,686,1439,1464]" class="Insecta" family="Carabidae" genus="Schizogenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FDBF15952AA48CB2" box="[533,686,1439,1464]" italics="true" pageId="1" pageNumber="62">Schizogenius</emphasis>
</taxonomicName>
, not hooked at apex. Gonocoxite straight and rounded at apex, distinguished from
<taxonomicName id="1ED54D04FFCD6841FDE415CD2ADD8CEA" box="[590,727,1479,1504]" class="Insecta" family="Carabidae" genus="Halocoryza" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FDE415CD2ADD8CEA" box="[590,727,1479,1504]" italics="true" pageId="1" pageNumber="62">Halocoryza</emphasis>
</taxonomicName>
or
<taxonomicName id="1ED54D04FFCD6841FD5415CD2B9D8CEA" box="[766,919,1479,1504]" class="Insecta" family="Carabidae" genus="Schizogenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FD5415CD2B9D8CEA" box="[766,919,1479,1504]" italics="true" pageId="1" pageNumber="62">Schizogenius</emphasis>
</taxonomicName>
by the absence of setae.
</paragraph>
</subSubSection>
<subSubSection id="91CF650CFFCD6843FE9215E42B2288F4" lastPageId="3" lastPageNumber="64" pageId="1" pageNumber="62" type="etymology">
<paragraph id="D96A3687FFCD6841FE9215E42B038F8B" blockId="1.[264,1323,918,1905]" pageId="1" pageNumber="62">
<emphasis id="EBA1EA95FFCD6841FE9215E429CE8F02" bold="true" box="[312,452,1518,1544]" pageId="1" pageNumber="62">Etymology.</emphasis>
<taxonomicName id="1ED54D04FFCD6841FE7A15E52A898F02" box="[464,643,1519,1544]" class="Insecta" family="Carabidae" genus="Psammocoryza" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FE7A15E52A898F02" box="[464,643,1519,1544]" italics="true" pageId="1" pageNumber="62">Psammocoryza</emphasis>
</taxonomicName>
is derived from the existing genus group name
<taxonomicName id="1ED54D04FFCD6841FB7C15E529698F3B" authority="Putzeys" authorityName="Putzeys" class="Insecta" family="Carabidae" genus="Coryza" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FB7C15E52D218F02" box="[1238,1323,1519,1544]" italics="true" pageId="1" pageNumber="62">Coryza</emphasis>
Putzeys
</taxonomicName>
(probably a latinised form of the Greek feminine noun ‘
<emphasis id="EBA1EA95FFCD6841FC5D161D2C3D8F3A" box="[1015,1079,1559,1584]" italics="true" pageId="1" pageNumber="62">korys</emphasis>
’, meaning helmet or crest) and the Greek feminine noun ‘
<emphasis id="EBA1EA95FFCD6841FD6F16352B388F52" box="[709,818,1599,1624]" italics="true" pageId="1" pageNumber="62">psammos</emphasis>
’, meaning sand, in reference to the likely habitat of this new genus. Gender feminine.
</paragraph>
<paragraph id="D96A3687FFCD6842FE9216842A3B8854" blockId="1.[264,1323,918,1905]" lastBlockId="2.[264,1323,284,350]" lastPageId="2" lastPageNumber="63" pageId="1" pageNumber="62">
<emphasis id="EBA1EA95FFCD6841FE9216842A3E8FA2" bold="true" box="[312,564,1678,1704]" pageId="1" pageNumber="62">Taxonomic position.</emphasis>
In the absence of a comprehensive analysis of the relationships among the genera of the
<taxonomicName id="1ED54D04FFCD6841FD9916BD2A948FDB" box="[563,670,1719,1745]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="tribe" tribe="Clivinini">Clivinini</taxonomicName>
, comments on the placement of
<taxonomicName id="1ED54D04FFCD6841FB8E16BD2CDD8FDA" box="[1060,1239,1719,1744]" class="Insecta" family="Carabidae" genus="Psammocoryza" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FB8E16BD2CDD8FDA" box="[1060,1239,1719,1744]" italics="true" pageId="1" pageNumber="62">Psammocoryza</emphasis>
</taxonomicName>
within the tribe are necessarily tentative. Despite this, the multiple longitudinal carinae of the frons (‘frontal carinae’) and trituberculate clypeal margin of
<taxonomicName id="1ED54D04FFCD6841FBA2170D2CB18E2A" box="[1032,1211,1799,1824]" class="Insecta" family="Carabidae" genus="Psammocoryza" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FBA2170D2CB18E2A" box="[1032,1211,1799,1824]" italics="true" pageId="1" pageNumber="62">Psammocoryza</emphasis>
</taxonomicName>
are also shared with
<taxonomicName id="1ED54D04FFCD6841FE0A17252A238E42" box="[416,553,1839,1864]" class="Insecta" family="Carabidae" genus="Halocoryza" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FE0A17252A238E42" box="[416,553,1839,1864]" italics="true" pageId="1" pageNumber="62">Halocoryza</emphasis>
</taxonomicName>
and
<taxonomicName id="1ED54D04FFCD6841FDC017252B098E42" box="[618,771,1839,1864]" class="Insecta" family="Carabidae" genus="Schizogenius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FDC017252B098E42" box="[618,771,1839,1864]" italics="true" pageId="1" pageNumber="62">Schizogenius</emphasis>
</taxonomicName>
and these three genera probably constitute a monophyletic group. The old world genus
<taxonomicName id="1ED54D04FFCD6841FD54175D2BF78E7B" authority="Alluaud" authorityName="Alluaud" box="[766,1021,1879,1905]" class="Insecta" family="Carabidae" genus="Lophocoryza" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCD6841FD54175D2B9D8E7A" box="[766,919,1879,1904]" italics="true" pageId="1" pageNumber="62">Lophocoryza</emphasis>
Alluaud
</taxonomicName>
is also characterised by a tuberculate clypleal margin and paired carinae of the frons (
<bibRefCitation id="BD444B76FFCE6842FC4A11162CBA883C" author="Basilewsky" box="[992,1200,284,310]" pageId="2" pageNumber="68" refString="Basilewsky, P. (1973) Insectes Coleopteres Carabidae Scaritinae. Faune de Madagascar, 37, 1 - 322." type="journal article" year="1973">Basilewsky 1973</bibRefCitation>
) and may also belong in this group.
</paragraph>
<caption id="8DAA660FFFCE6842FEA216292D218F8A" httpUri="https://zenodo.org/record/174319/files/figure.png" pageId="2" pageNumber="63" targetBox="[264,1323,414,1531]" targetPageId="2">
<paragraph id="D96A3687FFCE6842FEA216292D218F8A" blockId="2.[264,1323,1571,1664]" pageId="2" pageNumber="63">
<emphasis id="EBA1EA95FFCE6842FEA2162929BD8F31" bold="true" box="[264,439,1571,1595]" pageId="2" pageNumber="63">FIGURES 1–4.</emphasis>
<taxonomicName id="1ED54D04FFCE6842FE6B162E2B3F8F31" authority="Hogan" authorityName="Hogan" box="[449,821,1572,1595]" class="Insecta" family="Carabidae" genus="Psammocoryza" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="vanemdeni" status="gen. nov.">
<emphasis id="EBA1EA95FFCE6842FE6B162E2AEB8F31" box="[449,737,1572,1595]" italics="true" pageId="2" pageNumber="63">Psammocoryza vanemdeni</emphasis>
Hogan
</taxonomicName>
,
<taxonomicNameLabel id="F09257EEFFCE6842FCE9162E2BBD8F31" box="[835,951,1572,1595]" pageId="2" pageNumber="63" rank="genus">new genus</taxonomicNameLabel>
and species, holotype, female. 1, habitus, latero-dorsal aspect (total length of specimen 3.64mm); 2, labium, ventral aspect, showing palpi and ligula; 3, left maxilla, ventral aspect; 4, ovipositor styli, dorsal aspect, Scale bar = 0.2mm.
</paragraph>
</caption>
<paragraph id="D96A3687FFCE6843FE9216A42B6788A4" blockId="2.[264,1323,1710,1896]" lastBlockId="3.[264,1322,284,510]" lastPageId="3" lastPageNumber="64" pageId="2" pageNumber="63">
The close relationship of
<taxonomicName id="1ED54D04FFCE6842FDCD16A52AFA8FC2" box="[615,752,1711,1736]" class="Insecta" family="Carabidae" genus="Halocoryza" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCE6842FDCD16A52AFA8FC2" box="[615,752,1711,1736]" italics="true" pageId="2" pageNumber="63">Halocoryza</emphasis>
</taxonomicName>
to
<taxonomicName id="1ED54D04FFCE6842FCB316A52BB88FC2" box="[793,946,1711,1736]" class="Insecta" family="Carabidae" genus="Schizogenius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCE6842FCB316A52BB88FC2" box="[793,946,1711,1736]" italics="true" pageId="2" pageNumber="63">Schizogenius</emphasis>
</taxonomicName>
has been discussed in detail by
<bibRefCitation id="BD444B76FFCE6842FEA216DC29EB8FFA" author="Whitehead" box="[264,481,1750,1776]" pageId="2" pageNumber="68" refString="Whitehead, D. R. (1966) A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera: Carabidae). Psyche, 73, 217 - 228." type="journal article" year="1966">Whitehead (1966)</bibRefCitation>
. Based on adult morphology,
<taxonomicName id="1ED54D04FFCE6842FCEA16DD2BF98FFA" box="[832,1011,1751,1776]" class="Insecta" family="Carabidae" genus="Psammocoryza" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCE6842FCEA16DD2BF98FFA" box="[832,1011,1751,1776]" italics="true" pageId="2" pageNumber="63">Psammocoryza</emphasis>
</taxonomicName>
would appear to be closer to
<taxonomicName id="1ED54D04FFCE6842FE8616F529CF8E12" box="[300,453,1791,1816]" class="Insecta" family="Carabidae" genus="Schizogenius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCE6842FE8616F529CF8E12" box="[300,453,1791,1816]" italics="true" pageId="2" pageNumber="63">Schizogenius</emphasis>
</taxonomicName>
than
<taxonomicName id="1ED54D04FFCE6842FDBA16F52A938E12" box="[528,665,1791,1816]" class="Insecta" family="Carabidae" genus="Halocoryza" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCE6842FDBA16F52A938E12" box="[528,665,1791,1816]" italics="true" pageId="2" pageNumber="63">Halocoryza</emphasis>
</taxonomicName>
. Both
<taxonomicName id="1ED54D04FFCE6842FD5916F52BAC8E12" box="[755,934,1791,1816]" class="Insecta" family="Carabidae" genus="Psammocoryza" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCE6842FD5916F52BAC8E12" box="[755,934,1791,1816]" italics="true" pageId="2" pageNumber="63">Psammocoryza</emphasis>
</taxonomicName>
and
<taxonomicName id="1ED54D04FFCE6842FC4316F52C888E12" box="[1001,1154,1791,1816]" class="Insecta" family="Carabidae" genus="Schizogenius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCE6842FC4316F52C888E12" box="[1001,1154,1791,1816]" italics="true" pageId="2" pageNumber="63">Schizogenius</emphasis>
</taxonomicName>
share similar setation of the lacinia, the clypeal suture is at least partially engraved and the gula narrow.
<taxonomicName id="1ED54D04FFCE6842FEA2174529B18E62" box="[264,443,1871,1896]" class="Insecta" family="Carabidae" genus="Psammocoryza" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCE6842FEA2174529B18E62" box="[264,443,1871,1896]" italics="true" pageId="2" pageNumber="63">Psammocoryza</emphasis>
</taxonomicName>
differs from
<taxonomicName id="1ED54D04FFCE6842FDC517452B028E62" box="[623,776,1871,1896]" class="Insecta" family="Carabidae" genus="Schizogenius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCE6842FDC517452B028E62" box="[623,776,1871,1896]" italics="true" pageId="2" pageNumber="63">Schizogenius</emphasis>
</taxonomicName>
in the arrangement of the frontal carinae, macrosculpture of the pronotum, tuberculate femora and the shape and setation of the gonocoxite. These two genera also show a difference in habitat preference. All
<taxonomicName id="1ED54D04FFCF6843FEA2116629AB888F" box="[264,417,364,389]" class="Insecta" family="Carabidae" genus="Schizogenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCF6843FEA2116629AB888F" box="[264,417,364,389]" italics="true" pageId="3" pageNumber="64">Schizogenius</emphasis>
</taxonomicName>
occupy inland freshwater habitats while the two known specimens of
<taxonomicName id="1ED54D04FFCF6843FEA2119E29B188A7" box="[264,443,404,429]" class="Insecta" family="Carabidae" genus="Psammocoryza" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCF6843FEA2119E29B188A7" box="[264,443,404,429]" italics="true" pageId="3" pageNumber="64">Psammocoryza</emphasis>
</taxonomicName>
were collected from a coastal beach.
</paragraph>
<paragraph id="D96A3687FFCF6843FE9211B62B2288F4" blockId="3.[264,1322,284,510]" pageId="3" pageNumber="64">
Diagnostic characters of
<taxonomicName id="1ED54D04FFCF6843FDF411B62AED88DF" box="[606,743,444,469]" class="Insecta" family="Carabidae" genus="Halocoryza" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCF6843FDF411B62AED88DF" box="[606,743,444,469]" italics="true" pageId="3" pageNumber="64">Halocoryza</emphasis>
</taxonomicName>
and
<taxonomicName id="1ED54D04FFCF6843FC8811B62BB188DF" box="[802,955,444,469]" class="Insecta" family="Carabidae" genus="Schizogenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCF6843FC8811B62BB188DF" box="[802,955,444,469]" italics="true" pageId="3" pageNumber="64">Schizogenius</emphasis>
</taxonomicName>
(from
<bibRefCitation id="BD444B76FFCF6843FBA411B62CD388DC" author="Whitehead" box="[1038,1241,444,470]" pageId="3" pageNumber="68" refString="Whitehead, D. R. (1966) A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera: Carabidae). Psyche, 73, 217 - 228." type="journal article" year="1966">Whitehead, 1966</bibRefCitation>
,
<bibRefCitation id="BD444B76FFCF6843FB4C11B62D2888DC" author="Whitehead" box="[1254,1314,444,470]" pageId="3" pageNumber="68" refString="Whitehead, D. R. (1972) Classification, phylogeny and zoogeography of Schizogenius Putzeys (Coleoptera: Carabidae: Scaritini). Quaestiones Entomologicae, 8, 131 - 348." type="journal article" year="1972">1972</bibRefCitation>
) and
<taxonomicName id="1ED54D04FFCF6843FE9011EE29E788F7" box="[314,493,484,509]" class="Insecta" family="Carabidae" genus="Psammocoryza" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="EBA1EA95FFCF6843FE9011EE29E788F7" box="[314,493,484,509]" italics="true" pageId="3" pageNumber="64">Psammocoryza</emphasis>
</taxonomicName>
are summarised in table 1.
</paragraph>
</subSubSection>
</treatment>
</document>
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