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<mods:title id="00086CA5860757576675C79D3624CA2D">On the synonymy of two Acantholycosa species (Araneae, Lycosidae) from the Altai</mods:title>
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<mods:namePart id="938792C1E44340106251C033213AFAD1">Fomichev, Alexander A.</mods:namePart>
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<mods:namePart id="F36FC0B2C328EDE7AA129149FD114340">Marusik, Yuri M.</mods:namePart>
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<mods:namePart id="1833958BDCFF27C7CCFC5EA14B179669">Koponen, Seppo</mods:namePart>
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<treatment id="5174476420361CDDEC3C2E345A7ABF2B" ID-GBIF-Taxon="127870891" LSID="urn:lsid:plazi:treatment:5174476420361CDDEC3C2E345A7ABF2B" httpUri="http://treatment.plazi.org/id/5174476420361CDDEC3C2E345A7ABF2B" lastPageId="3" lastPageNumber="154" pageId="1" pageNumber="152">
<subSubSection id="82D210FB07D309496399E05E114F24E7" pageId="1" pageNumber="152" type="multiple">
<paragraph id="97EA65CE8BE579371033315673BCA0B6" pageId="1" pageNumber="152">Taxon classification Animalia Araneae Lycosidae</paragraph>
</subSubSection>
<subSubSection id="55990DBA683FCD23F80B4B6CD59C28BF" pageId="1" pageNumber="152" type="nomenclature">
<paragraph id="14FDFA1C3183BF51E05597F112479C8C" pageId="1" pageNumber="152">
<taxonomicName id="DB9EDBD9BE5CD01516D17E2150E32970" ID-CoL="64GCH" authority="Marusik, Azarkina &amp; Koponen, 2004" authorityName="Marusik, Azarkina &amp; Koponen" authorityYear="2004" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acantholycosa katunensis" order="Araneae" pageId="1" pageNumber="152" phylum="Arthropoda" rank="species" species="katunensis">Acantholycosa katunensis Marusik, Azarkina &amp; Koponen, 2004</taxonomicName>
Figs 1-2, 3-10
</paragraph>
</subSubSection>
<subSubSection id="04DD06DD5AFB54CF03749B3D6753D5DE" pageId="1" pageNumber="152" type="reference_group">
<paragraph id="1C4BB638EB0B6DE61455AFC33DB9871E" pageId="1" pageNumber="152">
<taxonomicName id="89F2D50CC58736333B18260845117AFB" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acantholycosa katunensis" order="Araneae" pageId="1" pageNumber="152" phylum="Arthropoda" rank="species" species="katunensis">Acantholycosa katunensis</taxonomicName>
Marusik, Azarkina &amp; Koponen, 2004: 107, figs 21-23 (♂).
</paragraph>
<paragraph id="3A1118B1F349E540368DDD6D98CA978B" pageId="1" pageNumber="152">
<taxonomicName id="F83C4C71C6478A95B5043ACF56DA9C08" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acantholycosa kurchumensis" order="Araneae" pageId="1" pageNumber="152" phylum="Arthropoda" rank="species" species="kurchumensis">Acantholycosa kurchumensis</taxonomicName>
Marusik, Azarkina &amp; Koponen, 2004: 119, figs 82-83 (♀). Syn. n.
</paragraph>
</subSubSection>
<subSubSection id="B98C723E8BD368E64330CF748BE54FBF" pageId="1" pageNumber="152" type="materials_examined">
<paragraph id="AFBCD1C1DD78466FC1248BB8487A8CA7" pageId="1" pageNumber="152">Material examined.</paragraph>
<paragraph id="1C3914A9231D3ED09A3B2999D24087D0" pageId="1" pageNumber="152">
RUSSIA, Altai Republic, Kosh-Agach District: 3♂ (ISEA), Karagemskyi Mt. Range (
<geoCoordinate id="AE779206CF68B1AB40055FBD934F7AF5" direction="north" orientation="latitude" precision="925" value="49.866665">49°52'N</geoCoordinate>
;
<geoCoordinate id="71FCAEBA4F1B33360AAD63B4348387C3" direction="east" orientation="longitude" precision="925" value="87.11667">87°07'E</geoCoordinate>
), 2500-2900 m,
<normalizedToken id="27181C964500526A5AF0F95284D0EA8F" originalValue="“kurums”">&quot;kurums&quot;</normalizedToken>
(=scree formed by huge boulders) and alpine meadow, 27.06.2014 (A.A. Fomichev); 1♀ (ISEA), Karagem River valley (
<geoCoordinate id="1E350D0C6712B83C84FBC2A8F2E6E2ED" direction="north" orientation="latitude" precision="925" value="49.883335">49°53'N</geoCoordinate>
;
<geoCoordinate id="E652489491B238A837B94A946C1204AF" direction="east" orientation="longitude" precision="925" value="87.183334">87°11'E</geoCoordinate>
), 1360 m, stony steppe slope, 28.06.2014 (A.A. Fomichev); KAZAKHSTAN, East Kazakhstan Area: 1♂ (holotype of
<taxonomicName id="D9FD0F9820070D1A4D1F4C7B879675AB" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acantholycosa katunensis" order="Araneae" pageId="1" pageNumber="152" phylum="Arthropoda" rank="species" species="katunensis">Acantholycosa katunensis</taxonomicName>
) (ISEA) South Altai, south part of
<normalizedToken id="C98C73333812490A4C2532527628BD09" originalValue="Katun’">Katun'</normalizedToken>
Mt. Range, 5 km SE of Rakhmanovskiye Klyuchi (=Springs), 2100-2500 m, alpine zone, 26.06.1997 (R.Yu. Dudko and V.K. Zinchenko); 1♀ (holotype of
<taxonomicName id="D9800D7DBABF8A78270BBEF8D904C9C2" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acantholycosa kurchumensis" order="Araneae" pageId="1" pageNumber="152" phylum="Arthropoda" rank="species" species="kurchumensis">Acantholycosa kurchumensis</taxonomicName>
) (ISEA) Kurchum Mt. Range, Kurchum River, upper flow, 23.08.1990 (V.K. Zinchenko).
</paragraph>
</subSubSection>
<subSubSection id="F83DB4A6141A626ABB8270F14A925D73" lastPageId="2" lastPageNumber="153" pageId="1" pageNumber="152" type="diagnosis">
<paragraph id="65C459F355E53F20A7C8F49572FFD6A8" pageId="1" pageNumber="152">Diagnosis.</paragraph>
<paragraph id="6A6FB03EE66453B4AEC956AE3EC84568" lastPageId="2" lastPageNumber="153" pageId="1" pageNumber="152">
<taxonomicName id="2E9511A81DA80B47EC90436EF2E97BC0" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acantholycosa katunensis" order="Araneae" pageId="1" pageNumber="152" phylum="Arthropoda" rank="species" species="katunensis">Acantholycosa katunensis</taxonomicName>
is most similar to
<taxonomicName id="6EF6DCF32935C585790F4D85CDD7C6C0" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acantholycosa dudkorum" order="Araneae" pageId="1" pageNumber="152" phylum="Arthropoda" rank="species" species="dudkorum">Acantholycosa dudkorum</taxonomicName>
Marusik, Azarkina &amp; Koponen, 2004 by having a similarly shaped tegular apophysis that lacks an apical arm, a similar conductor, a wide apical pocket and a thin septum. The two species can be separated by the shape of the embolus, which tapers toward the tip in
<taxonomicName id="D6037C8464808459866BCBD2CD815CF3" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acantholycosa katunensis" order="Araneae" pageId="1" pageNumber="152" phylum="Arthropoda" rank="species" species="katunensis">Acantholycosa katunensis</taxonomicName>
(Figs 3-5) and widens in
<taxonomicName id="5AA108A6602676A334D2E18DA952D559" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acantholycosa dudkorum" order="Araneae" pageId="1" pageNumber="152" phylum="Arthropoda" rank="species" species="dudkorum">Acantholycosa dudkorum</taxonomicName>
(cf.
<bibRefCitation id="6D03DAC600C427378CD79229A991E96A" author="Marusik, YM" journalOrPublisher="Arthropoda Selecta" pageId="4" pageNumber="155" pagination="101 - 148" title="A survey of east Palaearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera." volume="12" year="2004">Marusik et al. 2004</bibRefCitation>
: figs 73, 75); the shape of
<pageBreakToken id="972E6F09F4A6EAB851F469E73B131F08" pageId="2" pageNumber="153" start="start">the</pageBreakToken>
palea: in
<taxonomicName id="D5C8586199354F0B706BE646C20C5DBA" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acantholycosa dudkorum" order="Araneae" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="dudkorum">Acantholycosa dudkorum</taxonomicName>
(cf.
<bibRefCitation id="8CFF84B53B0D8EA5F41233D59CD12671" author="Marusik, YM" journalOrPublisher="Arthropoda Selecta" pageId="4" pageNumber="155" pagination="101 - 148" title="A survey of east Palaearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera." volume="12" year="2004">Marusik et al. 2004</bibRefCitation>
: figs 73, 75), the paleal process is thin with a hollow on the prolateral side, and it is unmodified in
<taxonomicName id="554893FD2D7C9732A23810E4F3782B38" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acantholycosa katunensis" order="Araneae" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="katunensis">Acantholycosa katunensis</taxonomicName>
(Figs 4-5, 9). Females of the two species can be distinguished by the shape of septum: long (starting from the pocket) with a subparallel base in
<taxonomicName id="7EBA52409BBAAD0225FAF1F548717F55" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acantholycosa katunensis" order="Araneae" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="katunensis">Acantholycosa katunensis</taxonomicName>
and short (starting from fovea) and widened at the base in
<taxonomicName id="13CCD912ACE381C49DA6031FC8A3C090" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acantholycosa dudkorum" order="Araneae" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="dudkorum">Acantholycosa dudkorum</taxonomicName>
(cf.
<bibRefCitation id="BA1170D809810A8EDD79AFDCC58B57E8" author="Marusik, YM" journalOrPublisher="Arthropoda Selecta" pageId="4" pageNumber="155" pagination="101 - 148" title="A survey of east Palaearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera." volume="12" year="2004">Marusik et al. 2004</bibRefCitation>
: fig. 78).
</paragraph>
<caption id="D194B9602F6BCABDF64B784D8C83A0EE" pageId="2" pageNumber="153">
<paragraph id="61E79F20DAAF33C5D7CA92637CAF46A2" pageId="2" pageNumber="153">
Figures 1-2. Habitus of
<taxonomicName id="ACA057C253194BEB5282C9DFE9FA96E8" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acantholycosa katunensis" order="Araneae" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="katunensis">Acantholycosa katunensis</taxonomicName>
. 1 female 2 male. Scale: 2 mm.
</paragraph>
</caption>
<caption id="07C7DB2F60BFA16C067752C4D2A0A664" pageId="2" pageNumber="153">
<paragraph id="E5C7D13C82568A07EFBB2F581E55DE54" pageId="2" pageNumber="153">
Figures 3-10. Copulatory organs of
<taxonomicName id="058BDB47178B99102660D079C6030872" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acantholycosa katunensis" order="Araneae" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="katunensis">Acantholycosa katunensis</taxonomicName>
. 3 male palp, ventral 4-5 embolic division, ventral and anterior 6-7 epigyne, ventral and dorsal (left head of receptacle is broken) 8-10 bulb, ventral, anterior and retrolateral. Scale: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="17A41008EE10BCA5E06CC6AE60B024A8" pageId="2" pageNumber="153" type="description">
<paragraph id="1520E6C3E9DFB9E2C1561C52DFCE86E5" pageId="2" pageNumber="153">Description.</paragraph>
<paragraph id="4A75D9773028C7FE9FAC45316CE91C18" pageId="2" pageNumber="153">
See
<bibRefCitation id="2AA12A7390C14543E6CE7F0B0691BEFF" author="Marusik, YM" journalOrPublisher="Arthropoda Selecta" pageId="4" pageNumber="155" pagination="101 - 148" title="A survey of east Palaearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera." volume="12" year="2004">Marusik et al. (2004)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0ABBC11098F14BC31BA5B903EFB89E5A" lastPageId="3" lastPageNumber="154" pageId="2" pageNumber="153" type="distribution">
<paragraph id="193325CC2B056F0CAC4B7735FC489FBB" pageId="2" pageNumber="153">Distribution.</paragraph>
<paragraph id="E92A029E57308D53424D0D017596504A" lastPageId="3" lastPageNumber="154" pageId="2" pageNumber="153">
So far, this species is known from three localities. The most distant localities, the upper reaches of the Kurchum River (locality 1, Map 1) and the Karagemskyi Mt Range (locality 3) are approximately 190 km apart (Map 1). Rakhmanovskiye Klyuchi Village (locality 2), the type locality of
<taxonomicName id="BED89C7F72C27909647C4310339CC873" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acantholycosa katunensis" order="Araneae" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="katunensis">Acantholycosa katunensis</taxonomicName>
, is located between two extreme distribution records (Map 1). Listings for the species by Platnick
<pageBreakToken id="63D08CC750601F595DFFF629032BB0AB" pageId="3" pageNumber="154" start="start">(</pageBreakToken>
2004-2014) and the
<bibRefCitation id="43756D38B7CEBBCA0AEAD2B3EF40885D" author="Platnick, NI" journalOrPublisher="Natural History Museum Bern" pageId="4" pageNumber="155" title="World Spider Catalog" url="http://wsc.nmbe.ch" year="2015">World Spider Catalog (2015)</bibRefCitation>
for Russia were erroneous based on wrong country data provided by
<bibRefCitation id="C2615018035E678B25A515225FA2918E" author="Marusik, YM" journalOrPublisher="Arthropoda Selecta" pageId="4" pageNumber="155" pagination="101 - 148" title="A survey of east Palaearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera." volume="12" year="2004">Marusik et al. (2004)</bibRefCitation>
. Type locality of
<taxonomicName id="F2742DBF36ECE7A6F967C92B6AEEB19F" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acantholycosa katunensis" order="Araneae" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="katunensis">Acantholycosa katunensis</taxonomicName>
was mistakenly assigned to Russia (Altai) by
<bibRefCitation id="616C1E470E93FC0E9006BED8A0BE2FA4" author="Marusik, YM" journalOrPublisher="Arthropoda Selecta" pageId="4" pageNumber="155" pagination="101 - 148" title="A survey of east Palaearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera." volume="12" year="2004">Marusik et al (2004)</bibRefCitation>
instead of Kazakhstan (East Kazakhstan Area). The new record from the Karagemskyi Mt Range is the first locality record of this species in Russia.
</paragraph>
<caption id="6D91D31CD951FBCC824D95413044FA30" pageId="3" pageNumber="154">
<paragraph id="4AE9D8D984178C3D8B2CD56089FBA95A" pageId="3" pageNumber="154">
Map 1. Distribution records of
<taxonomicName id="4C8DEE4D54B5CE0C16B6782C25EF3A1F" class="Arachnida" family="Lycosidae" genus="Acantholycosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acantholycosa katunensis" order="Araneae" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="katunensis">Acantholycosa katunensis</taxonomicName>
. 1 Kurchum River 2 Rakhmanovskiye Klyuchi Village 3 Karagemskyi Mt. Range.
</paragraph>
</caption>
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</treatment>
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