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<document id="76F6FD7A221AA758FBE33399B08820E6" ID-CLB-Dataset="21948" ID-DOI="10.5852/ejt.2017.376" ID-GBIF-Dataset="4298f5cf-d85e-48fb-afe8-f12c99ad65b3" ID-ISSN="2118-9773" ID-Zenodo-Dep="3838544" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590088443336" checkinUser="carolina" docAuthor="Aharon, Shlomi, Huber, Bernhard A. &amp; Gavish-Regev, Efrat" docDate="2017" docId="516F87BB1327BC5CFD55BFACFD4FEF26" docLanguage="en" docName="ejt-2017-376.pdf.imf" docOrigin="European Journal of Taxonomy 376 (376)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Artema Walckenaer 1837" docType="treatment" docVersion="9" lastPageNumber="44" masterDocId="AD56FFC3130CBC77FFD0BC5EFF85E94D" masterDocTitle="Daddy-long-leg giants: revision of the spider genus Artema Walckenaer, 1837 (Araneae, Pholcidae)" masterLastPageNumber="57" masterPageNumber="1" pageNumber="44" updateTime="1698840747163" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="CCA6E3EA8EE0877FC4576F5706D16327">Daddy-long-leg giants: revision of the spider genus Artema Walckenaer, 1837 (Araneae, Pholcidae)</mods:title>
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<mods:namePart id="50FD56EFC7148F2B42389730B2E2CC83">Aharon, Shlomi</mods:namePart>
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<mods:affiliation id="772D30E56E0B54962ECE4C91F841CDD5">Blaustein Institutes for Desert Research, Ben-Gurion University of the Negev, Sede Boqer Campus, Midreshet Ben-Gurion, Israel. Alexander Koenig Research Museum of Zoology, Adenauerallee 160, 53113 Bonn, Germany. The Arachnid National Natural History Collection, The Hebrew University of Jerusalem, Edmond J. Safra Campus, Givat Ram, Jerusalem, Israel.</mods:affiliation>
<mods:nameIdentifier id="31763566C2754A1944D6263574D940BC" type="email">shlomi.aharon@gmail.com</mods:nameIdentifier>
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<mods:namePart id="FEF5F39457DB3D1E12E4C75FBEDD39F0">Huber, Bernhard A.</mods:namePart>
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<mods:affiliation id="F01740740A0A37BF17C3CF907A9795B6">Alexander Koenig Research Museum of Zoology, Adenauerallee 160, 53113 Bonn, Germany. &amp; Email: b. huber @ zfmk. de &amp; urn: lsid: zoobank. org: author: 33607 F 65 - 19 BF- 4 DC 9 - 94 FD- 4 BB 88 CED 455 F</mods:affiliation>
<mods:nameIdentifier id="177FFC4D8187BBC70F5D527F4F627D6B" type="email">b.huber@zfmk.de</mods:nameIdentifier>
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<mods:namePart id="78FCC311C08A27A90E90645546AB9C3F">Gavish-Regev, Efrat</mods:namePart>
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<mods:affiliation id="45A79223076D31C6D2F95FAB2C00BA1D">The Arachnid National Natural History Collection, The Hebrew University of Jerusalem, Edmond J. Safra Campus, Givat Ram, Jerusalem, Israel. &amp; urn: lsid: zoobank. org: author: FC 073 F 19 - 2202 - 4 C 89 - 8 B 43 - CEA 4 CC 5 E 2 D 50 &amp; Corresponding author: efrat. gavish-regev @ mail. huji. ac. il</mods:affiliation>
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<mods:title id="5950FA07E56F08A813D9C80117A0E289">European Journal of Taxonomy</mods:title>
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<mods:date id="9AD185C08960998F3F0574E4BA2C36C4">2017</mods:date>
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<treatment id="516F87BB1327BC5CFD55BFACFD4FEF26" ID-DOI="http://doi.org/10.5281/zenodo.3852328" ID-GBIF-Taxon="164246579" ID-Zenodo-Dep="3852328" LSID="urn:lsid:plazi:treatment:516F87BB1327BC5CFD55BFACFD4FEF26" httpUri="http://treatment.plazi.org/id/516F87BB1327BC5CFD55BFACFD4FEF26" lastPageNumber="44" pageId="43" pageNumber="44">
<subSubSection id="91DC65261327BC5CFD55BFACFC2AED41" box="[645,943,1010,1036]" pageId="43" pageNumber="44" type="nomenclature">
<paragraph id="D97936AD1327BC5CFD55BFACFC2AED41" blockId="43.[645,943,1010,1071]" box="[645,943,1010,1036]" pageId="43" pageNumber="44">
<heading id="823181C11327BC5CFD55BFACFC2AED41" box="[645,943,1010,1036]" centered="true" fontSize="11" level="2" pageId="43" pageNumber="44" reason="2">
<emphasis id="EBB2EABF1327BC5CFD55BFACFC9AED41" bold="true" box="[645,799,1010,1036]" pageId="43" pageNumber="44">
<taxonomicName id="1EC64D2E1327BC5CFD55BFACFD5AED41" ID-CoL="62CGD" ID-ENA="280364" authorityName="Walckenaer" authorityYear="1837" box="[645,735,1010,1036]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="EBB2EABF1327BC5CFD55BFACFD5AED41" bold="true" box="[645,735,1010,1036]" italics="true" pageId="43" pageNumber="44">Artema</emphasis>
</taxonomicName>
sp. a
</emphasis>
from
<collectingCountry id="A1D1763D1327BC5CFCB7BFACFC2AED41" box="[871,943,1010,1036]" name="Oman" pageId="43" pageNumber="44">Oman</collectingCountry>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="91DC65261327BC5CFD39B84BFCCEED62" box="[745,843,1045,1071]" pageId="43" pageNumber="44" type="description">
<paragraph id="D97936AD1327BC5CFD39B84BFCCEED62" blockId="43.[645,943,1010,1071]" box="[745,843,1045,1071]" pageId="43" pageNumber="44">
<figureCitation id="41FD2A281327BC5CFD39B84BFCCEED62" box="[745,843,1045,1071]" captionStart="Figs 170179. 170" captionStartId="44.[189,241,1718,1744]" captionTargetBox="[198,1391,246,1677]" captionTargetId="figure@44.[189,1399,231,1693]" captionTargetPageId="44" captionText="Figs 170179. 170. Artema sp. a from Oman Jebal Akhdar, E of Al Hamra, Al Ghubrat (NHMW), epigynum, ventral view. 171179. Artema sp. b from Algeria, El Goléa (NHMW). 171173. Male left pedipalp: prolateral, dorsal, and retrolateral views. 174176. Male left bulb and procursus: prolateral, dorsal, and retrolateral views. 177179. Male chelicerae: lateral and frontal views. Scale lines: 170, 174179 = 0.5 mm; 171173 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838592" httpUri="https://zenodo.org/record/3838592/files/figure.png" pageId="43" pageNumber="44">Fig. 170</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="91DC65261327BC5CFF6DB809FDEFED8E" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="D97936AD1327BC5CFF6DB809FE2FED3C" blockId="43.[189,426,1111,1137]" box="[189,426,1111,1137]" pageId="43" pageNumber="44">
<heading id="823181C11327BC5CFF6DB809FE2FED3C" bold="true" box="[189,426,1111,1137]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="EBB2EABF1327BC5CFF6DB809FE2FED3C" bold="true" box="[189,426,1111,1137]" pageId="43" pageNumber="44">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="D97936AD1327BC5CFF6DB8D8FDEFED8E" blockId="43.[189,1398,1157,1219]" pageId="43" pageNumber="44">
<materialsCitation id="69AE3CF01327BC5CFF6DB8D8FDD8ED8E" ID-GBIF-Occurrence="3763211420" collectingDate="2001-11" collectionCode="NHMW" country="Oman" latitude="23.1" location="Al Hamra" longLatPrecision="7548" longitude="57.4" municipality="Jebal Akhdar" pageId="43" pageNumber="44" specimenCount="1" specimenCount-female="1">
<collectingCountry id="A1D1763D1327BC5CFF6DB8D8FE9EEDED" box="[189,283,1158,1184]" name="Oman" pageId="43" pageNumber="44">OMAN</collectingCountry>
:
<specimenCount id="CFC0FD241327BC5CFEF6B8D8FED6EDED" box="[294,339,1158,1184]" pageId="43" pageNumber="44" type="female">1 ♀</specimenCount>
,
<collectingMunicipality id="391DACD71327BC5CFEB2B8DBFE7BEDD2" box="[354,510,1157,1183]" pageId="43" pageNumber="44">Jebal Akhdar</collectingMunicipality>
, E of
<location id="DC1960761327BC5CFD91B8DBFD3CEDED" LSID="urn:lsid:plazi:treatment:516F87BB1327BC5CFD55BFACFD4FEF26:DC1960761327BC5CFD91B8DBFD3CEDED" box="[577,697,1157,1184]" country="Oman" latitude="23.1" longLatPrecision="7548" longitude="57.4" municipality="Jebal Akhdar" name="Al Hamra" pageId="43" pageNumber="44">Al Hamra</location>
,
<location id="DC1960761327BC5CFD15B8DBFCD0EDD2" LSID="urn:lsid:plazi:treatment:516F87BB1327BC5CFD55BFACFD4FEF26:DC1960761327BC5CFD15B8DBFCD0EDD2" box="[709,853,1157,1183]" country="Oman" latitude="23.1" longLatPrecision="7548" longitude="57.4" municipality="Jebal Akhdar" name="Wadi Fallah" pageId="43" pageNumber="44">Wadi Fallah</location>
,
<location id="DC1960761327BC5CFCB0B8DBFC60EDD2" LSID="urn:lsid:plazi:treatment:516F87BB1327BC5CFD55BFACFD4FEF26:DC1960761327BC5CFCB0B8DBFC60EDD2" box="[864,997,1157,1183]" country="Oman" latitude="23.1" longLatPrecision="7548" longitude="57.4" municipality="Jebal Akhdar" name="Al Ghubrat" pageId="43" pageNumber="44">Al Ghubrat</location>
[
<geoCoordinate id="BCF2506A1327BC5CFC24B8D8FBD0EDED" box="[1012,1109,1158,1184]" degrees="23.1" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="5555" value="23.1">23.1° N</geoCoordinate>
,
<geoCoordinate id="BCF2506A1327BC5CFBB1B8DBFB3BEDED" box="[1121,1214,1157,1184]" degrees="57.4" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="5555" value="57.4">57.4° E</geoCoordinate>
], “
<location id="DC1960761327BC5CFB0DB8DBFAEDEDD2" LSID="urn:lsid:plazi:treatment:516F87BB1327BC5CFD55BFACFD4FEF26:DC1960761327BC5CFB0DB8DBFAEDEDD2" box="[1245,1384,1157,1183]" country="Oman" latitude="23.1" longLatPrecision="7548" longitude="57.4" municipality="Jebal Akhdar" name="Quellhohle" pageId="43" pageNumber="44">Quellhöhle</location>
”,
<date id="AD78106D1327BC5CFF6DB8F7FEBCED8E" box="[189,313,1193,1219]" pageId="43" pageNumber="44" value="2001-11">
<collectingDate id="BD3CE9851327BC5CFF6DB8F7FEBCED8E" box="[189,313,1193,1219]" pageId="43" pageNumber="44" value="2001-11">Nov. 2001</collectingDate>
</date>
,
<location id="DC1960761327BC5CFE96B8F7FE30ED8E" LSID="urn:lsid:plazi:treatment:516F87BB1327BC5CFD55BFACFD4FEF26:DC1960761327BC5CFE96B8F7FE30ED8E" box="[326,437,1193,1219]" country="Oman" latitude="23.1" longLatPrecision="7548" longitude="57.4" municipality="Jebal Akhdar" name="Sattmann" pageId="43" pageNumber="44">Sattmann</location>
leg. (
<collectionCode id="BFD7AE681327BC5CFE26B8F7FDD8ED8E" box="[502,605,1193,1219]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="43" pageNumber="44" type="Museum">NHMW</collectionCode>
</materialsCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="91DC65261327BC5CFF6DB8B4FD4FEF26" pageId="43" pageNumber="44" type="discussion">
<paragraph id="D97936AD1327BC5CFF6DB8B4FE86EC49" blockId="43.[189,259,1258,1284]" box="[189,259,1258,1284]" pageId="43" pageNumber="44">
<heading id="823181C11327BC5CFF6DB8B4FE86EC49" bold="true" box="[189,259,1258,1284]" fontSize="11" level="3" pageId="43" pageNumber="44" reason="3">
<emphasis id="EBB2EABF1327BC5CFF6DB8B4FE86EC49" bold="true" box="[189,259,1258,1284]" pageId="43" pageNumber="44">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="D97936AD1327BC5CFF6DB947FCC6EC8D" blockId="43.[189,1398,1305,1473]" pageId="43" pageNumber="44">
Female distinguished from all other
<taxonomicName id="1EC64D2E1327BC5CFD59B947FD5AEC7E" authorityName="Walckenaer" authorityYear="1837" box="[649,735,1305,1331]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="EBB2EABF1327BC5CFD59B947FD5AEC7E" box="[649,735,1305,1331]" italics="true" pageId="43" pageNumber="44">Artema</emphasis>
</taxonomicName>
females by simple rectangular epigynum (
<figureCitation id="41FD2A281327BC5CFB2CB947FAECEC7E" box="[1276,1385,1305,1331]" captionStart="Figs 170179. 170" captionStartId="44.[189,241,1718,1744]" captionTargetBox="[198,1391,246,1677]" captionTargetId="figure@44.[189,1399,231,1693]" captionTargetPageId="44" captionText="Figs 170179. 170. Artema sp. a from Oman Jebal Akhdar, E of Al Hamra, Al Ghubrat (NHMW), epigynum, ventral view. 171179. Artema sp. b from Algeria, El Goléa (NHMW). 171173. Male left pedipalp: prolateral, dorsal, and retrolateral views. 174176. Male left bulb and procursus: prolateral, dorsal, and retrolateral views. 177179. Male chelicerae: lateral and frontal views. Scale lines: 170, 174179 = 0.5 mm; 171173 = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3838592" httpUri="https://zenodo.org/record/3838592/files/figure.png" pageId="43" pageNumber="44">Fig. 170</figureCitation>
). Stridulatory files laterally on chelicerae present. Epigynum almost twice as wide as long, posterior margin almost straight, epigynal plate consisting of two gently sclerotized lateral areas that appear swollen posteriorly, pale median area wider anteriorly, dark median sclerite absent. Pair of projections anterior to epigynum (AEP) present but not prominent.
</paragraph>
<paragraph id="D97936AD1327BC5CFF6DB9B6FD4FEF26" blockId="43.[189,1399,1512,1644]" pageId="43" pageNumber="44">
Female genitalia in the genus
<taxonomicName id="1EC64D2E1327BC5CFDFCB9B6FD07EF4F" authorityName="Walckenaer" authorityYear="1837" box="[556,642,1512,1538]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="EBB2EABF1327BC5CFDFCB9B6FD07EF4F" box="[556,642,1512,1538]" italics="true" pageId="43" pageNumber="44">Artema</emphasis>
</taxonomicName>
often show high intra-specific variation, e.g., as in
<taxonomicName id="1EC64D2E1327BC5CFB29B9B6FAF2EF4F" authority="Aharon &amp; Huber &amp; Gavish-Regev, 2017" authorityName="Aharon &amp; Huber &amp; Gavish-Regev" authorityYear="2017" box="[1273,1399,1512,1538]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="nephilit" status="sp. nov.">
<emphasis id="EBB2EABF1327BC5CFB29B9B6FAF2EF4F" box="[1273,1399,1512,1538]" italics="true" pageId="43" pageNumber="44">A. nephilit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F08157C41327BC5CFF6DBA55FE9DEF68" box="[189,280,1547,1573]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="41FD2A281327BC5CFEF7BA55FE2BEF6B" box="[295,430,1547,1574]" captionStart="Figs 4554" captionStartId="20.[190,242,1857,1883]" captionTargetBox="[197,1393,889,1786]" captionTargetId="figure@20.[189,1399,880,1801]" captionTargetPageId="20" captionText="Figs 4554. Artema nephilit sp. nov. 4550. Epigyna, ventral views, females from Berniki caves (45, 49: HUJ Ara 15996-7),En Tamar (46: HUJ Ara 16019), Samar (47: HUJ Ara 16020), Petra (48: ZFMK Ar 15227) and HaTanur Waterfall (50: CSZ). 5154. Female abdomens, dorsal and lateral views, from Petra (5152: ZFMK Ar 15227), and En Tamar (5354: HUJ Ara 16019). Figs 45-47, 49-50, 53-54 Israel; 48, 51-52 Jordan. Scale lines: 4550, 54 = 1 mm; 5153 = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3838560" httpUri="https://zenodo.org/record/3838560/files/figure.png" pageId="43" pageNumber="44">Figs 4554</figureCitation>
) and in
<taxonomicName id="1EC64D2E1327BC5CFDDDBA55FDFEEF68" authorityName="Thorell" authorityYear="1881" box="[525,635,1547,1573]" class="Arachnida" family="Pholcidae" genus="Artema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="doriae">
<emphasis id="EBB2EABF1327BC5CFDDDBA55FDFEEF68" box="[525,635,1547,1573]" italics="true" pageId="43" pageNumber="44">A. doriae</emphasis>
</taxonomicName>
(
<figureCitation id="41FD2A281327BC5CFD5ABA55FCB5EF68" box="[650,816,1547,1573]" captionStart="Figs 103114" captionStartId="32.[189,241,1822,1848]" captionTargetBox="[197,1391,962,1774]" captionTargetId="figure@32.[189,1399,954,1783]" captionTargetPageId="32" captionText="Figs 103114. Artema doriae Thorell, 1881, intraspecific variation of epigynum. 103. Iran, Tochal (ZFMK Ar 15233). 104. Iran, Esfahan (FMNH). 105. Iran, Lorestan, Maamulan (MHNG). 106. Iran, Tehran (MNHN AR 10180). 107. Iran, 10 km NE of Bafq (SMF). 108109. Turkey, Urfa-Gaziantep (XUM). 110. Turkey, Diyarbakır Prov., Çermik (ZFMK Ar 15931). 111. Israel, Upper Galilee, Ramot Naftali (CSZ). 112. Israel, Jericho (HUJ Ara 16035). 113. Israel, Memorial Ha-Biqa, NE Pezael (HUJ Ara 16033). 114. Israel, Modiin (SMNH). Scale lines: 103, 105113 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3838574" httpUri="https://zenodo.org/record/3838574/files/figure.png" pageId="43" pageNumber="44">Figs 103114</figureCitation>
), both of which occur close to
<collectingCountry id="A1D1763D1327BC5CFB4ABA55FB67EF68" box="[1178,1250,1547,1573]" name="Oman" pageId="43" pageNumber="44">Oman</collectingCountry>
(
<collectingCountry id="A1D1763D1327BC5CFB21BA55FAC0EF68" box="[1265,1349,1547,1573]" name="Yemen" pageId="43" pageNumber="44">Yemen</collectingCountry>
and the
<collectingCountry id="A1D1763D1327BC5CFF37BA70FE6FEF04" box="[231,490,1582,1609]" name="United Arab Emirates" pageId="43" pageNumber="44">United Arab Emirates</collectingCountry>
). Therefore, we prefer not to describe this putative species formally until more individuals (females and males) are avilable.
</paragraph>
</subSubSection>
</treatment>
</document>