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<document id="C7A013436E91F9924337B4DEDE9D86D3" ID-DOI="10.11646/zootaxa.4087.1.1" ID-GBIF-Dataset="17164bda-a943-496a-b044-1958db25c495" ID-ISSN="1175-5326" ID-Zenodo-Dep="399511" ID-ZooBank="C510CF71-0039-478A-91ED-BFD65B6FE0BE" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457771752054" checkinUser="plazi" docAuthor="Elena P. Kameneva &amp; Valery A. Kroneyev" docDate="2016" docId="5101BA35FF83FFDAFF1AE3EAD463F9DE" docLanguage="en" docName="15545-5067-1-PB.pdf.imf" docOrigin="Zootaxa 4087 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Physiphora euphorbiana Krivosheina &amp; Krivosheina 1997" docType="treatment" docVersion="9" lastPageNumber="36" masterDocId="AD38C24DFFA2FFF9FF8DE104D70CFF88" masterDocTitle="Revision of the Genus Physiphora Fallén 1810 (Diptera: Ulidiidae: Ulidiinae)" masterLastPageNumber="88" masterPageNumber="1" pageNumber="34" updateTime="1698184504803" updateUser="plazi">
<mods:mods id="EA4567B6788BEAB10179DDE5B21F26B9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B84FF366A9B6905650380887A1B2F436">
<mods:title id="EECC24B1C998ACEA0176DA7C1FF95BDB">Revision of the Genus Physiphora Fallén 1810 (Diptera: Ulidiidae: Ulidiinae)</mods:title>
</mods:titleInfo>
<mods:name id="0D2D8538F3BC1AD33403DBABEF554B73" type="personal">
<mods:role id="FD5C7BD5C122BAFBB22D4454EFD42268">
<mods:roleTerm id="25C551EA7C6E3F1E2258D6B3D3FCDD6D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="10B1CD4CACC05E62509C987F9CDD1422">Elena P. Kameneva</mods:namePart>
</mods:name>
<mods:name id="2585C86F165C751EFD849D0D8546B5B2" type="personal">
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<mods:namePart id="2BF3D30EF7AE92A47810B614D37AA736">Valery A. Kroneyev</mods:namePart>
</mods:name>
<mods:typeOfResource id="F83D453F9CBC5812263A7AE23C7D8F74">text</mods:typeOfResource>
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<mods:title id="B140A09E3B79DC84D48C6611168D4237">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B084B0EC7C82777EC0B475642CFA86B3">
<mods:date id="A00FB8080E205D0AF797FE87DD6FC2C2">2016</mods:date>
<mods:detail id="ECBB2D87996C2CAC82D7619006EA57B5" type="volume">
<mods:number id="D3C88CD87A2D3D8720FBB7C5C856B777">4087</mods:number>
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<mods:detail id="78F9E57B6B1BA486CB084E17D7BD16C0" type="issue">
<mods:number id="9C32DC9B6CE5860053FE6BFD399C4818">1</mods:number>
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<mods:classification id="E571A0AA5F18CC819AC36C1D3B5D8659">journal article</mods:classification>
<mods:identifier id="7AF680719F33D69B61D0AFCD0C7B51EC" type="DOI">10.11646/zootaxa.4087.1.1</mods:identifier>
<mods:identifier id="EC80DE2DAF3E9CCBBCB34B13FC5694F0" type="GBIF-Dataset">17164bda-a943-496a-b044-1958db25c495</mods:identifier>
<mods:identifier id="7C498F0B56DB645A611D1F5E963C67DD" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="35781142A70EA0FC0BDA713FF05E0508" type="Zenodo-Dep">399511</mods:identifier>
<mods:identifier id="C5A4DC40A96F449748E36985D84C641E" type="ZooBank">C510CF71-0039-478A-91ED-BFD65B6FE0BE</mods:identifier>
</mods:mods>
<treatment id="5101BA35FF83FFDAFF1AE3EAD463F9DE" ID-DOI="http://doi.org/10.5281/zenodo.6066595" ID-GBIF-Taxon="117897985" ID-Zenodo-Dep="6066595" LSID="urn:lsid:plazi:treatment:5101BA35FF83FFDAFF1AE3EAD463F9DE" httpUri="http://treatment.plazi.org/id/5101BA35FF83FFDAFF1AE3EAD463F9DE" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34">
<subSubSection id="91B258A8FF83FFD8FF1AE3EAD427FCE7" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="D9170B23FF83FFD8FF1AE3EAD46AFC80" blockId="33.[151,870,750,810]" box="[151,870,750,776]" pageId="33" pageNumber="34">
<heading id="825FBC4FFF83FFD8FF1AE3EAD46AFC80" bold="true" box="[151,870,750,776]" fontSize="11" level="1" pageId="33" pageNumber="34" reason="1">
<taxonomicName id="1EA870A0FF83FFD8FF1AE3EAD46AFC80" ID-CoL="77HZT" authority="Krivosheina &amp; Krivosheina 1997" authorityName="Krivosheina &amp; Krivosheina" authorityYear="1997" box="[151,870,750,776]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="euphorbiana">
<emphasis id="EBDCD731FF83FFD8FF1AE3EAD46AFC80" bold="true" box="[151,870,750,776]" pageId="33" pageNumber="34">
<emphasis id="EBDCD731FF83FFD8FF1AE3EAD6C8FC80" bold="true" box="[151,452,750,776]" italics="true" pageId="33" pageNumber="34">Physiphora euphorbiana</emphasis>
Krivosheina &amp; Krivosheina 1997
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="D9170B23FF83FFD8FF1AE215D657FCA2" blockId="33.[151,870,750,810]" box="[151,347,785,810]" pageId="33" pageNumber="34">
<figureCitation id="419317A6FF83FFD8FF1AE215D65BFCA2" box="[151,343,785,810]" captionStart-0="FIGURES 124 – 130" captionStart-1="FIGURES 131 – 137" captionStartId-0="34.[151,264,1423,1444]" captionStartId-1="35.[151,264,961,982]" captionTargetBox-0="[172,1416,221,1401]" captionTargetBox-1="[162,1433,196,909]" captionTargetId-0="figure@34.[151,1436,193,1401]" captionTargetId-1="figure@35.[151,1436,193,939]" captionTargetPageId-0="34" captionTargetPageId-1="35" captionText-0="FIGURES 124 – 130. Physiphora euphorbiana: 124, habitus, left; 125, habitus, left, anterior and dorsal; 126,127, head and thorax, dorsal and left; 128, head, anterior; 129, wing; 130, fore leg." captionText-1="FIGURES 131 – 137. Physiphora euphorbiana: 131, ♂ phallus; 132, glans, enlarged; 133, epandrium and hypandrium, right; 134, epandrium, anterior; 135, epandrium, posterior; 136, hypandrium, ventral; 137, ejaculatory apodeme." httpUri-0="https://zenodo.org/record/399586/files/figure.png" httpUri-1="https://zenodo.org/record/399587/files/figure.png" pageId="33" pageNumber="34">Figures 124–137</figureCitation>
.
</paragraph>
<paragraph id="D9170B23FF83FFD8FF1AE25DD427FCE7" blockId="33.[151,1393,857,910]" box="[151,811,857,879]" pageId="33" pageNumber="34">
<taxonomicName id="1EA870A0FF83FFD8FF1AE25DD42BFCE7" authority="Krivosheina &amp; Krivosheina, 1997: 464" authorityName="Krivosheina &amp; Krivosheina" authorityPageNumber="464" authorityYear="1997" box="[151,807,857,879]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="euphorbiana">
<emphasis id="EBDCD731FF83FFD8FF1AE25DD69FFCE7" box="[151,403,857,879]" italics="true" pageId="33" pageNumber="34">Physiphora euphorbiana</emphasis>
Krivosheina &amp; Krivosheina, 1997: 464
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="91B258A8FF83FFD8FF1AE27CD27DFC06" box="[151,1393,888,910]" pageId="33" pageNumber="34" type="description">
<paragraph id="D9170B23FF83FFD8FF1AE27CD27DFC06" blockId="33.[151,1393,857,910]" box="[151,1393,888,910]" pageId="33" pageNumber="34">
<taxonomicName id="1EA870A0FF83FFD8FF1AE27CD605FC06" box="[151,265,888,910]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Physiphora</taxonomicName>
euphorbiae: Kameneva &amp; Korneyev, 2010: 622 (unavailable; incorrect subsequent spelling of
<taxonomicName id="1EA870A0FF83FFD8FB45E27DD26FFC06" box="[1224,1379,888,910]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="euphorbiana">
<emphasis id="EBDCD731FF83FFD8FB45E27DD26FFC06" box="[1224,1379,888,910]" italics="true" pageId="33" pageNumber="34">P. euphorbiana</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="91B258A8FF83FFD8FF1AE2BED403FBEB" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="D9170B23FF83FFD8FF1AE2BED403FBEB" blockId="33.[151,1437,954,2026]" pageId="33" pageNumber="34">
<emphasis id="EBDCD731FF83FFD8FF1AE2BED7F3FC5B" bold="true" box="[151,255,954,979]" pageId="33" pageNumber="34">Material</emphasis>
.
<emphasis id="EBDCD731FF83FFD8FE9FE2BFD647FC5A" bold="true" box="[274,331,955,978]" pageId="33" pageNumber="34">
<typeStatus id="0613B581FF83FFD8FE9FE2BFD647FC5A" box="[274,331,955,978]" pageId="33" pageNumber="34">Type</typeStatus>
</emphasis>
.
<typeStatus id="0613B581FF83FFD8FEEDE2BED6C4FC5B" box="[352,456,954,979]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
♂ and
<typeStatus id="0613B581FF83FFD8FDA6E2BFD59AFC5A" box="[555,662,955,978]" pageId="33" pageNumber="34" type="paratype">paratypes</typeStatus>
<specimenCount id="CFAEC0AAFF83FFD8FD2EE2BED5C6FC5B" box="[675,714,954,979]" pageId="33" pageNumber="34" type="generic">2♂</specimenCount>
:
<emphasis id="EBDCD731FF83FFD8FD57E2BED416FC5B" bold="true" box="[730,794,954,979]" pageId="33" pageNumber="34">
<collectingCountry id="A1BF4BB3FF83FFD8FD57E2BED416FC5B" box="[730,794,954,979]" name="India" pageId="33" pageNumber="34">India</collectingCountry>
</emphasis>
: Radjasthan, Jodhpur, from larvae collected in rotting
<taxonomicName id="1EA870A0FF83FFD8FF1AE2DAD603FC7F" box="[151,271,990,1015]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="genus">
<emphasis id="EBDCD731FF83FFD8FF1AE2DAD603FC7F" box="[151,271,990,1015]" italics="true" pageId="33" pageNumber="34">Euphorbia</emphasis>
</taxonomicName>
sp. (A. Kompantsev) (ZMUM; not examined).
<emphasis id="EBDCD731FF83FFD8FCDEE2DBD4CDFC7E" bold="true" box="[851,961,991,1014]" pageId="33" pageNumber="34">Non-type</emphasis>
.
<emphasis id="EBDCD731FF83FFD8FC5BE2DAD332FC7F" bold="true" box="[982,1086,990,1015]" pageId="33" pageNumber="34">
<collectingCountry id="A1BF4BB3FF83FFD8FC5BE2DAD332FC7F" box="[982,1086,990,1015]" name="Pakistan" pageId="33" pageNumber="34">Pakistan</collectingCountry>
</emphasis>
: Kahuta,
<geoCoordinate id="BC9C6DE4FF83FFD8FB34E2DBD22DFC7E" box="[1209,1313,991,1015]" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="925" value="33.583332">33º35′ N</geoCoordinate>
<geoCoordinate id="BC9C6DE4FF83FFD8FAA2E2DBD29BFC7E" box="[1327,1431,991,1015]" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="925" value="73.38333">73º23′ E</geoCoordinate>
, “maggot in a rottened stem of
<emphasis id="EBDCD731FF83FFD8FD84E507D516FB92" box="[521,538,1027,1050]" italics="true" pageId="33" pageNumber="34">E</emphasis>
.
<emphasis id="EBDCD731FF83FFD8FDACE506D401FB93" box="[545,781,1026,1051]" italics="true" pageId="33" pageNumber="34">
[uphorbia]
<taxonomicName id="1EA870A0FF83FFD8FD27E506D401FB93" box="[682,781,1026,1051]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="species" species="royleana">royleana</taxonomicName>
</emphasis>
, 30.
<date id="AD162DE3FF83FFD8FCCCE506D4CDFB93" box="[833,961,1026,1051]" pageId="33" pageNumber="34" value="1961-07-03">vii.1961, 3</date>
♀ (BMNH); Lasbella Prov.,
<quantity id="1E50A6C6FF83FFD8FA9EE507D254FB93" box="[1299,1368,1026,1051]" metricMagnitude="4" metricUnit="m" metricValue="6.9201792000000015" pageId="33" pageNumber="34" unit="mi" value="43.0">43 mi</quantity>
N of Karachi, 28.iii–1.
<date id="AD162DE3FF83FFD8FEDAE522D6C5FBB7" box="[343,457,1062,1087]" pageId="33" pageNumber="34" value="1965-04-05">
iv.1965,
<specimenCount id="CFAEC0AAFF83FFD8FE36E522D6C5FBB7" box="[443,457,1062,1087]" pageId="33" pageNumber="34" type="generic">5</specimenCount>
</date>
♂, 2♀ (Neal) (USNM);
<emphasis id="EBDCD731FF83FFD8FD59E522D418FBB7" bold="true" box="[724,788,1062,1087]" pageId="33" pageNumber="34">
<collectingCountry id="A1BF4BB3FF83FFD8FD59E522D418FBB7" box="[724,788,1062,1087]" name="India" pageId="33" pageNumber="34">India</collectingCountry>
</emphasis>
: Tamil Nadu, Dohnavur,
<geoCoordinate id="BC9C6DE4FF83FFD8FBBCE523D388FBB6" box="[1073,1156,1063,1087]" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="925" value="8.466666">8º28′ N</geoCoordinate>
<geoCoordinate id="BC9C6DE4FF83FFD8FB06E523D3E7FBB6" box="[1163,1259,1063,1087]" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="925" value="77.583336">77º35′ E</geoCoordinate>
, Donevelly Dt., 350′, 30.
<date id="AD162DE3FF83FFD8FF7AE54ED66AFBEB" box="[247,358,1098,1123]" pageId="33" pageNumber="34" value="1938-09-01">
ix.1938,
<specimenCount id="CFAEC0AAFF83FFD8FED6E54ED66AFBEB" box="[347,358,1098,1123]" pageId="33" pageNumber="34" type="generic">1</specimenCount>
</date>
♂, 1♀ (collector unknown) (BMNH).
</paragraph>
</subSubSection>
<subSubSection id="91B258A8FF83FFD8FF4AE56AD23AFA46" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="D9170B23FF83FFD8FF4AE56AD23AFA46" blockId="33.[151,1437,954,2026]" pageId="33" pageNumber="34">
<emphasis id="EBDCD731FF83FFD8FF4AE56AD636FB0F" bold="true" box="[199,314,1134,1159]" pageId="33" pageNumber="34">Diagnosis</emphasis>
. This species differs from other
<taxonomicName id="1EA870A0FF83FFD8FD31E56AD431FB0F" box="[700,829,1134,1159]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="EBDCD731FF83FFD8FD31E56AD431FB0F" box="[700,829,1134,1159]" italics="true" pageId="33" pageNumber="34">Physiphora</emphasis>
</taxonomicName>
by the combination of entirely brown or black fore basitarsomere, pair of large subrectangular microtrichose parafrontal areas on satin black frons with violet tinge, face with entire transverse microtrichose area and epistome black medially but yellow laterally, wing with cell r4+5 closed but not petiolate, and male genitalia with bilobate surstyli, very wide, partly fused nipple-like structures of cerci (
<figureCitation id="419317A6FF83FFD8FF50E405D68CFA92" box="[221,384,1281,1306]" captionStart="FIGURES 131 – 137" captionStartId="35.[151,264,961,982]" captionTargetBox="[162,1433,196,909]" captionTargetId="figure@35.[151,1436,193,939]" captionTargetPageId="35" captionText="FIGURES 131 – 137. Physiphora euphorbiana: 131, ♂ phallus; 132, glans, enlarged; 133, epandrium and hypandrium, right; 134, epandrium, anterior; 135, epandrium, posterior; 136, hypandrium, ventral; 137, ejaculatory apodeme." httpUri="https://zenodo.org/record/399587/files/figure.png" pageId="33" pageNumber="34">Figs. 134–135</figureCitation>
) and hypandrium with symmetrical phallapodeme having widely separated, parallel posterior branches and equal vanes of phallapodeme; left gonite entirely lacking (
<figureCitation id="419317A6FF83FFD8FC79E421D354FAB6" box="[1012,1112,1317,1342]" captionStart="FIGURES 131 – 137" captionStartId="35.[151,264,961,982]" captionTargetBox="[162,1433,196,909]" captionTargetId="figure@35.[151,1436,193,939]" captionTargetPageId="35" captionText="FIGURES 131 – 137. Physiphora euphorbiana: 131, ♂ phallus; 132, glans, enlarged; 133, epandrium and hypandrium, right; 134, epandrium, anterior; 135, epandrium, posterior; 136, hypandrium, ventral; 137, ejaculatory apodeme." httpUri="https://zenodo.org/record/399587/files/figure.png" pageId="33" pageNumber="34">Fig. 136</figureCitation>
) and phallus with preglans conspicuously longer than wide, leaf-like stipe (
<figureCitation id="419317A6FF83FFD8FD35E44DD414FAEA" box="[696,792,1353,1378]" captionStart="FIGURES 131 – 137" captionStartId="35.[151,264,961,982]" captionTargetBox="[162,1433,196,909]" captionTargetId="figure@35.[151,1436,193,939]" captionTargetPageId="35" captionText="FIGURES 131 – 137. Physiphora euphorbiana: 131, ♂ phallus; 132, glans, enlarged; 133, epandrium and hypandrium, right; 134, epandrium, anterior; 135, epandrium, posterior; 136, hypandrium, ventral; 137, ejaculatory apodeme." httpUri="https://zenodo.org/record/399587/files/figure.png" pageId="33" pageNumber="34">Fig. 131</figureCitation>
) and glans with short, inconspicuous lobes (
<figureCitation id="419317A6FF83FFD8FA9EE44DD279FAEA" box="[1299,1397,1353,1378]" captionStart="FIGURES 131 – 137" captionStartId="35.[151,264,961,982]" captionTargetBox="[162,1433,196,909]" captionTargetId="figure@35.[151,1436,193,939]" captionTargetPageId="35" captionText="FIGURES 131 – 137. Physiphora euphorbiana: 131, ♂ phallus; 132, glans, enlarged; 133, epandrium and hypandrium, right; 134, epandrium, anterior; 135, epandrium, posterior; 136, hypandrium, ventral; 137, ejaculatory apodeme." httpUri="https://zenodo.org/record/399587/files/figure.png" pageId="33" pageNumber="34">Fig. 132</figureCitation>
). It differs from the other species with widely microtrichose frons (
<taxonomicName id="1EA870A0FF83FFD8FC28E46AD344FA0E" box="[933,1096,1389,1414]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="leucotricha">
<emphasis id="EBDCD731FF83FFD8FC28E46AD344FA0E" box="[933,1096,1389,1414]" italics="true" pageId="33" pageNumber="34">P. leucotricha</emphasis>
</taxonomicName>
,
<taxonomicName id="1EA870A0FF83FFD8FBD2E46AD3EEFA0E" box="[1119,1250,1389,1414]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="steyskali" status="sp. nov.">
<emphasis id="EBDCD731FF83FFD8FBD2E46AD3EEFA0E" box="[1119,1250,1389,1414]" italics="true" pageId="33" pageNumber="34">P. steyskali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0EF6A4AFF83FFD8FB7FE46AD23AFA0E" box="[1266,1334,1389,1414]" pageId="33" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
) by the combination of entirely black fore metatarsus and entire microtrichose crossband in the upper half of frons (fore metatarsus mostly white and face with λ-shaped microtrichose area on facial carina in compared species).
</paragraph>
</subSubSection>
<subSubSection id="91B258A8FF83FFDAFF4AE4DDD534FA2B" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34" type="description">
<paragraph id="D9170B23FF83FFD8FF4AE4DDD62BF941" blockId="33.[151,1437,954,2026]" pageId="33" pageNumber="34">
<emphasis id="EBDCD731FF83FFD8FF4AE4DDD65DFA7A" bold="true" box="[199,337,1497,1522]" pageId="33" pageNumber="34">Description</emphasis>
. Head (
<figureCitation id="419317A6FF83FFD8FE21E4DDD543FA7A" box="[428,591,1497,1522]" captionStart="FIGURES 124 – 130" captionStartId="34.[151,264,1423,1444]" captionTargetBox="[172,1416,221,1401]" captionTargetId="figure@34.[151,1436,193,1401]" captionTargetPageId="34" captionText="FIGURES 124 – 130. Physiphora euphorbiana: 124, habitus, left; 125, habitus, left, anterior and dorsal; 126,127, head and thorax, dorsal and left; 128, head, anterior; 129, wing; 130, fore leg." httpUri="https://zenodo.org/record/399586/files/figure.png" pageId="33" pageNumber="34">Figs. 127–128</figureCitation>
) black, with brown sides of face, gena and postgena; rarely frons, sides of face, postgena and middle of vertex brownish yellow. Frons 1.3 times as long as wide, black or brown, medially with matt black triangle with violet tinge, shining black anterolateral corners and wide parafrontal microtichose area almost reaching antero-medial margin (
<figureCitation id="419317A6FF83FFD8FD19E741D5FBF9D6" box="[660,759,1605,1630]" captionStart="FIGURES 124 – 130" captionStartId="34.[151,264,1423,1444]" captionTargetBox="[172,1416,221,1401]" captionTargetId="figure@34.[151,1436,193,1401]" captionTargetPageId="34" captionText="FIGURES 124 – 130. Physiphora euphorbiana: 124, habitus, left; 125, habitus, left, anterior and dorsal; 126,127, head and thorax, dorsal and left; 128, head, anterior; 129, wing; 130, fore leg." httpUri="https://zenodo.org/record/399586/files/figure.png" pageId="33" pageNumber="34">Fig. 128</figureCitation>
), and poorly differentiated calluses posterior to its middle; sides of frons with numerous moderately long and thick setulae inserted into small pits. Vertical plates and ocellar triangle shining black with greenish sheen; 2 pairs of black orbital setae; ocellar setae lateroclinate, as long as orbital setae.
</paragraph>
<paragraph id="D9170B23FF83FFD8FF4AE7D2D224F862" blockId="33.[151,1437,954,2026]" pageId="33" pageNumber="34">
Face mostly yellow, facial carina smooth, yellow, with entire transverse white microtrichose area covering also antennal grooves; epistome and ventral part of antennal grooves black; lateral parts of face yellow. Gena high, half as high as eye; parafacial and facial ridge mostly brown, each with narrow white microtrichose stripe; gena posteriorly with arcuate white microtrichose mark separating it from anteriorly shining orange and posteriorly black postgena (
<figureCitation id="419317A6FF83FFD8FED9E661D6B4F8F6" box="[340,440,1893,1918]" captionStart="FIGURES 124 – 130" captionStartId="34.[151,264,1423,1444]" captionTargetBox="[172,1416,221,1401]" captionTargetId="figure@34.[151,1436,193,1401]" captionTargetPageId="34" captionText="FIGURES 124 – 130. Physiphora euphorbiana: 124, habitus, left; 125, habitus, left, anterior and dorsal; 126,127, head and thorax, dorsal and left; 128, head, anterior; 129, wing; 130, fore leg." httpUri="https://zenodo.org/record/399586/files/figure.png" pageId="33" pageNumber="34">Fig. 127</figureCitation>
). Occiput black. Medial vertical seta half as long as frons width, 1.1 times as long as lateral vertical and 4–5 times as long as ocellar, orbital and postocellar setae. Antenna yellow, except falgellomere 1 brown apex, rounded apically, twice as long as wide, greyish microtrichose; arista bare, brownish yellow in basal 1/5, remainder black. Clypeus black. Palp black, grey microtrichose and black setose. Mouthparts black.
</paragraph>
<caption id="8DD75BABFF80FFDBFF1AE48BD444FA4B" httpUri="https://zenodo.org/record/399586/files/figure.png" pageId="34" pageNumber="35" targetBox="[172,1416,221,1401]" targetPageId="34">
<paragraph id="D9170B23FF80FFDBFF1AE48BD444FA4B" blockId="34.[151,1435,1423,1475]" pageId="34" pageNumber="35">
<emphasis id="EBDCD731FF80FFDBFF1AE48BD663FA2C" bold="true" box="[151,367,1423,1444]" pageId="34" pageNumber="35">FIGURES 124–130.</emphasis>
<taxonomicName id="1EA870A0FF80FFDBFEF4E48BD574FA2D" box="[377,632,1423,1445]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="euphorbiana">
<emphasis id="EBDCD731FF80FFDBFEF4E48BD574FA2D" box="[377,632,1423,1445]" italics="true" pageId="34" pageNumber="35">Physiphora euphorbiana</emphasis>
</taxonomicName>
: 124, habitus, left; 125, habitus, left, anterior and dorsal; 126,127, head and thorax, dorsal and left; 128, head, anterior; 129, wing; 130, fore leg.
</paragraph>
</caption>
<paragraph id="D9170B23FF80FFDBFF4AE4F4D4AAF8A1" blockId="34.[151,1436,1520,2022]" pageId="34" pageNumber="35">
Thorax (
<figureCitation id="419317A6FF80FFDBFEA7E4F4D6DFF981" box="[298,467,1520,1545]" captionStart="FIGURES 124 – 130" captionStartId="34.[151,264,1423,1444]" captionTargetBox="[172,1416,221,1401]" captionTargetId="figure@34.[151,1436,193,1401]" captionTargetPageId="34" captionText="FIGURES 124 – 130. Physiphora euphorbiana: 124, habitus, left; 125, habitus, left, anterior and dorsal; 126,127, head and thorax, dorsal and left; 128, head, anterior; 129, wing; 130, fore leg." httpUri="https://zenodo.org/record/399586/files/figure.png" pageId="34" pageNumber="35">Figs. 125–126</figureCitation>
). Scutum and scutellum black, densely rugulose, almost matt, with green tinge and variable, red to violet reflections; antepronotum, postpronotal lobe, posterior surface of notopleural triangle, supraalar and all prescutellar parts of scutum, anterior half of anepisternum and katepisternum, and whole anepimeron shining black, usually with blue sheen; posterodorsal parts of anepisternum and katepisternum shagreened; supraalar area and tympanal fossa brown to black, postscutellum black, gray microtrichose; postero-ventral margin of scutellum without microtrichose area; anatergite, katatergite and lateroventral parts of mediotergite grey microtrichose. Mesonotal scutum with acrostichal rows of setulae as well as acrostichal seta indistinguishable, dorsocentral setulae more visible; dorsocentral setae hair-like; other setae moderately long, black: one postprononal, 2 postsutural supra-alar, one intra-alar and one postalar.
</paragraph>
<paragraph id="D9170B23FF80FFDBFF4AE630D32BF8F8" blockId="34.[151,1436,1520,2022]" pageId="34" pageNumber="35">Scutellum conspicuously swollen, sparsely rugulose, with slight golden reflection, apparently with short setulae over disk (1/10 times as long as setae) and 2 pairs of black scutellar setae.</paragraph>
<paragraph id="D9170B23FF80FFDBFF4AE678D44AF869" blockId="34.[151,1436,1520,2022]" pageId="34" pageNumber="35">
Wing. Entirely hyaline, with yellow veins; cell r4+5 narrowly closed, apical section of M arcuate (
<figureCitation id="419317A6FF80FFDBFAA6E678D283F81D" box="[1323,1423,1916,1941]" captionStart="FIGURES 124 – 130" captionStartId="34.[151,264,1423,1444]" captionTargetBox="[172,1416,221,1401]" captionTargetId="figure@34.[151,1436,193,1401]" captionTargetPageId="34" captionText="FIGURES 124 – 130. Physiphora euphorbiana: 124, habitus, left; 125, habitus, left, anterior and dorsal; 126,127, head and thorax, dorsal and left; 128, head, anterior; 129, wing; 130, fore leg." httpUri="https://zenodo.org/record/399586/files/figure.png" pageId="34" pageNumber="35">Fig. 129</figureCitation>
). Postero-apical extension of cell cup 1.5 times as long as vein A1+CuA2, and 3 times as long as transverse section of vein CuA2. Calypters with white fringe. Length: 3.2–3.5 mm.
</paragraph>
<caption id="8DD75BABFF81FFDAFF1AE2C5D3C7FC7D" httpUri="https://zenodo.org/record/399587/files/figure.png" pageId="35" pageNumber="36" targetBox="[162,1433,196,909]" targetPageId="35">
<paragraph id="D9170B23FF81FFDAFF1AE2C5D3C7FC7D" blockId="35.[151,1435,960,1013]" pageId="35" pageNumber="36">
<emphasis id="EBDCD731FF81FFDAFF1AE2C5D661FC5E" bold="true" box="[151,365,961,982]" pageId="35" pageNumber="36">FIGURES 131–137.</emphasis>
<taxonomicName id="1EA870A0FF81FFDAFEFBE2C5D578FC5F" box="[374,628,961,983]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="euphorbiana">
<emphasis id="EBDCD731FF81FFDAFEFBE2C5D578FC5F" box="[374,628,961,983]" italics="true" pageId="35" pageNumber="36">Physiphora euphorbiana</emphasis>
</taxonomicName>
: 131, ♂ phallus; 132, glans, enlarged; 133, epandrium and hypandrium, right; 134, epandrium, anterior; 135, epandrium, posterior; 136, hypandrium, ventral; 137, ejaculatory apodeme.
</paragraph>
</caption>
<paragraph id="D9170B23FF81FFDAFF4AE527D5D7FB0B" blockId="35.[151,1436,1058,1623]" pageId="35" pageNumber="36">Legs. Black including whole fore tarsus, mid- and hind tibiae brownish yellow or basally yellow, apically brown; mid- and hind tarsi yellow except two apical tarsomeres brown or black; all setae black; fore femur posteroventrally with 5 thickened short setae in apical half.</paragraph>
<paragraph id="D9170B23FF81FFDAFF4AE58AD379FB43" blockId="35.[151,1436,1058,1623]" pageId="35" pageNumber="36">Abdomen. Both tergites and sternites black, finely rugulose, with golden (blue on posterior tergites) reflection; all setulae black; female abdominal tergite 2 with pair of dimple-like structures laterally.</paragraph>
<paragraph id="D9170B23FF81FFDAFF4AE5D2D265FAF7" blockId="35.[151,1436,1058,1623]" pageId="35" pageNumber="36">
<collectingCountry id="A1BF4BB3FF81FFDAFF4AE5D2D60DFB67" box="[199,257,1238,1263]" name="Malta" pageId="35" pageNumber="36">Male</collectingCountry>
postabdomen brown to black; epandrium as on
<figureCitation id="419317A6FF81FFDAFC93E5D2D4CDFB67" box="[798,961,1238,1263]" captionStart="FIGURES 131 – 137" captionStartId="35.[151,264,961,982]" captionTargetBox="[162,1433,196,909]" captionTargetId="figure@35.[151,1436,193,939]" captionTargetPageId="35" captionText="FIGURES 131 – 137. Physiphora euphorbiana: 131, ♂ phallus; 132, glans, enlarged; 133, epandrium and hypandrium, right; 134, epandrium, anterior; 135, epandrium, posterior; 136, hypandrium, ventral; 137, ejaculatory apodeme." httpUri="https://zenodo.org/record/399587/files/figure.png" pageId="35" pageNumber="36">Figs. 133–135</figureCitation>
, surstyli bilobate, nipple-like structures of cerci very wide, partly fused; phallus with widened, leaf-like stipe conspicuously shorter than preglans and glans; caecum almost three times as long as stipe width; preglans with very long wrinkled membranous area (
<figureCitation id="419317A6FF81FFDAFAA1E41AD282FABF" box="[1324,1422,1310,1335]" captionStart="FIGURES 131 – 137" captionStartId="35.[151,264,961,982]" captionTargetBox="[162,1433,196,909]" captionTargetId="figure@35.[151,1436,193,939]" captionTargetPageId="35" captionText="FIGURES 131 – 137. Physiphora euphorbiana: 131, ♂ phallus; 132, glans, enlarged; 133, epandrium and hypandrium, right; 134, epandrium, anterior; 135, epandrium, posterior; 136, hypandrium, ventral; 137, ejaculatory apodeme." httpUri="https://zenodo.org/record/399587/files/figure.png" pageId="35" pageNumber="36">Fig. 132</figureCitation>
); glans with 3–4 very short lobes (
<figureCitation id="419317A6FF81FFDAFDACE446D5C7FAD3" box="[545,715,1346,1371]" captionStart="FIGURES 131 – 137" captionStartId="35.[151,264,961,982]" captionTargetBox="[162,1433,196,909]" captionTargetId="figure@35.[151,1436,193,939]" captionTargetPageId="35" captionText="FIGURES 131 – 137. Physiphora euphorbiana: 131, ♂ phallus; 132, glans, enlarged; 133, epandrium and hypandrium, right; 134, epandrium, anterior; 135, epandrium, posterior; 136, hypandrium, ventral; 137, ejaculatory apodeme." httpUri="https://zenodo.org/record/399587/files/figure.png" pageId="35" pageNumber="36">Figs. 131–132</figureCitation>
). Hypandrium with symmetrical phallapodeme having widely separated, parallel posterior branches and equal vanes of phallapodeme; left gonite entirely lacking (
<figureCitation id="419317A6FF81FFDAFB76E462D250FAF7" box="[1275,1372,1382,1407]" captionStart="FIGURES 131 – 137" captionStartId="35.[151,264,961,982]" captionTargetBox="[162,1433,196,909]" captionTargetId="figure@35.[151,1436,193,939]" captionTargetPageId="35" captionText="FIGURES 131 – 137. Physiphora euphorbiana: 131, ♂ phallus; 132, glans, enlarged; 133, epandrium and hypandrium, right; 134, epandrium, anterior; 135, epandrium, posterior; 136, hypandrium, ventral; 137, ejaculatory apodeme." httpUri="https://zenodo.org/record/399587/files/figure.png" pageId="35" pageNumber="36">Fig. 136</figureCitation>
).
</paragraph>
<paragraph id="D9170B23FF81FFDAFF4AE48ED534FA2B" blockId="35.[151,1436,1058,1623]" box="[199,568,1418,1443]" pageId="35" pageNumber="36">Female terminalia not examined.</paragraph>
</subSubSection>
<subSubSection id="91B258A8FF81FFDAFF4AE4AAD540FA4F" box="[199,588,1454,1479]" pageId="35" pageNumber="36" type="distribution">
<paragraph id="D9170B23FF81FFDAFF4AE4AAD540FA4F" blockId="35.[151,1436,1058,1623]" box="[199,588,1454,1479]" pageId="35" pageNumber="36">
<emphasis id="EBDCD731FF81FFDAFF4AE4AAD655FA4F" bold="true" box="[199,345,1454,1479]" pageId="35" pageNumber="36">Distribution</emphasis>
. Indian subcontinent.
</paragraph>
</subSubSection>
<subSubSection id="91B258A8FF81FFDAFF4AE4D6D4AAFA63" box="[199,934,1490,1515]" pageId="35" pageNumber="36" type="biology_ecology">
<paragraph id="D9170B23FF81FFDAFF4AE4D6D4AAFA63" blockId="35.[151,1436,1058,1623]" box="[199,934,1490,1515]" pageId="35" pageNumber="36">
<emphasis id="EBDCD731FF81FFDAFF4AE4D6D613FA63" bold="true" box="[199,287,1490,1515]" pageId="35" pageNumber="36">Biology</emphasis>
. Reared from rotting stems of
<taxonomicName id="1EA870A0FF81FFDAFDF5E4D6D4AAFA63" authority="Boiss." authorityName="Boiss." box="[632,934,1490,1515]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="royleana">
<emphasis id="EBDCD731FF81FFDAFDF5E4D6D454FA63" box="[632,856,1490,1515]" italics="true" pageId="35" pageNumber="36">Euphorbia royleana</emphasis>
Boiss.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="91B258A8FF81FFDAFF4AE4F3D463F9DE" pageId="35" pageNumber="36" type="discussion">
<paragraph id="D9170B23FF81FFDAFF4AE4F3D463F9DE" blockId="35.[151,1436,1058,1623]" pageId="35" pageNumber="36">
<emphasis id="EBDCD731FF81FFDAFF4AE4F3D638F986" bold="true" box="[199,308,1527,1550]" pageId="35" pageNumber="36">Remarks</emphasis>
. This species possesses aberrant structure of epandrium and cerci, which showns no similarities in Ulidiini. It reminds
<taxonomicName id="1EA870A0FF81FFDAFE1DE71FD519F9BB" box="[400,533,1562,1587]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="steyskali" status="sp. nov.">
<emphasis id="EBDCD731FF81FFDAFE1DE71FD519F9BB" box="[400,533,1562,1587]" italics="true" pageId="35" pageNumber="36">P. steyskali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0EF6A4AFF81FFDAFDA8E71FD560F9BA" box="[549,620,1563,1586]" pageId="35" pageNumber="36" rank="species">sp. n.</taxonomicNameLabel>
, which shares widely microtrichose frons and similar structure of phallapodeme, and might be somehow related to
<taxonomicName id="1EA870A0FF81FFDAFD30E73BD464F9DF" box="[701,872,1598,1623]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="euphorbiana">
<emphasis id="EBDCD731FF81FFDAFD30E73BD464F9DF" box="[701,872,1598,1623]" italics="true" pageId="35" pageNumber="36">P. euphorbiana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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