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<document id="42C2C640B42322559BFBFF547C058740" ID-DOI="10.5852/ejt.2014.71" ID-GBIF-Dataset="376ad5e1-afef-42db-b017-593e1bcd01ac" ID-ISSN="2118-9773" ID-Zenodo-Dep="3834958" ID-ZooBank="F667B43B-86A3-4D7D-9A22-021E52CDA30F" IM.bibliography_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1589938823289" checkinUser="tatiana" docAuthor="Blandin, Patrick, Bristow, Roger, Neild, Andrew, Sousa, Juan Carlos De, Gareca, Yuvinka &amp; Huertas, Blanca" docDate="2014" docId="506287BEFFACFFD9FE36FE2B6C809536" docLanguage="en" docName="ejt-2014-71.pdf.imf" docOrigin="European Journal of Taxonomy 71 (71)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Eryphanis zolvizora subsp. casagrande Blandin &amp; Bristow &amp; Neild &amp; Sousa &amp; Gareca &amp; Huertas 2014, ssp. nov." docType="treatment" docVersion="14" lastPageNumber="36" masterDocId="AC5BFFC6FF8DFFFAFFC6FF9D6E549103" masterDocTitle="Revisiting the Andean butterfly Eryphanis zolvizora group (Lepidoptera, Nymphalidae): one or several species?" masterLastPageNumber="66" masterPageNumber="1" pageNumber="34" updateTime="1737116849075" updateUser="carolina" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="14D1BEFBDC21D78285C4DFD73DAFD8C3">Revisiting the Andean butterfly Eryphanis zolvizora group (Lepidoptera, Nymphalidae): one or several species?</mods:title>
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<mods:namePart id="A4E76DDAD5DEC8C76E1D1AF221E16D0A">Blandin, Patrick</mods:namePart>
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<mods:affiliation id="D2D380A8C31EB24F6BAC706CBDCDE4B8">Muséum National dHistoire Naturelle, Entomologie, 57 rue Cuvier, 75005 Paris, France E-mail: patrick. blandin @ yahoo. fr (corresponding author) &amp; urn: lsid: zoobank. org: author: F 79 EECF 9 - 4 E 10 - 4826 - BA 99 - 01 DE 1 EB 0 DD 0 E</mods:affiliation>
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<mods:namePart id="A5F1B53EDB99B6231C38AFAEC3AEA4CF">Bristow, Roger</mods:namePart>
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<mods:affiliation id="18AD700F5AF4EB83DFAE1447A9F31389">2 Sid Bank, Sid Lane, Sidmouth, Devon EX 10 9 AW, UK E-mail: devonbristows @ btinternet. com &amp; urn: lsid: zoobank. org: author: E 9 AD 6 AA 1 - 2 C 47 - 4358 - B 606 - 0 A 84042 D 872 D</mods:affiliation>
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<mods:namePart id="608A1773188D7F3558F9B06D66824720">Neild, Andrew</mods:namePart>
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<mods:affiliation id="D7DAEFC9EF9E20D52CC22D58582D8E8C">The Natural History Museum, Entomology Department, Cromwell Road, SW 7 5 BD, London, UK E-mails: andrew. neild @ blueyonder. co. uk, B. Huertas @ nhm. ac. uk &amp; urn: lsid: zoobank. org: author: 268 E 6 AD 0 - 5 E 7 A- 4073 - B 530 - 6 CE 39 D 963474</mods:affiliation>
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<mods:namePart id="1732AA5B656ED6A3E4AB7437283E258D">Sousa, Juan Carlos De</mods:namePart>
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<mods:affiliation id="1277416EB93156F48A11F2C78D6436C6">Museo del Instituto de Zoología Agrícola, Facultad de Agronomía, Universidad Central de Venezuela, Maracay, Apdo.</mods:affiliation>
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<mods:namePart id="FDDE9560FF1C533C5BE57AB38B91970E">Gareca, Yuvinka</mods:namePart>
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<mods:affiliation id="882E7DE1E5E2379F65065903B5F6A8BB">Museo de Historia Natural Alcide dOrbigny, Entomologia, CP 4324, Av. Potosi N ° 1458 (esq. Av.</mods:affiliation>
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<mods:affiliation id="F5D1B5433B7A0119162D2D6854DD4046">The Natural History Museum, Entomology Department, Cromwell Road, SW 7 5 BD, London, UK E-mails: andrew. neild @ blueyonder. co. uk, B. Huertas @ nhm. ac. uk &amp; urn: lsid: zoobank. org: author: 268 E 6 AD 0 - 5 E 7 A- 4073 - B 530 - 6 CE 39 D 963474 &amp; urn: lsid: zoobank. org: author: F 96 FCD 0 A- 4 A 22 - 4782 - B 077 - 4588 E 193134 A</mods:affiliation>
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<mods:title id="9A80A55EEEBEEB968531AE144BD42EFB">European Journal of Taxonomy</mods:title>
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<mods:date id="5B9F6FEB77128FB14F588C83C78742EB">2014</mods:date>
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<mods:number id="BE4F75DA338BFAD4639387A514EE2656">71</mods:number>
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<treatment id="506287BEFFACFFD9FE36FE2B6C809536" ID-DOI="http://doi.org/10.5281/zenodo.3851872" ID-GBIF-Taxon="164241150" ID-Zenodo-Dep="3851872" LSID="urn:lsid:plazi:treatment:506287BEFFACFFD9FE36FE2B6C809536" httpUri="http://treatment.plazi.org/id/506287BEFFACFFD9FE36FE2B6C809536" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34" scope_class="Hexapoda" scope_family="Nymphalidae" scope_order="Lepidoptera" scope_subFamily="Morphinae" scope_superFamily="Papilionoidea" scope_tribe="Brassolini">
<subSubSection id="90D16523FFACFFDBFE36FE2B6A1090D2" box="[496,1092,438,465]" pageId="33" pageNumber="34" type="nomenclature">
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<heading id="833C81C4FFACFFDBFE36FE2B6A1090D2" box="[496,1092,438,465]" centered="true" fontSize="11" level="2" pageId="33" pageNumber="34" reason="2">
<taxonomicName id="1FCB4D2BFFACFFDBFE36FE2B6D8390D2" authority="Bristow" authorityName="Blandin &amp; Bristow &amp; Neild &amp; Sousa &amp; Gareca &amp; Huertas" authorityYear="2014" box="[496,983,438,465]" class="Hexapoda" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="subSpecies" species="zolvizora" status="subsp. nov." subSpecies="casagrande">
<emphasis id="EABFEABAFFACFFDBFE36FE2B6D2790D2" bold="true" box="[496,883,438,465]" italics="true" pageId="33" pageNumber="34">Eryphanis zolvizora casagrande</emphasis>
Bristow
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<taxonomicNameLabel id="F18C57C1FFACFFDBFC18FE2A6A1090D2" box="[990,1092,439,465]" pageId="33" pageNumber="34" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
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<subSubSection id="90D16523FFACFFDBFDABFE476D9290F6" box="[621,966,474,501]" pageId="33" pageNumber="34" type="description">
<paragraph id="D87436A8FFACFFDBFDABFE476D9290F6" blockId="33.[496,1092,438,501]" box="[621,966,474,501]" pageId="33" pageNumber="34">
<figureCitation id="40F02A2DFFACFFDBFDABFE476CBF90F6" box="[621,747,474,501]" captionStart="Fig" captionStartId="11.[189,232,1824,1850]" captionTargetBox="[225,1364,264,1787]" captionTargetId="figure@11.[224,1363,265,1787]" captionTargetPageId="11" captionText="Fig. 5. Habitus of males. Red spot: holotype (HT) or lectotype (LT). A. Eryphanis zolvizora greeneyi Penz &amp; DeVries, 2008, stat. rev., HT (Yanayacu, Napo, Ecuador; BMNH). B. E. z. greeneyi, specimen with increased forewing spots (Baeza, Napo, Ecuador, Pinas). C. Eryphanis zolvizora casagrande ssp. nov., HT (Altaquer, Nariño, Colombia; ICNUN). D. E. z. casagrande from Western Ecuador (Las Gralarias, Pichincha; FLMNH). E. Eryphanis zolvizora opimus (Staudinger, 1887), LT (Manizales, Colombia; ZMHB). F. E. z. opimus, specimen with reduced spots (Cauca, Colombia; MNHN, CG)." figureDoi="http://doi.org/10.5281/zenodo.3834968" httpUri="https://zenodo.org/record/3834968/files/figure.png" pageId="33" pageNumber="34">Figs 5C-D</figureCitation>
,
<figureCitation id="40F02A2DFFACFFDBFD31FE476D6D90F7" box="[759,825,474,500]" captionStart="Fig" captionStartId="16.[189,232,1790,1816]" captionTargetBox="[224,1363,264,1737]" captionTargetId="figure@16.[224,1363,265,1737]" captionTargetPageId="16" captionText="Fig. 7. Habitus of females. Yellow spot: paratype (PT). A. Eryphanis zolvizora zolvizora (Hewitson, 1877) (San Jacinto, Bolivia; BMNH). B. Eryphanis zolvizora chachapoya ssp. nov., PT (San José de Molinopampa, Amazonas, Peru; MNHN, PBGL 198). C. Eryphanis zolvizora greeneyi Penz &amp; DeVries, 2008, stat. rev. (Ecuador; JFLC). D. Eryphanis zolvizora casagrande ssp. nov., PT (La Planada, Nariño, Colombia; IAvH). E. E. z. casagrande ssp. nov. (Huigra, Chimborazo, Ecuador; ANSP). F. Eryphanis zolvizora opimus (Staudinger, 1887) (Río Agua (presumably Río Aguacatal), Valle del Cauca, Colombia; BMNH)." figureDoi="http://doi.org/10.5281/zenodo.3834972" httpUri="https://zenodo.org/record/3834972/files/figure.png" pageId="33" pageNumber="34">7D-E</figureCitation>
,
<figureCitation id="40F02A2DFFACFFDBFC83FE476DC690F7" box="[837,914,474,500]" captionStart="Fig" captionStartId="20.[189,232,1720,1746]" captionTargetBox="[188,1399,961,1678]" captionTargetId="figure@20.[189,1398,961,1678]" captionTargetPageId="20" captionText="Fig. 13. A-C. Male genitalia of Eryphanis zolvizora greeneyi Penz &amp; DeVries, 2008, stat. rev. A. Specimen from Rio Blanco, Ecuador (MNHN, PBB 729). B. Specimen from Rio Blanco, Ecuador (MNHN, PBB 537). C. Specimen from Rio Blanco, Ecuador (MNHN, PBB 257). D-F. Male genitalia of Eryphanis zolvizora casagrande ssp. nov. D. HT from Altaquer, Nariño, Colombia (ICNUN). E. PT from Ricaurte, Nariño, Colombia (IAvH; drawing by Jean-François Le Crom, JFL 336). F. Specimen from Las Gralarias, Pichincha, Ecuador (FLMNH). G-I. Male genitalia of Eryphanis zolvizora opimus (Staudinger, 1887). G. Specimen from Manizales, Caldas, Colombia (BMNH 8228). H. Specimen from Arménia, Quindio, Colombia (MNHN, CG). I. Specimen from Pereira, Risaralda, Colombia (BMNH 8227)." figureDoi="http://doi.org/10.5281/zenodo.3834984" httpUri="https://zenodo.org/record/3834984/files/figure.png" pageId="33" pageNumber="34">13D-F</figureCitation>
,
<figureCitation id="40F02A2DFFACFFDBFC58FE476D9290F6" box="[926,966,474,501]" captionStart="Fig" captionStartId="22.[189,232,1578,1604]" captionTargetBox="[188,1399,264,1515]" captionTargetId="figure@22.[189,1398,277,1514]" captionTargetPageId="22" captionText="Fig. 15. Female sterigma. A. Eryphanis zolvizora zolvizora (Hewitson, 1877) (Bolivia; MNHN, CG). B-H. Eryphanis zolvizora chachapoya ssp. nov. B. PT from Cushi, Pasco, Peru (BMNH 8226). C. PT from Carpish, Huánuco, Peru (MNHN, PBB 2285). D. Specimen from Mallqui, Huánuco, Peru (MNHN, PBB 2327). E. PT from San José de Molinopampa, Amazonas, Peru (MNHN, PBGL 198). F. PT from Nieva, Amazonas, Peru (MNHN, PBGL 596). G. PT from Alto Nieva, Amazonas, Peru (MNHN, PBGL 519). H. Specimen from Alto Nieva, Amazonas, Peru (MNHN, PBGL 520). I. Eryphanis zolvizora greeneyi Penz &amp; DeVries, 2008, stat. rev., specimen fromValladolid, Zamora-Chinchipe, Ecuador (MNHN, PBB 2288). J. Eryphanis zolvizora casagrande ssp. nov., PT from La Planada, Nariño, Colombia (IAvH; drawing by Jean-François Le Crom, JFL 337). K. E. zolvizora opimus (Staudinger, 1887), specimen from Colombie (MNHN, CG). L. Eryphanis zolvizora reyi ssp. nov., PT from La Mina, Barinas, Venezuela (R, 116-JCSC). M-O. E. zolvizora isabelae ssp. nov. M. PT from Choroní, Aragua, Venezuela (R, 114-JCSC). N. PT from Rancho Grande, Aragua, Venezuela (MIZA, 118-JCSC). O. PT from La Llanada, Distrito Federal, Venezuela (MNHN, PBB 2322)." figureDoi="http://doi.org/10.5281/zenodo.3834988" httpUri="https://zenodo.org/record/3834988/files/figure.png" pageId="33" pageNumber="34">15J</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="90D16523FFACFFDBFF7BFD866C139258" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="D87436A8FFACFFDBFF7BFD866F629336" blockId="33.[189,310,539,565]" box="[189,310,539,565]" pageId="33" pageNumber="34">
<heading id="833C81C4FFACFFDBFF7BFD866F629336" bold="true" box="[189,310,539,565]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="EABFEABAFFACFFDBFF7BFD866F629336" bold="true" box="[189,310,539,565]" pageId="33" pageNumber="34">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="D87436A8FFACFFDBFF7BFDD66C139258" blockId="33.[189,1399,586,859]" pageId="33" pageNumber="34">
HW projection only poorly developed; no R5-M1 orange mark on DFW (
<figureCitation id="40F02A2DFFACFFDBFBC9FDD76A3D9366" box="[1039,1129,586,613]" captionStart="Fig" captionStartId="11.[189,232,1824,1850]" captionTargetBox="[225,1364,264,1787]" captionTargetId="figure@11.[224,1363,265,1787]" captionTargetPageId="11" captionText="Fig. 5. Habitus of males. Red spot: holotype (HT) or lectotype (LT). A. Eryphanis zolvizora greeneyi Penz &amp; DeVries, 2008, stat. rev., HT (Yanayacu, Napo, Ecuador; BMNH). B. E. z. greeneyi, specimen with increased forewing spots (Baeza, Napo, Ecuador, Pinas). C. Eryphanis zolvizora casagrande ssp. nov., HT (Altaquer, Nariño, Colombia; ICNUN). D. E. z. casagrande from Western Ecuador (Las Gralarias, Pichincha; FLMNH). E. Eryphanis zolvizora opimus (Staudinger, 1887), LT (Manizales, Colombia; ZMHB). F. E. z. opimus, specimen with reduced spots (Cauca, Colombia; MNHN, CG)." figureDoi="http://doi.org/10.5281/zenodo.3834968" httpUri="https://zenodo.org/record/3834968/files/figure.png" pageId="33" pageNumber="34">Fig. 5C</figureCitation>
); there is a wide bridge in space Cu2-2A on the VFW formed by a complete white connection between the white stripes (
<figureCitation id="40F02A2DFFACFFDBFAD8FDF06B269384" box="[1310,1394,621,647]" captionStart="Fig" captionStartId="5.[189,232,1823,1849]" captionTargetBox="[188,1399,681,1783]" captionTargetId="figure@5.[189,1398,682,1783]" captionTargetPageId="5" captionText="Fig. 2. Characters studied on the habitus (verso), and their major variations. f. Ground colour of the ventral surface and of the median area. g. Isolated white spot on FW cell. h. White spot at the angle formed by the cubital vein of the FW cell and Cu2. i. Bridge between white stripes in FW cell Cu2-2A. j. Length on HW of posterior extension of the white stripes. k. Black designs in the HW cell, and black curved line anterior to the Cu1-Cu2 ocellus. l. Ring around the HW costal ocellus. m. Widest diameter Φ of the HW Cu1-Cu2 ocellus." figureDoi="http://doi.org/10.5281/zenodo.3834962" httpUri="https://zenodo.org/record/3834962/files/figure.png" pageId="33" pageNumber="34">Figs 2i</figureCitation>
,
<figureCitation id="40F02A2DFFACFFDBFF7BFD0D6EB693A8" box="[189,226,656,683]" captionStart="Fig" captionStartId="11.[189,232,1824,1850]" captionTargetBox="[225,1364,264,1787]" captionTargetId="figure@11.[224,1363,265,1787]" captionTargetPageId="11" captionText="Fig. 5. Habitus of males. Red spot: holotype (HT) or lectotype (LT). A. Eryphanis zolvizora greeneyi Penz &amp; DeVries, 2008, stat. rev., HT (Yanayacu, Napo, Ecuador; BMNH). B. E. z. greeneyi, specimen with increased forewing spots (Baeza, Napo, Ecuador, Pinas). C. Eryphanis zolvizora casagrande ssp. nov., HT (Altaquer, Nariño, Colombia; ICNUN). D. E. z. casagrande from Western Ecuador (Las Gralarias, Pichincha; FLMNH). E. Eryphanis zolvizora opimus (Staudinger, 1887), LT (Manizales, Colombia; ZMHB). F. E. z. opimus, specimen with reduced spots (Cauca, Colombia; MNHN, CG)." figureDoi="http://doi.org/10.5281/zenodo.3834968" httpUri="https://zenodo.org/record/3834968/files/figure.png" pageId="33" pageNumber="34">5C</figureCitation>
); the isolated white spot in the VFW cell is quite large and semicircular; the narrow outer white stripe on the VHW only just extends down to the smaller of the lower ocelli; the broader inner white stripe only extends as far as the larger of the lower ocelli (
<figureCitation id="40F02A2DFFACFFDBFD12FD4A6D7A93F1" box="[724,814,727,754]" captionStart="Fig" captionStartId="11.[189,232,1824,1850]" captionTargetBox="[225,1364,264,1787]" captionTargetId="figure@11.[224,1363,265,1787]" captionTargetPageId="11" captionText="Fig. 5. Habitus of males. Red spot: holotype (HT) or lectotype (LT). A. Eryphanis zolvizora greeneyi Penz &amp; DeVries, 2008, stat. rev., HT (Yanayacu, Napo, Ecuador; BMNH). B. E. z. greeneyi, specimen with increased forewing spots (Baeza, Napo, Ecuador, Pinas). C. Eryphanis zolvizora casagrande ssp. nov., HT (Altaquer, Nariño, Colombia; ICNUN). D. E. z. casagrande from Western Ecuador (Las Gralarias, Pichincha; FLMNH). E. Eryphanis zolvizora opimus (Staudinger, 1887), LT (Manizales, Colombia; ZMHB). F. E. z. opimus, specimen with reduced spots (Cauca, Colombia; MNHN, CG)." figureDoi="http://doi.org/10.5281/zenodo.3834968" httpUri="https://zenodo.org/record/3834968/files/figure.png" pageId="33" pageNumber="34">Fig. 5C</figureCitation>
); the dark brown lines in the VHW cell and curved line above the Cu1-Cu2 ocellus are well developed, the distal lines form two conjoined elongate circles (
<figureCitation id="40F02A2DFFACFFDBFF00FC836F76923B" box="[198,290,797,824]" captionStart="Fig" captionStartId="11.[189,232,1824,1850]" captionTargetBox="[225,1364,264,1787]" captionTargetId="figure@11.[224,1363,265,1787]" captionTargetPageId="11" captionText="Fig. 5. Habitus of males. Red spot: holotype (HT) or lectotype (LT). A. Eryphanis zolvizora greeneyi Penz &amp; DeVries, 2008, stat. rev., HT (Yanayacu, Napo, Ecuador; BMNH). B. E. z. greeneyi, specimen with increased forewing spots (Baeza, Napo, Ecuador, Pinas). C. Eryphanis zolvizora casagrande ssp. nov., HT (Altaquer, Nariño, Colombia; ICNUN). D. E. z. casagrande from Western Ecuador (Las Gralarias, Pichincha; FLMNH). E. Eryphanis zolvizora opimus (Staudinger, 1887), LT (Manizales, Colombia; ZMHB). F. E. z. opimus, specimen with reduced spots (Cauca, Colombia; MNHN, CG)." figureDoi="http://doi.org/10.5281/zenodo.3834968" httpUri="https://zenodo.org/record/3834968/files/figure.png" pageId="33" pageNumber="34">Fig. 5C</figureCitation>
). The valva axis has no swelling; the dorsal ridge has a row of more than 6 spines, diminishing towards the extremity (
<figureCitation id="40F02A2DFFACFFDBFE0BFCDC6C6E9258" box="[461,570,833,859]" captionStart="Fig" captionStartId="20.[189,232,1720,1746]" captionTargetBox="[188,1399,961,1678]" captionTargetId="figure@20.[189,1398,961,1678]" captionTargetPageId="20" captionText="Fig. 13. A-C. Male genitalia of Eryphanis zolvizora greeneyi Penz &amp; DeVries, 2008, stat. rev. A. Specimen from Rio Blanco, Ecuador (MNHN, PBB 729). B. Specimen from Rio Blanco, Ecuador (MNHN, PBB 537). C. Specimen from Rio Blanco, Ecuador (MNHN, PBB 257). D-F. Male genitalia of Eryphanis zolvizora casagrande ssp. nov. D. HT from Altaquer, Nariño, Colombia (ICNUN). E. PT from Ricaurte, Nariño, Colombia (IAvH; drawing by Jean-François Le Crom, JFL 336). F. Specimen from Las Gralarias, Pichincha, Ecuador (FLMNH). G-I. Male genitalia of Eryphanis zolvizora opimus (Staudinger, 1887). G. Specimen from Manizales, Caldas, Colombia (BMNH 8228). H. Specimen from Arménia, Quindio, Colombia (MNHN, CG). I. Specimen from Pereira, Risaralda, Colombia (BMNH 8227)." figureDoi="http://doi.org/10.5281/zenodo.3834984" httpUri="https://zenodo.org/record/3834984/files/figure.png" pageId="33" pageNumber="34">Fig. 13D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="90D16523FFACFFDBFF7BFC1E6C8D92ED" pageId="33" pageNumber="34" type="etymology">
<paragraph id="D87436A8FFACFFDBFF7BFC1E6F10929E" blockId="33.[189,324,899,925]" box="[189,324,899,925]" pageId="33" pageNumber="34">
<heading id="833C81C4FFACFFDBFF7BFC1E6F10929E" bold="true" box="[189,324,899,925]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="EABFEABAFFACFFDBFF7BFC1E6F10929E" bold="true" box="[189,324,899,925]" pageId="33" pageNumber="34">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="D87436A8FFACFFDBFF7BFC2C6C8D92ED" blockId="33.[189,1399,945,1007]" pageId="33" pageNumber="34">
Named after Prof. Dr. Mirna Casagrande (Universidade Federal do
<collectingRegion id="1A0FF84AFFACFFDBFC1FFC2C6A7892C8" box="[985,1068,945,971]" country="Brazil" name="Parana" pageId="33" pageNumber="34">Paraná</collectingRegion>
, Curitiba,
<collectingRegion id="1A0FF84AFFACFFDBFB61FC2C6AAE92C8" box="[1191,1274,945,971]" country="Brazil" name="Parana" pageId="33" pageNumber="34">Paraná</collectingRegion>
,
<collectingCountry id="A0DC7638FFACFFDBFAC0FC2C6B1B92CF" box="[1286,1359,945,972]" name="Brazil" pageId="33" pageNumber="34">Brazil</collectingCountry>
) in recognition of all her work on the brassolines.
</paragraph>
</subSubSection>
<subSubSection id="90D16523FFACFFD8FF7BFB8B6F8A9072" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="D87436A8FFACFFDBFF7BFB8B6C439532" blockId="33.[189,535,1046,1073]" box="[189,535,1046,1073]" pageId="33" pageNumber="34">
<heading id="833C81C4FFACFFDBFF7BFB8B6F389533" box="[189,364,1046,1072]" pageId="33" pageNumber="34">
<emphasis id="EABFEABAFFACFFDBFF7BFB8B6F389533" bold="true" box="[189,364,1046,1072]" pageId="33" pageNumber="34">Type material</emphasis>
</heading>
(
<specimenCount id="CECDFD21FFACFFDBFEBBFB8B6C5B9532" box="[381,527,1046,1073]" pageId="33" pageNumber="34" type="generic">3 specimens</specimenCount>
)
</paragraph>
<paragraph id="D87436A8FFACFFDBFF2CFBD86F08955C" blockId="33.[189,1399,1093,1224]" box="[234,348,1093,1119]" pageId="33" pageNumber="34">
<emphasis id="EABFEABAFFACFFDBFF2CFBD86F08955C" bold="true" box="[234,348,1093,1119]" pageId="33" pageNumber="34">
<typeStatus id="0770880AFFACFFDBFF2CFBD86F08955C" box="[234,348,1093,1119]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="D87436A8FFACFFDBFF7BFBF56C8E95CB" blockId="33.[189,1399,1093,1224]" pageId="33" pageNumber="34">
<materialsCitation id="68A33CF5FFACFFDBFF7BFBF56C8E95CB" ID-GBIF-Occurrence="2620871308" collectingDate="1995-07-24" collectionCode="ICNO" collectorName="G. Andrade-C." country="Colombia" county="Barbacoas" elevation="1425" latitude="1.25" location="Reserva" longLatPrecision="1308" longitude="-78.11667" municipality="Altaquer" pageId="33" pageNumber="34" specimenCode="ICN-MHN-L 14791" specimenCount="1" specimenCount-male="1" stateProvince="Narino" typeStatus="holotype">
<collectingCountry id="A0DC7638FFACFFDBFF7BFBF56F0A9581" box="[189,350,1128,1154]" name="Colombia" pageId="33" pageNumber="34">COLOMBIA</collectingCountry>
:
<specimenCount id="CECDFD21FFACFFDBFEAEFBF56FD49581" box="[360,384,1128,1154]" pageId="33" pageNumber="34" type="male"></specimenCount>
, four white labels printed in black: //
<collectingRegion id="1A0FF84AFFACFFDBFCF7FBF56DD19581" box="[817,901,1128,1154]" country="Colombia" name="Narino" pageId="33" pageNumber="34">Nariño</collectingRegion>
,
<collectingCounty id="31154E24FFACFFDBFC56FBF56A599581" box="[912,1037,1128,1154]" pageId="33" pageNumber="34">Barbacoas</collectingCounty>
,
<collectingMunicipality id="3810ACD2FFACFFDBFBD0FBF56A2B9581" box="[1046,1151,1128,1154]" pageId="33" pageNumber="34">Altaquer</collectingMunicipality>
,
<location id="DD146073FFACFFDBFB4CFBF56ABC9581" LSID="urn:lsid:plazi:treatment:506287BEFFACFFD9FE36FE2B6C809536:DD146073FFACFFDBFB4CFBF56ABC9581" box="[1162,1256,1128,1154]" country="Colombia" county="Barbacoas" latitude="1.25" longLatPrecision="1308" longitude="-78.11667" municipality="Altaquer" name="Reserva" pageId="33" pageNumber="34" stateProvince="Narino">Reserva</location>
<location id="DD146073FFACFFDBFB28FBF56B239581" LSID="urn:lsid:plazi:treatment:506287BEFFACFFD9FE36FE2B6C809536:DD146073FFACFFDBFB28FBF56B239581" box="[1262,1399,1126,1154]" country="Colombia" county="Barbacoas" latitude="1.25" longLatPrecision="1308" longitude="-78.11667" municipality="Altaquer" name="Rio Nambi." pageId="33" pageNumber="34" stateProvince="Narino">Río Ñambi.</location>
[
<geoCoordinate id="BDFF506FFFACFFDBFF03FB166F4995A5" box="[197,285,1163,1190]" degrees="1" direction="north" minutes="15" orientation="latitude" pageId="33" pageNumber="34" precision="925" value="1.25">1°15N</geoCoordinate>
<geoCoordinate id="BDFF506FFFACFFDBFEE2FB176FC795A6" box="[292,403,1162,1189]" degrees="78" direction="west" minutes="07" orientation="longitude" pageId="33" pageNumber="34" precision="925" value="-78.11667">78°07W</geoCoordinate>
] Julio 24 de 1995 [
<date id="AC751068FFACFFDBFDB6FB166D5095A6" box="[624,772,1162,1189]" pageId="33" pageNumber="34" value="1995-07-24">
<collectingDate id="BC31E980FFACFFDBFDB6FB166D5095A6" box="[624,772,1162,1189]" pageId="33" pageNumber="34" value="1995-07-24">24 Jul. 1995</collectingDate>
</date>
]
<collectorName id="753E537EFFACFFDBFCD4FB166DEA95A6" box="[786,958,1163,1189]" pageId="33" pageNumber="34">G. Andrade-C.</collectorName>
Leg. Altitud:
<quantity id="1F339B4DFFACFFDBFBA7FB166AE895A5" box="[1121,1212,1163,1190]" metricMagnitude="3" metricUnit="m" metricValue="1.425" pageId="33" pageNumber="34" unit="m" value="1425.0">
<elevation id="53E6D19BFFACFFDBFBA7FB166AE895A5" box="[1121,1212,1163,1190]" metricMagnitude="3" metricUnit="m" metricValue="1.425" pageId="33" pageNumber="34" unit="m" value="1425.0">1425 m</elevation>
</quantity>
. // GAC: 7615//
<specimenCode id="886D9ED3FFACFFDBFF7BFB336FFD95CB" box="[189,425,1198,1224]" pageId="33" pageNumber="34">ICN-MHN-L 14791</specimenCode>
// ICN 011101 // (
<collectionCode id="BEDAAE6DFFACFFDBFDB9FB336C9995CB" box="[639,717,1198,1224]" pageId="33" pageNumber="34">ICNO</collectionCode>
).
</materialsCitation>
</paragraph>
<materialsCitation id="68A33CF5FFACFFDBFF2CFB6D6A0D9453" collectingDate="1994-09-08" collectionCode="IAvH" collectorName="Julian Salazar" country="Colombia" elevation="1800" latitude="1.085" location="Ricaurte La Planada" longLatPrecision="21" longitude="-77.884995" municipality="Ricaurte La Planada" pageId="33" pageNumber="34" specimenCode="IAvH 8238" specimenCount="1" specimenCount-male="1" stateProvince="Narino" typeStatus="paratype">
<paragraph id="D87436A8FFACFFDBFF2CFB6D6F339409" blockId="33.[189,1399,1264,1431]" box="[234,359,1264,1290]" pageId="33" pageNumber="34">
<emphasis id="EABFEABAFFACFFDBFF2CFB6D6F339409" bold="true" box="[234,359,1264,1290]" pageId="33" pageNumber="34">
<typeStatus id="0770880AFFACFFDBFF2CFB6D6F339409" box="[234,359,1264,1290]" pageId="33" pageNumber="34" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="D87436A8FFACFFDBFF7BFA8E6CA59494" blockId="33.[189,1399,1264,1431]" pageId="33" pageNumber="34">
<collectingCountry id="A0DC7638FFACFFDBFF7BFA8E6F0A942E" box="[189,350,1299,1325]" name="Colombia" pageId="33" pageNumber="34">COLOMBIA</collectingCountry>
:
<specimenCount id="CECDFD21FFACFFDBFEAAFA896FD6942D" box="[364,386,1300,1326]" count="1" pageId="33" pageNumber="34" type="male"></specimenCount>
//
<collectingRegion id="1A0FF84AFFACFFDBFE61FA8E6FAC942E" box="[423,504,1299,1325]" country="Colombia" name="Narino" pageId="33" pageNumber="34">Nariño</collectingRegion>
<collectingMunicipality id="3810ACD2FFACFFDBFDC5FA8E6CA3942E" box="[515,759,1299,1326]" pageId="33" pageNumber="34">Ricaurte La Planada</collectingMunicipality>
<geoCoordinate id="BDFF506FFFACFFDBFCC7FA8E6DC5942D" box="[769,913,1299,1326]" degrees="01" direction="north" minutes="05" orientation="latitude" pageId="33" pageNumber="34" precision="15" seconds="06" value="1.085">01°0506”N</geoCoordinate>
<geoCoordinate id="BDFF506FFFACFFDBFC5DFA8E6A65942D" box="[923,1073,1299,1326]" degrees="77" direction="west" minutes="53" orientation="longitude" pageId="33" pageNumber="34" precision="15" seconds="06" value="-77.884995">77°5306”W</geoCoordinate>
,
<quantity id="1F339B4DFFACFFDBFB87FA8E6AC9942D" box="[1089,1181,1299,1326]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="33" pageNumber="34" unit="m" value="1800.0">
<elevation id="53E6D19BFFACFFDBFB87FA8E6AC9942D" box="[1089,1181,1299,1326]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="33" pageNumber="34" unit="m" value="1800.0">1800 m</elevation>
</quantity>
<date id="AC751068FFACFFDBFB61FA8E6B42942E" box="[1191,1302,1299,1325]" pageId="33" pageNumber="34" value="1994-09-08">
<collectingDate id="BC31E980FFACFFDBFB61FA8E6B42942E" box="[1191,1302,1299,1325]" pageId="33" pageNumber="34" value="1994-09-08">8.ix.1994</collectingDate>
</date>
[
<date id="AC751068FFACFFDBFAEFFA8E6EA99452" pageId="33" pageNumber="34" value="1994-09-08">
<collectingDate id="BC31E980FFACFFDBFAEFFA8E6EA99452" pageId="33" pageNumber="34" value="1994-09-08">8 Sep. 1994</collectingDate>
</date>
],
<collectorName id="753E537EFFACFFDBFED7FAAB6FEC9453" box="[273,440,1334,1360]" pageId="33" pageNumber="34">Julian Salazar</collectorName>
, Leg. SM-4385 //
<specimenCode id="886D9ED3FFACFFDBFD54FAAA6D429453" box="[658,790,1334,1361]" pageId="33" pageNumber="34">IAvH 8238</specimenCode>
// genit JFL 336 // (
<collectionCode id="BEDAAE6DFFACFFDBFBC3FAAB6A189453" box="[1029,1100,1334,1360]" pageId="33" pageNumber="34">IAvH</collectionCode>
);
<materialsCitation id="68A33CF5FFACFFDBFBA7FAAA6CB89494" collectingDate="2000-10-16" collectingDateMax="2000-11-02" collectingDateMin="2000-10-16" collectingMethod="Malaise" collectionCode="IAvH" collectorName="G. Oliva Leg. M." elevation="1930" latitude="1.25" location="La Planada Via Hondon" longLatPrecision="1308" longitude="-78.25" municipality="La Planada Via Hondon" pageId="33" pageNumber="34" specimenCode="IAvH-E 67776" specimenCount="1" specimenCount-female="1" stateProvince="Narino" typeStatus="paratype">
<specimenCount id="CECDFD21FFACFFDBFBA7FAAA6A239452" box="[1121,1143,1335,1361]" count="1" pageId="33" pageNumber="34" type="female"></specimenCount>
//
<collectingRegion id="1A0FF84AFFACFFDBFB5EFAAB6ABD9453" box="[1176,1257,1334,1360]" country="Colombia" name="Narino" pageId="33" pageNumber="34">Nariño</collectingRegion>
<collectingMunicipality id="3810ACD2FFACFFDBFB34FAAA6F1F9470" pageId="33" pageNumber="34">La Planada Via Hondon</collectingMunicipality>
<geoCoordinate id="BDFF506FFFACFFDBFE97FAC46FFC9477" box="[337,424,1369,1396]" degrees="1" direction="north" minutes="15" orientation="latitude" pageId="33" pageNumber="34" precision="925" value="1.25">1°15N</geoCoordinate>
<geoCoordinate id="BDFF506FFFACFFDBFE69FAC46C4E9477" box="[431,538,1369,1396]" degrees="78" direction="west" minutes="15" orientation="longitude" pageId="33" pageNumber="34" precision="925" value="-78.25">78°15W</geoCoordinate>
<quantity id="1F339B4DFFACFFDBFDE6FAC76C2D9477" box="[544,633,1370,1396]" metricMagnitude="3" metricUnit="m" metricValue="1.93" pageId="33" pageNumber="34" unit="m" value="1930.0">
<elevation id="53E6D19BFFACFFDBFDE6FAC76C2D9477" box="[544,633,1370,1396]" metricMagnitude="3" metricUnit="m" metricValue="1.93" pageId="33" pageNumber="34" unit="m" value="1930.0">1930 m</elevation>
</quantity>
<collectingMethod id="018A4EBFFFACFFDBFDB9FAC46C889470" box="[639,732,1369,1395]" pageId="33" pageNumber="34">Malaise</collectingMethod>
<date id="AC751068FFACFFDBFD25FAC46D6C9470" box="[739,824,1369,1396]" pageId="33" pageNumber="34" value="2016-10-02">16 x. 2.</date>
xi. 2000 [
<date id="AC751068FFACFFDBFC61FAC46AC99477" box="[935,1181,1369,1396]" pageId="33" pageNumber="34" value="2000-10-16" valueMax="2000-11-02" valueMin="2000-10-16">
<collectingDate id="BC31E980FFACFFDBFC61FAC46AC99477" box="[935,1181,1369,1396]" pageId="33" pageNumber="34" value="2000-10-16" valueMax="2000-11-02" valueMin="2000-10-16">16 Oct.-2 Nov. 2000</collectingDate>
</date>
].
<collectorName id="753E537EFFACFFDBFB69FAC76B239477" box="[1199,1399,1369,1396]" pageId="33" pageNumber="34">G. Oliva Leg. M.</collectorName>
139 //
<specimenCode id="886D9ED3FFACFFDBFECEFAE06FE19494" box="[264,437,1405,1431]" pageId="33" pageNumber="34">IAvH-E 67776</specimenCode>
// genit JFL 337 // (
<collectionCode id="BEDAAE6DFFACFFDBFD5BFAE06CB09494" box="[669,740,1405,1431]" pageId="33" pageNumber="34">IAvH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</materialsCitation>
<paragraph id="D87436A8FFACFFDBFF7BFA236FAD94DB" blockId="33.[189,505,1470,1496]" box="[189,505,1470,1496]" pageId="33" pageNumber="34">
<heading id="833C81C4FFACFFDBFF7BFA236FAD94DB" bold="true" box="[189,505,1470,1496]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="EABFEABAFFACFFDBFF7BFA236FAD94DB" bold="true" box="[189,505,1470,1496]" pageId="33" pageNumber="34">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="D87436A8FFACFFDBFF7BFA706A97974E" blockId="33.[189,1399,1517,1614]" pageId="33" pageNumber="34">
Because so few specimens have been seen from the west of
<collectingCountry id="A0DC7638FFACFFDBFCB0FA706DBE9704" box="[886,1002,1517,1543]" name="Colombia" pageId="33" pageNumber="34">Colombia</collectingCountry>
(
<specimenCount id="CECDFD21FFACFFDBFC3FFA706A699704" box="[1017,1085,1517,1543]" pageId="33" pageNumber="34" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CECDFD21FFACFFDBFB8FFA706A279704" box="[1097,1139,1517,1543]" pageId="33" pageNumber="34" type="female">1 ♀</specimenCount>
) and
<collectingCountry id="A0DC7638FFACFFDBFB70FA706B4E9704" box="[1206,1306,1517,1543]" name="Ecuador" pageId="33" pageNumber="34">Ecuador</collectingCountry>
, (
<specimenCount id="CECDFD21FFACFFDBFAF6FA706B089704" box="[1328,1372,1517,1543]" pageId="33" pageNumber="34" type="male">1 ♂</specimenCount>
,
<specimenCount id="CECDFD21FFACFFDBFAAEFA706EB39728" pageId="33" pageNumber="34" type="female">2 ♀♀</specimenCount>
) it is possible that, when more specimens are examined, the Ecuadorian population may prove to be a separate taxon. Therefore, the following specimens are not included in the
<typeStatus id="0770880AFFACFFDBFBF8F9A96A25974D" box="[1086,1137,1588,1614]" pageId="33" pageNumber="34">type</typeStatus>
series:
</paragraph>
<paragraph id="D87436A8FFACFFDBFF7BF9E86D4C97FA" blockId="33.[189,1399,1652,1785]" pageId="33" pageNumber="34">
<collectingCountry id="A0DC7638FFACFFDBFF7BF9E86F04978C" box="[189,336,1653,1679]" name="Ecuador" pageId="33" pageNumber="34">ECUADOR</collectingCountry>
:
<specimenCount id="CECDFD21FFACFFDBFE9CF9EB6F269793" box="[346,370,1654,1680]" pageId="33" pageNumber="34" type="male"></specimenCount>
, //
<collectingRegion id="1A0FF84AFFACFFDBFE52F9E86C5F978C" box="[404,523,1653,1679]" country="Ecuador" name="Pichincha" pageId="33" pageNumber="34">Pichincha</collectingRegion>
, Reserva Las Galarias [sic], Nunbird fr. 0°034”S, 78°4350”W,
<date id="AC751068FFACFFDBFB3AF9E86B22978C" box="[1276,1398,1653,1679]" pageId="33" pageNumber="34" value="2011-04-27">27.iv.2011</date>
[
<date id="AC751068FFACFFDBFF02F9056F2597B0" box="[196,369,1688,1715]" pageId="33" pageNumber="34" value="2011-04-27">27 Apr. 2011</date>
]/T Kell/#008/FLMNH-MGCL-15-11-26/DNA Voucher lep-04376 (FLMNH);
<specimenCount id="CECDFD21FFACFFDBFAFFF9046B1B97B0" box="[1337,1359,1689,1715]" pageId="33" pageNumber="34" type="female"></specimenCount>
, // Huigra,
<quantity id="1F339B4DFFACFFDBFEDAF9216F2397D5" box="[284,375,1724,1750]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="33" pageNumber="34" unit="m" value="1220.0">1220 m</quantity>
, [2°18S 78°59W]
<date id="AC751068FFACFFDBFDA7F9216C8897D6" box="[609,732,1723,1750]" pageId="33" pageNumber="34" value="1926-11">Nov. 1926</date>
, Coxey // (ANSP);
<specimenCount id="CECDFD21FFACFFDBFC79F9216D8197D5" box="[959,981,1724,1750]" pageId="33" pageNumber="34" type="female"></specimenCount>
, //
<collectingRegion id="1A0FF84AFFACFFDBFC3FF9266AE097D5" box="[1017,1204,1723,1750]" country="Ecuador" name="Pichincha" pageId="33" pageNumber="34">Pichincha Prov.</collectingRegion>
, Santo Domingo de los Colorados, alt.
<quantity id="1F339B4DFFACFFDBFE7BF9426C5D97FA" box="[445,521,1759,1785]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="33" pageNumber="34" unit="m" value="200.0">200 m</quantity>
,
<date id="AC751068FFACFFDBFDD2F9426CDE97FA" box="[532,650,1759,1785]" pageId="33" pageNumber="34" value="1982-04">Apr. 1982</date>
// (LACM).
</paragraph>
<paragraph id="D87436A8FFACFFDBFF7BF8BC6DE996E8" blockId="33.[189,1399,1824,2027]" pageId="33" pageNumber="34">
A male labelled “EKWADOR [sic], Prov.
<collectingRegion id="1A0FF84AFFACFFDBFD73F8BD6DC29639" box="[693,918,1824,1850]" country="Ecuador" name="Zamora-Chinchipe" pageId="33" pageNumber="34">Zamora-Chinchipe</collectingRegion>
, Valladolid,
<date id="AC751068FFACFFDBFBEDF8BD6AE79639" box="[1067,1203,1824,1851]" pageId="33" pageNumber="34" value="1998-05-17">17 05 1998</date>
[
<date id="AC751068FFACFFDBFB05F8BD6B3F9638" box="[1219,1387,1824,1851]" pageId="33" pageNumber="34" value="1998-05-17">17 May 1998</date>
], leg. K. Jasinski” (MZUJ), exhibits the white bridge on VFW, and the broader inner white stripe on VHW only extends as far as the larger of the lower ocelli, as in
<emphasis id="EABFEABAFFACFFDBFCA0F8F56A789682" box="[870,1068,1895,1921]" italics="true" pageId="33" pageNumber="34">E. z. casagrande</emphasis>
ssp. nov. Other features are consistent with the above diagnosis. Keith Willmott (pers. comm.) commented that the locality may be not reliable, and we suspect an inversion of labels with the
<emphasis id="EABFEABAFFACFFDBFC4DF8336A7A96C4" box="[907,1070,1966,1991]" italics="true" pageId="33" pageNumber="34">
E. z.
<taxonomicName id="1FCB4D2BFFACFFDBFC0CF8336A7A96C4" authorityName="Penz &amp; DeVries" authorityYear="2008" baseAuthorityName="Blandin &amp; Bristow &amp; Neild &amp; Sousa &amp; Gareca &amp; Huertas" baseAuthorityYear="2008" box="[970,1070,1966,1991]" class="Hexapoda" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="greeneyi">greeneyi</taxonomicName>
</emphasis>
-like specimen, in the same collection, which bears a label from the western
<collectingRegion id="1A0FF84AFFACFFDBFD30F84C6DED96E8" box="[758,953,2001,2027]" country="Ecuador" name="El Oro" pageId="33" pageNumber="34">El Oro Province</collectingRegion>
.
</paragraph>
<paragraph id="D87436A8FFAFFFD8FF7BFE916F8A9072" blockId="34.[189,1399,266,370]" pageId="34" pageNumber="35">
In addition, we have seen a web image of a male at Tandayapa Lodge (
<quantity id="1F339B4DFFAFFFD8FBE2FE966AD09025" box="[1060,1156,267,294]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="34" pageNumber="35" unit="m" value="1600.0">1600 m</quantity>
,
<date id="AC751068FFAFFFD8FB55FE916B1C9026" box="[1171,1352,266,294]" pageId="34" pageNumber="35" value="2007-11-28">
Nov. 28
<superScript id="2FBE9BE0FFAFFFD8FB35FE976B54901B" attach="left" box="[1267,1280,266,280]" fontSize="6" pageId="34" pageNumber="35">th</superScript>
2007
</date>
, W. Cook), and there is a picture of a male (presumably) in
<bibRefCitation id="BC5A4B59FFAFFFD8FC83FEAC6D8D9048" author="Silva" box="[837,985,305,331]" firstAuthor="Silva" pageId="34" pageNumber="35" refId="ref30845" refString="Silva X. 2011. Ecologia de Mariposas del Ecuador. Universidad San Francisco, Quito." type="book" year="2011">Silva (2011)</bibRefCitation>
, taken at the Río Guajalito Reserve (
<quantity id="1F339B4DFFAFFFD8FF03FECA6F769071" box="[197,290,343,370]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="34" pageNumber="35" unit="m" value="1900.0">1900 m</quantity>
,
<date id="AC751068FFAFFFD8FEE8FECA6F869072" box="[302,466,343,370]" pageId="34" pageNumber="35" value="2009-11-12">12 Nov. 2009</date>
).
</paragraph>
</subSubSection>
<subSubSection id="90D16523FFAFFFD8FF7BFE066C639675" pageId="34" pageNumber="35" type="description">
<paragraph id="D87436A8FFAFFFD8FF7BFE066F1A90B6" blockId="34.[189,334,411,437]" box="[189,334,411,437]" pageId="34" pageNumber="35">
<heading id="833C81C4FFAFFFD8FF7BFE066F1A90B6" bold="true" box="[189,334,411,437]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="EABFEABAFFAFFFD8FF7BFE066F1A90B6" bold="true" box="[189,334,411,437]" pageId="34" pageNumber="35">Description</emphasis>
</heading>
</paragraph>
<paragraph id="D87436A8FFAFFFD8FF2CFE506F7E90E4" blockId="34.[189,1399,461,1320]" box="[234,298,461,487]" pageId="34" pageNumber="35">
<emphasis id="EABFEABAFFAFFFD8FF2CFE506F7E90E4" bold="true" box="[234,298,461,487]" pageId="34" pageNumber="35">Male</emphasis>
</paragraph>
<paragraph id="D87436A8FFAFFFD8FF7BFE6F6CC39424" blockId="34.[189,1399,461,1320]" pageId="34" pageNumber="35">
The FW length in the
<typeStatus id="0770880AFFAFFFD8FE7CFE6F6C75930F" box="[442,545,498,524]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
is
<quantity id="1F339B4DFFAFFFD8FD87FE6F6CC1930E" box="[577,661,498,525]" metricMagnitude="-2" metricUnit="m" metricValue="5.8" pageId="34" pageNumber="35" unit="mm" value="58.0">58 mm</quantity>
, in the
<typeStatus id="0770880AFFAFFFD8FD21FE6E6D18930E" box="[743,844,499,525]" pageId="34" pageNumber="35" type="paratype">paratype</typeStatus>
<quantity id="1F339B4DFFAFFFD8FC94FE6F6DF3930E" box="[850,935,498,525]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="34" pageNumber="35" unit="mm" value="62.0">62 mm</quantity>
. The values of the
<taxonomicName id="1FCB4D2BFFAFFFD8FBB9FE6E6A81930F" box="[1151,1237,498,525]" class="Hexapoda" family="Nymphalidae" genus="Pavonia" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="index">P index</taxonomicName>
are estimated at
<quantity id="1F339B4DFFAFFFD8FF1DFD856F639330" box="[219,311,536,563]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="34" pageNumber="35" unit="mm" value="5.5">5.5 mm</quantity>
in the
<typeStatus id="0770880AFFAFFFD8FE48FD856FA19331" box="[398,501,536,562]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
and
<quantity id="1F339B4DFFAFFFD8FDF7FD856CD99330" box="[561,653,536,563]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="34" pageNumber="35" unit="mm" value="4.4">4.4 mm</quantity>
in the Ecuadorian specimen from Las Gralarias. These values are close to those observed in
<emphasis id="EABFEABAFFAFFFD8FDE0FDA26C93935B" box="[550,711,575,600]" italics="true" pageId="34" pageNumber="35">
E. z.
<taxonomicName id="1FCB4D2BFFAFFFD8FDA5FDA26C93935B" authorityName="Penz &amp; DeVries" authorityYear="2008" baseAuthorityName="Blandin &amp; Bristow &amp; Neild &amp; Sousa &amp; Gareca &amp; Huertas" baseAuthorityYear="2008" box="[611,711,575,600]" class="Hexapoda" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="greeneyi">greeneyi</taxonomicName>
</emphasis>
; possibly, the HW projection should be more pronounced than in
<emphasis id="EABFEABAFFAFFFD8FED5FDF86FF5937D" box="[275,417,613,638]" italics="true" pageId="34" pageNumber="35">
E. z.
<taxonomicName id="1FCB4D2BFFAFFFD8FE8AFDF86FF5937D" authorityName="Staudinger" authorityYear="1887" box="[332,417,613,638]" class="Hexapoda" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="opimus">opimus</taxonomicName>
</emphasis>
. There is no R5-M1 orange mark on DFW; the orange DFW band is either moderate (HT;
<figureCitation id="40F02A2DFFAFFFD8FF3AFD146F0193A7" box="[252,341,649,676]" captionStart="Fig" captionStartId="11.[189,232,1824,1850]" captionTargetBox="[225,1364,264,1787]" captionTargetId="figure@11.[224,1363,265,1787]" captionTargetPageId="11" captionText="Fig. 5. Habitus of males. Red spot: holotype (HT) or lectotype (LT). A. Eryphanis zolvizora greeneyi Penz &amp; DeVries, 2008, stat. rev., HT (Yanayacu, Napo, Ecuador; BMNH). B. E. z. greeneyi, specimen with increased forewing spots (Baeza, Napo, Ecuador, Pinas). C. Eryphanis zolvizora casagrande ssp. nov., HT (Altaquer, Nariño, Colombia; ICNUN). D. E. z. casagrande from Western Ecuador (Las Gralarias, Pichincha; FLMNH). E. Eryphanis zolvizora opimus (Staudinger, 1887), LT (Manizales, Colombia; ZMHB). F. E. z. opimus, specimen with reduced spots (Cauca, Colombia; MNHN, CG)." figureDoi="http://doi.org/10.5281/zenodo.3834968" httpUri="https://zenodo.org/record/3834968/files/figure.png" pageId="34" pageNumber="35">Fig. 5C</figureCitation>
) or well developed and composed of five spots of variable size which are just contiguous, being separated only by the veins; in that way,
<emphasis id="EABFEABAFFAFFFD8FD1BFD2D6DF493CA" box="[733,928,687,713]" italics="true" pageId="34" pageNumber="35">E. z. casagrande</emphasis>
ssp. nov. may differ from
<emphasis id="EABFEABAFFAFFFD8FB14FD2D6B3B93CA" box="[1234,1391,688,713]" italics="true" pageId="34" pageNumber="35">
E. z.
<taxonomicName id="1FCB4D2BFFAFFFD8FACDFD2D6B3B93CA" authorityName="Penz &amp; DeVries" authorityYear="2008" baseAuthorityName="Blandin &amp; Bristow &amp; Neild &amp; Sousa &amp; Gareca &amp; Huertas" baseAuthorityYear="2008" box="[1291,1391,688,713]" class="Hexapoda" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="greeneyi">greeneyi</taxonomicName>
</emphasis>
, where the spots are often separated. The iridescence is extremely faint. The androconial patch of the
<typeStatus id="0770880AFFAFFFD8FF7BFD666F709216" box="[189,292,763,789]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
is less lemon-coloured than in
<emphasis id="EABFEABAFFAFFFD8FD63FD616D6F9216" box="[677,827,764,789]" italics="true" pageId="34" pageNumber="35">
E. z.
<taxonomicName id="1FCB4D2BFFAFFFD8FD20FD616D6F9216" authorityName="Staudinger" authorityYear="1887" box="[742,827,764,789]" class="Hexapoda" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="opimus">opimus</taxonomicName>
</emphasis>
(
<figureCitation id="40F02A2DFFAFFFD8FC88FD666DF99215" box="[846,941,763,790]" captionStart="Fig" captionStartId="11.[189,232,1824,1850]" captionTargetBox="[225,1364,264,1787]" captionTargetId="figure@11.[224,1363,265,1787]" captionTargetPageId="11" captionText="Fig. 5. Habitus of males. Red spot: holotype (HT) or lectotype (LT). A. Eryphanis zolvizora greeneyi Penz &amp; DeVries, 2008, stat. rev., HT (Yanayacu, Napo, Ecuador; BMNH). B. E. z. greeneyi, specimen with increased forewing spots (Baeza, Napo, Ecuador, Pinas). C. Eryphanis zolvizora casagrande ssp. nov., HT (Altaquer, Nariño, Colombia; ICNUN). D. E. z. casagrande from Western Ecuador (Las Gralarias, Pichincha; FLMNH). E. Eryphanis zolvizora opimus (Staudinger, 1887), LT (Manizales, Colombia; ZMHB). F. E. z. opimus, specimen with reduced spots (Cauca, Colombia; MNHN, CG)." figureDoi="http://doi.org/10.5281/zenodo.3834968" httpUri="https://zenodo.org/record/3834968/files/figure.png" pageId="34" pageNumber="35">Fig. 5C</figureCitation>
); however, the
<typeStatus id="0770880AFFAFFFD8FBADFD616A849215" box="[1131,1232,764,790]" pageId="34" pageNumber="35" type="paratype">paratype</typeStatus>
male and the single Ecuadorian male examined have a very pale cream androconial patch (
<figureCitation id="40F02A2DFFAFFFD8FBB7FCBC6A87923F" box="[1137,1235,801,828]" captionStart="Fig" captionStartId="11.[189,232,1824,1850]" captionTargetBox="[225,1364,264,1787]" captionTargetId="figure@11.[224,1363,265,1787]" captionTargetPageId="11" captionText="Fig. 5. Habitus of males. Red spot: holotype (HT) or lectotype (LT). A. Eryphanis zolvizora greeneyi Penz &amp; DeVries, 2008, stat. rev., HT (Yanayacu, Napo, Ecuador; BMNH). B. E. z. greeneyi, specimen with increased forewing spots (Baeza, Napo, Ecuador, Pinas). C. Eryphanis zolvizora casagrande ssp. nov., HT (Altaquer, Nariño, Colombia; ICNUN). D. E. z. casagrande from Western Ecuador (Las Gralarias, Pichincha; FLMNH). E. Eryphanis zolvizora opimus (Staudinger, 1887), LT (Manizales, Colombia; ZMHB). F. E. z. opimus, specimen with reduced spots (Cauca, Colombia; MNHN, CG)." figureDoi="http://doi.org/10.5281/zenodo.3834968" httpUri="https://zenodo.org/record/3834968/files/figure.png" pageId="34" pageNumber="35">Fig. 5D</figureCitation>
). The ground colour of the verso is lighter, more yellowish than in
<emphasis id="EABFEABAFFAFFFD8FC8DFCD56A4E9262" box="[843,1050,839,865]" italics="true" pageId="34" pageNumber="35">E. z. chachapoya</emphasis>
ssp. nov. The isolated white spot in the VFW cell is quite large and semicircular. On the VFW, there is no white spot in the angle formed by the cubital vein and Cu2; the complete white connection between the white stripes in space Cu2-2A delimits half-a-circle (up against Cu2) with a chestnut-brown inner core and a dark brown outer ring (
<figureCitation id="40F02A2DFFAFFFD8FF3BFC436F0E92FA" box="[253,346,990,1017]" captionStart="Fig" captionStartId="11.[189,232,1824,1850]" captionTargetBox="[225,1364,264,1787]" captionTargetId="figure@11.[224,1363,265,1787]" captionTargetPageId="11" captionText="Fig. 5. Habitus of males. Red spot: holotype (HT) or lectotype (LT). A. Eryphanis zolvizora greeneyi Penz &amp; DeVries, 2008, stat. rev., HT (Yanayacu, Napo, Ecuador; BMNH). B. E. z. greeneyi, specimen with increased forewing spots (Baeza, Napo, Ecuador, Pinas). C. Eryphanis zolvizora casagrande ssp. nov., HT (Altaquer, Nariño, Colombia; ICNUN). D. E. z. casagrande from Western Ecuador (Las Gralarias, Pichincha; FLMNH). E. Eryphanis zolvizora opimus (Staudinger, 1887), LT (Manizales, Colombia; ZMHB). F. E. z. opimus, specimen with reduced spots (Cauca, Colombia; MNHN, CG)." figureDoi="http://doi.org/10.5281/zenodo.3834968" httpUri="https://zenodo.org/record/3834968/files/figure.png" pageId="34" pageNumber="35">Fig. 5C</figureCitation>
). The narrow outer white stripe on the VHW only just extends down to the smaller of the lower ocelli; the broader inner white stripe only extends as far as the larger of the lower ocelli. The dark brown lines in the VHW cell and curved line above the Cu1-Cu2 ocellus are well developed; the distal lines form two conjoined elongate circles (
<figureCitation id="40F02A2DFFAFFFD8FD71FBCD6D419568" box="[695,789,1104,1131]" captionStart="Fig" captionStartId="11.[189,232,1824,1850]" captionTargetBox="[225,1364,264,1787]" captionTargetId="figure@11.[224,1363,265,1787]" captionTargetPageId="11" captionText="Fig. 5. Habitus of males. Red spot: holotype (HT) or lectotype (LT). A. Eryphanis zolvizora greeneyi Penz &amp; DeVries, 2008, stat. rev., HT (Yanayacu, Napo, Ecuador; BMNH). B. E. z. greeneyi, specimen with increased forewing spots (Baeza, Napo, Ecuador, Pinas). C. Eryphanis zolvizora casagrande ssp. nov., HT (Altaquer, Nariño, Colombia; ICNUN). D. E. z. casagrande from Western Ecuador (Las Gralarias, Pichincha; FLMNH). E. Eryphanis zolvizora opimus (Staudinger, 1887), LT (Manizales, Colombia; ZMHB). F. E. z. opimus, specimen with reduced spots (Cauca, Colombia; MNHN, CG)." figureDoi="http://doi.org/10.5281/zenodo.3834968" httpUri="https://zenodo.org/record/3834968/files/figure.png" pageId="34" pageNumber="35">Fig. 5C</figureCitation>
). The costal ocellus on the VHW has a narrow, offwhite outer ring; the black ring is almost complete. In the
<typeStatus id="0770880AFFAFFFD8FCB4FBEB6D889593" box="[882,988,1142,1168]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
, on the VHW, the largest diameter of Cu1-Cu2 ocellus measures
<quantity id="1F339B4DFFAFFFD8FDDBFB016CD195B5" box="[541,645,1180,1206]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="34" pageNumber="35" unit="mm" value="10.1">10.1 mm</quantity>
(cf.
<tableCitation id="95490313FFAFFFD8FD7CFB066D4695B5" box="[698,786,1179,1206]" captionStart="Table 5" captionStartId="14.[189,258,1093,1119]" captionTargetId="graphics@14.[189,1398,1184,1861]" captionTargetPageId="14" captionText="Table 5. Variations of the larger diameter (Φ) of the Cu1-Cu2 ocellus on male ventral hindwing. (n = number of individuals on which measurements have been made, directly or on scaled pictures)." httpUri="http://table.plazi.org/id/8CB46620FF83FFF4FF7BFBD86B4E9580" pageId="34" pageNumber="35" tableUuid="8CB46620FF83FFF4FF7BFBD86B4E9580">Table 5</tableCitation>
); it is weakly larger in the
<typeStatus id="0770880AFFAFFFD8FB8CFB016AE795B5" box="[1098,1203,1180,1206]" pageId="34" pageNumber="35" type="paratype">paratype</typeStatus>
; thus, there is no indication of a difference with either
<emphasis id="EABFEABAFFAFFFD8FDA8FB5F6D5F95D8" box="[622,779,1218,1243]" italics="true" pageId="34" pageNumber="35">
E. z.
<taxonomicName id="1FCB4D2BFFAFFFD8FD61FB5F6D5F95D8" authorityName="Penz &amp; DeVries" authorityYear="2008" baseAuthorityName="Blandin &amp; Bristow &amp; Neild &amp; Sousa &amp; Gareca &amp; Huertas" baseAuthorityYear="2008" box="[679,779,1218,1243]" class="Hexapoda" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="greeneyi">greeneyi</taxonomicName>
</emphasis>
or
<emphasis id="EABFEABAFFAFFFD8FCF7FB5F6DEB95D8" box="[817,959,1218,1243]" italics="true" pageId="34" pageNumber="35">
E. z.
<taxonomicName id="1FCB4D2BFFAFFFD8FCACFB5F6DEB95D8" authorityName="Staudinger" authorityYear="1887" box="[874,959,1218,1243]" class="Hexapoda" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="opimus">opimus</taxonomicName>
</emphasis>
. Genitalia are similar to those of
<emphasis id="EABFEABAFFAFFFD8FA82FB5F6F759402" italics="true" pageId="34" pageNumber="35">
E. z.
<taxonomicName id="1FCB4D2BFFAFFFD8FF7BFB756F759402" authorityName="Penz &amp; DeVries" authorityYear="2008" baseAuthorityName="Blandin &amp; Bristow &amp; Neild &amp; Sousa &amp; Gareca &amp; Huertas" baseAuthorityYear="2008" box="[189,289,1256,1281]" class="Hexapoda" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="greeneyi">greeneyi</taxonomicName>
</emphasis>
and other subspecies from the Northern Andes, while they strongly differ from the genitalia of
<emphasis id="EABFEABAFFAFFFD8FF7BFA936FD29424" box="[189,390,1293,1319]" italics="true" pageId="34" pageNumber="35">E. z. chachapoya</emphasis>
ssp. nov. (
<figureCitation id="40F02A2DFFAFFFD8FDC4FA906CDF9424" box="[514,651,1293,1319]" captionStart="Fig" captionStartId="20.[189,232,1720,1746]" captionTargetBox="[188,1399,961,1678]" captionTargetId="figure@20.[189,1398,961,1678]" captionTargetPageId="20" captionText="Fig. 13. A-C. Male genitalia of Eryphanis zolvizora greeneyi Penz &amp; DeVries, 2008, stat. rev. A. Specimen from Rio Blanco, Ecuador (MNHN, PBB 729). B. Specimen from Rio Blanco, Ecuador (MNHN, PBB 537). C. Specimen from Rio Blanco, Ecuador (MNHN, PBB 257). D-F. Male genitalia of Eryphanis zolvizora casagrande ssp. nov. D. HT from Altaquer, Nariño, Colombia (ICNUN). E. PT from Ricaurte, Nariño, Colombia (IAvH; drawing by Jean-François Le Crom, JFL 336). F. Specimen from Las Gralarias, Pichincha, Ecuador (FLMNH). G-I. Male genitalia of Eryphanis zolvizora opimus (Staudinger, 1887). G. Specimen from Manizales, Caldas, Colombia (BMNH 8228). H. Specimen from Arménia, Quindio, Colombia (MNHN, CG). I. Specimen from Pereira, Risaralda, Colombia (BMNH 8227)." figureDoi="http://doi.org/10.5281/zenodo.3834984" httpUri="https://zenodo.org/record/3834984/files/figure.png" pageId="34" pageNumber="35">Fig. 13D-F</figureCitation>
).
</paragraph>
<paragraph id="D87436A8FFAFFFD8FF7BFACF6DC0972A" blockId="34.[189,1399,1361,1577]" pageId="34" pageNumber="35">
<emphasis id="EABFEABAFFAFFFD8FF7BFACF6EA09468" box="[189,244,1362,1387]" italics="true" pageId="34" pageNumber="35">Note</emphasis>
. One striking feature of the male from Las Gralarias,
<collectingCountry id="A0DC7638FFAFFFD8FCBBFACC6DB59468" box="[893,993,1361,1387]" name="Ecuador" pageId="34" pageNumber="35">Ecuador</collectingCountry>
, is the circular orange spot with a large dark brown core in space M3-Cu1 of the dorsal forewing band (
<figureCitation id="40F02A2DFFAFFFD8FC37FAEA6A049491" box="[1009,1104,1399,1426]" captionStart="Fig" captionStartId="11.[189,232,1824,1850]" captionTargetBox="[225,1364,264,1787]" captionTargetId="figure@11.[224,1363,265,1787]" captionTargetPageId="11" captionText="Fig. 5. Habitus of males. Red spot: holotype (HT) or lectotype (LT). A. Eryphanis zolvizora greeneyi Penz &amp; DeVries, 2008, stat. rev., HT (Yanayacu, Napo, Ecuador; BMNH). B. E. z. greeneyi, specimen with increased forewing spots (Baeza, Napo, Ecuador, Pinas). C. Eryphanis zolvizora casagrande ssp. nov., HT (Altaquer, Nariño, Colombia; ICNUN). D. E. z. casagrande from Western Ecuador (Las Gralarias, Pichincha; FLMNH). E. Eryphanis zolvizora opimus (Staudinger, 1887), LT (Manizales, Colombia; ZMHB). F. E. z. opimus, specimen with reduced spots (Cauca, Colombia; MNHN, CG)." figureDoi="http://doi.org/10.5281/zenodo.3834968" httpUri="https://zenodo.org/record/3834968/files/figure.png" pageId="34" pageNumber="35">Fig. 5D</figureCitation>
). This is unique amongst all the
<taxonomicName id="1FCB4D2BFFAFFFD8FED6FA036FCA94B4" box="[272,414,1437,1463]" class="Hexapoda" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="zolvizora">
<emphasis id="EABFEABAFFAFFFD8FED6FA036FCA94B4" box="[272,414,1437,1463]" italics="true" pageId="34" pageNumber="35">E. zolvizora</emphasis>
</taxonomicName>
specimens examined. However, a smaller dark brown point is present in one male from the Río Aguacatal (eastern slope of the Colombian western cordillera), collected by Fassl (USNM), a male from “Cali” in MNHN (PBB 1406), and two males of
<emphasis id="EABFEABAFFAFFFD8FC4BFA746A019701" box="[909,1109,1512,1538]" italics="true" pageId="34" pageNumber="35">E. z. chachapoya</emphasis>
ssp. nov. from the upper Río Nieva (Amazonas,
<collectingCountry id="A0DC7638FFAFFFD8FE16F9926C5C972A" box="[464,520,1551,1577]" name="Peru" pageId="34" pageNumber="35">Peru</collectingCountry>
) in MNHN (PBGL 471 and 518).
</paragraph>
<paragraph id="D87436A8FFAFFFD8FF2CF9CE6F11976E" blockId="34.[189,1399,1619,1910]" box="[234,325,1619,1645]" pageId="34" pageNumber="35">
<emphasis id="EABFEABAFFAFFFD8FF2CF9CE6F11976E" bold="true" box="[234,325,1619,1645]" pageId="34" pageNumber="35">Female</emphasis>
</paragraph>
<paragraph id="D87436A8FFAFFFD8FF7BF9E56C639675" blockId="34.[189,1399,1619,1910]" pageId="34" pageNumber="35">
The
<typeStatus id="0770880AFFAFFFD8FF34F9E46F039790" box="[242,343,1657,1683]" pageId="34" pageNumber="35" type="paratype">paratype</typeStatus>
female FWL is
<quantity id="1F339B4DFFAFFFD8FDD3F9E56C3F9790" box="[533,619,1656,1683]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="34" pageNumber="35" unit="mm" value="62.0">62 mm</quantity>
; Ecuadorian females range from
<quantity id="1F339B4DFFAFFFD8FC36F9E56ADD9790" box="[1008,1161,1656,1683]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="6.7" metricValueMin="6.0" pageId="34" pageNumber="35" unit="mm" value="63.5" valueMax="67.0" valueMin="60.0">60 to 67 mm</quantity>
. The ventral surface shows the same diagnostic features as the male. However, on VHW, the inner white stripe goes slightly beyond the ocelli, but remains shorter than in all other subspecies (
<figureCitation id="40F02A2DFFAFFFD8FC0AF9596A7F97DC" box="[972,1067,1732,1759]" captionStart="Fig" captionStartId="16.[189,232,1790,1816]" captionTargetBox="[224,1363,264,1737]" captionTargetId="figure@16.[224,1363,265,1737]" captionTargetPageId="16" captionText="Fig. 7. Habitus of females. Yellow spot: paratype (PT). A. Eryphanis zolvizora zolvizora (Hewitson, 1877) (San Jacinto, Bolivia; BMNH). B. Eryphanis zolvizora chachapoya ssp. nov., PT (San José de Molinopampa, Amazonas, Peru; MNHN, PBGL 198). C. Eryphanis zolvizora greeneyi Penz &amp; DeVries, 2008, stat. rev. (Ecuador; JFLC). D. Eryphanis zolvizora casagrande ssp. nov., PT (La Planada, Nariño, Colombia; IAvH). E. E. z. casagrande ssp. nov. (Huigra, Chimborazo, Ecuador; ANSP). F. Eryphanis zolvizora opimus (Staudinger, 1887) (Río Agua (presumably Río Aguacatal), Valle del Cauca, Colombia; BMNH)." figureDoi="http://doi.org/10.5281/zenodo.3834972" httpUri="https://zenodo.org/record/3834972/files/figure.png" pageId="34" pageNumber="35">Fig. 7D</figureCitation>
). The markings in the VHW cell of the Huigra female are rather faint, but this specimen is fairly old and worn (
<figureCitation id="40F02A2DFFAFFFD8FB4CF9776AB19607" box="[1162,1253,1770,1796]" captionStart="Fig" captionStartId="16.[189,232,1790,1816]" captionTargetBox="[224,1363,264,1737]" captionTargetId="figure@16.[224,1363,265,1737]" captionTargetPageId="16" captionText="Fig. 7. Habitus of females. Yellow spot: paratype (PT). A. Eryphanis zolvizora zolvizora (Hewitson, 1877) (San Jacinto, Bolivia; BMNH). B. Eryphanis zolvizora chachapoya ssp. nov., PT (San José de Molinopampa, Amazonas, Peru; MNHN, PBGL 198). C. Eryphanis zolvizora greeneyi Penz &amp; DeVries, 2008, stat. rev. (Ecuador; JFLC). D. Eryphanis zolvizora casagrande ssp. nov., PT (La Planada, Nariño, Colombia; IAvH). E. E. z. casagrande ssp. nov. (Huigra, Chimborazo, Ecuador; ANSP). F. Eryphanis zolvizora opimus (Staudinger, 1887) (Río Agua (presumably Río Aguacatal), Valle del Cauca, Colombia; BMNH)." figureDoi="http://doi.org/10.5281/zenodo.3834972" httpUri="https://zenodo.org/record/3834972/files/figure.png" pageId="34" pageNumber="35">Fig. 7E</figureCitation>
). The dorsal branches of the inward projections of the sterigma are similar to those of
<emphasis id="EABFEABAFFAFFFD8FBE9F88C6A869629" box="[1071,1234,1809,1834]" italics="true" pageId="34" pageNumber="35">
E. z.
<taxonomicName id="1FCB4D2BFFAFFFD8FBA8F88C6A869629" authorityName="Penz &amp; DeVries" authorityYear="2008" baseAuthorityName="Blandin &amp; Bristow &amp; Neild &amp; Sousa &amp; Gareca &amp; Huertas" baseAuthorityYear="2008" box="[1134,1234,1809,1834]" class="Hexapoda" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="greeneyi">greeneyi</taxonomicName>
</emphasis>
, as they have irregular outlines, but they are more distant from the internal side of the arch, as in
<emphasis id="EABFEABAFFAFFFD8FBBBF8AB6B109653" box="[1149,1348,1846,1872]" italics="true" pageId="34" pageNumber="35">E. z. chachapoya</emphasis>
ssp. nov. and
<emphasis id="EABFEABAFFAFFFD8FEE1F8C16FE39676" box="[295,439,1884,1909]" italics="true" pageId="34" pageNumber="35">
E. z.
<taxonomicName id="1FCB4D2BFFAFFFD8FEA4F8C16FE39676" authorityName="Staudinger" authorityYear="1887" box="[354,439,1884,1909]" class="Hexapoda" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="opimus">opimus</taxonomicName>
</emphasis>
(
<figureCitation id="40F02A2DFFAFFFD8FE01F8C66C7E9675" box="[455,554,1883,1910]" captionStart="Fig" captionStartId="22.[189,232,1578,1604]" captionTargetBox="[188,1399,264,1515]" captionTargetId="figure@22.[189,1398,277,1514]" captionTargetPageId="22" captionText="Fig. 15. Female sterigma. A. Eryphanis zolvizora zolvizora (Hewitson, 1877) (Bolivia; MNHN, CG). B-H. Eryphanis zolvizora chachapoya ssp. nov. B. PT from Cushi, Pasco, Peru (BMNH 8226). C. PT from Carpish, Huánuco, Peru (MNHN, PBB 2285). D. Specimen from Mallqui, Huánuco, Peru (MNHN, PBB 2327). E. PT from San José de Molinopampa, Amazonas, Peru (MNHN, PBGL 198). F. PT from Nieva, Amazonas, Peru (MNHN, PBGL 596). G. PT from Alto Nieva, Amazonas, Peru (MNHN, PBGL 519). H. Specimen from Alto Nieva, Amazonas, Peru (MNHN, PBGL 520). I. Eryphanis zolvizora greeneyi Penz &amp; DeVries, 2008, stat. rev., specimen fromValladolid, Zamora-Chinchipe, Ecuador (MNHN, PBB 2288). J. Eryphanis zolvizora casagrande ssp. nov., PT from La Planada, Nariño, Colombia (IAvH; drawing by Jean-François Le Crom, JFL 337). K. E. zolvizora opimus (Staudinger, 1887), specimen from Colombie (MNHN, CG). L. Eryphanis zolvizora reyi ssp. nov., PT from La Mina, Barinas, Venezuela (R, 116-JCSC). M-O. E. zolvizora isabelae ssp. nov. M. PT from Choroní, Aragua, Venezuela (R, 114-JCSC). N. PT from Rancho Grande, Aragua, Venezuela (MIZA, 118-JCSC). O. PT from La Llanada, Distrito Federal, Venezuela (MNHN, PBB 2322)." figureDoi="http://doi.org/10.5281/zenodo.3834988" httpUri="https://zenodo.org/record/3834988/files/figure.png" pageId="34" pageNumber="35">Fig. 15J</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="90D16523FFAFFFD8FF7BF8026AE996E8" pageId="34" pageNumber="35" type="biology_ecology">
<paragraph id="D87436A8FFAFFFD8FF7BF8026F3496BA" blockId="34.[189,352,1951,1977]" box="[189,352,1951,1977]" pageId="34" pageNumber="35">
<heading id="833C81C4FFAFFFD8FF7BF8026F3496BA" bold="true" box="[189,352,1951,1977]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="EABFEABAFFAFFFD8FF7BF8026F3496BA" bold="true" box="[189,352,1951,1977]" pageId="34" pageNumber="35">Flight period</emphasis>
</heading>
</paragraph>
<paragraph id="D87436A8FFAFFFD8FF7BF84C6AE996E8" blockId="34.[189,1213,2001,2027]" box="[189,1213,2001,2027]" pageId="34" pageNumber="35">Specimens have been caught or seen in April, July, September, October and November.</paragraph>
</subSubSection>
<subSubSection id="90D16523FFAEFFD9FF7BFE966F7290FC" pageId="35" pageNumber="36" type="distribution">
<paragraph id="D87436A8FFAEFFD9FF7BFE966F029026" blockId="35.[189,342,267,293]" box="[189,342,267,293]" pageId="35" pageNumber="36">
<heading id="833C81C4FFAEFFD9FF7BFE966F029026" bold="true" box="[189,342,267,293]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="EABFEABAFFAEFFD9FF7BFE966F029026" bold="true" box="[189,342,267,293]" pageId="35" pageNumber="36">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="D87436A8FFAEFFD9FF7BFEA46ADC9057" blockId="35.[189,1160,313,340]" box="[189,1160,313,340]" pageId="35" pageNumber="36">
<collectingCountry id="A0DC7638FFAEFFD9FF7BFEA46F619050" box="[189,309,313,339]" name="Colombia" pageId="35" pageNumber="36">Colombia</collectingCountry>
.
<collectingRegion id="1A0FF84AFFAEFFD9FEF9FEA46C769057" box="[319,546,313,340]" country="Colombia" name="Narino" pageId="35" pageNumber="36">Nariño Department</collectingRegion>
of south-west
<collectingCountry id="A0DC7638FFAEFFD9FD16FEA46D109050" box="[720,836,313,339]" name="Colombia" pageId="35" pageNumber="36">Colombia</collectingCountry>
between
<date id="AC751068FFAEFFD9FC72FEA76A379057" box="[948,1123,313,340]" pageId="35" pageNumber="36" value="1380" valueMax="1930">
1380 and
<quantity id="1F339B4DFFAEFFD9FBEEFEA76A379057" box="[1064,1123,314,340]" metricMagnitude="3" metricUnit="m" metricValue="1.93" pageId="35" pageNumber="36" unit="m" value="1930.0">1930</quantity>
</date>
m.
</paragraph>
<paragraph id="D87436A8FFAEFFD9FF7BFEE66F7290FC" blockId="35.[189,1399,379,511]" pageId="35" pageNumber="36">
<collectingCountry id="A0DC7638FFAEFFD9FF7BFEE66F759096" box="[189,289,379,405]" name="Ecuador" pageId="35" pageNumber="36">Ecuador</collectingCountry>
.
<emphasis id="EABFEABAFFAEFFD9FEF6FEE16FA99096" box="[304,509,379,405]" italics="true" pageId="35" pageNumber="36">E. z. casagrande</emphasis>
ssp. nov. presumably extends in suitable habitat throughout the western slopes from the Colombian border to the extreme south of the country at heights of
<quantity id="1F339B4DFFAEFFD9FB50FE036ABB90BA" box="[1174,1263,414,441]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="35" pageNumber="36" unit="m" value="1200.0">1200 m</quantity>
to
<quantity id="1F339B4DFFAEFFD9FAD3FE036B2590BA" box="[1301,1393,414,441]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="35" pageNumber="36" unit="m" value="1800.0">1800 m</quantity>
; the Santo Domingo specimen at
<quantity id="1F339B4DFFAEFFD9FDFAFE5F6CD290DF" box="[572,646,450,476]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="36" unit="m" value="200.0">200 m</quantity>
is an anomaly and may be from a higher altitude along the road to Quito.
</paragraph>
</subSubSection>
<subSubSection id="90D16523FFAEFFD9FF7BFDBA6C809536" pageId="35" pageNumber="36" type="discussion">
<paragraph id="D87436A8FFAEFFD9FF7BFDBA6F7B9342" blockId="35.[189,303,551,577]" box="[189,303,551,577]" pageId="35" pageNumber="36">
<heading id="833C81C4FFAEFFD9FF7BFDBA6F7B9342" bold="true" box="[189,303,551,577]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="EABFEABAFFAEFFD9FF7BFDBA6F7B9342" bold="true" box="[189,303,551,577]" pageId="35" pageNumber="36">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="D87436A8FFAEFFD9FF7BFDC86F3792AE" blockId="35.[189,1399,597,941]" pageId="35" pageNumber="36">
Specimens of
<taxonomicName id="1FCB4D2BFFAEFFD9FEADFDCB6FA8936C" box="[363,508,597,623]" class="Hexapoda" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="zolvizora">
<emphasis id="EABFEABAFFAEFFD9FEADFDCB6FA8936C" box="[363,508,597,623]" italics="true" pageId="35" pageNumber="36">E. zolvizora</emphasis>
</taxonomicName>
from south-western
<collectingCountry id="A0DC7638FFAEFFD9FD3CFDC86D3A936C" box="[762,878,597,623]" name="Colombia" pageId="35" pageNumber="36">Colombia</collectingCountry>
and western
<collectingCountry id="A0DC7638FFAEFFD9FBD0FDC86A2C936C" box="[1046,1144,597,623]" name="Ecuador" pageId="35" pageNumber="36">Ecuador</collectingCountry>
are characterized by a wide bridge in space Cu2-2A on the VFW formed by a complete white connection between the white stripes, delimiting half-a-circle (up against Cu2) with a chestnut-brown inner core and a dark brown outer ring; in that way,
<emphasis id="EABFEABAFFAEFFD9FDE1FD5D6CB993DA" box="[551,749,703,729]" italics="true" pageId="35" pageNumber="36">E. z. casagrande</emphasis>
ssp. nov. differs from its near neighbours
<emphasis id="EABFEABAFFAEFFD9FB23FD5D6B2393DA" box="[1253,1399,704,729]" italics="true" pageId="35" pageNumber="36">
E. z.
<taxonomicName id="1FCB4D2BFFAEFFD9FAE4FD5D6B2393DA" authorityName="Staudinger" authorityYear="1887" box="[1314,1399,704,729]" class="Hexapoda" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="opimus">opimus</taxonomicName>
</emphasis>
in the
<collectingRegion id="1A0FF84AFFAEFFD9FECFFD7F6F0693FF" box="[265,338,738,764]" country="Colombia" name="Cauca" pageId="35" pageNumber="36">Cauca</collectingRegion>
Valley,
<collectingCountry id="A0DC7638FFAEFFD9FE72FD7F6C7F93FF" box="[436,555,738,764]" name="Colombia" pageId="35" pageNumber="36">Colombia</collectingCountry>
, and
<emphasis id="EABFEABAFFAEFFD9FDADFD7E6D5993FF" box="[619,781,739,764]" italics="true" pageId="35" pageNumber="36">
E. z.
<taxonomicName id="1FCB4D2BFFAEFFD9FD6FFD7E6D5993FF" authorityName="Penz &amp; DeVries" authorityYear="2008" baseAuthorityName="Blandin &amp; Bristow &amp; Neild &amp; Sousa &amp; Gareca &amp; Huertas" baseAuthorityYear="2008" box="[681,781,739,764]" class="Hexapoda" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="greeneyi">greeneyi</taxonomicName>
</emphasis>
in the Oriente of
<collectingCountry id="A0DC7638FFAEFFD9FC22FD7F6A1C93FF" box="[996,1096,738,764]" name="Ecuador" pageId="35" pageNumber="36">Ecuador</collectingCountry>
, as well as from all other subspecies, except
<emphasis id="EABFEABAFFAEFFD9FE62FC9B6C20921C" box="[420,628,773,799]" italics="true" pageId="35" pageNumber="36">E. z. chachapoya</emphasis>
ssp. nov., where we know 3 (of 35) specimens having a white bridge (MNHN: PBB 2310, PBGL 521 and PBGL 705). We consider that the presence of a wide, white bridge is the principal diagnostic character of
<emphasis id="EABFEABAFFAEFFD9FD12FCD06DC29265" box="[724,918,844,870]" italics="true" pageId="35" pageNumber="36">E. z. casagrande</emphasis>
ssp. nov., but suggesting a closer affinity with
<emphasis id="EABFEABAFFAEFFD9FF3FFCED6F97928A" box="[249,451,879,905]" italics="true" pageId="35" pageNumber="36">E. z. chachapoya</emphasis>
ssp. nov. than
<emphasis id="EABFEABAFFAEFFD9FDB5FCED6D46928A" box="[627,786,880,905]" italics="true" pageId="35" pageNumber="36">
E. z.
<taxonomicName id="1FCB4D2BFFAEFFD9FD68FCED6D46928A" authorityName="Penz &amp; DeVries" authorityYear="2008" baseAuthorityName="Blandin &amp; Bristow &amp; Neild &amp; Sousa &amp; Gareca &amp; Huertas" baseAuthorityYear="2008" box="[686,786,880,905]" class="Hexapoda" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="greeneyi">greeneyi</taxonomicName>
</emphasis>
. However, the male genitalia are similar to those of
<emphasis id="EABFEABAFFAEFFD9FF7BFC0E6F0892AF" box="[189,348,915,940]" italics="true" pageId="35" pageNumber="36">
E. z.
<taxonomicName id="1FCB4D2BFFAEFFD9FF3EFC0E6F0892AF" authorityName="Penz &amp; DeVries" authorityYear="2008" baseAuthorityName="Blandin &amp; Bristow &amp; Neild &amp; Sousa &amp; Gareca &amp; Huertas" baseAuthorityYear="2008" box="[248,348,915,940]" class="Hexapoda" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="greeneyi">greeneyi</taxonomicName>
</emphasis>
.
</paragraph>
<paragraph id="D87436A8FFAEFFD9FF7BFC496C809536" blockId="35.[189,1398,978,1077]" pageId="35" pageNumber="36">
There are specimens of
<taxonomicName id="1FCB4D2BFFAEFFD9FE14FC486C0992ED" box="[466,605,980,1006]" class="Hexapoda" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="zolvizora">
<emphasis id="EABFEABAFFAEFFD9FE14FC486C0992ED" box="[466,605,980,1006]" italics="true" pageId="35" pageNumber="36">E. zolvizora</emphasis>
</taxonomicName>
collected in the 19
<superScript id="2FBE9BE0FFAEFFD9FCFFFC4F6D1292E3" attach="left" box="[825,838,978,992]" fontSize="6" pageId="35" pageNumber="36">th</superScript>
and early 20
<superScript id="2FBE9BE0FFAEFFD9FC1AFC4F6DBD92E3" attach="left" box="[988,1001,978,992]" fontSize="6" pageId="35" pageNumber="36">th</superScript>
century labelled from the western Ecuadorian village of Balzapamba. Like many specimens supposedly from this locality, they are from eastern
<collectingCountry id="A0DC7638FFAEFFD9FED1FB866F2D9536" box="[279,377,1051,1077]" name="Ecuador" pageId="35" pageNumber="36">Ecuador</collectingCountry>
and are in fact
<emphasis id="EABFEABAFFAEFFD9FDE9FB866C999537" box="[559,717,1051,1076]" italics="true" pageId="35" pageNumber="36">
E. z.
<taxonomicName id="1FCB4D2BFFAEFFD9FDAFFB866C999537" authorityName="Penz &amp; DeVries" authorityYear="2008" baseAuthorityName="Blandin &amp; Bristow &amp; Neild &amp; Sousa &amp; Gareca &amp; Huertas" baseAuthorityYear="2008" box="[617,717,1051,1076]" class="Hexapoda" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="greeneyi">greeneyi</taxonomicName>
</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>