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151 lines
14 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.205.76790" ID-Pensoft-Pub="1314-2003-205-99" ID-Pensoft-UUID="4024A478048757B197E5CF8734331A9E" ModsDocID="1314-2003-205-99" checkinTime="1661230211668" checkinUser="pensoft" docAuthor="O'Donnell, Shawn A., Ringelberg, Jens J. & Lewis, Gwilym P." docDate="2022" docId="50461AFEF7615515B439D818B1CA837D" docLanguage="en" docName="PhytoKeys 205: 99-145" docOrigin="PhytoKeys 205" docPubDate="2022-08-22" docSource="http://dx.doi.org/10.3897/phytokeys.205.76790" docTitle="Entada arenaria Schinz, Mém. Herb. Boissier 8: 118. 1900." docType="treatment" docVersion="1" id="4024A478048757B197E5CF8734331A9E" lastPageNumber="99" masterDocId="4024A478048757B197E5CF8734331A9E" masterDocTitle="Re-circumscription of the mimosoid genus Entada including new combinations for all species of the phylogenetically nested Elephantorrhiza (Leguminosae, Caesalpinioideae, mimosoid clade)" masterLastPageNumber="145" masterPageNumber="99" pageNumber="99" updateTime="1661230211668" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Re-circumscription of the mimosoid genus Entada including new combinations for all species of the phylogenetically nested Elephantorrhiza (Leguminosae, Caesalpinioideae, mimosoid clade)</mods:title>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>O'Donnell, Shawn A.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0731-7425</mods:nameIdentifier>
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<mods:affiliation>Department of Geography and Environmental Sciences, Northumbria University, Newcastle upon Tyne, NE 1 8 ST, UK</mods:affiliation>
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<mods:nameIdentifier type="email">shawn.odonnell@cantab.net</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ringelberg, Jens J.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0567-5210</mods:nameIdentifier>
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<mods:affiliation>Department of Systematic and Evolutionary Botany, University of Zurich, 8008 Zurich, Switzerland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lewis, Gwilym P.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2599-4577</mods:nameIdentifier>
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<mods:affiliation>Accelerated Taxonomy Department, Royal Botanic Gardens, Kew, Richmond, TW 9 3 AE, UK</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-08-22</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>205</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>99</mods:start>
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<mods:end>145</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.205.76790</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.205.76790</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-205-99</mods:identifier>
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<mods:identifier type="Pensoft-UUID">4024A478048757B197E5CF8734331A9E</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:50461AFEF7615515B439D818B1CA837D" httpUri="http://treatment.plazi.org/id/50461AFEF7615515B439D818B1CA837D" lastPageNumber="99" pageId="0" pageNumber="99">
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<subSubSection pageId="0" pageNumber="99" type="nomenclature">
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<paragraph pageId="0" pageNumber="99">
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<taxonomicName LSID="50461AFE-F761-5515-B439-D818B1CA837D" authority="Schinz, Mem. Herb. Boissier 8: 118. 1900." authorityName="Schinz, Mém. Herb. Boissier 8: 118. 1900." class="Magnoliopsida" family="Fabaceae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada arenaria" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="arenaria">
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Entada arenaria Schinz,
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<normalizedToken originalValue="Mém">Mem</normalizedToken>
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. Herb. Boissier 8: 118. 1900.
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="99" type="materials_examined">
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<paragraph pageId="0" pageNumber="99">
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<typeStatus>Type</typeStatus>
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.
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</paragraph>
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<paragraph pageId="0" pageNumber="99">
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<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="NAMIBIA" location="Grootfontein District" municipality="Omuramba-Omatako River" specimenCount="1" typeStatus="holotype">
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<collectingCountry name="Namibia">NAMIBIA</collectingCountry>
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. Hereroland,
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<location LSID="urn:lsid:plazi:treatment:50461AFEF7615515B439D818B1CA837D:E2A90B462A24225C8E2627C29E645A0D" country="NAMIBIA" municipality="Omuramba-Omatako River" name="Grootfontein District">Grootfontein District</location>
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,
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<collectingMunicipality>Omuramba-Omatako River</collectingMunicipality>
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,
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<emphasis italics="true" pageId="0" pageNumber="99">Schinz 277</emphasis>
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(
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<typeStatus>holotype</typeStatus>
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: Z)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="99" type="description">
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<paragraph pageId="0" pageNumber="99">Description.</paragraph>
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<paragraph pageId="0" pageNumber="99">
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Geoxylic suffrutex with erect annual 5-120 cm stems, young stems densely pubescent (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Entada arenaria habit, vegetative and reproductive structures. A geoxylic suffrutex with erect annual stem, Angola (photo: D Goyder CC BY-NC 4.0, Ueda (2021) observation 35199077) B leaves and spiciform racemes pre-anthesis, Democratic Republic of Congo (photo: J Stevens, Dressler et al. (2014 a)) C open, pedicellate flowers, Democratic Republic of Congo (photo: W McCleland, all rights reserved, iNaturalist (2021) observation 95512918) D leaves and immature fruits, Zambia (photo: W McCleland, Dressler et al. (2014 a))." figureDoi="10.3897/phytokeys.205.76790.figure8" httpUri="https://binary.pensoft.net/fig/732703" pageId="0" pageNumber="99">8A</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="99">Leaves</emphasis>
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: petiole 6-12 cm long, grooved above, puberulous; rachis 4-17 cm long, grooved above; pinnae 2-4 pairs per leaf, 7.5-14 cm long, with 6-13 pairs of leaflets; leaflets (1.2-)2-3.5(-4)
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<normalizedToken originalValue="×">x</normalizedToken>
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0.7-2 cm, narrowly oblong to obovate-oblong, apex rounded to emarginate, base asymmetric, rounded to cordate on proximal margin, cuneate to cuneate-rounded on distal margin, lamina pubescent below at least on mid-rib and often throughout (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Entada arenaria habit, vegetative and reproductive structures. A geoxylic suffrutex with erect annual stem, Angola (photo: D Goyder CC BY-NC 4.0, Ueda (2021) observation 35199077) B leaves and spiciform racemes pre-anthesis, Democratic Republic of Congo (photo: J Stevens, Dressler et al. (2014 a)) C open, pedicellate flowers, Democratic Republic of Congo (photo: W McCleland, all rights reserved, iNaturalist (2021) observation 95512918) D leaves and immature fruits, Zambia (photo: W McCleland, Dressler et al. (2014 a))." figureDoi="10.3897/phytokeys.205.76790.figure8" httpUri="https://binary.pensoft.net/fig/732703" pageId="0" pageNumber="99">8A, B, D</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="99">Inflorescence</emphasis>
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: an axillary spiciform raceme 4-12 cm long, 1-3 per axil, rachis usually glabrous (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Entada arenaria habit, vegetative and reproductive structures. A geoxylic suffrutex with erect annual stem, Angola (photo: D Goyder CC BY-NC 4.0, Ueda (2021) observation 35199077) B leaves and spiciform racemes pre-anthesis, Democratic Republic of Congo (photo: J Stevens, Dressler et al. (2014 a)) C open, pedicellate flowers, Democratic Republic of Congo (photo: W McCleland, all rights reserved, iNaturalist (2021) observation 95512918) D leaves and immature fruits, Zambia (photo: W McCleland, Dressler et al. (2014 a))." figureDoi="10.3897/phytokeys.205.76790.figure8" httpUri="https://binary.pensoft.net/fig/732703" pageId="0" pageNumber="99">8A-C</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="99">Flowers</emphasis>
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: pale cream, pedicels 1-2 mm long; calyx campanulate, 1-2 mm long, shallowly toothed; petals 3-4 mm long; stamen filaments 5-6 mm long (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Entada arenaria habit, vegetative and reproductive structures. A geoxylic suffrutex with erect annual stem, Angola (photo: D Goyder CC BY-NC 4.0, Ueda (2021) observation 35199077) B leaves and spiciform racemes pre-anthesis, Democratic Republic of Congo (photo: J Stevens, Dressler et al. (2014 a)) C open, pedicellate flowers, Democratic Republic of Congo (photo: W McCleland, all rights reserved, iNaturalist (2021) observation 95512918) D leaves and immature fruits, Zambia (photo: W McCleland, Dressler et al. (2014 a))." figureDoi="10.3897/phytokeys.205.76790.figure8" httpUri="https://binary.pensoft.net/fig/732703" pageId="0" pageNumber="99">8C</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="99">Fruit</emphasis>
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: a torulose, laterally compressed, straight to distinctly falcate craspedium, 7.5-22
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<normalizedToken originalValue="×">x</normalizedToken>
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1.5-6 cm, with transverse septa between seeds dividing the fruit into one-seeded segments which, upon ripening, fall from the persistent replum (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Entada arenaria habit, vegetative and reproductive structures. A geoxylic suffrutex with erect annual stem, Angola (photo: D Goyder CC BY-NC 4.0, Ueda (2021) observation 35199077) B leaves and spiciform racemes pre-anthesis, Democratic Republic of Congo (photo: J Stevens, Dressler et al. (2014 a)) C open, pedicellate flowers, Democratic Republic of Congo (photo: W McCleland, all rights reserved, iNaturalist (2021) observation 95512918) D leaves and immature fruits, Zambia (photo: W McCleland, Dressler et al. (2014 a))." figureDoi="10.3897/phytokeys.205.76790.figure8" httpUri="https://binary.pensoft.net/fig/732703" pageId="0" pageNumber="99">8D</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="99">Seeds</emphasis>
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: 12.5
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<normalizedToken originalValue="×">x</normalizedToken>
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9 mm or smaller (see subsp.
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<taxonomicName class="Magnoliopsida" family="Leguminosae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada subsp. microcarpa" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="subSpecies" species="arenaria" subSpecies="microcarpa">Entada microcarpa</taxonomicName>
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below), dark brown, smooth.
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</paragraph>
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<caption doi="10.3897/phytokeys.205.76790.figure8" httpUri="https://binary.pensoft.net/fig/732703" pageId="0" pageNumber="99" start="Figure 8" startId="F8">
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<paragraph pageId="0" pageNumber="99">
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<emphasis bold="true" pageId="0" pageNumber="99">Figure 8.</emphasis>
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<taxonomicName class="Magnoliopsida" family="Leguminosae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada arenaria" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="arenaria">
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<emphasis italics="true" pageId="0" pageNumber="99">Entada arenaria</emphasis>
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</taxonomicName>
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habit, vegetative and reproductive structures.
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<emphasis bold="true" pageId="0" pageNumber="99">A</emphasis>
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geoxylic suffrutex with erect annual stem, Angola (photo: D Goyder
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<emphasis italics="true" pageId="0" pageNumber="99">CC BY-NC 4.0</emphasis>
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,
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<bibRefCitation author="Ueda, K" journalOrPublisher="Acta Phytotaxonomica et Geobotanica" pageId="0" pageNumber="99" publicationUrl="iNaturalist.org" refId="B59" refString="Ueda, K, 2021. iNaturalist Research-grade Observations. iNaturalist.org" title="iNaturalist Research-grade Observations." url="iNaturalist.org" year="2021">Ueda (2021)</bibRefCitation>
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observation 35199077)
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<emphasis bold="true" pageId="0" pageNumber="99">B</emphasis>
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leaves and spiciform racemes pre-anthesis, Democratic Republic of Congo (photo: J Stevens,
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<bibRefCitation author="Dressler, S" journalOrPublisher="Acacia), Caesalpiniaceae. CSIRO Australia, Melbourne" pageId="0" pageNumber="99" publicationUrl="www.africanplants.senckenberg.de" refId="B20" refString="Dressler, S, Schmidt, M, Zizka, G, 2014a. (continuously updated) African Plants - A Photo Guide. Forschungsinstitut Senckenberg, Frankfurt/Main, Germany. www.africanplants.senckenberg.de" title="(continuously updated) African Plants - A Photo Guide. Forschungsinstitut Senckenberg, Frankfurt / Main, Germany." url="www.africanplants.senckenberg.de" year="2014 a">Dressler et al. (2014a)</bibRefCitation>
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)
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<emphasis bold="true" pageId="0" pageNumber="99">C</emphasis>
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open, pedicellate flowers, Democratic Republic of Congo (photo: W McCleland, all rights reserved,
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<bibRefCitation author="Hyde, MA" journalOrPublisher="Institut Francais de Pondichery, Pondichery" pageId="0" pageNumber="99" publicationUrl="www.inaturalist.org" refId="B27" refString="2021. . www.inaturalist.org" url="www.inaturalist.org" year="2021">iNaturalist (2021)</bibRefCitation>
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observation 95512918)
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<emphasis bold="true" pageId="0" pageNumber="99">D</emphasis>
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leaves and immature fruits, Zambia (photo: W McCleland,
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<bibRefCitation author="Dressler, S" journalOrPublisher="Acacia), Caesalpiniaceae. CSIRO Australia, Melbourne" pageId="0" pageNumber="99" publicationUrl="www.africanplants.senckenberg.de" refId="B20" refString="Dressler, S, Schmidt, M, Zizka, G, 2014a. (continuously updated) African Plants - A Photo Guide. Forschungsinstitut Senckenberg, Frankfurt/Main, Germany. www.africanplants.senckenberg.de" title="(continuously updated) African Plants - A Photo Guide. Forschungsinstitut Senckenberg, Frankfurt / Main, Germany." url="www.africanplants.senckenberg.de" year="2014 a">Dressler et al. (2014a)</bibRefCitation>
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).
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |