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<mods:title id="EDE88769B3000955BFC01DDFF421217D">Identity of the millipede genus Pyrgodesmus Pocock, 1892, the type genus of the family Pyrgodesmidae (Diplopoda, Polydesmida)</mods:title>
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<treatment id="5038879DCE2A550590A12A4CFDDAFCFE" ID-DOI="http://doi.org/10.5281/zenodo.5709644" ID-GBIF-Taxon="190310265" ID-Zenodo-Dep="5709644" LSID="urn:lsid:plazi:treatment:5038879DCE2A550590A12A4CFDDAFCFE" httpUri="http://treatment.plazi.org/id/5038879DCE2A550590A12A4CFDDAFCFE" lastPageId="3" lastPageNumber="575" pageId="1" pageNumber="573">
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<paragraph id="D82E368BCE2A550790A12A4CFDE5F981" blockId="1.[151,592,1615,1678]" box="[151,592,1615,1642]" pageId="1" pageNumber="573">
<heading id="836681E7CE2A550790A12A4CFDE5F981" bold="true" box="[151,592,1615,1642]" fontSize="11" level="1" pageId="1" pageNumber="573" reason="1">
<taxonomicName id="1F914D08CE2A550790A12A4CFDE5F981" ID-CoL="4QT8C" authority="Pocock 1892" authorityName="Pocock" authorityYear="1892" box="[151,592,1615,1642]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="573" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="EAE5EA99CE2A550790A12A4CFDE5F981" bold="true" box="[151,592,1615,1642]" pageId="1" pageNumber="573">
<emphasis id="EAE5EA99CE2A550790A12A4CFE18F981" bold="true" box="[151,429,1615,1642]" italics="true" pageId="1" pageNumber="573">Pyrgodesmus obscurus</emphasis>
<bibRefCitation id="BC004B7ACE2A550791822A53FDE5F981" author="Pocock, R. I." box="[436,592,1616,1642]" pageId="1" pageNumber="573" pagination="131 - 174" refId="ref3370" refString="Pocock, R. I. (1892) Report upon two collections of Myriapoda sent from Ceylon by Mr. E. E. Green, and from various parts of southern India by Mr. Edgar Thurston, of the Government Central Museum, Madras. Journal of the Bombay Natural History Society, 7, 131 - 174. [https: // www. biodiversitylibrary. org / item / 95456 # page / 7 / mode / 1 up]" type="journal article" year="1892">Pocock 1892</bibRefCitation>
</emphasis>
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<subSubSection id="908B6500CE2A550790A12A77FF4DF965" box="[151,248,1652,1678]" pageId="1" pageNumber="573" type="description">
<paragraph id="D82E368BCE2A550790A12A77FF4DF965" blockId="1.[151,592,1615,1678]" box="[151,248,1652,1678]" pageId="1" pageNumber="573">
<figureCitation id="40AA2A0ECE2A550790A12A77FF4DF965" box="[151,248,1652,1678]" captionStart="FIGURES 18" captionStartId="2.[151,264,1857,1881]" captionTargetBox="[186,1401,181,1831]" captionTargetId="figure-15@2.[186,1401,181,1834]" captionTargetPageId="2" captionText="FIGURES 18. Pyrgodesmus obscurus Pocock, 1892, male syntypes. 1. Original labels. 2. Both syntypes pierced with glass minutiae, shown relative to the length of an insect pin. 3. Habitus of a syntype, lateral view (after Pocock (1892)). 4. Picture of the same, lateral view. 5. Body rings 57 and gonopods in situ, ventral view. 6. Body rings 69 and gonopods in situ, ventrolateral view. 7 &amp; 8. Enlarged gonopods in situ, ventral and ventrolateral views, respectively. Figs 13, 7 and 8 not to scale. Images courtesy Dmitry Telnov." figureDoi="http://doi.org/10.5281/zenodo.5709646" httpUri="https://zenodo.org/record/5709646/files/figure.png" pageId="1" pageNumber="573">Figs 18</figureCitation>
</paragraph>
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<paragraph id="D82E368BCE2A550790A12ABBFC84F93A" blockId="1.[151,1436,1720,1806]" box="[151,817,1720,1745]" pageId="1" pageNumber="573">
<treatmentCitationGroup id="F88111A5CE2A550790A12ABBFC84F93A" box="[151,817,1720,1745]" pageId="1" pageNumber="573">
<treatmentCitation id="5930109ACE2A550790A12ABBFDF3F93B" author="Pocock, R. I." box="[151,582,1720,1745]" page="155" pageId="1" pageNumber="573" year="1892">
<taxonomicName id="1F914D08CE2A550790A12ABBFDF3F93B" ID-CoL="4QT8C" authority="Pocock, 1892: 155" authorityName="Pocock" authorityPageNumber="155" authorityYear="1892" box="[151,582,1720,1745]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="573" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="EAE5EA99CE2A550790A12ABBFE35F93B" box="[151,384,1720,1744]" italics="true" pageId="1" pageNumber="573">Pyrgodesmus obscurus</emphasis>
<bibRefCitation id="BC004B7ACE2A550791B12ABBFDF3F93B" author="Pocock, R. I." box="[391,582,1720,1745]" pageId="1" pageNumber="573" pagination="131 - 174" refId="ref3370" refString="Pocock, R. I. (1892) Report upon two collections of Myriapoda sent from Ceylon by Mr. E. E. Green, and from various parts of southern India by Mr. Edgar Thurston, of the Government Central Museum, Madras. Journal of the Bombay Natural History Society, 7, 131 - 174. [https: // www. biodiversitylibrary. org / item / 95456 # page / 7 / mode / 1 up]" type="journal article" year="1892">Pocock, 1892: 155</bibRefCitation>
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, plate 2, figs 11b (D).
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<taxonomicName id="1F914D08CE2A550790A12AD4FECAF904" ID-CoL="4QT8C" authorityName="Pocock" authorityYear="1892" box="[151,383,1751,1775]" class="Diplopoda" family="Pyrgodesmidae" genus="Pyrgodesmus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="573" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="EAE5EA99CE2A550790A12AD4FECAF904" box="[151,383,1751,1775]" italics="true" pageId="1" pageNumber="573">Pyrgodesmus obscurus</emphasis>
</taxonomicName>
<treatmentCitation id="5930109ACE2A550791AC2AD4FDE1F904" author="Attems, C." box="[410,596,1751,1775]" page="174" pageId="1" pageNumber="573" year="1914">
<bibRefCitation id="BC004B7ACE2A550791AC2AD4FDE1F904" author="Attems, C." box="[410,596,1751,1775]" pageId="1" pageNumber="573" pagination="1 - 398" refId="ref2787" refString="Attems, C. (1914) Die indo-australischen Myriopoden. Archiv fur Naturgeschichte, 80 A (4), 1 - 398." type="journal article" year="1914">Attems, 1914: 174</bibRefCitation>
</treatmentCitation>
(R); 1936: 246 (R); 1940: 261, fig. 370 (D, R);
<treatmentCitation id="5930109ACE2A550794022AD4FB51F904" author="Jeekel, C. A. W." box="[1076,1252,1751,1775]" page="350" pageId="1" pageNumber="573" year="1971">
<bibRefCitation id="BC004B7ACE2A550794022AD4FB51F904" author="Jeekel, C. A. W." box="[1076,1252,1751,1775]" pageId="1" pageNumber="573" pagination="1 - 412" refId="ref3252" refString="Jeekel, C. A. W. (1971) Nomenclator generum et familiarum Diplopodorum: A list of the genus and family-group names in the Class Diplopoda from the 10 th edition of Linnaeus, 1758, to the end of 1957. Monografieёn van de Nederlandse Entomolo- gische Veneniging, 5, i - xii + 1 - 412. [for 1970]" type="journal article" year="1971">Jeekel, 1971: 350</bibRefCitation>
</treatmentCitation>
(R, M);
<treatmentCitation id="5930109ACE2A5507950D2ADBFED8F8E5" ID-DOI="http://doi.org/10.5281/zenodo.6059185" ID-Zenodo-Dep="6059185" author="De Zoysa, H. K. S. &amp; Nguyen, D. A. &amp; Wickramasinghe, S." httpUri="http://treatment.plazi.org/id/1E1387ACFFD11B68FF6F41431D84F9F7" page="477" pageId="1" pageNumber="573" year="2016">
<bibRefCitation id="BC004B7ACE2A5507950D2ADBFED8F8E5" author="De Zoysa, H. K. S. &amp; Nguyen, D. A. &amp; Wickramasinghe, S." pageId="1" pageNumber="573" pagination="451 - 482" refId="ref2941" refString="De Zoysa, H. K. S., Nguyen, D. A. &amp; Wickramasinghe, S. (2016) Annotated checklist of millipedes (Myriapoda: Diplopoda) of Sri Lanka. Zootaxa, 4061 (5), 451 - 482. https: // doi. org / 10.11646 / zootaxa. 4061.5.1" type="journal article" year="2016">
De Zoysa
<emphasis id="EAE5EA99CE2A550790F12AF4FF4FF8E5" box="[199,250,1782,1806]" italics="true" pageId="1" pageNumber="573">et al.</emphasis>
, 2016: 477
</bibRefCitation>
</treatmentCitation>
(R, M);
<treatmentCitation id="5930109ACE2A550791F22AF5FD7FF8E5" author="Aswathy, M. D. &amp; Golovatch, S. I. &amp; Sudhikumar, A. V." box="[452,714,1782,1806]" page="373" pageId="1" pageNumber="573" year="2021">
<bibRefCitation id="BC004B7ACE2A550791F22AF5FD7FF8E5" author="Aswathy, M. D. &amp; Golovatch, S. I. &amp; Sudhikumar, A. V." box="[452,714,1782,1806]" pageId="1" pageNumber="573" pagination="373 - 382" refId="ref2712" refString="Aswathy, M. D., Golovatch, S. I. &amp; Sudhikumar, A. V. (2021) The millipede genus Klimakodesmus Carl, 1932, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Pyrgodesmidae). Zootaxa, 4980 (2), 373 - 382. https: // doi. org / 10.11646 / zootaxa. 4980.2.8" type="journal article" year="2021">
Aswathy
<emphasis id="EAE5EA99CE2A550792122AF4FDEDF8E5" box="[548,600,1782,1806]" italics="true" pageId="1" pageNumber="573">et al.</emphasis>
, 2021: 373
</bibRefCitation>
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(D, R).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="908B6500CE2A550590A12B3FFE77FD1A" lastPageId="3" lastPageNumber="575" pageId="1" pageNumber="573" type="discussion">
<paragraph id="D82E368BCE2A550790A12B3FFCA8F891" blockId="1.[151,1436,1852,1987]" pageId="1" pageNumber="573">
<emphasis id="EAE5EA99CE2A550790A12B3FFEB5F8BD" bold="true" box="[151,256,1852,1878]" pageId="1" pageNumber="573">Remark.</emphasis>
As noted above, the original description of this species was rather superficial and incomplete, especially as regards the crucial gonopodal structure (
<bibRefCitation id="BC004B7ACE2A5507924F2B63FCA5F891" author="Pocock, R. I." box="[633,784,1888,1914]" pageId="1" pageNumber="573" pagination="131 - 174" refId="ref3370" refString="Pocock, R. I. (1892) Report upon two collections of Myriapoda sent from Ceylon by Mr. E. E. Green, and from various parts of southern India by Mr. Edgar Thurston, of the Government Central Museum, Madras. Journal of the Bombay Natural History Society, 7, 131 - 174. [https: // www. biodiversitylibrary. org / item / 95456 # page / 7 / mode / 1 up]" type="journal article" year="1892">Pocock 1892</bibRefCitation>
).
</paragraph>
<paragraph id="D82E368BCE2A550790F12B87FBB2F828" blockId="1.[151,1436,1852,1987]" pageId="1" pageNumber="573">
<emphasis id="EAE5EA99CE2A550790F12B87FE05F875" bold="true" box="[199,432,1924,1950]" pageId="1" pageNumber="573">Brief redescription.</emphasis>
Body with 20 rings (19+T), length
<emphasis id="EAE5EA99CE2A550793742B86FCE9F875" box="[834,860,1925,1950]" italics="true" pageId="1" pageNumber="573">ca</emphasis>
<quantity id="1F699B6ECE2A550793552B87FC7FF874" box="[867,970,1924,1951]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="573" unit="mm" value="10.5">10.5 mm</quantity>
. Coloration mostly dark grey-brown with contrasting pallid legs, tegument heavily coated with an earth crust (
<figureCitation id="40AA2A0ECE2A550793A22BABFC4FF828" box="[916,1018,1960,1987]" captionStart="FIGURES 18" captionStartId="2.[151,264,1857,1881]" captionTargetBox="[186,1401,181,1831]" captionTargetId="figure-15@2.[186,1401,181,1834]" captionTargetPageId="2" captionText="FIGURES 18. Pyrgodesmus obscurus Pocock, 1892, male syntypes. 1. Original labels. 2. Both syntypes pierced with glass minutiae, shown relative to the length of an insect pin. 3. Habitus of a syntype, lateral view (after Pocock (1892)). 4. Picture of the same, lateral view. 5. Body rings 57 and gonopods in situ, ventral view. 6. Body rings 69 and gonopods in situ, ventrolateral view. 7 &amp; 8. Enlarged gonopods in situ, ventral and ventrolateral views, respectively. Figs 13, 7 and 8 not to scale. Images courtesy Dmitry Telnov." figureDoi="http://doi.org/10.5281/zenodo.5709646" httpUri="https://zenodo.org/record/5709646/files/figure.png" pageId="1" pageNumber="573">Figs 46</figureCitation>
).
</paragraph>
<paragraph id="D82E368BCE28550590F12C94FE79FDF2" blockId="3.[151,1437,150,789]" pageId="3" pageNumber="575">
Head fully covered from above by a flabellate anterior brim of a dome-shaped collum, the latter with 5+5 equal, distinct, rounded lobulations at anterior margin. Antennae C-shaped, short and clavate, antennomere 5 being the largest and subequal in length to 2
<superScript id="2FE49BC3CE28550592162CDDFD85FF07" attach="none" box="[544,560,222,236]" fontSize="6" pageId="3" pageNumber="575">nd</superScript>
. Paramedian tubercles/crests (
<emphasis id="EAE5EA99CE28550593BD2CDCFC02FF12" bold="true" box="[907,951,223,249]" pageId="3" pageNumber="575">PM</emphasis>
) especially strongly developed, high, each tightly fused medially almost all along and only apically slightly divided, inclined anteriad on collum and rings 24, subvertical until ring 16, thereafter declined increasingly caudad (
<figureCitation id="40AA2A0ECE28550593B72D24FBB5FEAA" box="[897,1024,295,321]" captionStart="FIGURES 18" captionStartId="2.[151,264,1857,1881]" captionTargetBox="[186,1401,181,1831]" captionTargetId="figure-15@2.[186,1401,181,1834]" captionTargetPageId="2" captionText="FIGURES 18. Pyrgodesmus obscurus Pocock, 1892, male syntypes. 1. Original labels. 2. Both syntypes pierced with glass minutiae, shown relative to the length of an insect pin. 3. Habitus of a syntype, lateral view (after Pocock (1892)). 4. Picture of the same, lateral view. 5. Body rings 57 and gonopods in situ, ventral view. 6. Body rings 69 and gonopods in situ, ventrolateral view. 7 &amp; 8. Enlarged gonopods in situ, ventral and ventrolateral views, respectively. Figs 13, 7 and 8 not to scale. Images courtesy Dmitry Telnov." figureDoi="http://doi.org/10.5281/zenodo.5709646" httpUri="https://zenodo.org/record/5709646/files/figure.png" pageId="3" pageNumber="575">Figs 3 &amp; 4</figureCitation>
).
<emphasis id="EAE5EA99CE28550594202D24FBF7FEAA" bold="true" box="[1046,1090,295,321]" pageId="3" pageNumber="575">PM</emphasis>
mostly thickened and vaguely bilobate apically, only last few
<emphasis id="EAE5EA99CE28550591C82D48FD9FFE8E" bold="true" box="[510,554,331,357]" pageId="3" pageNumber="575">PM</emphasis>
trilobate (
<figureCitation id="40AA2A0ECE28550592972D48FD5DFE8E" box="[673,744,331,357]" captionStart="FIGURES 18" captionStartId="2.[151,264,1857,1881]" captionTargetBox="[186,1401,181,1831]" captionTargetId="figure-15@2.[186,1401,181,1834]" captionTargetPageId="2" captionText="FIGURES 18. Pyrgodesmus obscurus Pocock, 1892, male syntypes. 1. Original labels. 2. Both syntypes pierced with glass minutiae, shown relative to the length of an insect pin. 3. Habitus of a syntype, lateral view (after Pocock (1892)). 4. Picture of the same, lateral view. 5. Body rings 57 and gonopods in situ, ventral view. 6. Body rings 69 and gonopods in situ, ventrolateral view. 7 &amp; 8. Enlarged gonopods in situ, ventral and ventrolateral views, respectively. Figs 13, 7 and 8 not to scale. Images courtesy Dmitry Telnov." figureDoi="http://doi.org/10.5281/zenodo.5709646" httpUri="https://zenodo.org/record/5709646/files/figure.png" pageId="3" pageNumber="575">Fig. 4</figureCitation>
). Paraterga very low, oblique and strongly declined, vaguely bi- or trilobate at lateral margin (
<figureCitation id="40AA2A0ECE28550592272D6CFDCCFE61" box="[529,633,367,394]" captionStart="FIGURES 18" captionStartId="2.[151,264,1857,1881]" captionTargetBox="[186,1401,181,1831]" captionTargetId="figure-15@2.[186,1401,181,1834]" captionTargetPageId="2" captionText="FIGURES 18. Pyrgodesmus obscurus Pocock, 1892, male syntypes. 1. Original labels. 2. Both syntypes pierced with glass minutiae, shown relative to the length of an insect pin. 3. Habitus of a syntype, lateral view (after Pocock (1892)). 4. Picture of the same, lateral view. 5. Body rings 57 and gonopods in situ, ventral view. 6. Body rings 69 and gonopods in situ, ventrolateral view. 7 &amp; 8. Enlarged gonopods in situ, ventral and ventrolateral views, respectively. Figs 13, 7 and 8 not to scale. Images courtesy Dmitry Telnov." figureDoi="http://doi.org/10.5281/zenodo.5709646" httpUri="https://zenodo.org/record/5709646/files/figure.png" pageId="3" pageNumber="575">Figs 36</figureCitation>
). Pore formula normal (5, 7, 9, 10, 12, 13, 1519), ozopores borne on distinct porosteles on rings 5, 7, 9, 10, 12, 13 and 15, but opening flush on dorsal surface at base of caudal lobulation on rings 1619 (
<figureCitation id="40AA2A0ECE28550591792DB4FE03FE39" box="[335,438,439,466]" captionStart="FIGURES 18" captionStartId="2.[151,264,1857,1881]" captionTargetBox="[186,1401,181,1831]" captionTargetId="figure-15@2.[186,1401,181,1834]" captionTargetPageId="2" captionText="FIGURES 18. Pyrgodesmus obscurus Pocock, 1892, male syntypes. 1. Original labels. 2. Both syntypes pierced with glass minutiae, shown relative to the length of an insect pin. 3. Habitus of a syntype, lateral view (after Pocock (1892)). 4. Picture of the same, lateral view. 5. Body rings 57 and gonopods in situ, ventral view. 6. Body rings 69 and gonopods in situ, ventrolateral view. 7 &amp; 8. Enlarged gonopods in situ, ventral and ventrolateral views, respectively. Figs 13, 7 and 8 not to scale. Images courtesy Dmitry Telnov." figureDoi="http://doi.org/10.5281/zenodo.5709646" httpUri="https://zenodo.org/record/5709646/files/figure.png" pageId="3" pageNumber="575">Figs 36</figureCitation>
). Surface below
<emphasis id="EAE5EA99CE28550592442DB4FD2BFE3A" bold="true" box="[626,670,439,465]" pageId="3" pageNumber="575">PM</emphasis>
microgranulate, with neither
<emphasis id="EAE5EA99CE28550593DD2DB4FBA7FE3A" bold="true" box="[1003,1042,439,465]" pageId="3" pageNumber="575">DL</emphasis>
nor
<emphasis id="EAE5EA99CE28550594732DB5FBF8FE3B" bold="true" box="[1093,1101,438,464]" pageId="3" pageNumber="575">i</emphasis>
, nor
<emphasis id="EAE5EA99CE28550594BE2DB5FB01FE3B" bold="true" box="[1160,1204,438,464]" pageId="3" pageNumber="575">Am</emphasis>
, nor
<emphasis id="EAE5EA99CE28550594D92DB5FAAEFE3B" bold="true" box="[1263,1307,438,464]" pageId="3" pageNumber="575">Cm</emphasis>
discernible beneath an earth crust.
<emphasis id="EAE5EA99CE28550591AF2DD8FE70FE1E" bold="true" box="[409,453,475,501]" pageId="3" pageNumber="575">PM</emphasis>
19 subhorizontal, fused medially and forming a deep caudal notch, tip of epiproct being slightly visible from above.
</paragraph>
<paragraph id="D82E368BCE28550590F12E20FE77FD1A" blockId="3.[151,1437,150,789]" pageId="3" pageNumber="575">
Legs robust, densely setose, each prefemur with a particularly long distoventral seta; claw small and simple (
<figureCitation id="40AA2A0ECE28550590A92E44FEA9FD89" box="[159,284,582,610]" captionStart="FIGURES 18" captionStartId="2.[151,264,1857,1881]" captionTargetBox="[186,1401,181,1831]" captionTargetId="figure-15@2.[186,1401,181,1834]" captionTargetPageId="2" captionText="FIGURES 18. Pyrgodesmus obscurus Pocock, 1892, male syntypes. 1. Original labels. 2. Both syntypes pierced with glass minutiae, shown relative to the length of an insect pin. 3. Habitus of a syntype, lateral view (after Pocock (1892)). 4. Picture of the same, lateral view. 5. Body rings 57 and gonopods in situ, ventral view. 6. Body rings 69 and gonopods in situ, ventrolateral view. 7 &amp; 8. Enlarged gonopods in situ, ventral and ventrolateral views, respectively. Figs 13, 7 and 8 not to scale. Images courtesy Dmitry Telnov." figureDoi="http://doi.org/10.5281/zenodo.5709646" httpUri="https://zenodo.org/record/5709646/files/figure.png" pageId="3" pageNumber="575">Figs 5 &amp; 6</figureCitation>
). Gonopods (
<figureCitation id="40AA2A0ECE28550591832E44FD85FD8A" box="[437,560,583,609]" captionStart="FIGURES 18" captionStartId="2.[151,264,1857,1881]" captionTargetBox="[186,1401,181,1831]" captionTargetId="figure-15@2.[186,1401,181,1834]" captionTargetPageId="2" captionText="FIGURES 18. Pyrgodesmus obscurus Pocock, 1892, male syntypes. 1. Original labels. 2. Both syntypes pierced with glass minutiae, shown relative to the length of an insect pin. 3. Habitus of a syntype, lateral view (after Pocock (1892)). 4. Picture of the same, lateral view. 5. Body rings 57 and gonopods in situ, ventral view. 6. Body rings 69 and gonopods in situ, ventrolateral view. 7 &amp; 8. Enlarged gonopods in situ, ventral and ventrolateral views, respectively. Figs 13, 7 and 8 not to scale. Images courtesy Dmitry Telnov." figureDoi="http://doi.org/10.5281/zenodo.5709646" httpUri="https://zenodo.org/record/5709646/files/figure.png" pageId="3" pageNumber="575">Figs 7 &amp; 8</figureCitation>
) relatively simple,
<emphasis id="EAE5EA99CE285505933B2E44FCE7FD8B" box="[781,850,583,608]" italics="true" pageId="3" pageNumber="575">in situ</emphasis>
held almost parallel to each other, with only tips of telopodites (= solenomeres), both slightly curved ventrad and poorly bifid, crossing medially; each gonopod consisting of a large, globose, densely microsetose and micropapillate coxite and a unipartite, rather long, ribbon-shaped, distally attenuating telopodite/solenomere curved caudad, strongly sunken inside a deep gonocoel and terminating the seminal groove on top.
</paragraph>
</subSubSection>
<subSubSection id="908B6500CE28550590F12EF9FDDAFCFE" box="[199,623,762,789]" pageId="3" pageNumber="575" type="distribution">
<paragraph id="D82E368BCE28550590F12EF9FDDAFCFE" blockId="3.[151,1437,150,789]" box="[199,623,762,789]" pageId="3" pageNumber="575">
<emphasis id="EAE5EA99CE28550590F12EF9FED4FCFF" bold="true" box="[199,353,762,788]" pageId="3" pageNumber="575">Distribution.</emphasis>
Pundaluoya,
<collectingCountry id="A086761BCE28550591CC2EF8FDDEFCFE" box="[506,619,763,789]" name="Sri Lanka" pageId="3" pageNumber="575">Sri Lanka</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>