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<document ID-DOI="http://dx.doi.org/10.3897/zse.99.102586" ID-GBIF-Dataset="b028f1ea-32d8-4aa6-990c-474b0d2cc3ff" ID-Pensoft-Pub="1860-0743-2-285" ID-Pensoft-UUID="EADD100D16075520AFD7FBF99E77DAD6" ID-ZooBank="6856760F3F9E4B278AC2B59D25845F67" ModsDocID="1860-0743-99-2-285" checkinTime="1683904577099" checkinUser="pensoft" docAuthor="Zhang, Le-Jia, Du, Li-Na &amp; von Rintelen, Thomas" docDate="2023" docId="4F71F8C5709B5EE097E5B5C7E25A4C5B" docLanguage="en" docName="ZoosystEvol 99(2): 285-297" docOrigin="Zoosystematics and Evolution 99 (2)" docPubDate="2023-05-11" docSource="http://dx.doi.org/10.3897/zse.99.102586" docTitle="Dalipaludina oxytropoides Zhang &amp; Du &amp; von Rintelen 2023, comb. nov." docType="treatment" docVersion="4" id="EADD100D16075520AFD7FBF99E77DAD6" lastPageNumber="285" masterDocId="EADD100D16075520AFD7FBF99E77DAD6" masterDocTitle="A new genus of river snails, Dalipaludina (Gastropoda, Viviparidae), endemic to the Yunnan Plateau of SW China" masterLastPageNumber="297" masterPageNumber="285" pageNumber="285" updateTime="1732710123990" updateUser="ExternalLinkService">
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<mods:title id="0431E8F2137A8AF7B50D59025260AC52">A new genus of river snails, Dalipaludina (Gastropoda, Viviparidae), endemic to the Yunnan Plateau of SW China</mods:title>
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<mods:namePart id="A338348A6FFB2825C5A3FE684956B916">Zhang, Le-Jia</mods:namePart>
<mods:affiliation id="D8C8B4F0A8845A1DC855020D4A5015A1">Museum fuer Naturkunde, Leibniz, Institut fuer Evolutions und Biodiversitaetsforschung, Berlin 10115, Germany</mods:affiliation>
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<mods:namePart id="F12AA16C3BFDC71F267A5F71BD822441">Du, Li-Na</mods:namePart>
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<mods:affiliation id="6C2203869C6BC482FDC488900C814F83">Key Laboratory of Ecology of Rare and Endangered Species and Environmental Protection, Guangxi Normal University, Ministry of Education, Guilin, Guangxi 541004, China</mods:affiliation>
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<mods:namePart id="6922F0EC5DF457F766B32B4731A73135">von Rintelen, Thomas</mods:namePart>
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<mods:affiliation id="CB580B08D31E156CED51086C3861E402">Museum fuer Naturkunde, Leibniz, Institut fuer Evolutions und Biodiversitaetsforschung, Berlin 10115, Germany</mods:affiliation>
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<mods:title id="E43413A4B1C8B93636012A957D617A19">Zoosystematics and Evolution</mods:title>
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<mods:date id="9E2CD36D8D970BEC2051CF4432FE4C82">2023</mods:date>
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<treatment id="4F71F8C5709B5EE097E5B5C7E25A4C5B" ID-GBIF-Taxon="209619128" LSID="urn:lsid:plazi:treatment:4F71F8C5709B5EE097E5B5C7E25A4C5B" httpUri="http://treatment.plazi.org/id/4F71F8C5709B5EE097E5B5C7E25A4C5B" lastPageNumber="285" pageId="0" pageNumber="285" scope_class="Gastropoda" scope_family="Viviparidae" scope_subFamily="Bellamyinae">
<subSubSection id="22DE9DF8997BA54E5FE5DDE0AC202EE7" pageId="0" pageNumber="285" type="nomenclature">
<paragraph id="73123600A6236702100B2836B02CD821" pageId="0" pageNumber="285">
<taxonomicName id="382095A021FD52EA83E0DAFF95539B22" LSID="4F71F8C5-709B-5EE0-97E5-B5C7E25A4C5B" authority="(Heude, 1889)" authorityName="Zhang &amp; Du &amp; von Rintelen" authorityYear="2023" baseAuthorityName="Heude" baseAuthorityYear="1889" class="Gastropoda" family="Viviparidae" genus="Dalipaludina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dalipaludina oxytropoides" order="Architaenioglossa" pageId="0" pageNumber="285" phylum="Mollusca" rank="species" species="oxytropoides" status="comb. nov.">Dalipaludina oxytropoides (Heude, 1889)</taxonomicName>
<taxonomicNameLabel id="D320D1B159B0138C710BB65DEF8B231A" pageId="0" pageNumber="285">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="AD7079C427CD16F59DCC38A055D67014" pageId="0" pageNumber="285" type="reference_group">
<paragraph id="D9CD2DD139D5D18CDB437816FDDB875A" pageId="0" pageNumber="285">
<taxonomicName id="E321C4F7BCC0DFF62812F571A278060D" authorityName="Zhang &amp; Du &amp; von Rintelen" authorityYear="2023" baseAuthorityName="Zhang &amp; Du &amp; von Rintelen" baseAuthorityYear="2023" class="Gastropoda" family="Viviparidae" genus="Paludina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paludina oxytropoides" order="Architaenioglossa" pageId="0" pageNumber="285" phylum="Mollusca" rank="species" species="oxytropoides" status="comb. nov.">Paludina oxytropoides</taxonomicName>
Heude, 1889: 176 ('lacu prope
<normalizedToken id="9C8E5C6AA2460403CCF4178C59AF2DF9" originalValue="Tchao-tong">Tchao-tong'</normalizedToken>
= lake near Zhaotong City,Yunnan Province, China).
</paragraph>
<paragraph id="FE91EE109E79ED2DF70D313EA73D5637" pageId="0" pageNumber="285">
<taxonomicName id="1ADB00284C6BFF296D9BB464B00B19D7" authorityName="Zhang &amp; Du &amp; von Rintelen" authorityYear="2023" class="Gastropoda" family="Viviparidae" genus="Vivipara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vivipara oxytropoides" order="Architaenioglossa" pageId="0" pageNumber="285" phylum="Mollusca" rank="species" species="oxytropoides" status="comb. nov.">Vivipara oxytropoides</taxonomicName>
-
<bibRefCitation id="5AB061EF056814C191C93502BCA85D12" author="Kobelt, W" journalOrPublisher="Bauer &amp; Raspe, Nuremberg" pageId="0" pageNumber="285" refId="B7" refString="Kobelt, W, 1909. Systematisches Conchylien-Cabinet von Martini und Chemnitz. Die Raublungenschnecken (Agnatha). Zweite Abtheilung: Streptaxidae und Daudebardiidae. Bauer &amp; Raspe, Nuremberg" title="Systematisches Conchylien-Cabinet von Martini und Chemnitz. Die Raublungenschnecken (Agnatha). Zweite Abtheilung: Streptaxidae und Daudebardiidae." year="1909">Kobelt 1909</bibRefCitation>
: 128.
</paragraph>
<paragraph id="FCE927DCD111B35425661266D117D71B" pageId="0" pageNumber="285">
<taxonomicName id="8ACD2D5E88312D0CECBCBE90A259965A" authorityName="Zhang &amp; Du &amp; von Rintelen" authorityYear="2023" baseAuthorityName="Zhang &amp; Du &amp; von Rintelen" baseAuthorityYear="2023" class="Gastropoda" family="Viviparidae" genus="Margarya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Margarya" order="Architaenioglossa" pageId="0" pageNumber="285" phylum="Mollusca" rank="genus" status="comb. nov.">Margarya</taxonomicName>
sp. -
<bibRefCitation id="8400CA1A30EEEFDEEE8303454F1AD841" author="Du, LN" journalOrPublisher="Journal of Hydroecology" pageId="0" pageNumber="285" pagination="44 - 49" refId="B3" refString="Du, LN, Chen, XY, Yang, JX, Aldridge, DC, 2012. A survey of Molluscan compositions in four plateau lakes, Northwest Yunnan [In Chinese]. China. Journal of Hydroecology 33: 44 - 49" title="A survey of Molluscan compositions in four plateau lakes, Northwest Yunnan [In Chinese]. China." volume="33" year="2012">Du et al. 2012</bibRefCitation>
: 45.
</paragraph>
<paragraph id="DFE11281DB93A604857FCC1F3B59E994" pageId="0" pageNumber="285">
<taxonomicName id="87EBCF02F02EDDFDDE674E18C92A3BC7" authorityName="Zhang &amp; Du &amp; von Rintelen" authorityYear="2023" baseAuthorityName="Zhang &amp; Du &amp; von Rintelen" baseAuthorityYear="2023" class="Gastropoda" family="Viviparidae" genus="Margarya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Margarya oxytropoides" order="Architaenioglossa" pageId="0" pageNumber="285" phylum="Mollusca" rank="species" species="oxytropoides" status="comb. nov.">Margarya oxytropoides</taxonomicName>
-
<bibRefCitation id="1CF80B94035B85EB5184AE7E4BAD12BE" DOI="https://doi.org/10.1111/zoj.12260" author="Zhang, LJ" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="285" pagination="760 - 800" refId="B25" refString="Zhang, LJ, Chen, SC, Yang, LT, Jin, L, Koehler, F, 2015. Systematic revision of the freshwater snail Margarya Nevill, 1877 (Mollusca: Viviparidae) endemic to the ancient lakes of Yunnan, China, with description of new taxa. Zoological Journal of the Linnean Society 174 (4): 760 - 800, DOI: https://doi.org/10.1111/zoj.12260" title="Systematic revision of the freshwater snail Margarya Nevill, 1877 (Mollusca: Viviparidae) endemic to the ancient lakes of Yunnan, China, with description of new taxa." url="https://doi.org/10.1111/zoj.12260" volume="174" year="2015">Zhang et al. 2015</bibRefCitation>
: 777;
<bibRefCitation id="885998F7D31591B16E7223EC3070817D" DOI="https://doi.org/10.1002/ece3.8622" author="Wiese, R" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="285" refId="B17" refString="Wiese, R, Harrington, K, Hartmann, K, Hethke, M, von Rintelen, T, Zhang, HC, Zhang, LJ, Riedel, F, 2022. Can fractal dimensions objectivize gastropod shell morphometrics? A case study from Lake Lugu (SW China). Ecology and Evolution 12(3): e8622. https://doi.org/10.1002/ece3.8622" title="Can fractal dimensions objectivize gastropod shell morphometrics? A case study from Lake Lugu (SW China). Ecology and Evolution 12 (3): e 8622." url="https://doi.org/10.1002/ece3.8622" year="2022">Wiese et al. 2022</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8A83A47931FF7F78103F3861C856ADC8" pageId="0" pageNumber="285" type="materials_examined">
<paragraph id="4077C760DAD8798F4F9405018EAAE8D9" pageId="0" pageNumber="285">Material examined.</paragraph>
<paragraph id="584EE3BB1D6CA58C25024E7BDC3FC927" pageId="0" pageNumber="285">
<specimenCount id="235A80466FEE0B9F121229359F28BC09" type="generic" typeStatus="syntypes">
2
<emphasis id="74F942942DDB20BD56A47EFCB7F14FE6" bold="true" italics="true" pageId="0" pageNumber="285">syntypes</emphasis>
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, IZCAS-FG-492586, IZCAS-FG-492587;
<specimenCount id="DD87CA4B8A9AC728093A77BF796E7CE7" type="generic" typeStatus="paratype">
1
<emphasis id="31400CC1773AA10960E3EADBFC2EC7F9" bold="true" italics="true" pageId="0" pageNumber="285">paratype</emphasis>
</specimenCount>
, MCZ-Mala-167343;
<materialsCitation id="BAC2518CFC178F8082BCEEEADD987BBD" collectingDate="2014-09" collectingDateMax="2014-10-31" collectingDateMin="2014-09" collectorName="Frank Riedel" location="Lake Lugu" specimenCount="91">
<specimenCount id="26BAB9F62888695E3704FED41BC198ED" type="generic">91 specimens</specimenCount>
in ZMB, collected by
<collectorName id="24B89CC1E9EBC8D7A4C864B3B8240207">Frank Riedel</collectorName>
from around 1 to 6 metres depth in all over the
<location id="2605A372A58BBD3F6D9CD1E15395AB29" LSID="urn:lsid:plazi:treatment:4F71F8C5709B5EE097E5B5C7E25A4C5B:2605A372A58BBD3F6D9CD1E15395AB29" name="Lake Lugu">Lake Lugu</location>
in
<collectingDate id="60558EF00EE33EFF8F3B2350DA128515" value="2014-09" valueMax="2014-10-31" valueMin="2014-09-00">September and October 2014</collectingDate>
</materialsCitation>
;
<materialsCitation id="731C822A64976A47BDE323D01AB440F6" collectingDate="2022-04" collectorName="Le-Jia Zhang" location="Lake Lugu" specimenCount="5">
<specimenCount id="CA8DAB17750C2CA0C63DA51B5B08DECE" type="generic">5 specimens</specimenCount>
in ZMB, collected by
<collectorName id="3B561B31D4194C5B23487FF7154CE18A">Le-Jia Zhang</collectorName>
from 1 metre depth in the east shore of
<location id="4CCA9D440826CAA82A02F9FA9CE0053F" LSID="urn:lsid:plazi:treatment:4F71F8C5709B5EE097E5B5C7E25A4C5B:4CCA9D440826CAA82A02F9FA9CE0053F" name="Lake Lugu">Lake Lugu</location>
in
<collectingDate id="C917FC9FAA4340833512A01B9841DA44" value="2022-04">April 2022</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F18DC501F1CDFDA38DD874EF90E92FDE" pageId="0" pageNumber="285" type="description">
<paragraph id="359C0C8906908E11267F80AC05014126" pageId="0" pageNumber="285">Description.</paragraph>
<paragraph id="398123AEE4D29BAD496113EC1E6D65E9" pageId="0" pageNumber="285">
Shell (Fig.
<figureCitation id="0140812F11F8A406B5A19492E58277E6" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shell and operculum of Dalipaludina oxytropoides. A-C. Specimen collected from Lake Lugu, ZMB. Moll. 250844; D-F. Varieties collected from Lake Lugu, ZLJ; G. Syntype IZCAS-FG- 492587 from a lake in Zhaotong, Yunann, photo taken by Kaibaryer Meng, Institute of Zoology, Chinese Academy of Sciences; H. Paratype MCZ-Mala- 167343 from a lake in Zhaotong, Yunann, photo taken by Alana Rivera, Museum of Comparative Zoology, Harvard University, President and Fellows of Harvard College; I. Exterior (left) and interior (right) surface of operculum. Scale bar: 1 cm, B-F, H-I with the same scale bar of A." figureDoi="10.3897/zse.99.102586.figure4" httpUri="https://binary.pensoft.net/fig/849017" pageId="0" pageNumber="285">4A-H</figureCitation>
) large, conical, thin, greenish yellow to dark brown in colour; up to six whorls at adulthood, including one relatively smooth protoconch whorl, apex acute; each teleoconch whorls with one strong keel at suture, above suture with many rather weak spiral threads, sometimes with additional two to three strong cords, base of shell with many thin weak spiral threads, weak spiral cords and threads with short periostracum setae; aperture ovate, less than half of shell in height, lip thin and simple, umbilicus narrow, sometimes bordered by a keel.
</paragraph>
<caption id="5AD6E07E86F529379C6AAB7E4DE82552" doi="10.3897/zse.99.102586.figure4" httpUri="https://binary.pensoft.net/fig/849017" pageId="0" pageNumber="285" start="Figure 4" startId="F4">
<paragraph id="FD397E90EDE638BB2E9FB11A2A7A633E" pageId="0" pageNumber="285">
<emphasis id="CE12F1DFF13EC446C4C8EE8DC977F547" bold="true" pageId="0" pageNumber="285">Figure 4.</emphasis>
Shell and operculum of
<taxonomicName id="FC090B15C920C11A56E1387C3F240A66" authorityName="Zhang &amp; Du &amp; von Rintelen" authorityYear="2023" baseAuthorityName="Heude" baseAuthorityYear="1889" class="Gastropoda" family="Viviparidae" genus="Dalipaludina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dalipaludina oxytropoides" order="Architaenioglossa" pageId="0" pageNumber="285" phylum="Mollusca" rank="species" species="oxytropoides">
<emphasis id="B5EC3873E82784E85D0648379ADC7637" italics="true" pageId="0" pageNumber="285">Dalipaludina oxytropoides</emphasis>
</taxonomicName>
.
<emphasis id="067FE5E82D2FB1EAE42F87027D26C74F" bold="true" pageId="0" pageNumber="285">A-C.</emphasis>
Specimen collected from Lake Lugu, ZMB.Moll. 250844;
<emphasis id="A96F51418DC362838080B2B5832FD6C8" bold="true" pageId="0" pageNumber="285">D-F.</emphasis>
Varieties collected from Lake Lugu,ZLJ;
<emphasis id="5AA28D2B5B585F6845FA51FAB476D518" bold="true" pageId="0" pageNumber="285">G.</emphasis>
Syntype IZCAS-FG-492587 from a lake in Zhaotong, Yunann, photo taken by Kaibaryer Meng, Institute of Zoology, Chinese Academy of Sciences;
<emphasis id="A3A2EE9552896D4D62FBA903796FB29B" bold="true" pageId="0" pageNumber="285">H.</emphasis>
Paratype MCZ-Mala-167343 from a lake in Zhaotong, Yunann, photo taken by Alana Rivera, Museum of Comparative Zoology, Harvard University, President and Fellows of Harvard College;
<emphasis id="C476447C972DF38DA51EE0E489092758" bold="true" pageId="0" pageNumber="285">I.</emphasis>
Exterior (left) and interior (right) surface of operculum. Scale bar: 1 cm,
<emphasis id="73C744FF45BA0049D2FAB8037426CA6F" bold="true" pageId="0" pageNumber="285">B-F, H-I</emphasis>
with the same scale bar of
<emphasis id="95C2654480D434A463BCF8F4F07A1A59" bold="true" pageId="0" pageNumber="285">A</emphasis>
.
</paragraph>
</caption>
<paragraph id="707AC8C4D492685E856B54118389CDCC" pageId="0" pageNumber="285">
Operculum (Fig.
<figureCitation id="3AC42DDFA944CF22DD10C52FD2A93447" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shell and operculum of Dalipaludina oxytropoides. A-C. Specimen collected from Lake Lugu, ZMB. Moll. 250844; D-F. Varieties collected from Lake Lugu, ZLJ; G. Syntype IZCAS-FG- 492587 from a lake in Zhaotong, Yunann, photo taken by Kaibaryer Meng, Institute of Zoology, Chinese Academy of Sciences; H. Paratype MCZ-Mala- 167343 from a lake in Zhaotong, Yunann, photo taken by Alana Rivera, Museum of Comparative Zoology, Harvard University, President and Fellows of Harvard College; I. Exterior (left) and interior (right) surface of operculum. Scale bar: 1 cm, B-F, H-I with the same scale bar of A." figureDoi="10.3897/zse.99.102586.figure4" httpUri="https://binary.pensoft.net/fig/849017" pageId="0" pageNumber="285">4I</figureCitation>
) corneous, ovate, thin to relatively thick, yellow to reddish brown, with darker coloured nuclear region, exterior surface of operculum rather smooth, inner opercular region relatively small, nuclear region smooth or with small grains.
</paragraph>
<paragraph id="C0D6DECA27E28D3EEE0AA43AC940848C" pageId="0" pageNumber="285">
Radular (Fig.
<figureCitation id="4027B735625AF4A8A04CDCABFAB1B69B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. SEM photo of the radula and the protoconch of Dalipaludina species. A, B. Radula of Dalipaludina delavayana, A- ZMB. Moll. 181719, B- ZMB. Moll. 122707; C, D. Radula of Dalipaludina oxytropoides, ZMB. Moll. 250844; E, F. Radula of Dalipaludina occidentalis, ZMB. Moll. 122713. Abbreviations: ct-central teeth; lt-lateral teeth; imt-inner marginal teeth; omt-outer marginal teeth. Scale bars: 100 µm." figureDoi="10.3897/zse.99.102586.figure7" httpUri="https://binary.pensoft.net/fig/849020" pageId="0" pageNumber="285">7C, D</figureCitation>
) central tooth with one broad central denticle and three to four small sharp cusps on either side; lateral tooth with one broad central denticle and three to four small sharp cusps on either side; inner marginal tooth with one broad central denticle and four small sharp cusps on either side; each outer marginal tooth with 10 to 11 small sharp cusps.
</paragraph>
</subSubSection>
<subSubSection id="8A54403FCA67167EEF4F8D1821AFEDF3" pageId="0" pageNumber="285" type="remarks">
<paragraph id="F510BDA001B3F06C65E37D4F62D88826" pageId="0" pageNumber="285">Remarks.</paragraph>
<paragraph id="9B52C809D662409DAD4A6EC104D89B00" pageId="0" pageNumber="285">
This species has the most variable shell within
<taxonomicName id="033DC71403804702045D82D387A878EC" authorityName="Zhang" authorityYear="2023" class="Gastropoda" family="Viviparidae" genus="Dalipaludina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dalipaludina" order="Architaenioglossa" pageId="0" pageNumber="285" phylum="Mollusca" rank="genus">
<emphasis id="FA5BBA32CF4CF622202D482FA4870CDE" italics="true" pageId="0" pageNumber="285">Dalipaludina</emphasis>
</taxonomicName>
. The form with strong cords of this species can be easily distinguished from the other
<taxonomicName id="B1611B56231E5314C9746E2059BFA805" authorityName="Zhang" authorityYear="2023" class="Gastropoda" family="Viviparidae" genus="Dalipaludina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dalipaludina" order="Architaenioglossa" pageId="0" pageNumber="285" phylum="Mollusca" rank="genus">
<emphasis id="495F3C692DF687F064D80788A6DE80E3" italics="true" pageId="0" pageNumber="285">Dalipaludina</emphasis>
</taxonomicName>
species. It used to be considered as a species of
<taxonomicName id="4C10A035264535FF16F4B90280477A11" authorityName="Zhang &amp; Du &amp; von Rintelen" authorityYear="2023" baseAuthorityName="Zhang &amp; Du &amp; von Rintelen" baseAuthorityYear="2023" class="Gastropoda" family="Viviparidae" genus="Margarya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Margarya" order="Architaenioglossa" pageId="0" pageNumber="285" phylum="Mollusca" rank="genus">
<emphasis id="524FED5CF1793875280E749977B9F1F7" italics="true" pageId="0" pageNumber="285">Margarya</emphasis>
</taxonomicName>
with a distribution in Lake Dianchi as well (
<bibRefCitation id="9CBD65FC794D20B73DF778565600339B" DOI="https://doi.org/10.1111/zoj.12260" author="Zhang, LJ" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="285" pagination="760 - 800" refId="B25" refString="Zhang, LJ, Chen, SC, Yang, LT, Jin, L, Koehler, F, 2015. Systematic revision of the freshwater snail Margarya Nevill, 1877 (Mollusca: Viviparidae) endemic to the ancient lakes of Yunnan, China, with description of new taxa. Zoological Journal of the Linnean Society 174 (4): 760 - 800, DOI: https://doi.org/10.1111/zoj.12260" title="Systematic revision of the freshwater snail Margarya Nevill, 1877 (Mollusca: Viviparidae) endemic to the ancient lakes of Yunnan, China, with description of new taxa." url="https://doi.org/10.1111/zoj.12260" volume="174" year="2015">Zhang et al. 2015</bibRefCitation>
). However, the COI-based phylogeny showed that &quot;
<taxonomicName id="47F04BB343316EB622A20F25A7821BB2" class="Gastropoda" family="Viviparidae" genus="Margarya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Margarya oxytropides" order="Architaenioglossa" pageId="0" pageNumber="285" phylum="Mollusca" rank="species" species="oxytropides">
<emphasis id="4F6B6FAE396CC3FF86AFDFAAAA360AFE" italics="true" pageId="0" pageNumber="285">Margarya oxytropides</emphasis>
</taxonomicName>
&quot; from Dianchi is not a species of
<taxonomicName id="567B0B7C172CD013EB52C35E8DE5D9C7" authorityName="Zhang" authorityYear="2023" class="Gastropoda" family="Viviparidae" genus="Dalipaludina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dalipaludina" order="Architaenioglossa" pageId="0" pageNumber="285" phylum="Mollusca" rank="genus">
<emphasis id="D258363E5C147D2BE2882ECD403AA0AD" italics="true" pageId="0" pageNumber="285">Dalipaludina</emphasis>
</taxonomicName>
, but indeed a member of the
<taxonomicName id="52249371053C77FAAC516DD922EAAACA" authorityName="Hannibal" authorityYear="1912" class="Gastropoda" family="Viviparidae" genus="Cipangopaludina" kingdom="Animalia" lsidName="Cipangopaludina" order="Architaenioglossa" pageId="0" pageNumber="285" phylum="Mollusca" rank="genus">
<emphasis id="EDC3A0B7340407A3394D9026BA5747C0" italics="true" pageId="0" pageNumber="285">Cipangopaludina</emphasis>
</taxonomicName>
/
<taxonomicName id="EE2710BD2519CFE4608387C4365A64DB" authorityName="Zhang &amp; Du &amp; von Rintelen" authorityYear="2023" baseAuthorityName="Zhang &amp; Du &amp; von Rintelen" baseAuthorityYear="2023" class="Gastropoda" family="Viviparidae" genus="Margarya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Margarya" order="Architaenioglossa" pageId="0" pageNumber="285" phylum="Mollusca" rank="genus">
<emphasis id="649DE7507F1EBB78F27F8C2BB0DEDB7E" italics="true" pageId="0" pageNumber="285">Margarya</emphasis>
</taxonomicName>
complex. The syntypes of
<taxonomicName id="DE458E06C7F34BEAFCB648548C5091B3" authorityName="Zhang &amp; Du &amp; von Rintelen" authorityYear="2023" baseAuthorityName="Zhang &amp; Du &amp; von Rintelen" baseAuthorityYear="2023" class="Gastropoda" family="Viviparidae" genus="Paludina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paludina oxytropoides" order="Architaenioglossa" pageId="0" pageNumber="285" phylum="Mollusca" rank="species" species="oxytropoides">
<emphasis id="22643513FBC18779143BB9F98BC881BA" italics="true" pageId="0" pageNumber="285">Paludina oxytropoides</emphasis>
</taxonomicName>
Heude, 1889 from Zhaotong in Northeast Yunnan show a strong keel at the suture, two strong keels above the suture, a relatively smooth shell base with many weak spiral keels, and a narrow umbilicus with a keel around it. It is distinct from the &quot;
<taxonomicName id="A11034E56009C0FA0537A045D7F53DF3" class="Gastropoda" family="Viviparidae" genus="Margarya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Margarya oxytropides" order="Architaenioglossa" pageId="0" pageNumber="285" phylum="Mollusca" rank="species" species="oxytropides">
<emphasis id="F71AE6E0FCE64204D408E45BDB8C5DA4" italics="true" pageId="0" pageNumber="285">Margarya oxytropides</emphasis>
</taxonomicName>
&quot; from Lake Dianchi but quite similar to the strongly keeled form of
<taxonomicName id="AA7D2F19DF5DA09391D07DF027774872" lsidName="D. oxytropoides" pageId="0" pageNumber="285" rank="species" species="oxytropoides">
<emphasis id="8F3130A811D0EB0B2E3F37C66CE9B02E" italics="true" pageId="0" pageNumber="285">D. oxytropoides</emphasis>
</taxonomicName>
from Lake Lugu. Meanwhile, the shell morphology of &quot;
<taxonomicName id="AE97CE878A91713EE4EDF8A742690397" class="Gastropoda" family="Viviparidae" genus="Margarya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Margarya oxytropides" order="Architaenioglossa" pageId="0" pageNumber="285" phylum="Mollusca" rank="species" species="oxytropides">
<emphasis id="6E9068ED0B853F9079D4D3A83C7759A1" italics="true" pageId="0" pageNumber="285">Margarya oxytropides</emphasis>
</taxonomicName>
&quot; from Lake Dianchi matches the description and figure of
<taxonomicName id="0DB0535EB37D04584D7F6458BBF72EC4" authorityName="Zhang" authorityYear="1990" class="Gastropoda" family="Viviparidae" genus="Cipangopaludina" kingdom="Animalia" lsidName="Cipangopaludina dianchiensis" order="Architaenioglossa" pageId="0" pageNumber="285" phylum="Mollusca" rank="species" species="dianchiensis">
<emphasis id="141F35A03C836D206CEAF5C5D7677B58" italics="true" pageId="0" pageNumber="285">Cipangopaludina dianchiensis</emphasis>
</taxonomicName>
Zhang, 1990 endemic to Lake Dianchi (
<bibRefCitation id="2A4BFF7B7216E5BD937D092F57EF604B" author="Zhang, L" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="0" pageNumber="285" pagination="25 - 27" refId="B21" refString="Zhang, L, 1990. A new species of genus Cipangopaludina from Dianchi Lake in Yunnan Province, China. Acta Zootaxonomica Sinica 15 (1): 25 - 27" title="A new species of genus Cipangopaludina from Dianchi Lake in Yunnan Province, China." volume="15" year="1990">Zhang 1990</bibRefCitation>
). Therefore, instead of
<taxonomicName id="E0D2F8BDB033772296DA417B1F512BE2" lsidName="D. oxytropoides" pageId="0" pageNumber="285" rank="species" species="oxytropoides">
<emphasis id="D327B0B8CAC31D320FDF85C74B9BA625" italics="true" pageId="0" pageNumber="285">D. oxytropoides</emphasis>
</taxonomicName>
, &quot;
<taxonomicName id="9B1DFA022AA5FCDC64C1817D5D600E1E" class="Gastropoda" family="Viviparidae" genus="Margarya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Margarya oxytropides" order="Architaenioglossa" pageId="0" pageNumber="285" phylum="Mollusca" rank="species" species="oxytropides">
<emphasis id="3D28A630A25DF54167AB6BC766CEBF78" italics="true" pageId="0" pageNumber="285">Margarya oxytropides</emphasis>
</taxonomicName>
&quot; from Lake Dianchi is here recognised as
<taxonomicName id="0ABBDD30CE327BC9CD7D5A9A928A02E9" class="Gastropoda" family="Viviparidae" genus="Margarya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Margarya dianchiensis" order="Architaenioglossa" pageId="0" pageNumber="285" phylum="Mollusca" rank="species" species="dianchiensis">
<emphasis id="4D70AE7B61331D4622AF6D729BBAD3D3" italics="true" pageId="0" pageNumber="285">Margarya dianchiensis</emphasis>
</taxonomicName>
(Zhang, 1990) comb. nov.
<taxonomicName id="EF534D57B1C73A0B0D9FAB9A721718D1" authorityName="Zhang &amp; Du &amp; von Rintelen" authorityYear="2023" baseAuthorityName="Zhang &amp; Du &amp; von Rintelen" baseAuthorityYear="2023" class="Gastropoda" family="Viviparidae" genus="Paludina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paludina oxytropoides" order="Architaenioglossa" pageId="0" pageNumber="285" phylum="Mollusca" rank="species" species="oxytropoides">
<emphasis id="E30E4308A7AB3B7854C0CD692CE97675" italics="true" pageId="0" pageNumber="285">Paludina oxytropoides</emphasis>
</taxonomicName>
Heude, 1889 is reclassified as a member of
<taxonomicName id="59B99745736BC6B66EC839D71CB5374C" authorityName="Zhang" authorityYear="2023" class="Gastropoda" family="Viviparidae" genus="Dalipaludina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dalipaludina" order="Architaenioglossa" pageId="0" pageNumber="285" phylum="Mollusca" rank="genus">
<emphasis id="DD24F492653C26AEA4A282086DFED88A" italics="true" pageId="0" pageNumber="285">Dalipaludina</emphasis>
</taxonomicName>
. The recommended Chinese name of this species is
<normalizedToken id="53001FA115323C22948D029D5EC4599C" originalValue="“尖龙骨理田螺”">&quot;尖龙骨理田螺&quot;</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection id="66E59AFB100B4C429A210DF0C69A0863" pageId="0" pageNumber="285" type="habitat">
<paragraph id="C6ABD8C2DF01DE8AB1E9A2A36160D9FA" pageId="0" pageNumber="285">Habitat and distribution.</paragraph>
<paragraph id="4DEF7063BF150D3BC7CBCBBB0270BEC4" pageId="0" pageNumber="285">shallow to medium-depth water area with sandy bottoms in Lake Lugu, Yunnan/Sichuan, China; lakes in Zhaotong, Yunnan, China.</paragraph>
</subSubSection>
</treatment>
</document>