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<document id="BCDFCCA9B08B20E84DFAB4B0F0D31660" ID-DOI="10.11646/zootaxa.5216.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7403532" ID-ZooBank="F9C92B91-42CF-4D2D-B315-14FF801815D4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1670314409974" checkinUser="plazi" docAuthor="Piovesan, Mônica, Casagrande, Mirna Martins &amp; Mielke, Olaf Hermann Hendrik" docDate="2022" docId="4F391911FF61FF0DC3EFFED8BB5D4975" docLanguage="en" docName="zootaxa.5216.1.1.pdf" docOrigin="Zootaxa 5216 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Opsiphanes jacobsorum Chacon &amp; Montero 2012" docType="treatment" docVersion="3" lastPageNumber="206" masterDocId="B3006169FFADFFC0C378FFB6B8684A61" masterDocTitle="Systematics of Opsiphanes Doubleday, [1849] (Lepidoptera: Nymphalidae, Satyrinae, Brassolini): an integrative approach" masterLastPageNumber="278" masterPageNumber="1" pageNumber="205" updateTime="1699401028458" updateUser="plazi">
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<mods:title id="38CF8997D211C4F5A273DBFD2E70CCD4">Systematics of Opsiphanes Doubleday, [1849] (Lepidoptera: Nymphalidae, Satyrinae, Brassolini): an integrative approach</mods:title>
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<mods:namePart id="DC6A0951B150677B0140BDE0D179D63F">Mielke, Olaf Hermann Hendrik</mods:namePart>
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<paragraph id="C72FA807FF61FF0CC3EFFED8BA974BE9" blockId="204.[151,767,366,429]" box="[151,767,366,393]" pageId="204" pageNumber="205">
<heading id="9C671F6BFF61FF0CC3EFFED8BA974BE9" bold="true" box="[151,767,366,393]" fontSize="11" level="2" pageId="204" pageNumber="205" reason="4">
<taxonomicName id="0090D384FF61FF0CC3EFFED8BA974BE9" authority="Chacon &amp; Montero, 2012" authorityName="Chacon &amp; Montero" authorityYear="2012" box="[151,767,366,393]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="jacobsorum">
<emphasis id="F5E47415FF61FF0CC3EFFED8BA974BE9" bold="true" box="[151,767,366,393]" pageId="204" pageNumber="205">
<emphasis id="F5E47415FF61FF0CC3EFFED8B9D24BE8" bold="true" box="[151,442,366,393]" italics="true" pageId="204" pageNumber="205">Opsiphanes jacobsorum</emphasis>
Chacón &amp; Montero, 2012
</emphasis>
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<subSubSection id="8F8AFB8CFF61FF0CC3EFFE25B92B4BCC" box="[151,323,403,429]" pageId="204" pageNumber="205" type="description">
<paragraph id="C72FA807FF61FF0CC3EFFE25B92B4BCC" blockId="204.[151,767,366,429]" box="[151,323,403,429]" pageId="204" pageNumber="205">
(
<figureCitation id="5FABB482FF61FF0CC3E7FE25B9534BCC" box="[159,315,403,429]" captionStart-0="FIGURE 187" captionStart-1="FIGURE 188" captionStart-2="FIGURE 189" captionStart-3="FIGURE 190" captionStart-4="FIGURE 191" captionStartId-0="205.[151,250,1923,1947]" captionStartId-1="206.[151,250,1025,1049]" captionStartId-2="206.[151,250,1965,1989]" captionStartId-3="207.[151,250,1107,1131]" captionStartId-4="207.[151,250,1987,2011]" captionTargetBox-0="[187,1379,823,1879]" captionTargetBox-1="[554,1027,183,987]" captionTargetBox-2="[353,1235,1117,1940]" captionTargetBox-3="[287,1320,269,1065]" captionTargetBox-4="[229,1359,1176,1963]" captionTargetId-0="figure-271@205.[175,1412,797,1899]" captionTargetId-1="figure-28@206.[338,1249,166,1001]" captionTargetId-2="figure-53@206.[353,1235,1094,1940]" captionTargetId-3="figure-19@207.[250,1336,166,1083]" captionTargetId-4="figure-44@207.[229,1359,1176,1963]" captionTargetPageId-0="205" captionTargetPageId-1="206" captionTargetPageId-2="206" captionTargetPageId-3="207" captionTargetPageId-4="207" captionText-0="FIGURE 187. Opsiphanes jacobsorum Chacón &amp; Montero, 2012: Male, dorsal and ventral views: Costa Rica: a. (USN- MENT01590011-USNM) Guanacaste, b. (USNMENT01590012-USNM) Alajuela; Panama: c. (USNMENT01590040-USNM) Panamá, d. (FLMNH MGCL 1107067-MGCL) Piña. White arrows indicate diagnostic characters of the species." captionText-1="FIGURE 188. Opsiphanes jacobsorum Chacón &amp; Montero, 2012: Female, dorsal and ventral views: Costa Rica: a. (USN- MENT01590014-USNM), b. (USNMENT01590784-USNM) Guanacaste." captionText-2="FIGURE 189. Opsiphanes jacobsorum Chacón &amp; Montero, 2012: Male: a. abdomen, lateral view; genitalia: b. dorsal view, c. ventral view, d. lateral view (USNMENT01590012-USNM)." captionText-3="FIGURE 190. Opsiphanes jacobsorum Chacón &amp; Montero, 2012: Female genitalia: a. lateral view; b. detail of the signa; c. ventral view (USNMENT01590014-USNM)." captionText-4="FIGURE 191. Geographical distribution of Opsiphanes jacobsorum Chacón &amp; Montero, 2012." figureDoi-0="http://doi.org/10.5281/zenodo.7403985" figureDoi-1="http://doi.org/10.5281/zenodo.7403987" figureDoi-2="http://doi.org/10.5281/zenodo.7403989" figureDoi-3="http://doi.org/10.5281/zenodo.7403991" figureDoi-4="http://doi.org/10.5281/zenodo.7403993" httpUri-0="https://zenodo.org/record/7403985/files/figure.png" httpUri-1="https://zenodo.org/record/7403987/files/figure.png" httpUri-2="https://zenodo.org/record/7403989/files/figure.png" httpUri-3="https://zenodo.org/record/7403991/files/figure.png" httpUri-4="https://zenodo.org/record/7403993/files/figure.png" pageId="204" pageNumber="205">Figs 187–191</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="8F8AFB8CFF61FF0CC3EFFE61BD5548C6" pageId="204" pageNumber="205" type="reference_group">
<paragraph id="C72FA807FF61FF0CC3EFFE61BD5548C6" blockId="204.[151,1437,471,679]" pageId="204" pageNumber="205">
<taxonomicName id="0090D384FF61FF0CC3EFFE61BAF24B8E" authority="Chacon &amp; Montero, 2012" authorityName="Chacon &amp; Montero" authorityYear="2012" box="[151,666,471,495]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="jacobsorum">
<emphasis id="F5E47415FF61FF0CC3EFFE61B9E54B8E" box="[151,397,471,495]" italics="true" pageId="204" pageNumber="205">Opsiphanes jacobsorum</emphasis>
Chacón &amp; Montero, 2012
</taxonomicName>
,
<emphasis id="F5E47415FF61FF0CC1DCFE6EBAD04B8E" box="[676,696,472,495]" italics="true" pageId="204" pageNumber="205">in</emphasis>
Chacón
<emphasis id="F5E47415FF61FF0CC06AFE6EBB574B8E" box="[786,831,471,495]" italics="true" pageId="204" pageNumber="205">et al</emphasis>
.
<emphasis id="F5E47415FF61FF0CC033FE61BC404B8E" bold="true" box="[843,1064,471,495]" pageId="204" pageNumber="205">Bull. Allyn Mus. 166</emphasis>
: 3, figs 1–2 (male d, v), 3–4 (female d, v), 9–11 (male gen.), 12,13 (female gen.), 14 (scent organs), 15–26 (larva, pupa), 27 (genetic distance), 28 (geogr. distr.);
<typeStatus id="182B16A5FF61FF0CC3BFFDA2B948484D" box="[199,288,532,556]" pageId="204" pageNumber="205" type="holotype">holotype</typeStatus>
male, larva found
<date id="B32E8EC7FF61FF0CC29EFDA2BA37484D" box="[486,607,532,556]" pageId="204" pageNumber="205" value="2006-09-11">11-IX-2006</date>
, prepupa
<date id="B32E8EC7FF61FF0CC1BCFDA2BB56484D" box="[708,830,532,556]" pageId="204" pageNumber="205" value="2006-11-12">12-XI-2006</date>
, adult eclosed
<date id="B32E8EC7FF61FF0CC0A1FDA2BC27484D" box="[985,1103,532,556]" pageId="204" pageNumber="205" value="2006-12-08">8-XII-2006</date>
, Pasmompa, Sector Pitilla, Area de Conservación
<collectingRegion id="055466E5FF61FF0CC200FD85B987482A" box="[376,495,563,587]" country="Costa Rica" name="Guanacaste" pageId="204" pageNumber="205">Guanacaste</collectingRegion>
,
<collectingRegion id="055466E5FF61FF0CC282FD85BA19482A" box="[506,625,563,587]" country="Costa Rica" name="Guanacaste" pageId="204" pageNumber="205">Guanacaste</collectingRegion>
,
<collectingCountry id="BF87E897FF61FF0CC104FD85BA86482A" box="[636,750,563,587]" name="Costa Rica" pageId="204" pageNumber="205">Costa Rica</collectingCountry>
, lat 11.01926, long -85.40997,
<quantity id="006805E2FF61FF0CC754FD85BC07482A" box="[1068,1135,563,587]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="204" pageNumber="205" unit="m" value="440.0">440 m</quantity>
, Manuel Ríos
<emphasis id="F5E47415FF61FF0CC67CFD85BD4B482A" box="[1284,1315,563,587]" italics="true" pageId="204" pageNumber="205">leg</emphasis>
.—
<bibRefCitation id="A301D5F6FF61FF0CC639FD82B938480B" author="Neves, D. A. &amp; Paluch, M." pageId="204" pageNumber="205" pagination="13 - 21" refId="ref179012" refString="Neves, D. A. &amp; Paluch, M. (2016) Biology and External Morphology of Immatures of Opsiphanes quiteria meridionalis Staudinger (Lepidoptera: Nymphalidae). Neotropical Entomology, 45 (1), 13 - 21. https: // doi. org / 10.1007 / s 13744 - 015 - 0323 - 4" type="journal article" year="2016">Neves &amp; Paluch, 2016</bibRefCitation>
.
<emphasis id="F5E47415FF61FF0CC222FDE4BA66480B" bold="true" box="[346,526,594,618]" pageId="204" pageNumber="205">Neotrop. Ent. 45</emphasis>
(1): 13, 17, 18; cit.—S. Miller
<emphasis id="F5E47415FF61FF0CC034FDE5BB11480B" box="[844,889,594,618]" italics="true" pageId="204" pageNumber="205">et al</emphasis>
., 2016.
<emphasis id="F5E47415FF61FF0CC0B5FDE4BCB3480B" bold="true" box="[973,1243,593,618]" pageId="204" pageNumber="205">Phil. Trans. r. Soc. B 371</emphasis>
(1702)(20150339): 2.—Shirai
<emphasis id="F5E47415FF61FF0CC240FDC7B90E48E9" box="[312,358,624,648]" italics="true" pageId="204" pageNumber="205">et al</emphasis>
., 2017.
<emphasis id="F5E47415FF61FF0CC2C2FDC6BAA548E9" bold="true" box="[442,717,624,649]" pageId="204" pageNumber="205">Ann. Ent. Soc. Amer. 110</emphasis>
(2): 149, 155; biol., phylog.—
<bibRefCitation id="A301D5F6FF61FF0CC77AFDC6BC8248E8" author="Checa, M. F. &amp; Torres, K." box="[1026,1258,624,649]" pageId="204" pageNumber="205" pagination="56 - 61" refId="ref175457" refString="Checa, M. F. &amp; Torres, K. (2019) Notes on the natural history of six nymphalid butterfly species from an Ecuadorian dry forest. Tropical Lepidoptera Research, 29 (1), 56 - 61." type="journal article" year="2019">Checa &amp; Torres, 2019</bibRefCitation>
.
<emphasis id="F5E47415FF61FF0CC78CFDC7B88848C6" bold="true" pageId="204" pageNumber="205">Trop. Lep. Res. 29</emphasis>
(1): 59; cit.—
<bibRefCitation id="A301D5F6FF61FF0CC211FD39BA1848C6" author="Matos-Maravi, P. &amp; Wahlberg, N. &amp; Freitas, A. V. L. &amp; DeVries, P. &amp; Antonelli, A. &amp; Penz, C. M." box="[361,624,655,679]" pageId="204" pageNumber="205" pagination="704 - 724" refId="ref178428" refString="Matos-Maravi, P., Wahlberg, N., Freitas, A. V. L., DeVries, P., Antonelli, A. &amp; Penz, C. M. (2021) Mesoamerica is a cradle and the Atlantic Forest is a museum of Neotropical butterfly diversity: insights from the evolution and biogeography of Brassolini (Lepidoptera: Nymphalidae). Biological Journal of the Linnean Society, 133 (3), 704 - 724. https: // doi. org / 10.1093 / biolinnean / blab 034" type="journal article" year="2021">
Matos-Maraví
<emphasis id="F5E47415FF61FF0CC17AFD26BA4748C6" box="[514,559,655,679]" italics="true" pageId="204" pageNumber="205">et al</emphasis>
. 2021
</bibRefCitation>
,
<emphasis id="F5E47415FF61FF0CC103FD39BBE148C6" bold="true" box="[635,905,655,679]" pageId="204" pageNumber="205">Biol. Jour. Linn. Soc. 133</emphasis>
(3): 711, fig. 2 (phylogenetic tree); phylog.
</paragraph>
</subSubSection>
<subSubSection id="8F8AFB8CFF61FF0CC3EFFD63B9ED491E" pageId="204" pageNumber="205" type="materials_examined">
<paragraph id="C72FA807FF61FF0CC3EFFD63B9ED491E" blockId="204.[151,1437,725,2012]" pageId="204" pageNumber="205">
<emphasis id="F5E47415FF61FF0CC3EFFD63B92D488E" bold="true" box="[151,325,725,751]" pageId="204" pageNumber="205">Type material.</emphasis>
<taxonomicName id="0090D384FF61FF0CC234FD63BBE9488E" authority="Chacon &amp; Montero, 2012" authorityName="Chacon &amp; Montero" authorityYear="2012" box="[332,897,725,752]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="jacobsorum">
<emphasis id="F5E47415FF61FF0CC234FD63BA33488E" box="[332,603,725,751]" italics="true" pageId="204" pageNumber="205">Opsiphanes jacobsorum</emphasis>
<materialsCitation id="77F8A25AFF61FF0CC11AFD63BBE9488E" box="[610,897,725,752]" collectingDate="2006-09-11" collectingDateMax="2006-12-08" collectingDateMin="2006-09-11" collectionCode="INBio" collectorName="de Conservacion &amp; Sector Pitilla &amp; Pasmompa" country="Costa Rica" elevation="440" latitude="11.01926" location="Area de Conservacion" longLatPrecision="1" longitude="-85.40997" pageId="204" pageNumber="205" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="holotype">Chacón &amp; Montero, 2012</materialsCitation>
</taxonomicName>
was described based on the
<typeStatus id="182B16A5FF61FF0CC7B8FD63BD4A488E" box="[1216,1314,725,751]" pageId="204" pageNumber="205" type="holotype">holotype</typeStatus>
male from
<collectingCountry id="BF87E897FF61FF0CC3EFFD4FB97F4972" box="[151,279,761,787]" name="Costa Rica" pageId="204" pageNumber="205">Costa Rica</collectingCountry>
, deposited at INBio, with the following information:
<typeStatus id="182B16A5FF61FF0CC0FEFD4FBB864972" box="[902,1006,761,787]" pageId="204" pageNumber="205" type="holotype">Holotype</typeStatus>
<specimenCount id="D196638EFF61FF0CC080FD4CBC784975" box="[1016,1040,762,788]" pageId="204" pageNumber="205" type="male">♂</specimenCount>
:
<collectingCountry id="BF87E897FF61FF0CC767FD4FBCC94972" box="[1055,1185,761,787]" name="Costa Rica" pageId="204" pageNumber="205">Costa Rica</collectingCountry>
:
<collectingRegion id="055466E5FF61FF0CC7D6FD4FBD5C4972" box="[1198,1332,761,787]" country="Costa Rica" name="Guanacaste" pageId="204" pageNumber="205">Guanacaste</collectingRegion>
:
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Área
<collectorName id="6A65CDD1FF61FF0CC6FAFD4FB9594956" pageId="204" pageNumber="205">de Conservación</collectorName>
</location>
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,
<collectorName id="6A65CDD1FF61FF0CC2BEFCABBA324956" box="[454,602,797,823]" pageId="204" pageNumber="205">Sector Pitilla</collectorName>
,
<collectorName id="6A65CDD1FF61FF0CC11DFCA8BA8B4959" box="[613,739,798,824]" pageId="204" pageNumber="205">Pasmompa</collectorName>
,
<geoCoordinate id="A2A4CEC0FF61FF0CC195FCABBB304956" box="[749,856,797,823]" degrees="11.01926" direction="north" orientation="latitude" pageId="204" pageNumber="205" precision="1" value="11.01926">11.01926</geoCoordinate>
,
<geoCoordinate id="A2A4CEC0FF61FF0CC01BFCABBBB04959" box="[867,984,797,824]" degrees="85.40997" direction="west" orientation="longitude" pageId="204" pageNumber="205" precision="1" value="-85.40997">-85.40997</geoCoordinate>
,
<quantity id="006805E2FF61FF0CC09BFCABBC434959" box="[995,1067,797,824]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="204" pageNumber="205" unit="m" value="440.0">
<elevation id="4CBD4F34FF61FF0CC09BFCABBC434959" box="[995,1067,797,824]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="204" pageNumber="205" unit="m" value="440.0">440 m</elevation>
</quantity>
, larva found
<date id="B32E8EC7FF61FF0CC7C5FCABBDF04956" box="[1213,1432,797,823]" pageId="204" pageNumber="205" value="2006-09-11">
<collectingDate id="A36A772FFF61FF0CC7C5FCABBDF04956" box="[1213,1432,797,823]" pageId="204" pageNumber="205" value="2006-09-11">11 September 2006</collectingDate>
</date>
, prepupa
<date id="B32E8EC7FF61FF0CC380FCF7B9B9493A" box="[248,465,833,859]" pageId="204" pageNumber="205" value="2006-11-12">
<collectingDate id="A36A772FFF61FF0CC380FCF7B9B9493A" box="[248,465,833,859]" pageId="204" pageNumber="205" value="2006-11-12">12 November 2006</collectingDate>
</date>
, adult eclosed
<date id="B32E8EC7FF61FF0CC10CFCF7BB54493A" box="[628,828,833,859]" pageId="204" pageNumber="205" value="2006-12-08">
<collectingDate id="A36A772FFF61FF0CC10CFCF7BB54493A" box="[628,828,833,859]" pageId="204" pageNumber="205" value="2006-12-08">8 December 2006</collectingDate>
</date>
, Manuel Ríos, 6-SRNP-34098 (
<collectionCode id="A18130C2FF61FF0CC7E5FCF7BC81493A" box="[1181,1257,833,859]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="204" pageNumber="205">INBio</collectionCode>
); for images see
<bibRefCitation id="A301D5F6FF61FF0CC3EFFCD3B9E9491E" author="Chacon, I. A. &amp; Montero-Ramirez, J. &amp; Janzen, D. H. &amp; Hallwachs, W. &amp; Blandin, P. &amp; Bristow, C. R. &amp; Hajibabaei, M." box="[151,385,869,896]" pageId="204" pageNumber="205" pagination="1 - 15" refId="ref175372" refString="Chacon, I. A., Montero-Ramirez, J., Janzen, D. H., Hallwachs, W., Blandin, P., Bristow, C. R. &amp; Hajibabaei, M. (2012) A new species of Opsiphanes Doubleday, [1849] from Costa Rica (Nymphalidae: Morphinae: Brassolini), as revealed by its DNA barcodes and habitus. Bulletin of the Allyn Museum, 166, 1 - 15." type="journal article" year="2012">
Chacón
<emphasis id="F5E47415FF61FF0CC38CFCD0B94D491E" box="[244,293,869,895]" italics="true" pageId="204" pageNumber="205">et al</emphasis>
. (2012)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8F8AFB8CFF61FF0CC3BFFC3FBA674C66" pageId="204" pageNumber="205" type="diagnosis">
<paragraph id="C72FA807FF61FF0CC3BFFC3FBBC54F32" blockId="204.[151,1437,725,2012]" pageId="204" pageNumber="205">
<emphasis id="F5E47415FF61FF0CC3BFFC3FB92A49C2" bold="true" box="[199,322,905,931]" pageId="204" pageNumber="205">Diagnosis.</emphasis>
<taxonomicName id="0090D384FF61FF0CC231FC3FBA3149C2" authorityName="Chacon &amp; Montero" authorityYear="2012" box="[329,601,905,931]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="jacobsorum">
<emphasis id="F5E47415FF61FF0CC231FC3FBA3149C2" box="[329,601,905,931]" italics="true" pageId="204" pageNumber="205">Opsiphanes jacobsorum</emphasis>
</taxonomicName>
is sympatric with
<taxonomicName id="0090D384FF61FF0CC053FC3FBC0B49C2" baseAuthorityName="Boisduval" baseAuthorityYear="1870" box="[811,1123,905,931]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="subSpecies" species="fabricii" subSpecies="fabricii">
<emphasis id="F5E47415FF61FF0CC053FC3FBC0B49C2" box="[811,1123,905,931]" italics="true" pageId="204" pageNumber="205">Opsiphanes fabricii fabricii</emphasis>
</taxonomicName>
and
<taxonomicName id="0090D384FF61FF0CC7E4FC3FBDF549C2" authorityName="Piovesan &amp; Casagrande &amp; Mielke" authorityYear="2022" baseAuthorityName="Fruhstorfer" baseAuthorityYear="1912" box="[1180,1437,905,931]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="periphetes" status="stat. nov.">
<emphasis id="F5E47415FF61FF0CC7E4FC3FBDF549C2" box="[1180,1437,905,931]" italics="true" pageId="204" pageNumber="205">Opsiphanes periphetes</emphasis>
</taxonomicName>
<emphasis id="F5E47415FF61FF0CC3EFFC1BB96B49A6" bold="true" box="[151,259,941,967]" pageId="204" pageNumber="205">
<taxonomicNameLabel id="EED7C96EFF61FF0CC3EFFC1BB96B49A6" box="[151,259,941,967]" pageId="204" pageNumber="205">stat. nov.</taxonomicNameLabel>
</emphasis>
; males and females of
<taxonomicName id="0090D384FF61FF0CC175FC1BBB7649A6" authorityName="Chacon &amp; Montero" authorityYear="2012" box="[525,798,941,967]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="jacobsorum">
<emphasis id="F5E47415FF61FF0CC175FC1BBB7649A6" box="[525,798,941,967]" italics="true" pageId="204" pageNumber="205">Opsiphanes jacobsorum</emphasis>
</taxonomicName>
differ from both these species in having the HW with a crenulated outer margin, although this crenulation is not as marked as in the “
<taxonomicName id="0090D384FF61FF0CC759FC67BC36498A" authorityName="Doubleday" authorityYear="1849" box="[1057,1118,977,1003]" class="Insecta" family="Nymphalidae" genus="Caligo" kingdom="Animalia" order="Lepidoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="sallei">
<emphasis id="F5E47415FF61FF0CC759FC67BC36498A" box="[1057,1118,977,1003]" italics="true" pageId="204" pageNumber="205">sallei</emphasis>
</taxonomicName>
” and “
<taxonomicName id="0090D384FF61FF0CC7CBFC64BD64498A" baseAuthorityName="Stoll" baseAuthorityYear="1780" box="[1203,1292,978,1003]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="quiteria">
<emphasis id="F5E47415FF61FF0CC7CBFC64BD64498A" box="[1203,1292,978,1003]" italics="true" pageId="204" pageNumber="205">quiteria</emphasis>
</taxonomicName>
” subgroups. Additionally, the male of
<taxonomicName id="0090D384FF61FF0CC2C4FC43BAA44E6E" authorityName="Chacon &amp; Montero" authorityYear="2012" box="[444,716,1013,1039]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="jacobsorum">
<emphasis id="F5E47415FF61FF0CC2C4FC43BAA44E6E" box="[444,716,1013,1039]" italics="true" pageId="204" pageNumber="205">Opsiphanes jacobsorum</emphasis>
</taxonomicName>
differs from males of
<taxonomicName id="0090D384FF61FF0CC0B7FC43BD604E6E" baseAuthorityName="Boisduval" baseAuthorityYear="1870" box="[975,1288,1013,1039]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="subSpecies" species="fabricii" subSpecies="fabricii">
<emphasis id="F5E47415FF61FF0CC0B7FC43BD604E6E" box="[975,1288,1013,1039]" italics="true" pageId="204" pageNumber="205">Opsiphanes fabricii fabricii</emphasis>
</taxonomicName>
and
<taxonomicName id="0090D384FF61FF0CC63AFC43B92F4E52" authorityName="Piovesan &amp; Casagrande &amp; Mielke" authorityYear="2022" baseAuthorityName="Fruhstorfer" baseAuthorityYear="1912" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="periphetes" status="stat. nov.">
<emphasis id="F5E47415FF61FF0CC63AFC43B92F4E52" italics="true" pageId="204" pageNumber="205">Opsiphanes periphetes</emphasis>
</taxonomicName>
<emphasis id="F5E47415FF61FF0CC236FBAFB9D14E52" bold="true" box="[334,441,1049,1075]" pageId="204" pageNumber="205">
<taxonomicNameLabel id="EED7C96EFF61FF0CC236FBAFB9D14E52" box="[334,441,1049,1075]" pageId="204" pageNumber="205">stat. nov.</taxonomicNameLabel>
</emphasis>
in having the DFW with an irregular oblique band, which borders the discal cell at most until the anterior half of M
<subScript id="5B14AA42FF61FF0CC2B3FBE6B9BB4E3F" attach="left" box="[459,467,1104,1118]" fontSize="6" pageId="204" pageNumber="205">3</subScript>
-CuA
<subScript id="5B14AA42FF61FF0CC169FBE6BA714E3F" attach="left" box="[529,537,1104,1118]" fontSize="6" pageId="204" pageNumber="205">1</subScript>
and DHW with a submarginal, generally zigzag band, evident up to CuA
<subScript id="5B14AA42FF61FF0CC61BFBE6BD034E3F" attach="left" box="[1379,1387,1104,1118]" fontSize="6" pageId="204" pageNumber="205">1</subScript>
and two-thirds the width of the oblique band of DFW, on CuA
<subScript id="5B14AA42FF61FF0CC04BFBC2BB534EE3" attach="left" box="[819,827,1140,1154]" fontSize="6" pageId="204" pageNumber="205">1</subScript>
-CuA
<subScript id="5B14AA42FF61FF0CC001FBC2BBE94EE3" attach="left" box="[889,897,1140,1154]" fontSize="6" pageId="204" pageNumber="205">2</subScript>
. The female of
<taxonomicName id="0090D384FF61FF0CC742FBD7BD234E1A" authorityName="Chacon &amp; Montero" authorityYear="2012" box="[1082,1355,1121,1147]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="jacobsorum">
<emphasis id="F5E47415FF61FF0CC742FBD7BD234E1A" box="[1082,1355,1121,1147]" italics="true" pageId="204" pageNumber="205">Opsiphanes jacobsorum</emphasis>
</taxonomicName>
differs from the females of
<taxonomicName id="0090D384FF61FF0CC2F9FB33BAD44EFE" baseAuthorityName="Boisduval" baseAuthorityYear="1870" box="[385,700,1157,1183]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="subSpecies" species="fabricii" subSpecies="fabricii">
<emphasis id="F5E47415FF61FF0CC2F9FB33BAD44EFE" box="[385,700,1157,1183]" italics="true" pageId="204" pageNumber="205">Opsiphanes fabricii fabricii</emphasis>
</taxonomicName>
and
<taxonomicName id="0090D384FF61FF0CC18EFB33BB904EFE" authorityName="Piovesan &amp; Casagrande &amp; Mielke" authorityYear="2022" baseAuthorityName="Fruhstorfer" baseAuthorityYear="1912" box="[758,1016,1157,1183]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="periphetes" status="stat. nov.">
<emphasis id="F5E47415FF61FF0CC18EFB33BB904EFE" box="[758,1016,1157,1183]" italics="true" pageId="204" pageNumber="205">Opsiphanes periphetes</emphasis>
</taxonomicName>
<emphasis id="F5E47415FF61FF0CC779FB33BC054EFE" bold="true" box="[1025,1133,1157,1183]" pageId="204" pageNumber="205">
<taxonomicNameLabel id="EED7C96EFF61FF0CC779FB33BC054EFE" box="[1025,1133,1157,1183]" pageId="204" pageNumber="205">stat. nov.</taxonomicNameLabel>
</emphasis>
in having the DFW bands varying from whitish to yellowish and the DHW submarginal band entirely rusty red (
<figureCitation id="5FABB482FF61FF0CC72CFB1FBCAC4EA2" box="[1108,1220,1193,1219]" captionStart="FIGURE 187" captionStartId="205.[151,250,1923,1947]" captionTargetBox="[187,1379,823,1879]" captionTargetId="figure-271@205.[175,1412,797,1899]" captionTargetPageId="205" captionText="FIGURE 187. Opsiphanes jacobsorum Chacón &amp; Montero, 2012: Male, dorsal and ventral views: Costa Rica: a. (USN- MENT01590011-USNM) Guanacaste, b. (USNMENT01590012-USNM) Alajuela; Panama: c. (USNMENT01590040-USNM) Panamá, d. (FLMNH MGCL 1107067-MGCL) Piña. White arrows indicate diagnostic characters of the species." figureDoi="http://doi.org/10.5281/zenodo.7403985" httpUri="https://zenodo.org/record/7403985/files/figure.png" pageId="204" pageNumber="205">Figs 187a</figureCitation>
–
<figureCitation id="5FABB482FF61FF0CC7ACFB1FBD644EA2" box="[1236,1292,1193,1219]" captionStart="FIGURE 188" captionStartId="206.[151,250,1025,1049]" captionTargetBox="[554,1027,183,987]" captionTargetId="figure-28@206.[338,1249,166,1001]" captionTargetPageId="206" captionText="FIGURE 188. Opsiphanes jacobsorum Chacón &amp; Montero, 2012: Female, dorsal and ventral views: Costa Rica: a. (USN- MENT01590014-USNM), b. (USNMENT01590784-USNM) Guanacaste." figureDoi="http://doi.org/10.5281/zenodo.7403987" httpUri="https://zenodo.org/record/7403987/files/figure.png" pageId="204" pageNumber="205">188a</figureCitation>
). The female genitalia of
<taxonomicName id="0090D384FF61FF0CC266FB7BBA454E86" authorityName="Chacon &amp; Montero" authorityYear="2012" box="[286,557,1229,1255]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="jacobsorum">
<emphasis id="F5E47415FF61FF0CC266FB7BBA454E86" box="[286,557,1229,1255]" italics="true" pageId="204" pageNumber="205">Opsiphanes jacobsorum</emphasis>
</taxonomicName>
differ from other known
<taxonomicName id="0090D384FF61FF0CC034FB7BBC444E86" box="[844,1068,1229,1255]" pageId="204" pageNumber="205">
<emphasis id="F5E47415FF61FF0CC034FB7BBBB84E86" box="[844,976,1229,1255]" italics="true" pageId="204" pageNumber="205">Opsiphanes</emphasis>
species
</taxonomicName>
, in that the lamella antevaginalis is not connected to the lamella postvaginalis (
<figureCitation id="5FABB482FF61FF0CC1E5FB47BB684F6A" box="[669,768,1265,1291]" captionStart="FIGURE 190" captionStartId="207.[151,250,1107,1131]" captionTargetBox="[287,1320,269,1065]" captionTargetId="figure-19@207.[250,1336,166,1083]" captionTargetPageId="207" captionText="FIGURE 190. Opsiphanes jacobsorum Chacón &amp; Montero, 2012: Female genitalia: a. lateral view; b. detail of the signa; c. ventral view (USNMENT01590014-USNM)." figureDoi="http://doi.org/10.5281/zenodo.7403991" httpUri="https://zenodo.org/record/7403991/files/figure.png" pageId="204" pageNumber="205">Fig. 190</figureCitation>
). Molecular analyses recovered
<taxonomicName id="0090D384FF61FF0CC715FB47BD154F6A" authorityName="Chacon &amp; Montero" authorityYear="2012" box="[1133,1405,1265,1291]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="jacobsorum">
<emphasis id="F5E47415FF61FF0CC715FB47BD154F6A" box="[1133,1405,1265,1291]" italics="true" pageId="204" pageNumber="205">Opsiphanes jacobsorum</emphasis>
</taxonomicName>
as closely related to
<taxonomicName id="0090D384FF61FF0CC227FAA3BA544F4E" authorityName=", Boisduval" authorityYear="1870" box="[351,572,1301,1327]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="fabricii">
<emphasis id="F5E47415FF61FF0CC227FAA3BA544F4E" box="[351,572,1301,1327]" italics="true" pageId="204" pageNumber="205">Opsiphanes fabricii</emphasis>
</taxonomicName>
,
<taxonomicName id="0090D384FF61FF0CC133FAA3BB294F4E" authorityName="Stichel" authorityYear="1902" box="[587,833,1301,1327]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="merianae">
<emphasis id="F5E47415FF61FF0CC133FAA3BB294F4E" box="[587,833,1301,1327]" italics="true" pageId="204" pageNumber="205">Opsiphanes merianae</emphasis>
</taxonomicName>
<emphasis id="F5E47415FF61FF0CC030FAA3BBDD4F4E" bold="true" box="[840,949,1301,1327]" pageId="204" pageNumber="205">stat. rest.</emphasis>
and
<taxonomicName id="0090D384FF61FF0CC095FAA3BC854F4E" authorityName="Piovesan &amp; Casagrande &amp; Mielke" authorityYear="2022" baseAuthorityName="Fruhstorfer" baseAuthorityYear="1912" box="[1005,1261,1301,1327]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="periphetes" status="stat. nov.">
<emphasis id="F5E47415FF61FF0CC095FAA3BC854F4E" box="[1005,1261,1301,1327]" italics="true" pageId="204" pageNumber="205">Opsiphanes periphetes</emphasis>
</taxonomicName>
<emphasis id="F5E47415FF61FF0CC78CFAA3BD374F4E" bold="true" box="[1268,1375,1301,1327]" pageId="204" pageNumber="205">
<taxonomicNameLabel id="EED7C96EFF61FF0CC78CFAA3BD374F4E" box="[1268,1375,1301,1327]" pageId="204" pageNumber="205">stat. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="5FABB482FF61FF0CC617FAA3B8CE4F32" captionStart="FIGURE 2" captionStartId="7.[151,250,1853,1877]" captionTargetBox="[304,1282,181,1828]" captionTargetId="figure-19@7.[298,1290,181,1829]" captionTargetPageId="7" captionText="FIGURE 2. Phylogenetic tree (Maximum Likelihood) inferred with IQ-TREE based on six molecular markers: cytochrome c oxidase subunit I (COI) (mitochondrial), elongation factor 1-alpha (EF-1α), ribosomal protein S5 (RpS5), wingless (wg), Car- bamoylphosphate synthase domain protein (CAD), Glyceraldehyde-3-phosphate dehydrogenase (GAPDH) (nuclear), rooted in Catoblepia amphirhoe (Hübner, [1825]). Asterisk (*) indicates SH-aLRT&gt; = 80% and UFboot&gt; = 95%. The scale bar represents the expected number of nucleotide substitutions per site. Data of the specimens used are described in table 2." figureDoi="http://doi.org/10.5281/zenodo.7403536" httpUri="https://zenodo.org/record/7403536/files/figure.png" pageId="204" pageNumber="205">Fig. 2</figureCitation>
) (genetic distance 0.9–1.2%, 1.2–1.6%, and 1.4–1.7%, respectively).
</paragraph>
<paragraph id="C72FA807FF61FF0CC3BFFAEBBCF94FFA" blockId="204.[151,1437,725,2012]" pageId="204" pageNumber="205">
<emphasis id="F5E47415FF61FF0CC3BFFAEBB9114F16" bold="true" box="[199,377,1373,1399]" pageId="204" pageNumber="205">Male genitalia.</emphasis>
Tegumen in dorsal view with anterior margin anteriorly projected, shorter than the uncus; anterior projection of saccus in lateral view twice as long as dorsal arms of saccus (
<figureCitation id="5FABB482FF61FF0CC75AFA37BCEC4FFA" box="[1058,1156,1409,1435]" captionStart="FIGURE 189" captionStartId="206.[151,250,1965,1989]" captionTargetBox="[353,1235,1117,1940]" captionTargetId="figure-53@206.[353,1235,1094,1940]" captionTargetPageId="206" captionText="FIGURE 189. Opsiphanes jacobsorum Chacón &amp; Montero, 2012: Male: a. abdomen, lateral view; genitalia: b. dorsal view, c. ventral view, d. lateral view (USNMENT01590012-USNM)." figureDoi="http://doi.org/10.5281/zenodo.7403989" httpUri="https://zenodo.org/record/7403989/files/figure.png" pageId="204" pageNumber="205">Fig. 189</figureCitation>
).
</paragraph>
<paragraph id="C72FA807FF61FF0CC3BFFA13BA674C66" blockId="204.[151,1437,725,2012]" pageId="204" pageNumber="205">
<emphasis id="F5E47415FF61FF0CC3BFFA13B9FB4FDE" bold="true" box="[199,403,1445,1471]" pageId="204" pageNumber="205">Female genitalia.</emphasis>
Lamella antevaginalis as a narrow, sclerotized arch; lamella postvaginalis rounded medially, tapering in the distal region, and not fused to the lamella antevaginalis; ostium membranous; corpus bursae shorter than the ductus bursae (
<figureCitation id="5FABB482FF61FF0CC2D8FA5BBA6A4C66" box="[416,514,1517,1543]" captionStart="FIGURE 190" captionStartId="207.[151,250,1107,1131]" captionTargetBox="[287,1320,269,1065]" captionTargetId="figure-19@207.[250,1336,166,1083]" captionTargetPageId="207" captionText="FIGURE 190. Opsiphanes jacobsorum Chacón &amp; Montero, 2012: Female genitalia: a. lateral view; b. detail of the signa; c. ventral view (USNMENT01590014-USNM)." figureDoi="http://doi.org/10.5281/zenodo.7403991" httpUri="https://zenodo.org/record/7403991/files/figure.png" pageId="204" pageNumber="205">Fig. 190</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8F8AFB8CFF61FF0DC3BFF9A7BABE4AD1" lastPageId="205" lastPageNumber="206" pageId="204" pageNumber="205" type="description">
<paragraph id="C72FA807FF61FF0CC3BFF9A7BCDC4D62" blockId="204.[151,1437,725,2012]" pageId="204" pageNumber="205">
<emphasis id="F5E47415FF61FF0CC3BFF9A7B9574C4A" bold="true" box="[199,319,1553,1579]" pageId="204" pageNumber="205">Variation.</emphasis>
FW, length: male:
<quantity id="006805E2FF61FF0CC161F9A7BAFC4C4D" box="[537,660,1553,1580]" metricMagnitude="-2" metricUnit="m" metricValue="3.95" metricValueMax="4.2" metricValueMin="3.7" pageId="204" pageNumber="205" unit="mm" value="39.5" valueMax="42.0" valueMin="37.0">37–42 mm</quantity>
(n=5), female:
<quantity id="006805E2FF61FF0CC03DF9A7BBD74C4D" box="[837,959,1553,1580]" metricMagnitude="-2" metricUnit="m" metricValue="4.45" metricValueMax="4.5" metricValueMin="4.4" pageId="204" pageNumber="205" unit="mm" value="44.5" valueMax="45.0" valueMin="44.0">44–45 mm</quantity>
(n=3). DFW, the width of the band inside the discal cell, and the anterior and posterior margins of this band may have varied indentations; in females, this band ranges from defined to nearly imperceptible; the width and length of the oblique band; in females, the color of the bands ranges from whitish to yellowish, which may have different shades in the same specimen. DHW, the width and color of the submarginal band. VFW, the width, length, and color of the oblique band. VHW, the size and shape of the eyespot between Sc+R
<subScript id="5B14AA42FF61FF0CC15AF96EBA424C87" attach="left" box="[546,554,1752,1766]" fontSize="6" pageId="204" pageNumber="205">1</subScript>
and M
<subScript id="5B14AA42FF61FF0CC101F96EBAE94C87" attach="left" box="[633,641,1752,1766]" fontSize="6" pageId="204" pageNumber="205">1</subScript>
, and the width of its dark-brown ring; the size and shape of the eyespot between M
<subScript id="5B14AA42FF61FF0CC26CF94AB9744D6B" attach="left" box="[276,284,1788,1802]" fontSize="6" pageId="204" pageNumber="205">3</subScript>
and CuA
<subScript id="5B14AA42FF61FF0CC2F0F94AB9F84D6B" attach="left" box="[392,400,1788,1802]" fontSize="6" pageId="204" pageNumber="205">2</subScript>
; the zigzag EIII submarginal line is not always evident (
<figureCitation id="5FABB482FF61FF0CC771F95FBCC04D62" box="[1033,1192,1769,1795]" captionStart-0="FIGURE 187" captionStart-1="FIGURE 188" captionStartId-0="205.[151,250,1923,1947]" captionStartId-1="206.[151,250,1025,1049]" captionTargetBox-0="[187,1379,823,1879]" captionTargetBox-1="[554,1027,183,987]" captionTargetId-0="figure-271@205.[175,1412,797,1899]" captionTargetId-1="figure-28@206.[338,1249,166,1001]" captionTargetPageId-0="205" captionTargetPageId-1="206" captionText-0="FIGURE 187. Opsiphanes jacobsorum Chacón &amp; Montero, 2012: Male, dorsal and ventral views: Costa Rica: a. (USN- MENT01590011-USNM) Guanacaste, b. (USNMENT01590012-USNM) Alajuela; Panama: c. (USNMENT01590040-USNM) Panamá, d. (FLMNH MGCL 1107067-MGCL) Piña. White arrows indicate diagnostic characters of the species." captionText-1="FIGURE 188. Opsiphanes jacobsorum Chacón &amp; Montero, 2012: Female, dorsal and ventral views: Costa Rica: a. (USN- MENT01590014-USNM), b. (USNMENT01590784-USNM) Guanacaste." figureDoi-0="http://doi.org/10.5281/zenodo.7403985" figureDoi-1="http://doi.org/10.5281/zenodo.7403987" httpUri-0="https://zenodo.org/record/7403985/files/figure.png" httpUri-1="https://zenodo.org/record/7403987/files/figure.png" pageId="204" pageNumber="205">Figs 187–188</figureCitation>
).
</paragraph>
<paragraph id="C72FA807FF61FF0CC3BFF8BBB88C4D2D" blockId="204.[151,1437,725,2012]" pageId="204" pageNumber="205">
<emphasis id="F5E47415FF61FF0CC3BFF8BBB9ED4D46" bold="true" box="[199,389,1805,1831]" pageId="204" pageNumber="205">Natural history.</emphasis>
Chacón &amp; Montero (2012) described the last larval instar and the pupa of
<taxonomicName id="0090D384FF61FF0CC7B1F8BBB8B54D2A" authorityName="Chacon &amp; Montero" authorityYear="2012" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="jacobsorum">
<emphasis id="F5E47415FF61FF0CC7B1F8BBB8B54D2A" italics="true" pageId="204" pageNumber="205">Opsiphanes jacobsorum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C72FA807FF61FF0CC3BFF8E0BC5F4DF2" blockId="204.[151,1437,725,2012]" pageId="204" pageNumber="205">
Host plants:
<taxonomicName id="0090D384FF61FF0CC22DF8E0B9A64D11" box="[341,462,1878,1904]" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" pageId="204" pageNumber="205" phylum="Tracheophyta" rank="family">Arecaceae</taxonomicName>
:
<taxonomicName id="0090D384FF61FF0CC2A1F8E3BC604D0E" authority="(H. A. Wendl.) Bartlett" authorityName="Bartlett" baseAuthorityName="H. A. Wendl." box="[473,1032,1877,1904]" class="Liliopsida" family="Arecaceae" genus="Cryosophila" kingdom="Plantae" order="Arecales" pageId="204" pageNumber="205" phylum="Tracheophyta" rank="species" species="warscewiczii">
<emphasis id="F5E47415FF61FF0CC2A1F8E3BA944D0E" box="[473,764,1877,1903]" italics="true" pageId="204" pageNumber="205">Cryosophila warscewiczii</emphasis>
(H. A. Wendl.) Bartlett
</taxonomicName>
,
<taxonomicName id="0090D384FF61FF0CC76CF8E0BDF04D0E" authority="H. F. Loomis" authorityName="H. F. Loomis" box="[1044,1432,1877,1904]" class="Liliopsida" family="Arecaceae" genus="Astrocaryum" kingdom="Plantae" order="Arecales" pageId="204" pageNumber="205" phylum="Tracheophyta" rank="species" species="alatum">
<emphasis id="F5E47415FF61FF0CC76CF8E0BC914D0E" box="[1044,1273,1877,1904]" italics="true" pageId="204" pageNumber="205">Astrocaryum alatum</emphasis>
H. F. Loomis
</taxonomicName>
,
<taxonomicName id="0090D384FF61FF0CC3EFF8CCBC5B4DF2" authority="(H. Wendl. ex Burret) H. E. Moore (Chacon &amp; Montero 2012)" authorityName="H. E. Moore (Chacon &amp; Montero" authorityYear="2012" baseAuthorityName="H. Wendl. ex Burret" box="[151,1075,1913,1940]" class="Liliopsida" family="Arecaceae" genus="Prestoea" kingdom="Plantae" order="Arecales" pageId="204" pageNumber="205" phylum="Tracheophyta" rank="species" species="decurrens">
<emphasis id="F5E47415FF61FF0CC3EFF8CCB9184DF2" box="[151,368,1913,1939]" italics="true" pageId="204" pageNumber="205">Prestoea decurrens</emphasis>
(H. Wendl. ex Burret) H. E. Moore (Chacón &amp; Montero 2012)
</taxonomicName>
.
</paragraph>
<paragraph id="C72FA807FF61FF0CC3BFF82BBB364DBA" blockId="204.[151,1437,725,2012]" pageId="204" pageNumber="205">
<emphasis id="F5E47415FF61FF0CC3BFF82BB9874DD6" bold="true" box="[199,495,1949,1975]" pageId="204" pageNumber="205">Geographic distribution.</emphasis>
<materialsCitation id="77F8A25AFF61FF0CC28EF82BBBEB4DD6" box="[502,899,1949,1975]" country="Costa Rica" location="Guanacaste" pageId="204" pageNumber="205" specimenCount="1" stateProvince="Guanacaste">
<emphasis id="F5E47415FF61FF0CC28EF82BBA104DD6" bold="true" box="[502,632,1949,1975]" pageId="204" pageNumber="205">
<collectingCountry id="BF87E897FF61FF0CC28EF82BBA104DD6" box="[502,632,1949,1975]" name="Costa Rica" pageId="204" pageNumber="205">Costa Rica</collectingCountry>
</emphasis>
(
<collectingRegion id="055466E5FF61FF0CC1FFF82BBB654DD6" box="[647,781,1949,1975]" country="Costa Rica" name="Guanacaste" pageId="204" pageNumber="205">Guanacaste</collectingRegion>
,
<collectingRegion id="055466E5FF61FF0CC06FF82BBB134DD6" box="[791,891,1949,1975]" country="Costa Rica" name="Alajuela" pageId="204" pageNumber="205">Alajuela</collectingRegion>
)
</materialsCitation>
;
<materialsCitation id="77F8A25AFF61FF0CC0F6F82BBB324DBA" collectorName="Chacon &amp; Montero" country="Panama" elevation="640" location="It" pageId="204" pageNumber="205" specimenCount="1" stateProvince="Panama">
<emphasis id="F5E47415FF61FF0CC0F6F82BBB984DD6" bold="true" box="[910,1008,1949,1975]" pageId="204" pageNumber="205">
<collectingCountry id="BF87E897FF61FF0CC0F6F82BBB984DD6" box="[910,1008,1949,1975]" name="Panama" pageId="204" pageNumber="205">Panama</collectingCountry>
</emphasis>
(
<collectingRegion id="055466E5FF61FF0CC087F82BBC344DD6" box="[1023,1116,1949,1975]" country="Panama" name="Panama" pageId="204" pageNumber="205">Panamá</collectingRegion>
,
<collectingRegion id="055466E5FF61FF0CC71FF82BBCC54DD6" box="[1127,1197,1949,1975]" country="Panama" name="Colon" pageId="204" pageNumber="205">Colón</collectingRegion>
) (
<figureCitation id="5FABB482FF61FF0CC7BDF82BBD4E4DD6" box="[1221,1318,1949,1975]" captionStart="FIGURE 191" captionStartId="207.[151,250,1987,2011]" captionTargetBox="[229,1359,1176,1963]" captionTargetId="figure-44@207.[229,1359,1176,1963]" captionTargetPageId="207" captionText="FIGURE 191. Geographical distribution of Opsiphanes jacobsorum Chacón &amp; Montero, 2012." figureDoi="http://doi.org/10.5281/zenodo.7403993" httpUri="https://zenodo.org/record/7403993/files/figure.png" pageId="204" pageNumber="205">Fig. 191</figureCitation>
).
<location id="C24FFEDCFF61FF0CC643F828BD244DD9" LSID="urn:lsid:plazi:treatment:4F391911FF61FF0DC3EFFED8BB5D4975:C24FFEDCFF61FF0CC643F828BD244DD9" box="[1339,1356,1950,1976]" country="Panama" name="It" pageId="204" pageNumber="205" stateProvince="Panama">It</location>
occurs at altitudes between 380 and
<quantity id="006805E2FF61FF0CC2A5F877BA4C4DBD" box="[477,548,1985,2012]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="204" pageNumber="205" unit="m" value="640.0">
<elevation id="4CBD4F34FF61FF0CC2A5F877BA4C4DBD" box="[477,548,1985,2012]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="204" pageNumber="205" unit="m" value="640.0">640 m</elevation>
</quantity>
(
<collectorName id="6A65CDD1FF61FF0CC14AF877BAE14DBA" box="[562,649,1985,2011]" pageId="204" pageNumber="205">Chacón</collectorName>
&amp;
<collectorName id="6A65CDD1FF61FF0CC1D5F874BB654DBD" box="[685,781,1986,2012]" pageId="204" pageNumber="205">Montero</collectorName>
2012)
</materialsCitation>
.
</paragraph>
<paragraph id="C72FA807FF60FF0DC3BFFF20BABE4AD1" blockId="205.[151,1436,150,789]" box="[199,726,150,176]" pageId="205" pageNumber="206">
<emphasis id="F5E47415FF60FF0DC3BFFF20B9BF4AD1" bold="true" box="[199,471,150,176]" pageId="205" pageNumber="206">Temporal distribution.</emphasis>
March, May–October.
</paragraph>
</subSubSection>
<subSubSection id="8F8AFB8CFF60FF0DC3BFFF0CBB5D4975" pageId="205" pageNumber="206" type="etymology">
<paragraph id="C72FA807FF60FF0DC3BFFF0CB9F74A99" blockId="205.[151,1436,150,789]" pageId="205" pageNumber="206">
<emphasis id="F5E47415FF60FF0DC3BFFF0CB9264AB5" bold="true" box="[199,334,186,212]" pageId="205" pageNumber="206">Etymology.</emphasis>
Named in honor of Irwin and Joan Jacobs, who contributed funds to maintain the “Área de Conservación
<collectingRegion id="055466E5FF60FF0DC271FF68B9F84A99" box="[265,400,222,248]" country="Costa Rica" name="Guanacaste" pageId="205" pageNumber="206">Guanacaste</collectingRegion>
”.
</paragraph>
<paragraph id="C72FA807FF60FF0DC3BFFEB4BB5D4975" blockId="205.[151,1436,150,789]" pageId="205" pageNumber="206">
<emphasis id="F5E47415FF60FF0DC3BFFEB4B9AF4B7D" bold="true" box="[199,455,258,284]" pageId="205" pageNumber="206">Specimens examined.</emphasis>
<materialsCitation id="77F8A25AFF60FF0DC2B6FEB4BDFF4B21" collectingDate="2012-09-01" collectionCode="USNM" collectorName="R. Calero &amp; Sendero Cuestona" country="Costa Rica" elevation="415" latitude="11.01525" location="Coneja" longLatPrecision="1" longitude="-85.39766" municipality="Area de Conservacion Guanacaste" pageId="205" pageNumber="206" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste">
<emphasis id="F5E47415FF60FF0DC2B6FEB4BA134B7D" bold="true" box="[462,635,258,284]" pageId="205" pageNumber="206">
<collectingCountry id="BF87E897FF60FF0DC2B6FEB4BA134B7D" box="[462,635,258,284]" name="Costa Rica" pageId="205" pageNumber="206">COSTA RICA</collectingCountry>
</emphasis>
:
<emphasis id="F5E47415FF60FF0DC1F1FEB4BB7D4B7D" bold="true" box="[649,789,258,284]" pageId="205" pageNumber="206">
<collectingRegion id="055466E5FF60FF0DC1F1FEB4BB7D4B7D" box="[649,789,258,284]" country="Costa Rica" name="Guanacaste" pageId="205" pageNumber="206">Guanacaste</collectingRegion>
</emphasis>
—
<collectingMunicipality id="274B327DFF60FF0DC04AFEB2BCDC4B7D" box="[818,1204,258,287]" pageId="205" pageNumber="206">Área de Conservación Guanacaste</collectingMunicipality>
(
<emphasis id="F5E47415FF60FF0DC7BDFEB5BD7E4B7C" box="[1221,1302,259,285]" italics="true" pageId="205" pageNumber="206">
<location id="C24FFEDCFF60FF0DC7BDFEB5BD7E4B7C" LSID="urn:lsid:plazi:treatment:4F391911FF61FF0DC3EFFED8BB5D4975:C24FFEDCFF60FF0DC7BDFEB5BD7E4B7C" box="[1221,1302,259,285]" country="Costa Rica" latitude="11.01525" longLatPrecision="1" longitude="-85.39766" municipality="Area de Conservacion Guanacaste" name="Coneja" pageId="205" pageNumber="206" stateProvince="Guanacaste">Coneja</location>
</emphasis>
),
<geoCoordinate id="A2A4CEC0FF60FF0DC655FEB4BDF04B7C" box="[1325,1432,258,285]" degrees="11.01525" direction="north" orientation="latitude" pageId="205" pageNumber="206" precision="1" value="11.01525">11.01525</geoCoordinate>
,
<geoCoordinate id="A2A4CEC0FF60FF0DC3EFFE90B9654B20" box="[151,269,294,321]" degrees="85.39766" direction="west" orientation="longitude" pageId="205" pageNumber="206" precision="1" value="-85.39766">-85.39766</geoCoordinate>
,
<quantity id="006805E2FF60FF0DC266FE90B9064B20" box="[286,366,294,321]" metricMagnitude="2" metricUnit="m" metricValue="4.15" pageId="205" pageNumber="206" unit="m" value="415.0">
<elevation id="4CBD4F34FF60FF0DC266FE90B9064B20" box="[286,366,294,321]" metricMagnitude="2" metricUnit="m" metricValue="4.15" pageId="205" pageNumber="206" unit="m" value="415.0">415 m</elevation>
</quantity>
,
<date id="B32E8EC7FF60FF0DC2F9FE90B9924B21" box="[385,506,294,320]" pageId="205" pageNumber="206" value="2012-09-01">
<collectingDate id="A36A772FFF60FF0DC2F9FE90B9924B21" box="[385,506,294,320]" pageId="205" pageNumber="206" value="2012-09-01">1-IX-2012</collectingDate>
</date>
,
<specimenCount id="D196638EFF60FF0DC174FE90BA094B21" box="[524,609,294,320]" pageId="205" pageNumber="206" type="male">1 male</specimenCount>
,
<collectorName id="6A65CDD1FF60FF0DC10CFE91BA8E4B21" box="[628,742,294,321]" pageId="205" pageNumber="206">R. Calero</collectorName>
leg., USNMENT01589074 (
<collectionCode id="A18130C2FF60FF0DC747FE90BCFE4B21" box="[1087,1174,294,320]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="205" pageNumber="206" type="Museum">USNM</collectionCode>
), (
<emphasis id="F5E47415FF60FF0DC7C1FE90BDE44B20" box="[1209,1420,294,321]" italics="true" pageId="205" pageNumber="206">
<collectorName id="6A65CDD1FF60FF0DC7C1FE90BDE44B20" box="[1209,1420,294,321]" pageId="205" pageNumber="206">Sendero Cuestona</collectorName>
</emphasis>
)
</materialsCitation>
,
<materialsCitation id="77F8A25AFF60FF0DC3EFFEFCBDFF4B05" box="[151,1431,330,357]" collectingDate="2006-06-25" collectionCode="USNM" collectorName="C. Moraga &amp; Medrano" country="Costa Rica" elevation="640" latitude="10.99455" longLatPrecision="1" longitude="-85.41461" pageId="205" pageNumber="206" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste">
<geoCoordinate id="A2A4CEC0FF60FF0DC3EFFEFCB96B4B04" box="[151,259,330,357]" degrees="10.99455" direction="north" orientation="latitude" pageId="205" pageNumber="206" precision="1" value="10.99455">10.99455</geoCoordinate>
,
<geoCoordinate id="A2A4CEC0FF60FF0DC26AFEFCB9EF4B04" box="[274,391,330,357]" degrees="85.41461" direction="west" orientation="longitude" pageId="205" pageNumber="206" precision="1" value="-85.41461">-85.41461</geoCoordinate>
,
<quantity id="006805E2FF60FF0DC2EDFEFCB98A4B04" box="[405,482,330,357]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="205" pageNumber="206" unit="m" value="640.0">
<elevation id="4CBD4F34FF60FF0DC2EDFEFCB98A4B04" box="[405,482,330,357]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="205" pageNumber="206" unit="m" value="640.0">640 m</elevation>
</quantity>
,
<date id="B32E8EC7FF60FF0DC289FEFCBA1F4B04" box="[497,631,330,357]" pageId="205" pageNumber="206" value="2006-06-25">
<collectingDate id="A36A772FFF60FF0DC289FEFCBA1F4B04" box="[497,631,330,357]" pageId="205" pageNumber="206" value="2006-06-25">25-VI-2006</collectingDate>
</date>
,
<specimenCount id="D196638EFF60FF0DC1FEFEFCBAB04B05" box="[646,728,330,356]" pageId="205" pageNumber="206" type="male">1 male</specimenCount>
,
<collectorName id="6A65CDD1FF60FF0DC19EFEFCBB094B04" box="[742,865,330,357]" pageId="205" pageNumber="206">C. Moraga</collectorName>
leg., USNMENT01590011 (
<collectionCode id="A18130C2FF60FF0DC7D7FEFCBD6E4B05" box="[1199,1286,330,356]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="205" pageNumber="206" type="Museum">USNM</collectionCode>
), (
<emphasis id="F5E47415FF60FF0DC65DFEFCBDE44B05" box="[1317,1420,330,356]" italics="true" pageId="205" pageNumber="206">
<collectorName id="6A65CDD1FF60FF0DC65DFEFCBDE44B05" box="[1317,1420,330,356]" pageId="205" pageNumber="206">Medrano</collectorName>
</emphasis>
)
</materialsCitation>
,
<materialsCitation id="77F8A25AFF60FF0DC3EFFED8BDFF4BE9" box="[151,1431,366,393]" collectingDate="2006-06-25" collectionCode="USNM" collectorName="R. Calero &amp; Quebradona" country="Costa Rica" elevation="640" latitude="11.01602" longLatPrecision="1" longitude="-85.38053" pageId="205" pageNumber="206" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste">
<geoCoordinate id="A2A4CEC0FF60FF0DC3EFFED8B96A4BE9" box="[151,258,366,392]" degrees="11.01602" direction="north" orientation="latitude" pageId="205" pageNumber="206" precision="1" value="11.01602">11.01602</geoCoordinate>
,
<geoCoordinate id="A2A4CEC0FF60FF0DC274FED8B9EA4BE8" box="[268,386,366,393]" degrees="85.38053" direction="west" orientation="longitude" pageId="205" pageNumber="206" precision="1" value="-85.38053">-85.38053</geoCoordinate>
,
<quantity id="006805E2FF60FF0DC2F4FED8B9BC4BE8" box="[396,468,366,393]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="205" pageNumber="206" unit="m" value="640.0">
<elevation id="4CBD4F34FF60FF0DC2F4FED8B9BC4BE8" box="[396,468,366,393]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="205" pageNumber="206" unit="m" value="640.0">640 m</elevation>
</quantity>
,
<date id="B32E8EC7FF60FF0DC298FED8BA0E4BE8" box="[480,614,366,393]" pageId="205" pageNumber="206" value="2006-06-25">
<collectingDate id="A36A772FFF60FF0DC298FED8BA0E4BE8" box="[480,614,366,393]" pageId="205" pageNumber="206" value="2006-06-25">25-VI-2006</collectingDate>
</date>
,
<specimenCount id="D196638EFF60FF0DC109FED8BABC4BE9" box="[625,724,366,392]" pageId="205" pageNumber="206" type="female">1 female</specimenCount>
,
<collectorName id="6A65CDD1FF60FF0DC1A7FED9BB224BE9" box="[735,842,366,393]" pageId="205" pageNumber="206">R. Calero</collectorName>
leg., USNMENT01590013 (
<collectionCode id="A18130C2FF60FF0DC7F4FED8BC8B4BE9" box="[1164,1251,366,392]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="205" pageNumber="206" type="Museum">USNM</collectionCode>
), (
<emphasis id="F5E47415FF60FF0DC786FED8BDE44BE9" box="[1278,1420,366,392]" italics="true" pageId="205" pageNumber="206">
<collectorName id="6A65CDD1FF60FF0DC786FED8BDE44BE9" box="[1278,1420,366,392]" pageId="205" pageNumber="206">Quebradona</collectorName>
</emphasis>
)
</materialsCitation>
,
<materialsCitation id="77F8A25AFF60FF0DC3EFFE24BDFF4BCD" box="[151,1431,402,429]" collectingDate="2010-05-10" collectionCode="USNM" collectorName="R. Calero &amp; Pasmompa" country="Costa Rica" elevation="475" pageId="205" pageNumber="206" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste">
1099102, -85.39539,
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<elevation id="4CBD4F34FF60FF0DC2F4FE24B9BF4BCC" box="[396,471,402,429]" metricMagnitude="2" metricUnit="m" metricValue="4.75" pageId="205" pageNumber="206" unit="m" value="475.0">475 m</elevation>
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,
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<collectingDate id="A36A772FFF60FF0DC29DFE24BA084BCD" box="[485,608,402,428]" pageId="205" pageNumber="206" value="2010-05-10">10-V-2010</collectingDate>
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leg., USNMENT01590014 (
<collectionCode id="A18130C2FF60FF0DC7E2FE24BC994BCD" box="[1178,1265,402,428]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="205" pageNumber="206" type="Museum">USNM</collectionCode>
), (
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<collectorName id="6A65CDD1FF60FF0DC677FE25BDE34BCD" box="[1295,1419,403,428]" pageId="205" pageNumber="206">Pasmompa</collectorName>
</emphasis>
)
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,
<materialsCitation id="77F8A25AFF60FF0DC3EFFE00BCE94BB1" box="[151,1153,438,465]" collectingDate="2017-03-03" collectionCode="USNM" collectorName="M. Rios" country="Costa Rica" elevation="440" pageId="205" pageNumber="206" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste">
11.0193, -85.41,
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<elevation id="4CBD4F34FF60FF0DC22DFE00B9F64BB0" box="[341,414,438,465]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="205" pageNumber="206" unit="m" value="440.0">440 m</elevation>
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,
<date id="B32E8EC7FF60FF0DC2D1FE00BA484BB1" box="[425,544,438,464]" pageId="205" pageNumber="206" value="2017-03-03">
<collectingDate id="A36A772FFF60FF0DC2D1FE00BA484BB1" box="[425,544,438,464]" pageId="205" pageNumber="206" value="2017-03-03">3-III-2017</collectingDate>
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,
<collectorName id="6A65CDD1FF60FF0DC1FDFE01BAB64BB1" box="[645,734,438,465]" pageId="205" pageNumber="206">M. Ríos</collectorName>
leg., USNMENT01590780 (
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)
</materialsCitation>
,
<date id="B32E8EC7FF60FF0DC7F4FE00BD7B4BB1" box="[1164,1299,438,464]" pageId="205" pageNumber="206" value="2016-09-10">10-IX-2016</date>
,
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,
<materialsCitation id="77F8A25AFF60FF0DC60FFE00BC704B95" collectingDate="2016-09-12" collectionCode="USNM" collectorName="C. Moraga" country="Costa Rica" pageId="205" pageNumber="206" specimenCount="2" specimenCount-female="2" stateProvince="Guanacaste">
<date id="B32E8EC7FF60FF0DC60FFE00B8914B95" pageId="205" pageNumber="206" value="2016-09-12">
<collectingDate id="A36A772FFF60FF0DC60FFE00B8914B95" pageId="205" pageNumber="206" value="2016-09-12">12- IX-2016</collectingDate>
</date>
,
<specimenCount id="D196638EFF60FF0DC27CFE6CB91B4B95" box="[260,371,474,500]" pageId="205" pageNumber="206" type="female">2 females</specimenCount>
,
<collectorName id="6A65CDD1FF60FF0DC206FE6CB99E4B94" box="[382,502,474,501]" pageId="205" pageNumber="206">C. Moraga</collectorName>
leg., USNMENT01590783–01590785 (
<collectionCode id="A18130C2FF60FF0DC0C1FE6CBC784B95" box="[953,1040,474,500]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="205" pageNumber="206" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="77F8A25AFF60FF0DC75AFE6CB9A6485D" collectingDate="2013-06-11" collectionCode="USNM" collectorName="D. Briceno &amp; Sendero Huerta" country="Costa Rica" elevation="85359" latitude="11.0162" location="Piedrona" longLatPrecision="54" longitude="-85.359" municipality="Area de Conservacion Guanacaste" pageId="205" pageNumber="206" specimenCount="1" specimenCount-female="1" stateProvince="Alajuela">
<emphasis id="F5E47415FF60FF0DC75AFE6CBCEF4B95" bold="true" box="[1058,1159,474,500]" pageId="205" pageNumber="206">
<collectingRegion id="055466E5FF60FF0DC75AFE6CBCEF4B95" box="[1058,1159,474,500]" country="Costa Rica" name="Alajuela" pageId="205" pageNumber="206">Alajuela</collectingRegion>
</emphasis>
—
<collectingMunicipality id="274B327DFF60FF0DC7DCFE6AB9734879" pageId="205" pageNumber="206">Área de Conservación Guanacaste</collectingMunicipality>
, (
<emphasis id="F5E47415FF60FF0DC24AFE48B9F14879" box="[306,409,510,536]" italics="true" pageId="205" pageNumber="206">
<location id="C24FFEDCFF60FF0DC24AFE48B9F14879" LSID="urn:lsid:plazi:treatment:4F391911FF61FF0DC3EFFED8BB5D4975:C24FFEDCFF60FF0DC24AFE48B9F14879" box="[306,409,510,536]" country="Costa Rica" latitude="11.0162" longLatPrecision="54" longitude="-85.359" municipality="Area de Conservacion Guanacaste" name="Piedrona" pageId="205" pageNumber="206" stateProvince="Alajuela">Piedrona</location>
</emphasis>
),
<geoCoordinate id="A2A4CEC0FF60FF0DC2CBFE48BA784879" box="[435,528,510,536]" degrees="11.0162" direction="north" orientation="latitude" pageId="205" pageNumber="206" precision="5" value="11.0162">11.0162</geoCoordinate>
,
<geoCoordinate id="A2A4CEC0FF60FF0DC165FE48BA1E4878" box="[541,630,510,537]" degrees="85.359" direction="west" orientation="longitude" pageId="205" pageNumber="206" precision="55" value="-85.359">
-
<quantity id="006805E2FF60FF0DC15DFE48BA1E4878" box="[549,630,510,537]" metricMagnitude="4" metricUnit="m" metricValue="8.535934" pageId="205" pageNumber="206" unit="m" value="85359.34">
<elevation id="4CBD4F34FF60FF0DC15DFE48BA1E4878" box="[549,630,510,537]" metricMagnitude="4" metricUnit="m" metricValue="8.535934" pageId="205" pageNumber="206" unit="m" value="85359.34">85.359</elevation>
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, 340 m,
<date id="B32E8EC7FF60FF0DC1A5FE48BB0B4879" box="[733,867,510,536]" pageId="205" pageNumber="206" value="2013-06-11">
<collectingDate id="A36A772FFF60FF0DC1A5FE48BB0B4879" box="[733,867,510,536]" pageId="205" pageNumber="206" value="2013-06-11">11-VI-2013</collectingDate>
</date>
,
<specimenCount id="D196638EFF60FF0DC009FE48BBBF4879" box="[881,983,510,536]" pageId="205" pageNumber="206" type="female">1 female</specimenCount>
,
<collectorName id="6A65CDD1FF60FF0DC09DFE49BC0A4879" box="[997,1122,510,537]" pageId="205" pageNumber="206">D. Briceño</collectorName>
leg., USNMENT01590771 (
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), (
<emphasis id="F5E47415FF60FF0DC26AFD94B9AB485D" box="[274,451,546,572]" italics="true" pageId="205" pageNumber="206">
<collectorName id="6A65CDD1FF60FF0DC26AFD94B9AB485D" box="[274,451,546,572]" pageId="205" pageNumber="206">Sendero Huerta</collectorName>
</emphasis>
)
</materialsCitation>
,
<materialsCitation id="77F8A25AFF60FF0DC2A3FD94B9B24801" collectingDate="2016-08-29" collectionCode="USNM" collectorName="C. Cano &amp; Sendero Colegio" country="Costa Rica" elevation="520" latitude="10.9305" longLatPrecision="7" longitude="-85.3722" pageId="205" pageNumber="206" specimenCount="1" specimenCount-male="1" stateProvince="Alajuela">
<geoCoordinate id="A2A4CEC0FF60FF0DC2A3FD94BA51485C" box="[475,569,546,573]" degrees="10.9305" direction="north" orientation="latitude" pageId="205" pageNumber="206" precision="5" value="10.9305">10.9305</geoCoordinate>
,
<geoCoordinate id="A2A4CEC0FF60FF0DC13CFD94BAC4485C" box="[580,684,546,573]" degrees="85.3722" direction="west" orientation="longitude" pageId="205" pageNumber="206" precision="5" value="-85.3722">-85.3722</geoCoordinate>
,
<quantity id="006805E2FF60FF0DC1CFFD94BB68485C" box="[695,768,546,573]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="205" pageNumber="206" unit="m" value="520.0">
<elevation id="4CBD4F34FF60FF0DC1CFFD94BB68485C" box="[695,768,546,573]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="205" pageNumber="206" unit="m" value="520.0">520 m</elevation>
</quantity>
,
<date id="B32E8EC7FF60FF0DC074FD94BBCE485D" box="[780,934,546,572]" pageId="205" pageNumber="206" value="2016-08-29">
<collectingDate id="A36A772FFF60FF0DC074FD94BBCE485D" box="[780,934,546,572]" pageId="205" pageNumber="206" value="2016-08-29">29-VIII-2016</collectingDate>
</date>
,
<specimenCount id="D196638EFF60FF0DC0C9FD94BB97485D" box="[945,1023,546,572]" pageId="205" pageNumber="206" type="male">1 male</specimenCount>
,
<collectorName id="6A65CDD1FF60FF0DC772FD94BC0E485D" box="[1034,1126,546,572]" pageId="205" pageNumber="206">C. Cano</collectorName>
leg., USNMENT01590812 (
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), (
<emphasis id="F5E47415FF60FF0DC26BFDF0B9A74801" box="[275,463,582,608]" italics="true" pageId="205" pageNumber="206">
<collectorName id="6A65CDD1FF60FF0DC26BFDF0B9A74801" box="[275,463,582,608]" pageId="205" pageNumber="206">Sendero Colegio</collectorName>
</emphasis>
)
</materialsCitation>
,
<materialsCitation id="77F8A25AFF60FF0DC29FFDF0B98C48E5" collectingDate="2016-08-09" collectionCode="USNM" collectorName="E. Araya &amp; Estacion Llanura" country="Costa Rica" elevation="520" latitude="10.893" longLatPrecision="55" longitude="-85.3788" pageId="205" pageNumber="206" specimenCount="1" specimenCount-male="1" stateProvince="Alajuela">
<geoCoordinate id="A2A4CEC0FF60FF0DC29FFDF0BA5F4801" box="[487,567,582,608]" degrees="10.893" direction="north" orientation="latitude" pageId="205" pageNumber="206" precision="55" value="10.893">10.893</geoCoordinate>
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<geoCoordinate id="A2A4CEC0FF60FF0DC13BFDF0BAC34800" box="[579,683,582,609]" degrees="85.3788" direction="west" orientation="longitude" pageId="205" pageNumber="206" precision="5" value="-85.3788">-85.3788</geoCoordinate>
,
<quantity id="006805E2FF60FF0DC1CFFDF0BB694800" box="[695,769,582,609]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="205" pageNumber="206" unit="m" value="520.0">
<elevation id="4CBD4F34FF60FF0DC1CFFDF0BB694800" box="[695,769,582,609]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="205" pageNumber="206" unit="m" value="520.0">520 m</elevation>
</quantity>
,
<date id="B32E8EC7FF60FF0DC076FDF0BBF14801" box="[782,921,582,608]" pageId="205" pageNumber="206" value="2016-08-09">
<collectingDate id="A36A772FFF60FF0DC076FDF0BBF14801" box="[782,921,582,608]" pageId="205" pageNumber="206" value="2016-08-09">9-VIII-2016</collectingDate>
</date>
,
<specimenCount id="D196638EFF60FF0DC0DEFDF0BB9D4801" box="[934,1013,582,608]" pageId="205" pageNumber="206" type="male">1 male</specimenCount>
,
<collectorName id="6A65CDD1FF60FF0DC779FDF1BC0C4801" box="[1025,1124,582,609]" pageId="205" pageNumber="206">E. Araya</collectorName>
leg., USNMENT01590813 (
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), (
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<collectorName id="6A65CDD1FF60FF0DC26CFDDDB9B048E5" box="[276,472,618,645]" pageId="205" pageNumber="206">Estación Llanura</collectorName>
</emphasis>
)
</materialsCitation>
,
<materialsCitation id="77F8A25AFF60FF0DC28AFDDCBB6548C9" collectingDate="2012-03-22" collectionCode="USNM" collectorName="C. Umana &amp; Sendero Rincon" country="Costa Rica" elevation="135" latitude="10.93332" longLatPrecision="1" longitude="-85.25331" pageId="205" pageNumber="206" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Alajuela">
<geoCoordinate id="A2A4CEC0FF60FF0DC28AFDDCBA3648E5" box="[498,606,618,644]" degrees="10.93332" direction="north" orientation="latitude" pageId="205" pageNumber="206" precision="1" value="10.93332">10.93332</geoCoordinate>
,
<geoCoordinate id="A2A4CEC0FF60FF0DC112FDDCBA8848E4" box="[618,736,618,645]" degrees="85.25331" direction="west" orientation="longitude" pageId="205" pageNumber="206" precision="1" value="-85.25331">-85.25331</geoCoordinate>
,
<quantity id="006805E2FF60FF0DC194FDDCBB5F48E4" box="[748,823,618,645]" metricMagnitude="2" metricUnit="m" metricValue="1.35" pageId="205" pageNumber="206" unit="m" value="135.0">
<elevation id="4CBD4F34FF60FF0DC194FDDCBB5F48E4" box="[748,823,618,645]" metricMagnitude="2" metricUnit="m" metricValue="1.35" pageId="205" pageNumber="206" unit="m" value="135.0">135 m</elevation>
</quantity>
,
<date id="B32E8EC7FF60FF0DC03CFDDCBBA248E5" box="[836,970,618,644]" pageId="205" pageNumber="206" value="2012-03-22">
<collectingDate id="A36A772FFF60FF0DC03CFDDCBBA248E5" box="[836,970,618,644]" pageId="205" pageNumber="206" value="2012-03-22">22-III-2012</collectingDate>
</date>
,
<specimenCount id="D196638EFF60FF0DC0AFFDDCBC4E48E5" box="[983,1062,618,644]" pageId="205" pageNumber="206" type="male">1 male</specimenCount>
,
<specimenCount id="D196638EFF60FF0DC74CFDDCBCF148E5" box="[1076,1177,618,644]" pageId="205" pageNumber="206" type="female">1 female</specimenCount>
,
<collectorName id="6A65CDD1FF60FF0DC7DEFDDCBD7148E4" box="[1190,1305,618,645]" pageId="205" pageNumber="206">C. Umaña</collectorName>
leg., USN- MENT01590012, 01590015 (
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), (
<emphasis id="F5E47415FF60FF0DC12AFD38BB6A48C8" box="[594,770,654,681]" italics="true" pageId="205" pageNumber="206">
<collectorName id="6A65CDD1FF60FF0DC12AFD38BB6A48C8" box="[594,770,654,681]" pageId="205" pageNumber="206">Sendero Rincón</collectorName>
</emphasis>
)
</materialsCitation>
,
<materialsCitation id="77F8A25AFF60FF0DC060FD38BA5A48AD" collectingDate="2015-06-04" collectionCode="USNM" collectorName="A. Cordoba" country="Costa Rica" elevation="430" latitude="10.8962" longLatPrecision="7" longitude="-85.2777" pageId="205" pageNumber="206" specimenCount="1" specimenCount-male="1" stateProvince="Alajuela">
<geoCoordinate id="A2A4CEC0FF60FF0DC060FD38BB1E48C9" box="[792,886,654,680]" degrees="10.8962" direction="north" orientation="latitude" pageId="205" pageNumber="206" precision="5" value="10.8962">10.8962</geoCoordinate>
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<geoCoordinate id="A2A4CEC0FF60FF0DC0F8FD38BB8F48C8" box="[896,999,654,681]" degrees="85.2777" direction="west" orientation="longitude" pageId="205" pageNumber="206" precision="5" value="-85.2777">-85.2777</geoCoordinate>
,
<quantity id="006805E2FF60FF0DC089FD38BC5048C8" box="[1009,1080,654,681]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="205" pageNumber="206" unit="m" value="430.0">
<elevation id="4CBD4F34FF60FF0DC089FD38BC5048C8" box="[1009,1080,654,681]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="205" pageNumber="206" unit="m" value="430.0">430 m</elevation>
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,
<date id="B32E8EC7FF60FF0DC73BFD38BCD448C8" box="[1091,1212,654,681]" pageId="205" pageNumber="206" value="2015-06-04">
<collectingDate id="A36A772FFF60FF0DC73BFD38BCD448C8" box="[1091,1212,654,681]" pageId="205" pageNumber="206" value="2015-06-04">4-VI-2015</collectingDate>
</date>
,
<specimenCount id="D196638EFF60FF0DC7BEFD38BD7A48C9" box="[1222,1298,654,680]" pageId="205" pageNumber="206" type="male">1 male</specimenCount>
,
<collectorName id="6A65CDD1FF60FF0DC663FD38BDF448C9" box="[1307,1436,654,680]" pageId="205" pageNumber="206">A. Córdoba</collectorName>
leg., USNMENT01590799 (
<collectionCode id="A18130C2FF60FF0DC2ABFD04BA4248AD" box="[467,554,690,716]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="205" pageNumber="206" type="Museum">USNM</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="77F8A25AFF60FF0DC144FD04BD5648AD" box="[572,1342,690,717]" collectingDate="2015-05-06" collectionCode="USNM" collectorName="J. Hernandez" country="Costa Rica" pageId="205" pageNumber="206" specimenCount="1" specimenCount-male="1" stateProvince="Alajuela">
<date id="B32E8EC7FF60FF0DC144FD04BAC048AC" box="[572,680,690,717]" pageId="205" pageNumber="206" value="2015-05-06">
<collectingDate id="A36A772FFF60FF0DC144FD04BAC048AC" box="[572,680,690,717]" pageId="205" pageNumber="206" value="2015-05-06">6-V-2015</collectingDate>
</date>
,
<specimenCount id="D196638EFF60FF0DC1CBFD04BB6848AD" box="[691,768,690,716]" pageId="205" pageNumber="206" type="male">1 male</specimenCount>
,
<collectorName id="6A65CDD1FF60FF0DC073FD05BBF448AD" box="[779,924,690,717]" pageId="205" pageNumber="206">J. Hernández</collectorName>
leg., USNMENT01590801 (
<collectionCode id="A18130C2FF60FF0DC7A6FD04BD5D48AD" box="[1246,1333,690,716]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="205" pageNumber="206" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<emphasis id="F5E47415FF60FF0DC63FFD04B8AE4891" bold="true" pageId="205" pageNumber="206">PANA- MA</emphasis>
:
<materialsCitation id="77F8A25AFF60FF0DC3ACFD63BC9E4891" box="[212,1270,725,753]" collectingDate="1974-10-19" collectionCode="USNM" collectorName="G. B. Small" country="Panama" location="Lago Bayano" pageId="205" pageNumber="206" specimenCount="1" specimenCount-male="1" stateProvince="Alajuela">
<emphasis id="F5E47415FF60FF0DC3ACFD63B95E4891" bold="true" box="[212,310,725,752]" pageId="205" pageNumber="206">
<collectingCountry id="BF87E897FF60FF0DC3ACFD63B95E4891" box="[212,310,725,752]" name="Panama" pageId="205" pageNumber="206">Panamá</collectingCountry>
</emphasis>
—
<location id="C24FFEDCFF60FF0DC22BFD61B9814890" LSID="urn:lsid:plazi:treatment:4F391911FF61FF0DC3EFFED8BB5D4975:C24FFEDCFF60FF0DC22BFD61B9814890" box="[339,489,727,753]" country="Panama" name="Lago Bayano" pageId="205" pageNumber="206" stateProvince="Alajuela">Lago Bayano</location>
,
<date id="B32E8EC7FF60FF0DC28CFD60BA194891" box="[500,625,726,752]" pageId="205" pageNumber="206" value="1974-10-19">
<collectingDate id="A36A772FFF60FF0DC28CFD60BA194891" box="[500,625,726,752]" pageId="205" pageNumber="206" value="1974-10-19">19-X-1974</collectingDate>
</date>
,
<specimenCount id="D196638EFF60FF0DC104FD60BAA14891" box="[636,713,726,752]" pageId="205" pageNumber="206" type="male">1 male</specimenCount>
,
<collectorName id="6A65CDD1FF60FF0DC1ABFD60BB3D4891" box="[723,853,726,753]" pageId="205" pageNumber="206">G. B. Small</collectorName>
leg., USNMENT01590040 (
<collectionCode id="A18130C2FF60FF0DC7EFFD60BC864891" box="[1175,1262,726,752]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="205" pageNumber="206" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="77F8A25AFF60FF0DC678FD63BB594975" collectingDate="1970-07-01" collectionCode="MGCL" country="Panama" location="Pina" pageId="205" pageNumber="206" specimenCount="1" specimenCount-male="1" stateProvince="Colon">
<emphasis id="F5E47415FF60FF0DC678FD63BD204891" bold="true" box="[1280,1352,725,752]" pageId="205" pageNumber="206">
<collectingRegion id="055466E5FF60FF0DC678FD63BD204891" box="[1280,1352,725,752]" country="Panama" name="Colon" pageId="205" pageNumber="206">Colón</collectingRegion>
</emphasis>
—
<location id="C24FFEDCFF60FF0DC61DFD60BDF04891" LSID="urn:lsid:plazi:treatment:4F391911FF61FF0DC3EFFED8BB5D4975:C24FFEDCFF60FF0DC61DFD60BDF04891" box="[1381,1432,726,752]" country="Panama" name="Pina" pageId="205" pageNumber="206" stateProvince="Colon">Piña</location>
,
<date id="B32E8EC7FF60FF0DC3EFFD4CB9714975" box="[151,281,762,788]" pageId="205" pageNumber="206" value="1970-07-01">
<collectingDate id="A36A772FFF60FF0DC3EFFD4CB9714975" box="[151,281,762,788]" pageId="205" pageNumber="206" value="1970-07-01">1-VII-1970</collectingDate>
</date>
,
<specimenCount id="D196638EFF60FF0DC25DFD4CB91B4975" box="[293,371,762,788]" pageId="205" pageNumber="206" type="male">1 male</specimenCount>
, UFFLMNH MGCL 1107067 (
<collectionCode id="A18130C2FF60FF0DC1AAFD4CBB414975" box="[722,809,762,788]" pageId="205" pageNumber="206">MGCL</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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