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<document id="589DEEEE8F6FCC20A14EEB1509CCE315" ID-DOI="10.11646/zootaxa.5216.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7403532" ID-ZooBank="F9C92B91-42CF-4D2D-B315-14FF801815D4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1670314409974" checkinUser="plazi" docAuthor="Piovesan, Mônica, Casagrande, Mirna Martins &amp; Mielke, Olaf Hermann Hendrik" docDate="2022" docId="4F391911FF1BFF76C3EFFD04BCB84C31" docLanguage="en" docName="zootaxa.5216.1.1.pdf" docOrigin="Zootaxa 5216 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Opsiphanes fabricii" docType="treatment" docVersion="3" lastPageNumber="183" masterDocId="B3006169FFADFFC0C378FFB6B8684A61" masterDocTitle="Systematics of Opsiphanes Doubleday, [1849] (Lepidoptera: Nymphalidae, Satyrinae, Brassolini): an integrative approach" masterLastPageNumber="278" masterPageNumber="1" pageNumber="183" updateTime="1699401028458" updateUser="plazi">
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<mods:title id="A4D0AC1BDC0EBA1D5D27D628C3DBF316">Systematics of Opsiphanes Doubleday, [1849] (Lepidoptera: Nymphalidae, Satyrinae, Brassolini): an integrative approach</mods:title>
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<mods:namePart id="ED2B98DA5AE62539F31C90DC999819E0">Piovesan, Mônica</mods:namePart>
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<mods:namePart id="9B69960191737BBA7B1D5E9E7D864854">Casagrande, Mirna Martins</mods:namePart>
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<mods:namePart id="BB2D868E3173BA0DC1D3139CA24B37FF">Mielke, Olaf Hermann Hendrik</mods:namePart>
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<mods:date id="72FB54597ACA318BF24D979065E109B9">2022</mods:date>
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<mods:number id="B80B57A30D4AC9D8B3A49DFA87718E00">5216</mods:number>
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<treatment id="4F391911FF1BFF76C3EFFD04BCB84C31" ID-DOI="http://doi.org/10.5281/zenodo.7403629" ID-Zenodo-Dep="7403629" LSID="urn:lsid:plazi:treatment:4F391911FF1BFF76C3EFFD04BCB84C31" httpUri="http://treatment.plazi.org/id/4F391911FF1BFF76C3EFFD04BCB84C31" lastPageNumber="183" pageId="182" pageNumber="183">
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<paragraph id="C72FA807FF1BFF76C3EFFD04BA0C48AD" blockId="182.[151,612,690,752]" box="[151,612,690,717]" pageId="182" pageNumber="183">
<heading id="9C671F6BFF1BFF76C3EFFD04BA0C48AD" bold="true" box="[151,612,690,717]" fontSize="11" level="2" pageId="182" pageNumber="183" reason="4">
<taxonomicName id="0090D384FF1BFF76C3EFFD04BA0C48AD" ID-CoL="9C4ZC" authority="(Boisduval, 1870)" baseAuthorityName="Boisduval" baseAuthorityYear="1870" box="[151,612,690,717]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="fabricii">
<emphasis id="F5E47415FF1BFF76C3EFFD04BA0C48AD" bold="true" box="[151,612,690,717]" pageId="182" pageNumber="183">
<emphasis id="F5E47415FF1BFF76C3EFFD04B9EC48AC" bold="true" box="[151,388,690,717]" italics="true" pageId="182" pageNumber="183">Opsiphanes fabricii</emphasis>
(
<bibRefCitation id="A301D5F6FF1BFF76C2ECFD04BA3448AD" author="Boisduval, J. B. A. D. de" box="[404,604,690,716]" pageId="182" pageNumber="183" refId="ref174649" refString="Boisduval, J. B. A. D. de (1870) Considerations sur des lepidopteres envoyes du Guatemala a M. de l'Orza. Oberthur et fils., Rennes, Paris, 101 pp." type="book" year="1870">Boisduval, 1870</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="8F8AFB8CFF1BFF76C3EFFD60B92B4891" box="[151,323,726,752]" pageId="182" pageNumber="183" type="description">
<paragraph id="C72FA807FF1BFF76C3EFFD60B92B4891" blockId="182.[151,612,690,752]" box="[151,323,726,752]" pageId="182" pageNumber="183">
(
<figureCitation id="5FABB482FF1BFF76C3E7FD60B9534891" box="[159,315,726,752]" captionStart-0="FIGURE 169" captionStart-1="FIGURE 170" captionStart-10="FIGURE 179" captionStart-11="FIGURE 180" captionStart-12="FIGURE 181" captionStart-13="FIGURE 182" captionStart-14="FIGURE 183" captionStart-15="FIGURE 184" captionStart-16="FIGURE 185" captionStart-17="FIGURE 186" captionStart-2="FIGURE171" captionStart-3="FIGURE 172" captionStart-4="FIGURE 173" captionStart-5="FIGURE 174" captionStart-6="FIGURE 175" captionStart-7="FIGURE 176" captionStart-8="FIGURE 177" captionStart-9="FIGURE 178" captionStartId-0="185.[151,250,1786,1810]" captionStartId-1="186.[151,250,1739,1763]" captionStartId-10="199.[151,250,1888,1912]" captionStartId-11="200.[151,250,1069,1093]" captionStartId-12="200.[151,250,1969,1993]" captionStartId-13="201.[151,250,949,973]" captionStartId-14="201.[151,250,1960,1984]" captionStartId-15="202.[151,250,1029,1053]" captionStartId-16="202.[151,250,1961,1985]" captionStartId-17="203.[151,250,1345,1369]" captionStartId-2="187.[151,250,990,1014]" captionStartId-3="187.[151,250,1966,1990]" captionStartId-4="188.[151,250,1020,1044]" captionStartId-5="188.[151,250,1952,1976]" captionStartId-6="189.[151,250,1248,1272]" captionStartId-7="196.[151,250,1690,1714]" captionStartId-8="197.[151,250,1316,1340]" captionStartId-9="198.[151,250,1937,1961]" captionTargetBox-0="[204,1385,261,1747]" captionTargetBox-1="[208,1390,172,1701]" captionTargetBox-10="[200,1420,379,1851]" captionTargetBox-11="[250,1350,246,1025]" captionTargetBox-12="[214,1372,1139,1946]" captionTargetBox-13="[317,1247,184,908]" captionTargetBox-14="[317,1235,1073,1922]" captionTargetBox-15="[342,1208,182,989]" captionTargetBox-16="[341,1211,1098,1908]" captionTargetBox-17="[189,1363,169,1282]" captionTargetBox-2="[360,1241,170,949]" captionTargetBox-3="[565,1039,1101,1933]" captionTargetBox-4="[551,1046,170,987]" captionTargetBox-5="[562,1024,1104,1919]" captionTargetBox-6="[489,1095,183,1212]" captionTargetBox-7="[238,1355,722,1654]" captionTargetBox-8="[485,1113,183,1284]" captionTargetBox-9="[236,1353,925,1894]" captionTargetId-0="figure-39@185.[155,1432,258,1763]" captionTargetId-1="figure-28@186.[151,1437,166,1715]" captionTargetId-10="figure-115@199.[159,1428,366,1865]" captionTargetId-11="figure-28@200.[219,1367,166,1046]" captionTargetId-12="figure-53@200.[214,1372,1139,1946]" captionTargetId-13="figure-19@201.[317,1269,166,925]" captionTargetId-14="figure-42@201.[315,1271,1054,1936]" captionTargetId-15="figure-28@202.[342,1245,166,1005]" captionTargetId-16="figure-51@202.[341,1246,1098,1937]" captionTargetId-17="figure-19@203.[189,1398,166,1321]" captionTargetId-2="figure-19@187.[310,1276,166,967]" captionTargetId-3="figure-60@187.[299,1287,1096,1942]" captionTargetId-4="figure-28@188.[309,1279,166,996]" captionTargetId-5="figure-57@188.[330,1257,1089,1928]" captionTargetId-6="figure-19@189.[189,1398,166,1223]" captionTargetId-7="figure-273@196.[189,1398,689,1666]" captionTargetId-8="figure-19@197.[149,1437,166,1292]" captionTargetId-9="figure-336@198.[189,1398,910,1914]" captionTargetPageId-0="185" captionTargetPageId-1="186" captionTargetPageId-10="199" captionTargetPageId-11="200" captionTargetPageId-12="200" captionTargetPageId-13="201" captionTargetPageId-14="201" captionTargetPageId-15="202" captionTargetPageId-16="202" captionTargetPageId-17="203" captionTargetPageId-2="187" captionTargetPageId-3="187" captionTargetPageId-4="188" captionTargetPageId-5="188" captionTargetPageId-6="189" captionTargetPageId-7="196" captionTargetPageId-8="197" captionTargetPageId-9="198" captionText-0="FIGURE 169. Opsiphanes fabricii fabricii (Boisduval, 1870): Male, dorsal and ventral views: Mexico: a. (FLMNH MGCL 1092904-MGCL), b. (FLMNH MGCL 1092906-MGCL) Quintana Roo, c. (FLMNH MGCL 1092915-MGCL), d. (FLMNH MGCL 1092918-MGCL), e. (FLMNH MGCL 1092919-MGCL) Chiapas; Belize: f. (19.711-CGCM) Cayo District. White arrows indicate diagnostic characters of the species." captionText-1="FIGURE 170. Opsiphanes fabricii fabricii (Boisduval, 1870): Male, dorsal and ventral views: Guatemala: a. (AMNH_ IZC 00352751-AMNH) Alta Veracruz; Nicarágua: b. (FLMNH MGCL 1107041-MGCL) Manágua; Costa Rica: c. (USN- MENT01589023-USNM), d. (USNMENT01589048-USNM) e. (USNMENT01589075-USNM) Guanacaste; Panama: f. (US- NMENT01590034-USNM) Colón." captionText-10="FIGURE 179. Opsiphanes fabricii fabricii (Boisduval, 1870): Male: a. abdomen, lateral view, b. detail of the scent organ, lateral view; genitalia: c. dorsal view, d. ventral view, e. lateral view; aedeagus: f. dorsal view, g. ventral view, h. lateral view (19.711-CGCM); valva: i. (DZ 41.542-DZUP), j. (DZ 41.563-DZUP), k. (DZ 41.600-DZUP) variations of the dorso-distal portion." captionText-11="FIGURE 180. Opsiphanes fabricii fabricii (Boisduval, 1870): Female genitalia: a. lateral view; b. detail of the signa; c. ventral view (DZ 2475-DZUP)." captionText-12="FIGURE 181. Geographical distribution of Opsiphanes fabricii fabricii (Boisduval, 1870), Opsiphanes fabricii camposi Bristow, 1991 stat. nov. and Opsiphanes fabricii numatius Fruhstorfer, 1912 stat. nov." captionText-13="FIGURE 182. Opsiphanes fabricii fabricii (Boisduval, 1870). Lectotype of Opsiphanes fabricii fabricii (Boisduval, 1870); a. dorsal view, b. ventral view." captionText-14="FIGURE 183. Opsiphanes fabricii fabricii (Boisduval, 1870). Lectotype of Opsiphanes cassina chiriquensis Stichel, 1902 syn. nov.; a. dorsal view, b. ventral view." captionText-15="FIGURE 184. Opsiphanes fabricii fabricii (Boisduval, 1870). Holotype of Opsiphanes cassina aiellae Bristow, 1991; a. dorsal view, b. ventral view." captionText-16="FIGURE 185. Opsiphanes fabricii camposi Bristow, 1991 stat. nov. Holotype of Opsiphanes fabricii camposi Bristow, 1991 stat. nov.; a. dorsal view, b. ventral view." captionText-17="FIGURE 186. Opsiphanes fabricii numatius Fruhstorfer, 1912 stat. nov. Lectotype of Opsiphanes fabricii numatius Fruhstorfer, 1912 stat. nov.; a. dorsal view, b. ventral view." captionText-2="FIGURE171.Opsiphanesfabriciifabricii (Boisduval, 1870):Male,dorsal andventral views:Panama:a. (USNMENT01590042- USNM), b. (AMNH_IZC 00352768-AMNH) Colón. Female, dorsal and ventral views: Mexico: c. (USNMENT00941529- USNM) Oaxaca. White arrows indicate diagnostic characters of the species." captionText-3="FIGURE 172. Opsiphanes fabricii fabricii (Boisduval, 1870): Female, dorsal and ventral views: Mexico: a. (USN- MENT01590957-USNM) San Luíz Potosí, b. (FLMNH MGCL 1092711-MGCL) Oaxaca." captionText-4="FIGURE 173. Opsiphanes fabricii fabricii (Boisduval, 1870): Female, dorsal and ventral views: a. Guatemala: (FLMNH MGCL 1092766-MGCL) Petén; b. El Salvador (FLMNH MGCL 1092774-MGCL) San Salvador." captionText-5="FIGURE 174. Opsiphanes fabricii fabricii (Boisduval, 1870): Female, dorsal and ventral views: Costa Rica: a. (USN- MENT01589011-USNM), b. (USNMENT01589012-USNM) Guanacaste." captionText-6="FIGURE 175. Opsiphanes fabricii fabricii (Boisduval, 1870): Female, dorsal and ventral views: a. Costa Rica: (USN- MENT01589171-USNM) Guanacaste; b. Panama: (USNMENT01590049-USNM) Colón." captionText-7="FIGURE 176. Opsiphanes fabricii camposi Bristow, 1991 stat. nov.: Male, dorsal and ventral views: Ecuador: a. (FLMNH MGCL 1107114-MGCL), b. (USNMENT01590094-USNM), c. (USNMENT01590097-USNM), d. (USNMENT01590098- USNM) Guayaquil." captionText-8="FIGURE 177. Opsiphanes fabricii camposi Bristow, 1991 stat. nov.: Female, dorsal and ventral views: Ecuador: a. (USN- MENT01590101-USNM) Balzapamba, b. (FLMNH MGCL 1092866-MGCL) Pichincha." captionText-9="FIGURE 178. Opsiphanes fabricii numatius Fruhstorfer, 1912 stat. nov.: Male, dorsal and ventral views: Colombia: a. (USN- MENT01590066-USNM), b. (USNMENT01590067-USNM) Medelin. Female, dorsal and ventral views: Colombia: c. (USN- MENT01590070-USNM) Medelin." figureDoi-0="http://doi.org/10.5281/zenodo.7403945" figureDoi-1="http://doi.org/10.5281/zenodo.7403947" figureDoi-10="http://doi.org/10.5281/zenodo.7403965" figureDoi-11="http://doi.org/10.5281/zenodo.7403967" figureDoi-12="http://doi.org/10.5281/zenodo.7403969" figureDoi-13="http://doi.org/10.5281/zenodo.7403973" figureDoi-14="http://doi.org/10.5281/zenodo.7403975" figureDoi-15="http://doi.org/10.5281/zenodo.7403977" figureDoi-16="http://doi.org/10.5281/zenodo.7403981" figureDoi-17="http://doi.org/10.5281/zenodo.7403983" figureDoi-2="http://doi.org/10.5281/zenodo.7403949" figureDoi-3="http://doi.org/10.5281/zenodo.7403951" figureDoi-4="http://doi.org/10.5281/zenodo.7403953" figureDoi-5="http://doi.org/10.5281/zenodo.7403955" figureDoi-6="http://doi.org/10.5281/zenodo.7403957" figureDoi-7="http://doi.org/10.5281/zenodo.7403959" figureDoi-8="http://doi.org/10.5281/zenodo.7403961" figureDoi-9="http://doi.org/10.5281/zenodo.7403963" httpUri-0="https://zenodo.org/record/7403945/files/figure.png" httpUri-1="https://zenodo.org/record/7403947/files/figure.png" httpUri-10="https://zenodo.org/record/7403965/files/figure.png" httpUri-11="https://zenodo.org/record/7403967/files/figure.png" httpUri-12="https://zenodo.org/record/7403969/files/figure.png" httpUri-13="https://zenodo.org/record/7403973/files/figure.png" httpUri-14="https://zenodo.org/record/7403975/files/figure.png" httpUri-15="https://zenodo.org/record/7403977/files/figure.png" httpUri-16="https://zenodo.org/record/7403981/files/figure.png" httpUri-17="https://zenodo.org/record/7403983/files/figure.png" httpUri-2="https://zenodo.org/record/7403949/files/figure.png" httpUri-3="https://zenodo.org/record/7403951/files/figure.png" httpUri-4="https://zenodo.org/record/7403953/files/figure.png" httpUri-5="https://zenodo.org/record/7403955/files/figure.png" httpUri-6="https://zenodo.org/record/7403957/files/figure.png" httpUri-7="https://zenodo.org/record/7403959/files/figure.png" httpUri-8="https://zenodo.org/record/7403961/files/figure.png" httpUri-9="https://zenodo.org/record/7403963/files/figure.png" pageId="182" pageNumber="183">Figs 169–186</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="8F8AFB8CFF1BFF76C3EFFCA8BA8A4FA1" pageId="182" pageNumber="183" type="diagnosis">
<paragraph id="C72FA807FF1BFF76C3EFFCA8BC1D4F6D" blockId="182.[151,1437,798,1617]" pageId="182" pageNumber="183">
<emphasis id="F5E47415FF1BFF76C3EFFCA8B97A4959" bold="true" box="[151,274,798,824]" pageId="182" pageNumber="183">Diagnosis.</emphasis>
The male of
<taxonomicName id="0090D384FF1BFF76C2CAFCA8BAFB4959" baseAuthorityName="Boisduval" baseAuthorityYear="1870" box="[434,659,798,824]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="fabricii">
<emphasis id="F5E47415FF1BFF76C2CAFCA8BAFB4959" box="[434,659,798,824]" italics="true" pageId="182" pageNumber="183">Opsiphanes fabricii</emphasis>
</taxonomicName>
differs from the males of
<taxonomicName id="0090D384FF1BFF76C0B1FCA8BCB34959" authorityName="Chacon &amp; Montero" authorityYear="2012" box="[969,1243,798,824]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="jacobsorum">
<emphasis id="F5E47415FF1BFF76C0B1FCA8BCB34959" box="[969,1243,798,824]" italics="true" pageId="182" pageNumber="183">Opsiphanes jacobsorum</emphasis>
</taxonomicName>
and
<taxonomicName id="0090D384FF1BFF76C660FCA8B964493D" authorityName="Piovesan &amp; Casagrande &amp; Mielke" authorityYear="2022" baseAuthorityName="Fruhstorfer" baseAuthorityYear="1912" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="periphetes" status="stat. nov.">
<emphasis id="F5E47415FF1BFF76C660FCA8B964493D" italics="true" pageId="182" pageNumber="183">Opsiphanes periphetes</emphasis>
</taxonomicName>
<emphasis id="F5E47415FF1BFF76C26BFCF4B916493D" bold="true" box="[275,382,834,860]" pageId="182" pageNumber="183">
<taxonomicNameLabel id="EED7C96EFF1BFF76C26BFCF4B916493D" box="[275,382,834,860]" pageId="182" pageNumber="183">stat. nov.</taxonomicNameLabel>
</emphasis>
in the DHW with a defined submarginal band extending to the anal region, usually as wide as the DFW oblique band, between CuA
<subScript id="5B14AA42FF1BFF76C138FCCFBA2049E6" attach="left" box="[576,584,889,903]" fontSize="6" pageId="182" pageNumber="183">1</subScript>
and 2A. The female of
<taxonomicName id="0090D384FF1BFF76C02EFCD0BC5B49E1" baseAuthorityName="Boisduval" baseAuthorityYear="1870" box="[854,1075,870,896]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="fabricii">
<emphasis id="F5E47415FF1BFF76C02EFCD0BC5B49E1" box="[854,1075,870,896]" italics="true" pageId="182" pageNumber="183">Opsiphanes fabricii</emphasis>
</taxonomicName>
, on the other hand, differs from the female of
<taxonomicName id="0090D384FF1BFF76C24DFC3CBA2D49C5" authorityName="Chacon &amp; Montero" authorityYear="2012" box="[309,581,906,932]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="jacobsorum">
<emphasis id="F5E47415FF1BFF76C24DFC3CBA2D49C5" box="[309,581,906,932]" italics="true" pageId="182" pageNumber="183">Opsiphanes jacobsorum</emphasis>
</taxonomicName>
in the yellow DHW submarginal band, and from the female of
<taxonomicName id="0090D384FF1BFF76C660FC3CB96449A9" authorityName="Piovesan &amp; Casagrande &amp; Mielke" authorityYear="2022" baseAuthorityName="Fruhstorfer" baseAuthorityYear="1912" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="periphetes" status="stat. nov.">
<emphasis id="F5E47415FF1BFF76C660FC3CB96449A9" italics="true" pageId="182" pageNumber="183">Opsiphanes periphetes</emphasis>
</taxonomicName>
<emphasis id="F5E47415FF1BFF76C26BFC18B91749A9" bold="true" box="[275,383,942,968]" pageId="182" pageNumber="183">
<taxonomicNameLabel id="EED7C96EFF1BFF76C26BFC18B91749A9" box="[275,383,942,968]" pageId="182" pageNumber="183">stat. nov.</taxonomicNameLabel>
</emphasis>
in the continuous DHW submarginal band between the veins, from the costal margin to, usually, M
<subScript id="5B14AA42FF1BFF76C39FFC53B8874992" attach="left" box="[231,239,997,1011]" fontSize="6" pageId="182" pageNumber="183">2</subScript>
; the DHW submarginal band has the same width as the DFW oblique band (
<figureCitation id="5FABB482FF1BFF76C734FC64BCD6498D" box="[1100,1214,978,1004]" captionStart="FIGURE 169" captionStartId="185.[151,250,1786,1810]" captionTargetBox="[204,1385,261,1747]" captionTargetId="figure-39@185.[155,1432,258,1763]" captionTargetPageId="185" captionText="FIGURE 169. Opsiphanes fabricii fabricii (Boisduval, 1870): Male, dorsal and ventral views: Mexico: a. (FLMNH MGCL 1092904-MGCL), b. (FLMNH MGCL 1092906-MGCL) Quintana Roo, c. (FLMNH MGCL 1092915-MGCL), d. (FLMNH MGCL 1092918-MGCL), e. (FLMNH MGCL 1092919-MGCL) Chiapas; Belize: f. (19.711-CGCM) Cayo District. White arrows indicate diagnostic characters of the species." figureDoi="http://doi.org/10.5281/zenodo.7403945" httpUri="https://zenodo.org/record/7403945/files/figure.png" pageId="182" pageNumber="183">Figs 169a</figureCitation>
,
<figureCitation id="5FABB482FF1BFF76C7B2FC64BD6C498D" box="[1226,1284,978,1004]" captionStart="FIGURE171" captionStartId="187.[151,250,990,1014]" captionTargetBox="[360,1241,170,949]" captionTargetId="figure-19@187.[310,1276,166,967]" captionTargetPageId="187" captionText="FIGURE171.Opsiphanesfabriciifabricii (Boisduval, 1870):Male,dorsal andventral views:Panama:a. (USNMENT01590042- USNM), b. (AMNH_IZC 00352768-AMNH) Colón. Female, dorsal and ventral views: Mexico: c. (USNMENT00941529- USNM) Oaxaca. White arrows indicate diagnostic characters of the species." figureDoi="http://doi.org/10.5281/zenodo.7403949" httpUri="https://zenodo.org/record/7403949/files/figure.png" pageId="182" pageNumber="183">171a</figureCitation>
).
<taxonomicName id="0090D384FF1BFF76C660FC64B8814E71" baseAuthorityName="Boisduval" baseAuthorityYear="1870" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="fabricii">
<emphasis id="F5E47415FF1BFF76C660FC64B8814E71" italics="true" pageId="182" pageNumber="183">Opsiphanes fabricii</emphasis>
</taxonomicName>
and
<taxonomicName id="0090D384FF1BFF76C25AFC40BA704E71" authorityName="Stichel" authorityYear="1902" box="[290,536,1014,1040]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="merianae">
<emphasis id="F5E47415FF1BFF76C25AFC40BA704E71" box="[290,536,1014,1040]" italics="true" pageId="182" pageNumber="183">Opsiphanes merianae</emphasis>
</taxonomicName>
<emphasis id="F5E47415FF1BFF76C167FC40BAE54E71" bold="true" box="[543,653,1014,1040]" pageId="182" pageNumber="183">stat. rest.</emphasis>
are practically indistinguishable by the external morphology, and are differentiated by means of the genitalia. While the male of
<taxonomicName id="0090D384FF1BFF76C05FFBACBC6C4E55" baseAuthorityName="Boisduval" baseAuthorityYear="1870" box="[807,1028,1050,1076]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="fabricii">
<emphasis id="F5E47415FF1BFF76C05FFBACBC6C4E55" box="[807,1028,1050,1076]" italics="true" pageId="182" pageNumber="183">Opsiphanes fabricii</emphasis>
</taxonomicName>
has the tegumen in dorsal view with an anteriorly projected margin, in the male of
<taxonomicName id="0090D384FF1BFF76C1D8FB88BBFE4E39" authorityName="Stichel" authorityYear="1902" box="[672,918,1086,1112]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="merianae">
<emphasis id="F5E47415FF1BFF76C1D8FB88BBFE4E39" box="[672,918,1086,1112]" italics="true" pageId="182" pageNumber="183">Opsiphanes merianae</emphasis>
</taxonomicName>
<emphasis id="F5E47415FF1BFF76C0E6FB88BC644E39" bold="true" box="[926,1036,1086,1112]" pageId="182" pageNumber="183">stat. rest.</emphasis>
the anterior margin of the tegumen has a slight median concavity. The female of
<taxonomicName id="0090D384FF1BFF76C1E1FBD4BB1F4E1D" baseAuthorityName="Boisduval" baseAuthorityYear="1870" box="[665,887,1122,1148]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="fabricii">
<emphasis id="F5E47415FF1BFF76C1E1FBD4BB1F4E1D" box="[665,887,1122,1148]" italics="true" pageId="182" pageNumber="183">Opsiphanes fabricii</emphasis>
</taxonomicName>
has a C-shaped signum while
<taxonomicName id="0090D384FF1BFF76C7ABFBD4B8A54EC1" authorityName="Stichel" authorityYear="1902" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="merianae">
<emphasis id="F5E47415FF1BFF76C7ABFBD4B8A54EC1" italics="true" pageId="182" pageNumber="183">Opsiphanes merianae</emphasis>
</taxonomicName>
<emphasis id="F5E47415FF1BFF76C3AEFB30B92C4EC1" bold="true" box="[214,324,1158,1184]" pageId="182" pageNumber="183">stat. rest.</emphasis>
has two parallel signa. The geographical distribution can also be used to separate the two species; while
<taxonomicName id="0090D384FF1BFF76C3A3FB1CB9DF4EA5" baseAuthorityName="Boisduval" baseAuthorityYear="1870" box="[219,439,1194,1220]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="fabricii">
<emphasis id="F5E47415FF1BFF76C3A3FB1CB9DF4EA5" box="[219,439,1194,1220]" italics="true" pageId="182" pageNumber="183">Opsiphanes fabricii</emphasis>
</taxonomicName>
subspecies are trans-Andean,
<taxonomicName id="0090D384FF1BFF76C07EFB1CBB924EA5" authorityName="Stichel" authorityYear="1902" box="[774,1018,1194,1220]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="merianae">
<emphasis id="F5E47415FF1BFF76C07EFB1CBB924EA5" box="[774,1018,1194,1220]" italics="true" pageId="182" pageNumber="183">Opsiphanes merianae</emphasis>
</taxonomicName>
<emphasis id="F5E47415FF1BFF76C087FB1CBC034EA5" bold="true" box="[1023,1131,1194,1220]" pageId="182" pageNumber="183">stat. rest.</emphasis>
subspecies are cis-Andean.
<taxonomicName id="0090D384FF1BFF76C3EFFB78B91C4E89" baseAuthorityName="Boisduval" baseAuthorityYear="1870" box="[151,372,1230,1256]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="fabricii">
<emphasis id="F5E47415FF1BFF76C3EFFB78B91C4E89" box="[151,372,1230,1256]" italics="true" pageId="182" pageNumber="183">Opsiphanes fabricii</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="0090D384FF1BFF76C123FB78BB034E89" authorityName="Chacon &amp; Montero" authorityYear="2012" box="[603,875,1230,1256]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="jacobsorum">
<emphasis id="F5E47415FF1BFF76C123FB78BB034E89" box="[603,875,1230,1256]" italics="true" pageId="182" pageNumber="183">Opsiphanes jacobsorum</emphasis>
</taxonomicName>
,
<taxonomicName id="0090D384FF1BFF76C001FB78BC064E89" authorityName="Stichel" authorityYear="1902" box="[889,1134,1230,1256]" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="merianae">
<emphasis id="F5E47415FF1BFF76C001FB78BC064E89" box="[889,1134,1230,1256]" italics="true" pageId="182" pageNumber="183">Opsiphanes merianae</emphasis>
</taxonomicName>
<emphasis id="F5E47415FF1BFF76C70DFB78BC8A4E89" bold="true" box="[1141,1250,1230,1256]" pageId="182" pageNumber="183">stat. rest.</emphasis>
and
<taxonomicName id="0090D384FF1BFF76C660FB78B9644F6D" authorityName="Piovesan &amp; Casagrande &amp; Mielke" authorityYear="2022" baseAuthorityName="Fruhstorfer" baseAuthorityYear="1912" class="Insecta" family="Nymphalidae" genus="Opsiphanes" kingdom="Animalia" order="Lepidoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="periphetes" status="stat. nov.">
<emphasis id="F5E47415FF1BFF76C660FB78B9644F6D" italics="true" pageId="182" pageNumber="183">Opsiphanes periphetes</emphasis>
</taxonomicName>
<emphasis id="F5E47415FF1BFF76C26BFB44B9154F6D" bold="true" box="[275,381,1266,1292]" pageId="182" pageNumber="183">
<taxonomicNameLabel id="EED7C96EFF1BFF76C26BFB44B9154F6D" box="[275,381,1266,1292]" pageId="182" pageNumber="183">stat. nov.</taxonomicNameLabel>
</emphasis>
(genetic distance 0.9–1.2%, 1.2–1.9% and 1.0–1.7%, respectively).
</paragraph>
<paragraph id="C72FA807FF1BFF76C3BFFAA0BCF94F35" blockId="182.[151,1437,798,1617]" pageId="182" pageNumber="183">
<emphasis id="F5E47415FF1BFF76C3BFFAA0B9114F51" bold="true" box="[199,377,1302,1328]" pageId="182" pageNumber="183">Male genitalia.</emphasis>
Tegumen in dorsal view with anterior margin anteriorly projected, shorter than the uncus; anterior projection of saccus in lateral view twice as long as dorsal arms of saccus (
<figureCitation id="5FABB482FF1BFF76C75AFA8CBCEC4F35" box="[1058,1156,1338,1364]" captionStart="FIGURE 179" captionStartId="199.[151,250,1888,1912]" captionTargetBox="[200,1420,379,1851]" captionTargetId="figure-115@199.[159,1428,366,1865]" captionTargetPageId="199" captionText="FIGURE 179. Opsiphanes fabricii fabricii (Boisduval, 1870): Male: a. abdomen, lateral view, b. detail of the scent organ, lateral view; genitalia: c. dorsal view, d. ventral view, e. lateral view; aedeagus: f. dorsal view, g. ventral view, h. lateral view (19.711-CGCM); valva: i. (DZ 41.542-DZUP), j. (DZ 41.563-DZUP), k. (DZ 41.600-DZUP) variations of the dorso-distal portion." figureDoi="http://doi.org/10.5281/zenodo.7403965" httpUri="https://zenodo.org/record/7403965/files/figure.png" pageId="182" pageNumber="183">Fig. 179</figureCitation>
).
</paragraph>
<paragraph id="C72FA807FF1BFF76C3BFFAE8BA8A4FA1" blockId="182.[151,1437,798,1617]" pageId="182" pageNumber="183">
<emphasis id="F5E47415FF1BFF76C3BFFAE8B9FB4F19" bold="true" box="[199,403,1374,1400]" pageId="182" pageNumber="183">Female genitalia.</emphasis>
Lamella antevaginalis membranous medially and sclerotized laterally; lamella postvaginalis rounded medially; ostium with weaker sclerotization than lamella ante and postvaginalis; corpus bursae longer than the ductus bursae, with C-shaped signum (
<figureCitation id="5FABB482FF1BFF76C10BFA10BABD4FA1" box="[627,725,1446,1472]" captionStart="FIGURE 180" captionStartId="200.[151,250,1069,1093]" captionTargetBox="[250,1350,246,1025]" captionTargetId="figure-28@200.[219,1367,166,1046]" captionTargetPageId="200" captionText="FIGURE 180. Opsiphanes fabricii fabricii (Boisduval, 1870): Female genitalia: a. lateral view; b. detail of the signa; c. ventral view (DZ 2475-DZUP)." figureDoi="http://doi.org/10.5281/zenodo.7403967" httpUri="https://zenodo.org/record/7403967/files/figure.png" pageId="182" pageNumber="183">Fig. 180</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8F8AFB8CFF1BFF76C3BFFA7CBCB84C31" pageId="182" pageNumber="183" type="description">
<paragraph id="C72FA807FF1BFF76C3BFFA7CBCB84C31" blockId="182.[151,1437,798,1617]" pageId="182" pageNumber="183">
<emphasis id="F5E47415FF1BFF76C3BFFA7CB9574F85" bold="true" box="[199,319,1482,1508]" pageId="182" pageNumber="183">Variation.</emphasis>
Subspecies vary in the following characters: DFW, the width of the band inside the discal cell, and in the recesses of the anterior and posterior margins of this band; the width and length of the oblique band; due to variations in the width of these bands, the bifurcated appearence is not always continuous. DHW, the width of the submarginal band. VHW, the zigzag submarginal line EIII is not always evident (
<figureCitation id="5FABB482FF1BFF76C75CF980BCAB4C31" box="[1060,1219,1590,1616]" captionStart-0="FIGURE 169" captionStart-1="FIGURE 170" captionStart-2="FIGURE171" captionStart-3="FIGURE 172" captionStart-4="FIGURE 173" captionStart-5="FIGURE 174" captionStart-6="FIGURE 175" captionStart-7="FIGURE 176" captionStart-8="FIGURE 177" captionStart-9="FIGURE 178" captionStartId-0="185.[151,250,1786,1810]" captionStartId-1="186.[151,250,1739,1763]" captionStartId-2="187.[151,250,990,1014]" captionStartId-3="187.[151,250,1966,1990]" captionStartId-4="188.[151,250,1020,1044]" captionStartId-5="188.[151,250,1952,1976]" captionStartId-6="189.[151,250,1248,1272]" captionStartId-7="196.[151,250,1690,1714]" captionStartId-8="197.[151,250,1316,1340]" captionStartId-9="198.[151,250,1937,1961]" captionTargetBox-0="[204,1385,261,1747]" captionTargetBox-1="[208,1390,172,1701]" captionTargetBox-2="[360,1241,170,949]" captionTargetBox-3="[565,1039,1101,1933]" captionTargetBox-4="[551,1046,170,987]" captionTargetBox-5="[562,1024,1104,1919]" captionTargetBox-6="[489,1095,183,1212]" captionTargetBox-7="[238,1355,722,1654]" captionTargetBox-8="[485,1113,183,1284]" captionTargetBox-9="[236,1353,925,1894]" captionTargetId-0="figure-39@185.[155,1432,258,1763]" captionTargetId-1="figure-28@186.[151,1437,166,1715]" captionTargetId-2="figure-19@187.[310,1276,166,967]" captionTargetId-3="figure-60@187.[299,1287,1096,1942]" captionTargetId-4="figure-28@188.[309,1279,166,996]" captionTargetId-5="figure-57@188.[330,1257,1089,1928]" captionTargetId-6="figure-19@189.[189,1398,166,1223]" captionTargetId-7="figure-273@196.[189,1398,689,1666]" captionTargetId-8="figure-19@197.[149,1437,166,1292]" captionTargetId-9="figure-336@198.[189,1398,910,1914]" captionTargetPageId-0="185" captionTargetPageId-1="186" captionTargetPageId-2="187" captionTargetPageId-3="187" captionTargetPageId-4="188" captionTargetPageId-5="188" captionTargetPageId-6="189" captionTargetPageId-7="196" captionTargetPageId-8="197" captionTargetPageId-9="198" captionText-0="FIGURE 169. Opsiphanes fabricii fabricii (Boisduval, 1870): Male, dorsal and ventral views: Mexico: a. (FLMNH MGCL 1092904-MGCL), b. (FLMNH MGCL 1092906-MGCL) Quintana Roo, c. (FLMNH MGCL 1092915-MGCL), d. (FLMNH MGCL 1092918-MGCL), e. (FLMNH MGCL 1092919-MGCL) Chiapas; Belize: f. (19.711-CGCM) Cayo District. White arrows indicate diagnostic characters of the species." captionText-1="FIGURE 170. Opsiphanes fabricii fabricii (Boisduval, 1870): Male, dorsal and ventral views: Guatemala: a. (AMNH_ IZC 00352751-AMNH) Alta Veracruz; Nicarágua: b. (FLMNH MGCL 1107041-MGCL) Manágua; Costa Rica: c. (USN- MENT01589023-USNM), d. (USNMENT01589048-USNM) e. (USNMENT01589075-USNM) Guanacaste; Panama: f. (US- NMENT01590034-USNM) Colón." captionText-2="FIGURE171.Opsiphanesfabriciifabricii (Boisduval, 1870):Male,dorsal andventral views:Panama:a. (USNMENT01590042- USNM), b. (AMNH_IZC 00352768-AMNH) Colón. Female, dorsal and ventral views: Mexico: c. (USNMENT00941529- USNM) Oaxaca. White arrows indicate diagnostic characters of the species." captionText-3="FIGURE 172. Opsiphanes fabricii fabricii (Boisduval, 1870): Female, dorsal and ventral views: Mexico: a. (USN- MENT01590957-USNM) San Luíz Potosí, b. (FLMNH MGCL 1092711-MGCL) Oaxaca." captionText-4="FIGURE 173. Opsiphanes fabricii fabricii (Boisduval, 1870): Female, dorsal and ventral views: a. Guatemala: (FLMNH MGCL 1092766-MGCL) Petén; b. El Salvador (FLMNH MGCL 1092774-MGCL) San Salvador." captionText-5="FIGURE 174. Opsiphanes fabricii fabricii (Boisduval, 1870): Female, dorsal and ventral views: Costa Rica: a. (USN- MENT01589011-USNM), b. (USNMENT01589012-USNM) Guanacaste." captionText-6="FIGURE 175. Opsiphanes fabricii fabricii (Boisduval, 1870): Female, dorsal and ventral views: a. Costa Rica: (USN- MENT01589171-USNM) Guanacaste; b. Panama: (USNMENT01590049-USNM) Colón." captionText-7="FIGURE 176. Opsiphanes fabricii camposi Bristow, 1991 stat. nov.: Male, dorsal and ventral views: Ecuador: a. (FLMNH MGCL 1107114-MGCL), b. (USNMENT01590094-USNM), c. (USNMENT01590097-USNM), d. (USNMENT01590098- USNM) Guayaquil." captionText-8="FIGURE 177. Opsiphanes fabricii camposi Bristow, 1991 stat. nov.: Female, dorsal and ventral views: Ecuador: a. (USN- MENT01590101-USNM) Balzapamba, b. (FLMNH MGCL 1092866-MGCL) Pichincha." captionText-9="FIGURE 178. Opsiphanes fabricii numatius Fruhstorfer, 1912 stat. nov.: Male, dorsal and ventral views: Colombia: a. (USN- MENT01590066-USNM), b. (USNMENT01590067-USNM) Medelin. Female, dorsal and ventral views: Colombia: c. (USN- MENT01590070-USNM) Medelin." figureDoi-0="http://doi.org/10.5281/zenodo.7403945" figureDoi-1="http://doi.org/10.5281/zenodo.7403947" figureDoi-2="http://doi.org/10.5281/zenodo.7403949" figureDoi-3="http://doi.org/10.5281/zenodo.7403951" figureDoi-4="http://doi.org/10.5281/zenodo.7403953" figureDoi-5="http://doi.org/10.5281/zenodo.7403955" figureDoi-6="http://doi.org/10.5281/zenodo.7403957" figureDoi-7="http://doi.org/10.5281/zenodo.7403959" figureDoi-8="http://doi.org/10.5281/zenodo.7403961" figureDoi-9="http://doi.org/10.5281/zenodo.7403963" httpUri-0="https://zenodo.org/record/7403945/files/figure.png" httpUri-1="https://zenodo.org/record/7403947/files/figure.png" httpUri-2="https://zenodo.org/record/7403949/files/figure.png" httpUri-3="https://zenodo.org/record/7403951/files/figure.png" httpUri-4="https://zenodo.org/record/7403953/files/figure.png" httpUri-5="https://zenodo.org/record/7403955/files/figure.png" httpUri-6="https://zenodo.org/record/7403957/files/figure.png" httpUri-7="https://zenodo.org/record/7403959/files/figure.png" httpUri-8="https://zenodo.org/record/7403961/files/figure.png" httpUri-9="https://zenodo.org/record/7403963/files/figure.png" pageId="182" pageNumber="183">Figs 169–178</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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