293 lines
36 KiB
XML
293 lines
36 KiB
XML
<document id="FCF9E3F6C155E173624416BD20574CBA" ID-DOI="10.11646/zootaxa.3957.1.8" ID-GBIF-Dataset="d97787ca-3db9-4e8f-956f-d26b955e71d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="236432" ID-ZooBank="EBFD444B-8315-4908-866D-22D30AFB5557" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461059621670" checkinUser="plazi" docAuthor="Özbek, Hasan Hüseyin & Halliday, Bruce" docDate="2015" docId="4F248367DB5F5C349ADBFBAD3788821D" docLanguage="en" docName="zt03957p108.pdf" docOrigin="Zootaxa 3957 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pachyseius quadrigeminus Özbek & Halliday, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="104" masterDocId="B31DFB1FDB5C5C329A4CFFC934468300" masterDocTitle="Two new species of Pachyseius Berlese (Acari: Pachylaelapidae) from Turkey, with a key to the world species" masterLastPageNumber="108" masterPageNumber="98" pageNumber="101" updateTime="1698581885510" updateUser="plazi">
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<mods:mods id="2D95FCB8CF2C457F0EDFB064E7EEF529" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A9C8C1CB80FAAB7D2DD4AE5452EB5B91">
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<mods:title id="89E607E2E485BC7DEA5A8E86CF3DB17D">Two new species of Pachyseius Berlese (Acari: Pachylaelapidae) from Turkey, with a key to the world species</mods:title>
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</mods:titleInfo>
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<mods:name id="077C281E71FE905AAEB372926BA27D95" type="personal">
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<mods:role id="4E6DD14DF91D3827030BDC74E4EE2B28">
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<mods:roleTerm id="44E83165FC6E8E346C79E75BF41A66C9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="848E924C9445C09C832267B1D42FDA00">Özbek, Hasan Hüseyin</mods:namePart>
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</mods:name>
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<mods:name id="E28A0767E816F7978DD1C3E50067486C" type="personal">
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<mods:role id="2F725374AA27ACEBB09245C31C2248DF">
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<mods:roleTerm id="4B22C0B1C6DBC64B230B1A46CF81D554">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="43FB3F5BA60C4883F2AA343C24D0B4BC">Halliday, Bruce</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="FC35C0EDF22F85A48BF0CC309C03520D">text</mods:typeOfResource>
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<mods:titleInfo id="23911D1B0896BD5C2DC5DDCF5A3C29B9">
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<mods:title id="43CEDD39C66C070C8F3591824F9D0764">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="D6BCC4B1184405F2EE7C77C46098C0BB">
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<mods:date id="59283CD955C0060E0F15C8D8988DD469">2015</mods:date>
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<mods:detail id="495870A0E3C12F8B2A23A94D37597E1C" type="volume">
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<mods:number id="C25110D39567E482568CFA2F6E920BF3">3957</mods:number>
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</mods:detail>
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<mods:detail id="A77D9936EDB6245717813B8B0CA644BE" type="issue">
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<mods:number id="C097290ABCC3ED1245F7DC160693AC86">1</mods:number>
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</mods:detail>
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<mods:start id="71214A3AC02A0ED6BF4B5EC1E2831201">98</mods:start>
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<mods:end id="B2DB3A23E8459181763A7BCB07A5D1F8">108</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="2705D57457399BDF84957088171DA398">journal article</mods:classification>
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<mods:identifier id="DB1039932FA28B441D27CF035C3F5CE4" type="DOI">10.11646/zootaxa.3957.1.8</mods:identifier>
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<mods:identifier id="C2D992087FBF0ACEF72C270EC37A3113" type="GBIF-Dataset">d97787ca-3db9-4e8f-956f-d26b955e71d6</mods:identifier>
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<mods:identifier id="E5276FE93C0A486A84987BE8C39F8A75" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="4CB9FAC9CA3584684CA0BE0DD77E584A" type="Zenodo-Dep">236432</mods:identifier>
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<mods:identifier id="AB20B20C06C7D25CC0534C349BC22047" type="ZooBank">EBFD444B-8315-4908-866D-22D30AFB5557</mods:identifier>
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</mods:mods>
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<treatment id="4F248367DB5F5C349ADBFBAD3788821D" ID-DOI="http://doi.org/10.5281/zenodo.6093709" ID-GBIF-Taxon="119613791" ID-Zenodo-Dep="6093709" LSID="urn:lsid:plazi:treatment:4F248367DB5F5C349ADBFBAD3788821D" httpUri="http://treatment.plazi.org/id/4F248367DB5F5C349ADBFBAD3788821D" lastPageId="6" lastPageNumber="104" pageId="3" pageNumber="101">
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<subSubSection id="8F9761FADB5F5C319ADBFBAD3502879F" pageId="3" pageNumber="101" type="nomenclature">
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<paragraph id="C7323271DB5F5C319ADBFBAD3607877E" blockId="3.[151,577,1124,1183]" box="[151,577,1124,1150]" pageId="3" pageNumber="101">
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<heading id="9C7A851DDB5F5C319ADBFBAD3607877E" bold="true" box="[151,577,1124,1150]" fontSize="11" level="1" pageId="3" pageNumber="101" reason="1">
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<emphasis id="F5F9EE63DB5F5C319ADBFBAD3607877E" bold="true" box="[151,577,1124,1150]" pageId="3" pageNumber="101">
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<taxonomicName id="008D49F2DB5F5C319ADBFBAD359D877E" ID-CoL="8Y7NP" box="[151,475,1124,1150]" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="101" phylum="Arthropoda" rank="species" species="quadrigeminus" status="sp. nov.">
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<emphasis id="F5F9EE63DB5F5C319ADBFBAD359D877E" bold="true" box="[151,475,1124,1150]" italics="true" pageId="3" pageNumber="101">Pachyseius quadrigeminus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="EECA5318DB5F5C319BAFFBAD3607877E" box="[483,577,1124,1150]" pageId="3" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="C7323271DB5F5C319ADBFB4F3502879F" blockId="3.[151,577,1124,1183]" box="[151,324,1158,1183]" pageId="3" pageNumber="101">
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||
(
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<figureCitation id="5FB62EF4DB5F5C319AD3FB4F3554879F" box="[159,274,1158,1183]" captionStart-0="FIGURES 7 – 8" captionStart-1="FIGURES 9 – 13" captionStart-2="FIGURES 14 – 18" captionStartId-0="4.[151,264,1733,1756]" captionStartId-1="5.[151,264,1403,1426]" captionStartId-2="6.[151,264,1547,1570]" captionTargetBox-0="[165,1421,635,1704]" captionTargetBox-1="[174,1422,201,1378]" captionTargetBox-2="[162,1432,351,1514]" captionTargetId-0="figure@4.[151,1436,625,1712]" captionTargetId-1="figure@5.[151,1436,193,1382]" captionTargetId-2="figure@6.[151,1436,339,1526]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURES 7 – 8. Pachyseius quadrigeminus sp. nov., female. 7. Dorsal shield; 8. Ventral idiosoma." captionText-1="FIGURES 9 – 13. Pachyseius quadrigeminus sp. nov., female. 9. Ventral gnathosoma; 10. Epistome; 11. Chelicera; 12. Sperm access system; 13. Tarsus II." captionText-2="FIGURES 14 – 18. Pachyseius quadrigeminus sp. nov., male. 14. Ventral idiosoma; 15. Epistome; 16. Chelicera; 17. Ventral spurs on leg II (fem. = femur, gen. = genu, tib. = tibia); 18. Telotarsus II." httpUri-0="https://zenodo.org/record/236435/files/figure.png" httpUri-1="https://zenodo.org/record/236436/files/figure.png" httpUri-2="https://zenodo.org/record/236437/files/figure.png" pageId="3" pageNumber="101">Figs 7–18</figureCitation>
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; 22)
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</paragraph>
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</subSubSection>
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<subSubSection id="8F9761FADB5F5C319ADBFB0736F5852A" pageId="3" pageNumber="101" type="diagnosis">
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<paragraph id="C7323271DB5F5C319ADBFB0737128676" blockId="3.[151,1437,1230,2011]" pageId="3" pageNumber="101">
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<emphasis id="F5F9EE63DB5F5C319ADBFB07355787E7" bold="true" box="[151,273,1230,1255]" pageId="3" pageNumber="101">Diagnosis.</emphasis>
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In female, peritrematal shields fused with exopodal shields; ventri-anal shield bearing four pairs of preanal setae; pre-sternal shields absent; area between peritrematal shields and anterolateral margins of ventri-anal shield with two pairs of oval platelets, lateral and opisthogastric soft integument with seven pairs of setae. In male, holoventral shield present, opisthogastric region of shield with four pairs of ventral setae; spermatodactyl 90–110 long, and distally narrowed. Tarsus II with two spur-like setae.
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</paragraph>
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<paragraph id="C7323271DB5F5C319A8BFA4B36F5852A" blockId="3.[151,1437,1230,2011]" pageId="3" pageNumber="101">
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<emphasis id="F5F9EE63DB5F5C319A8BFA4B358E869B" bold="true" box="[199,456,1410,1435]" pageId="3" pageNumber="101">Specimens examined.</emphasis>
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<typeStatus id="18368CD3DB5F5C319B9CFA4B367E869B" box="[464,568,1410,1435]" pageId="3" pageNumber="101" type="holotype">Holotype</typeStatus>
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female, Gümüşhane Province, Çıkrıkdüzü Wold,
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<geoCoordinate id="A2B954B6DB5F5C319E3EFA4B3146869A" box="[1138,1280,1410,1435]" direction="north" orientation="latitude" pageId="3" pageNumber="101" precision="15" value="40.66611">40°39'58'' N</geoCoordinate>
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,
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<geoCoordinate id="A2B954B6DB5F5C319F41FA4B31D1869A" box="[1293,1431,1410,1435]" direction="east" orientation="longitude" pageId="3" pageNumber="101" precision="15" value="38.997223">38°59'50'' E</geoCoordinate>
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, alt.
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<quantity id="00759F94DB5F5C319A8EFA6E355D86BE" box="[194,283,1447,1471]" metricMagnitude="3" metricUnit="m" metricValue="1.9899999999999998" pageId="3" pageNumber="101" unit="m" value="1990.0">1990 m</quantity>
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,
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<date id="B33314B1DB5F5C319B64FA6E35AC86BF" box="[296,490,1446,1471]" pageId="3" pageNumber="101" value="2013-10-12">12 October 2013</date>
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, litter under
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<taxonomicName id="008D49F2DB5F5C319831FA6F36FD86BE" box="[637,699,1446,1470]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="101" phylum="Tracheophyta" rank="genus">
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<emphasis id="F5F9EE63DB5F5C319831FA6F36FD86BE" box="[637,699,1446,1470]" italics="true" pageId="3" pageNumber="101">Abies</emphasis>
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</taxonomicName>
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sp.
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<typeStatus id="18368CD3DB5F5C3198A0FA6E371886BF" box="[748,862,1447,1471]" pageId="3" pageNumber="101" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="D18BF9F8DB5F5C319925FA6E379C86BF" box="[873,986,1446,1471]" pageId="3" pageNumber="101" type="female">2 females</specimenCount>
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, same data as
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<typeStatus id="18368CD3DB5F5C319E33FA6F30A286BF" box="[1151,1252,1446,1471]" pageId="3" pageNumber="101" type="holotype">holotype</typeStatus>
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;
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<specimenCount id="D18BF9F8DB5F5C319EBEFA6E311286BF" box="[1266,1364,1446,1471]" pageId="3" pageNumber="101" type="female">1 female</specimenCount>
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and
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<specimenCount id="D18BF9F8DB5F5C319FC2FA6F349A86E3" pageId="3" pageNumber="101" type="male">6 males</specimenCount>
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; Gümüşhane Province, Kürtün Town, Örümcek Forest,
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<geoCoordinate id="A2B954B6DB5F5C31991CFA02379B86E2" box="[848,989,1483,1507]" direction="north" orientation="latitude" pageId="3" pageNumber="101" precision="15" value="40.663334">40°39'48'' N</geoCoordinate>
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,
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<geoCoordinate id="A2B954B6DB5F5C3199A5FA03303486E2" box="[1001,1138,1482,1507]" direction="east" orientation="longitude" pageId="3" pageNumber="101" precision="15" value="39.029446">39°01'46'' E</geoCoordinate>
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, alt.
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<quantity id="00759F94DB5F5C319EE5FA03314686E2" box="[1193,1280,1482,1507]" metricMagnitude="3" metricUnit="m" metricValue="1.455" pageId="3" pageNumber="101" unit="m" value="1455.0">1455 m</quantity>
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,
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<date id="B33314B1DB5F5C319F41FA0231DE86E3" box="[1293,1432,1483,1507]" pageId="3" pageNumber="101" value="2009-06-01">1 June 2009</date>
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, litter under
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<taxonomicName id="008D49F2DB5F5C319B56FA26351C8506" box="[282,346,1519,1542]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="101" phylum="Tracheophyta" rank="genus">
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<emphasis id="F5F9EE63DB5F5C319B56FA26351C8506" box="[282,346,1519,1542]" italics="true" pageId="3" pageNumber="101">Picea</emphasis>
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</taxonomicName>
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sp.;
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<specimenCount id="D18BF9F8DB5F5C319BDDFA2635B38507" box="[401,501,1518,1543]" pageId="3" pageNumber="101" type="female">1 female</specimenCount>
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, Gümüşhane Province, Kürtün Town, Kızılali Wold,
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<geoCoordinate id="A2B954B6DB5F5C319E07FA26309E8506" box="[1099,1240,1519,1543]" direction="north" orientation="latitude" pageId="3" pageNumber="101" precision="15" value="40.80472">40°48'17'' N</geoCoordinate>
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,
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<geoCoordinate id="A2B954B6DB5F5C319EA8FA26312B8506" box="[1252,1389,1519,1543]" direction="east" orientation="longitude" pageId="3" pageNumber="101" precision="15" value="39.067776">39°04'04'' E</geoCoordinate>
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, alt.
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<quantity id="00759F94DB5F5C319ADBF9DB34A8852A" box="[151,238,1554,1579]" metricMagnitude="3" metricUnit="m" metricValue="1.555" pageId="3" pageNumber="101" unit="m" value="1555.0">1555 m</quantity>
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,
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<date id="B33314B1DB5F5C319AB6F9DA35FF852B" box="[250,441,1554,1579]" pageId="3" pageNumber="101" value="2013-10-13">13 October 2013</date>
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, litter under
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<taxonomicName id="008D49F2DB5F5C31980AF9DA36CA852B" box="[582,652,1555,1579]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="3" pageNumber="101" phylum="Tracheophyta" rank="genus">
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<emphasis id="F5F9EE63DB5F5C31980AF9DA36CA852B" box="[582,652,1555,1579]" italics="true" pageId="3" pageNumber="101">Rubus</emphasis>
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</taxonomicName>
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sp.
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</paragraph>
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</subSubSection>
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<subSubSection id="8F9761FADB5F5C379A8BF9FF36868458" lastPageId="5" lastPageNumber="103" pageId="3" pageNumber="101" type="description">
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<paragraph id="C7323271DB5F5C319A8BF9FF36A285DE" blockId="3.[151,1437,1230,2011]" pageId="3" pageNumber="101">
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<emphasis id="F5F9EE63DB5F5C319A8BF9FF3563854F" bold="true" box="[199,293,1590,1615]" pageId="3" pageNumber="101">Female.</emphasis>
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<emphasis id="F5F9EE63DB5F5C319B62F9FE35CE854E" box="[302,392,1591,1614]" italics="true" pageId="3" pageNumber="101">Dorsum</emphasis>
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||
(
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<figureCitation id="5FB62EF4DB5F5C319BD6F9FF35A6854F" box="[410,480,1590,1615]" captionStart="FIGURES 7 – 8" captionStartId="4.[151,264,1733,1756]" captionTargetBox="[165,1421,635,1704]" captionTargetId="figure@4.[151,1436,625,1712]" captionTargetPageId="4" captionText="FIGURES 7 – 8. Pachyseius quadrigeminus sp. nov., female. 7. Dorsal shield; 8. Ventral idiosoma." httpUri="https://zenodo.org/record/236435/files/figure.png" pageId="3" pageNumber="101">Fig. 7</figureCitation>
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). Dorsal shield 600–676 long, 310–368 wide at widest point, oblong (length/width, 1.71–1.94), ornamented with reticulate pattern, carrying 30 pairs of setae, all smooth and pointed; setae
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<emphasis id="F5F9EE63DB5F5C319F69F99231788573" box="[1317,1342,1627,1651]" italics="true" pageId="3" pageNumber="101">z1</emphasis>
|
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shortest (8–10);
|
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<emphasis id="F5F9EE63DB5F5C319ABEF9B635418597" box="[242,263,1663,1687]" italics="true" pageId="3" pageNumber="101">
|
||
j
|
||
<date id="B33314B1DB5F5C319AB1F9B635418597" box="[253,263,1663,1687]" pageId="3" pageNumber="101" value="1915-01-20">1</date>
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</emphasis>
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15–20;
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<emphasis id="F5F9EE63DB5F5C319B2AF9B635398597" box="[358,383,1663,1687]" italics="true" pageId="3" pageNumber="101">z2</emphasis>
|
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,
|
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<emphasis id="F5F9EE63DB5F5C319BC3F9B635EE8597" box="[399,424,1663,1687]" italics="true" pageId="3" pageNumber="101">s2</emphasis>
|
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,
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<emphasis id="F5F9EE63DB5F5C319BFBF9B635978597" box="[439,465,1663,1687]" italics="true" pageId="3" pageNumber="101">r3</emphasis>
|
||
and
|
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<emphasis id="F5F9EE63DB5F5C319846F9B636638596" box="[522,549,1663,1686]" italics="true" pageId="3" pageNumber="101">J5</emphasis>
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distinctly longer (30–40), other dorsal setae 20–30 long; a pair of lyrifissures present between
|
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<emphasis id="F5F9EE63DB5F5C319B14F96A352885BB" box="[344,366,1699,1723]" italics="true" pageId="3" pageNumber="101">j2</emphasis>
|
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and
|
||
<emphasis id="F5F9EE63DB5F5C319BE6F96A358585BB" box="[426,451,1699,1723]" italics="true" pageId="3" pageNumber="101">z1</emphasis>
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; a pair of slit-like gland pores
|
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<emphasis id="F5F9EE63DB5F5C319962F96A372685BB" box="[814,864,1699,1723]" italics="true" pageId="3" pageNumber="101">gdj3</emphasis>
|
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present between seta
|
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<emphasis id="F5F9EE63DB5F5C319E2DF96A303F85BB" box="[1121,1145,1699,1723]" italics="true" pageId="3" pageNumber="101">z1</emphasis>
|
||
and
|
||
<emphasis id="F5F9EE63DB5F5C319EF9F96A308885BB" box="[1205,1230,1699,1723]" italics="true" pageId="3" pageNumber="101">z2</emphasis>
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. Anterior ends of peritremes reaching or extending beyond level of
|
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<emphasis id="F5F9EE63DB5F5C319888F90E369B85DF" box="[708,733,1735,1759]" italics="true" pageId="3" pageNumber="101">z1</emphasis>
|
||
.
|
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</paragraph>
|
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<paragraph id="C7323271DB5F5C369A8BF92230558265" blockId="3.[151,1437,1230,2011]" lastBlockId="4.[151,1437,151,573]" lastPageId="4" lastPageNumber="102" pageId="3" pageNumber="101">
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<emphasis id="F5F9EE63DB5F5C319A8BF922354A8402" box="[199,268,1771,1794]" italics="true" pageId="3" pageNumber="101">Venter</emphasis>
|
||
(
|
||
<figureCitation id="5FB62EF4DB5F5C319B51F92335278403" box="[285,353,1770,1795]" captionStart="FIGURES 7 – 8" captionStartId="4.[151,264,1733,1756]" captionTargetBox="[165,1421,635,1704]" captionTargetId="figure@4.[151,1436,625,1712]" captionTargetPageId="4" captionText="FIGURES 7 – 8. Pachyseius quadrigeminus sp. nov., female. 7. Dorsal shield; 8. Ventral idiosoma." httpUri="https://zenodo.org/record/236435/files/figure.png" pageId="3" pageNumber="101">Fig. 8</figureCitation>
|
||
). Pre-sternal shields absent. Sternal shield 140–152 long, 80–90 wide at level of coxae II, central region ornamented with punctate polygonal lines, carrying three pairs of smooth pointed setae and two pairs of lyrifissures, edges of shield strongly sclerotised. Length of setae
|
||
<emphasis id="F5F9EE63DB5F5C319921F8FA37EC844B" box="[877,938,1843,1867]" italics="true" pageId="3" pageNumber="101">
|
||
st
|
||
<date id="B33314B1DB5F5C3199CEF8FA37EC844B" box="[898,938,1843,1867]" pageId="3" pageNumber="101" value="1936-01-25" valueMax="1936-03-25" valueMin="1936-01-25">1–3</date>
|
||
</emphasis>
|
||
25–36. Metasternal shields very small, each bearing a metasternal seta (20–25) and a poroid. Epigynal shield 85–95 long, 100–110 wide at widest point, with punctate ornamentation; one pair of elongate platelets on its posterior margin; genital pores situated outside shield; genital setae length 23–27. Ventri-anal shield (
|
||
<figureCitation id="5FB62EF4DB5F5C3198E7F85736B884B7" box="[683,766,1950,1975]" captionStart="FIGURES 19 – 22" captionStartId="8.[151,264,1740,1763]" captionTargetBox="[327,1259,501,1719]" captionTargetId="figure@8.[327,1259,501,1720]" captionTargetPageId="8" captionText="FIGURES 19 – 22. Pachyseius spp., females. 19. P. destitutus sp. nov., anterior region of dorsal idiosoma; 20. P. destitutus sp. nov., ventri-anal shield; 21. P. destitutus sp. nov., ventral gnathosoma; 22. P. quadrigeminus sp. nov., ventri-anal shield. Not scaled." httpUri="https://zenodo.org/record/236438/files/figure.png" pageId="3" pageNumber="101">Fig. 22</figureCitation>
|
||
) longer than wide, 248–270 long, 208–233 wide at widest point (length/width 1.10–1.21), reticulated with punctate lines, and bearing four pairs of sub-equal pre-anal setae (20–30), a pair of longer para-anal setae (30–35) and a long post-anal seta (40–46). Peritrematal shields fused with exopodal shields, with two pairs of small post-stigmatal pores and truncated posterior margin, their posterior ends rounded, enclosing posterior margins of coxae IV and extending beyond level of genital setae. Soft integument between peritrematal shields and anterolateral margins of ventri-anal shield with two pairs of oval platelets. Metapodal shields elongate (60–65 long) and not well separated from anterolateral margins of ventri-anal shield. Lateral and opisthogastric soft integument bearing seven pairs of setae (15–25).
|
||
</paragraph>
|
||
<paragraph id="C7323271DB585C369A8BFEB936E2813C" blockId="4.[151,1437,151,573]" pageId="4" pageNumber="102">
|
||
<emphasis id="F5F9EE63DB585C369A8BFEB935108288" box="[199,342,368,392]" italics="true" pageId="4" pageNumber="102">Gnathosoma</emphasis>
|
||
(
|
||
<figureCitation id="5FB62EF4DB585C369B29FEA635EE8288" box="[357,424,367,392]" captionStart="FIGURES 9 – 13" captionStartId="5.[151,264,1403,1426]" captionTargetBox="[174,1422,201,1378]" captionTargetId="figure@5.[151,1436,193,1382]" captionTargetPageId="5" captionText="FIGURES 9 – 13. Pachyseius quadrigeminus sp. nov., female. 9. Ventral gnathosoma; 10. Epistome; 11. Chelicera; 12. Sperm access system; 13. Tarsus II." httpUri="https://zenodo.org/record/236436/files/figure.png" pageId="4" pageNumber="102">Fig. 9</figureCitation>
|
||
). Subcapitulum with four pairs of setae, all smooth and needle-like,
|
||
<emphasis id="F5F9EE63DB585C369EECFEB930FA8288" box="[1184,1212,368,392]" italics="true" pageId="4" pageNumber="102">h3</emphasis>
|
||
longest (48–52),
|
||
<emphasis id="F5F9EE63DB585C369FCCFEB931DD8288" box="[1408,1435,368,392]" italics="true" pageId="4" pageNumber="102">h2</emphasis>
|
||
shortest (12–15), setae
|
||
<emphasis id="F5F9EE63DB585C369BE4FE5C358582AC" box="[424,451,405,428]" italics="true" pageId="4" pageNumber="102">pc</emphasis>
|
||
18–22 long,
|
||
<emphasis id="F5F9EE63DB585C369828FE5D363982AC" box="[612,639,404,428]" italics="true" pageId="4" pageNumber="102">
|
||
h
|
||
<date id="B33314B1DB585C36983FFE5D363982AC" box="[627,639,404,428]" pageId="4" pageNumber="102" value="1926-01-30">1</date>
|
||
</emphasis>
|
||
26–30 long. Deutosternum wide, with five long transverse rows of denticles. Corniculi horn-like. Palp with 3-tined apotele. Anterior and lateral margins of tongue-shaped epistome denticulate (
|
||
<figureCitation id="5FB62EF4DB585C369B6EFE15353182F4" box="[290,375,476,501]" captionStart="FIGURES 9 – 13" captionStartId="5.[151,264,1403,1426]" captionTargetBox="[174,1422,201,1378]" captionTargetId="figure@5.[151,1436,193,1382]" captionTargetPageId="5" captionText="FIGURES 9 – 13. Pachyseius quadrigeminus sp. nov., female. 9. Ventral gnathosoma; 10. Epistome; 11. Chelicera; 12. Sperm access system; 13. Tarsus II." httpUri="https://zenodo.org/record/236436/files/figure.png" pageId="4" pageNumber="102">Fig. 10</figureCitation>
|
||
). Cheliceral digits slender, movable digit 54–58 long, with two subdistal teeth and a terminal hook; arthrodial brush short; fixed digit 40–45 long (to dorsal seta), with a single submedial tooth and bidentate terminal hook; pilus dentilis not seen (
|
||
<figureCitation id="5FB62EF4DB585C369809FDED36D0813C" box="[581,662,548,573]" captionStart="FIGURES 9 – 13" captionStartId="5.[151,264,1403,1426]" captionTargetBox="[174,1422,201,1378]" captionTargetId="figure@5.[151,1436,193,1382]" captionTargetPageId="5" captionText="FIGURES 9 – 13. Pachyseius quadrigeminus sp. nov., female. 9. Ventral gnathosoma; 10. Epistome; 11. Chelicera; 12. Sperm access system; 13. Tarsus II." httpUri="https://zenodo.org/record/236436/files/figure.png" pageId="4" pageNumber="102">Fig. 11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="93F262F9DB585C369ADBF90C30C685DB" box="[151,1152,1733,1756]" httpUri="https://zenodo.org/record/236435/files/figure.png" pageId="4" pageNumber="102" targetBox="[165,1421,635,1704]" targetPageId="4">
|
||
<paragraph id="C7323271DB585C369ADBF90C30C685DB" blockId="4.[151,1152,1733,1756]" box="[151,1152,1733,1756]" pageId="4" pageNumber="102">
|
||
<emphasis id="F5F9EE63DB585C369ADBF90C357F85DB" bold="true" box="[151,313,1733,1756]" pageId="4" pageNumber="102">FIGURES 7–8.</emphasis>
|
||
<taxonomicName id="008D49F2DB585C369B0CF90F361685DB" box="[320,592,1734,1755]" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="102" phylum="Arthropoda" rank="species" species="quadrigeminus" status="sp. nov.">
|
||
<emphasis id="F5F9EE63DB585C369B0CF90F361685DB" box="[320,592,1734,1755]" italics="true" pageId="4" pageNumber="102">Pachyseius quadrigeminus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EECA5318DB585C36981BF90F36E185DB" box="[599,679,1734,1755]" pageId="4" pageNumber="102" rank="species">
|
||
<emphasis id="F5F9EE63DB585C36981BF90F36E185DB" bold="true" box="[599,679,1734,1755]" pageId="4" pageNumber="102">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, female. 7. Dorsal shield; 8. Ventral idiosoma.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C7323271DB585C369A8BF8CA30B4841B" blockId="4.[151,1436,1795,2036]" box="[199,1266,1795,1820]" pageId="4" pageNumber="102">
|
||
<taxonomicName id="008D49F2DB585C369A8BF8CA359C841B" box="[199,474,1795,1819]" class="Arachnida" family="Pachylaelapidae" genus="Spermathecal" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="102" phylum="Arthropoda" rank="species" species="apparatus">
|
||
<emphasis id="F5F9EE63DB585C369A8BF8CA359C841B" box="[199,474,1795,1819]" italics="true" pageId="4" pageNumber="102">Spermathecal apparatus</emphasis>
|
||
</taxonomicName>
|
||
. Spermathecal tubes long and fine, associated with coxae III (
|
||
<figureCitation id="5FB62EF4DB585C369EDDF8CA30A2841B" box="[1169,1252,1795,1820]" captionStart="FIGURES 9 – 13" captionStartId="5.[151,264,1403,1426]" captionTargetBox="[174,1422,201,1378]" captionTargetId="figure@5.[151,1436,193,1382]" captionTargetPageId="5" captionText="FIGURES 9 – 13. Pachyseius quadrigeminus sp. nov., female. 9. Ventral gnathosoma; 10. Epistome; 11. Chelicera; 12. Sperm access system; 13. Tarsus II." httpUri="https://zenodo.org/record/236436/files/figure.png" pageId="4" pageNumber="102">Fig. 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C7323271DB585C369A8BF8E1371D84F4" blockId="4.[151,1436,1795,2036]" pageId="4" pageNumber="102">
|
||
<emphasis id="F5F9EE63DB585C369A8BF8E134BA843F" box="[199,252,1832,1855]" italics="true" pageId="4" pageNumber="102">Legs</emphasis>
|
||
. Chaetotaxy: Leg I. Coxa 0 0/2 0, trochanter 1 1/3 1, femur 2 5/4 2, genu 2 5/3 2, tibia 2 5/3 2. Leg II. Coxa 0 0/2 0, trochanter 1 0/3 1, femur 2 5/3 1, genu 2 5/2 2, tibia 2 4/2 2, tarsus 3 7/5 3. Leg III. Coxa 0 0/2 0, trochanter 1 0/3 1, femur 1 3/1 1, genu 2 4/1 1, tibia 1 3/2 1, tarsus 3 7/5 3. Leg IV. Coxa 0 0/1 0, trochanter 1 0/3 1, femur 1 3/1 1, genu 2 4/1 1, tibia 1 3/2 1, tarsus 3 7/5 3. Tarsus II with two large spur like setae (
|
||
<figureCitation id="5FB62EF4DB585C369E9CF85A316584AB" box="[1232,1315,1939,1964]" captionStart="FIGURES 9 – 13" captionStartId="5.[151,264,1403,1426]" captionTargetBox="[174,1422,201,1378]" captionTargetId="figure@5.[151,1436,193,1382]" captionTargetPageId="5" captionText="FIGURES 9 – 13. Pachyseius quadrigeminus sp. nov., female. 9. Ventral gnathosoma; 10. Epistome; 11. Chelicera; 12. Sperm access system; 13. Tarsus II." httpUri="https://zenodo.org/record/236436/files/figure.png" pageId="4" pageNumber="102">Fig. 13</figureCitation>
|
||
).
|
||
<emphasis id="F5F9EE63DB585C369A8BF87F354D84CF" bold="true" box="[199,267,1974,1999]" pageId="4" pageNumber="102">Male.</emphasis>
|
||
<emphasis id="F5F9EE63DB585C369B5EF871352A84CF" box="[274,364,1976,1999]" italics="true" pageId="4" pageNumber="102">Dorsum</emphasis>
|
||
. Dorsal shield 510–558 long, 313–342 wide at widest point, oblong (length/width, 1.59–1.66), ornamented with reticulate pattern. Other features as in female.
|
||
</paragraph>
|
||
<caption id="93F262F9DB595C379ADBFAB235F186B0" httpUri="https://zenodo.org/record/236436/files/figure.png" pageId="5" pageNumber="103" targetBox="[174,1422,201,1378]" targetPageId="5">
|
||
<paragraph id="C7323271DB595C379ADBFAB235F186B0" blockId="5.[151,1436,1403,1456]" pageId="5" pageNumber="103">
|
||
<emphasis id="F5F9EE63DB595C379ADBFAB235018691" bold="true" box="[151,327,1403,1426]" pageId="5" pageNumber="103">FIGURES 9–13.</emphasis>
|
||
<taxonomicName id="008D49F2DB595C379B02FAB536198691" box="[334,607,1404,1425]" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="103" phylum="Arthropoda" rank="species" species="quadrigeminus" status="sp. nov.">
|
||
<emphasis id="F5F9EE63DB595C379B02FAB536198691" box="[334,607,1404,1425]" italics="true" pageId="5" pageNumber="103">Pachyseius quadrigeminus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EECA5318DB595C37982AFAB536F18691" box="[614,695,1404,1425]" pageId="5" pageNumber="103" rank="species">
|
||
<emphasis id="F5F9EE63DB595C37982AFAB536F18691" bold="true" box="[614,695,1404,1425]" pageId="5" pageNumber="103">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, female. 9. Ventral gnathosoma; 10. Epistome; 11. Chelicera; 12. Sperm access system; 13. Tarsus II.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C7323271DB595C379A8BFA1036E28580" blockId="5.[151,1437,1495,2024]" pageId="5" pageNumber="103">
|
||
<emphasis id="F5F9EE63DB595C379A8BFA10354A86F0" box="[199,268,1497,1520]" italics="true" pageId="5" pageNumber="103">Venter</emphasis>
|
||
(
|
||
<figureCitation id="5FB62EF4DB595C379B51FA1E353686F0" box="[285,368,1495,1520]" captionStart="FIGURES 14 – 18" captionStartId="6.[151,264,1547,1570]" captionTargetBox="[162,1432,351,1514]" captionTargetId="figure@6.[151,1436,339,1526]" captionTargetPageId="6" captionText="FIGURES 14 – 18. Pachyseius quadrigeminus sp. nov., male. 14. Ventral idiosoma; 15. Epistome; 16. Chelicera; 17. Ventral spurs on leg II (fem. = femur, gen. = genu, tib. = tibia); 18. Telotarsus II." httpUri="https://zenodo.org/record/236437/files/figure.png" pageId="5" pageNumber="103">Fig. 14</figureCitation>
|
||
). Holoventral shield present, 423–464 long, 80–85 wide at level of coxae II; sternal region with punctate-reticulate ornamentation, and bearing five pairs of needle-like setae and three pairs of lyrifissures; opisthogastric portion of shield ornamented with punctate polygonal lines, with four pairs of setae (15–28); anal portion with a pair of para-anal setae (25–30) and longer post-anal seta (35–40). Peritrematal shields connected to holoventral shield. Other features as in female.
|
||
</paragraph>
|
||
<paragraph id="C7323271DB595C379A8BF945361F8410" blockId="5.[151,1437,1495,2024]" pageId="5" pageNumber="103">
|
||
<emphasis id="F5F9EE63DB595C379A8BF945351085A4" box="[199,342,1676,1700]" italics="true" pageId="5" pageNumber="103">Gnathosoma</emphasis>
|
||
. Epistome triangular, with denticulate anterior margin (
|
||
<figureCitation id="5FB62EF4DB595C37998EF942305385A4" box="[962,1045,1675,1700]" captionStart="FIGURES 14 – 18" captionStartId="6.[151,264,1547,1570]" captionTargetBox="[162,1432,351,1514]" captionTargetId="figure@6.[151,1436,339,1526]" captionTargetPageId="6" captionText="FIGURES 14 – 18. Pachyseius quadrigeminus sp. nov., male. 14. Ventral idiosoma; 15. Epistome; 16. Chelicera; 17. Ventral spurs on leg II (fem. = femur, gen. = genu, tib. = tibia); 18. Telotarsus II." httpUri="https://zenodo.org/record/236437/files/figure.png" pageId="5" pageNumber="103">Fig. 15</figureCitation>
|
||
). Movable digit of chelicera 60–65 long, with a larger subterminal tooth and bidentate terminal hook; spermatodactyl directed posteriorly, 90–110 long, and distally narrowed; fixed digit about 46 long (to dorsal seta), slender, edentate and with a minute pilus dentilis; arthrodial brush short (
|
||
<figureCitation id="5FB62EF4DB595C379BB4F93E360A8410" box="[504,588,1783,1808]" captionStart="FIGURES 14 – 18" captionStartId="6.[151,264,1547,1570]" captionTargetBox="[162,1432,351,1514]" captionTargetId="figure@6.[151,1436,339,1526]" captionTargetPageId="6" captionText="FIGURES 14 – 18. Pachyseius quadrigeminus sp. nov., male. 14. Ventral idiosoma; 15. Epistome; 16. Chelicera; 17. Ventral spurs on leg II (fem. = femur, gen. = genu, tib. = tibia); 18. Telotarsus II." httpUri="https://zenodo.org/record/236437/files/figure.png" pageId="5" pageNumber="103">Fig. 16</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C7323271DB595C379A8BF8D536868458" blockId="5.[151,1437,1495,2024]" pageId="5" pageNumber="103">
|
||
<emphasis id="F5F9EE63DB595C379A8BF8D535458433" box="[199,259,1820,1843]" italics="true" pageId="5" pageNumber="103">Legs.</emphasis>
|
||
Femur II with well-developed ventral spur, genu II and tibia II each with smaller ventral spur (
|
||
<figureCitation id="5FB62EF4DB595C379F76F8D231C88434" box="[1338,1422,1819,1844]" captionStart="FIGURES 14 – 18" captionStartId="6.[151,264,1547,1570]" captionTargetBox="[162,1432,351,1514]" captionTargetId="figure@6.[151,1436,339,1526]" captionTargetPageId="6" captionText="FIGURES 14 – 18. Pachyseius quadrigeminus sp. nov., male. 14. Ventral idiosoma; 15. Epistome; 16. Chelicera; 17. Ventral spurs on leg II (fem. = femur, gen. = genu, tib. = tibia); 18. Telotarsus II." httpUri="https://zenodo.org/record/236437/files/figure.png" pageId="5" pageNumber="103">Fig. 17</figureCitation>
|
||
); tarsus II with two spur-like dorsal setae (
|
||
<figureCitation id="5FB62EF4DB595C379813F8F636F58458" box="[607,691,1855,1880]" captionStart="FIGURES 14 – 18" captionStartId="6.[151,264,1547,1570]" captionTargetBox="[162,1432,351,1514]" captionTargetId="figure@6.[151,1436,339,1526]" captionTargetPageId="6" captionText="FIGURES 14 – 18. Pachyseius quadrigeminus sp. nov., male. 14. Ventral idiosoma; 15. Epistome; 16. Chelicera; 17. Ventral spurs on leg II (fem. = femur, gen. = genu, tib. = tibia); 18. Telotarsus II." httpUri="https://zenodo.org/record/236437/files/figure.png" pageId="5" pageNumber="103">Fig. 18</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8F9761FADB595C349A8BF8AA3788821D" lastPageId="6" lastPageNumber="104" pageId="5" pageNumber="103" type="etymology">
|
||
<paragraph id="C7323271DB595C379A8BF8AA3636849F" blockId="5.[151,1437,1495,2024]" pageId="5" pageNumber="103">
|
||
<emphasis id="F5F9EE63DB595C379A8BF8AA3508847C" bold="true" box="[199,334,1891,1916]" pageId="5" pageNumber="103">Etymology.</emphasis>
|
||
The name
|
||
<taxonomicName id="008D49F2DB595C379B85F8AD3637847C" box="[457,625,1892,1916]" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="103" phylum="Arthropoda" rank="species" species="quadrigeminus">
|
||
<emphasis id="F5F9EE63DB595C379B85F8AD3637847C" box="[457,625,1892,1916]" italics="true" pageId="5" pageNumber="103">quadrigeminus</emphasis>
|
||
</taxonomicName>
|
||
, four pairs, refers to the presence of four pairs of setae on the ventri-anal shield, in addition to the circumanal setae.
|
||
</paragraph>
|
||
<paragraph id="C7323271DB595C379A8BF86231DB84E8" blockId="5.[151,1437,1495,2024]" pageId="5" pageNumber="103">
|
||
<emphasis id="F5F9EE63DB595C379A8BF862355784C4" bold="true" box="[199,273,1963,1988]" pageId="5" pageNumber="103">Notes.</emphasis>
|
||
The new species is easily distinguishable from the other congeners by the ventri-anal shield with four pairs of pre-anal setae, peritrematal shields fused with exopodal shields, and the male with a holoventral shield.
|
||
</paragraph>
|
||
<paragraph id="C7323271DB5A5C349ADBFF5E3788821D" blockId="6.[151,1437,151,285]" pageId="6" pageNumber="104">
|
||
The ventri-anal shield in
|
||
<taxonomicName id="008D49F2DB5A5C349BFCFF51366883B0" box="[432,558,152,176]" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="104" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F5F9EE63DB5A5C349BFCFF51366883B0" box="[432,558,152,176]" italics="true" pageId="6" pageNumber="104">Pachyseius</emphasis>
|
||
</taxonomicName>
|
||
usually has two or three pairs of pre-anal setae. Only
|
||
<taxonomicName id="008D49F2DB5A5C349EC0FF5030B083AF" box="[1164,1270,152,176]" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="104" phylum="Arthropoda" rank="species" species="sinicus">
|
||
<emphasis id="F5F9EE63DB5A5C349EC0FF5030DC83B0" box="[1164,1178,153,176]" italics="true" pageId="6" pageNumber="104">P</emphasis>
|
||
.
|
||
<emphasis id="F5F9EE63DB5A5C349EE4FF5130B083AF" box="[1192,1270,152,175]" italics="true" pageId="6" pageNumber="104">sinicus</emphasis>
|
||
</taxonomicName>
|
||
has four pairs, but according to
|
||
<bibRefCitation id="A31C4F80DB5A5C349B19FF75363183D4" author="Ozbek" box="[341,631,188,213]" pageId="6" pageNumber="104" refString="Ozbek, H. H. & Halliday, B. (2014) Two new species of Pachyseius Berlese (Acari: Pachylaelapidae) from Turkey, with a key to the world species. Zootaxa, 3841 (1), 107 - 116. http: // dx. doi. org / 10.11646 / zootaxa. 3841.1.5" type="journal article" year="2014">Özbek & Halliday (2014)</bibRefCitation>
|
||
, two of these pairs are on the extreme lateral edges of the shield, where their positions may show intra-specific or artificial variation. All of our females of
|
||
<taxonomicName id="008D49F2DB5A5C349E7BFF2830BD83F8" box="[1079,1275,224,248]" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="104" phylum="Arthropoda" rank="species" species="quadrigeminus" status="sp. nov.">
|
||
<emphasis id="F5F9EE63DB5A5C349E7BFF28300383F8" box="[1079,1093,225,248]" italics="true" pageId="6" pageNumber="104">P</emphasis>
|
||
.
|
||
<emphasis id="F5F9EE63DB5A5C349E1FFF2930BD83F8" box="[1107,1275,224,248]" italics="true" pageId="6" pageNumber="104">quadrigeminus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EECA5318DB5A5C349F4EFF16311D83F8" box="[1282,1371,223,248]" pageId="6" pageNumber="104" rank="species">
|
||
<emphasis id="F5F9EE63DB5A5C349F4EFF16311D83F8" bold="true" box="[1282,1371,223,248]" pageId="6" pageNumber="104">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
carry four pairs of setae, and they are well inside the lateral edges of the shield.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |