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<document ID-CLB-Dataset="257398" ID-DOI="http://dx.doi.org/10.3897/asp.81.e102404" ID-GBIF-Dataset="b1c3ca9e-6c0d-4869-86b3-13da55d2d43d" ID-Pensoft-Pub="1864-8312-81-439" ID-Pensoft-UUID="ACB315A2B53250ACA5FC528F6F3916CC" ID-ZooBank="879AE99E69874A83B10FE38BF7D545BF" ModsDocID="1864-8312-81-439" checkinTime="1682710844061" checkinUser="pensoft" docAuthor="Simon Prazak, Jan, Fikacek, Martin, Prokop, Jakub &amp; Lackner, Tomas" docDate="2023" docId="4E1C441CB8C25040AF7AC8A471E329FB" docLanguage="en" docName="ArthropodSystPhyl 81: 439-453" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 81" docPubDate="2023-04-28" docSource="http://dx.doi.org/10.3897/asp.81.e102404" docTitle="Cretanapleus seideli Simon Prazak &amp; Lackner 2023, sp. nov." docType="treatment" docUuid="1E124B42-BF71-4DA4-9171-60B147356678" docUuidSource="ZooBank" docVersion="5" id="ACB315A2B53250ACA5FC528F6F3916CC" lastPageNumber="439" masterDocId="ACB315A2B53250ACA5FC528F6F3916CC" masterDocTitle="Under the Cretaceous bark: Fossil evidence for the ancient origin of subcortical lifestyle of clown beetles (Coleoptera: Histeridae)" masterLastPageNumber="453" masterPageNumber="439" pageNumber="439" updateTime="1732751914251" updateUser="ExternalLinkService">
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<mods:title id="D0FBE55285A42ECE3DA64952BAB3603B">Under the Cretaceous bark: Fossil evidence for the ancient origin of subcortical lifestyle of clown beetles (Coleoptera: Histeridae)</mods:title>
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<mods:name id="BBE7574E6675E7AFF83EFEB5D446C04F" type="personal">
<mods:role id="A16A035FCD5FEAA39A6B6E5DB529CE1F">
<mods:roleTerm id="11A43ADAC0123CC0FF3F66B1DCE2C72E">Author</mods:roleTerm>
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<mods:namePart id="6C848C3C609995BAAC1FE704B1568B9F">Simon Prazak, Jan</mods:namePart>
<mods:nameIdentifier id="BC294E4CAB3CF79ACEE32AC98DD0347F" type="ORCID">https://orcid.org/0000-0001-7438-2548</mods:nameIdentifier>
<mods:affiliation id="1A2E8EFB0E6826F9C09C6ED7FF9898AA">Charles University, Faculty of Science, Department of Zoology, Vinicna 7, 12800 Prague 2, Czech Republic &amp; Museum of Eastern Bohemia in Hradec Kralove, Eliscino nabrezi 465, 500 03 Hradec Kralove 3, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="DD78FB7B845DBD406FA494E652123987" type="email">Honza.prazak@email.cz</mods:nameIdentifier>
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<mods:namePart id="2B00FB8D069A9BA8008E790C0472B9C3">Fikacek, Martin</mods:namePart>
<mods:nameIdentifier id="84005D1F38EEF2A8A79575CB5BBD14FB" type="ORCID">https://orcid.org/0000-0002-2078-6798</mods:nameIdentifier>
<mods:affiliation id="82EB94862A5867082EF6A057BFC6A5D2">Department of Biological Sciences, National Sun Yat-sen University, No. 70, Lienhai Rd., Kaohsiung 80424, Taiwan &amp; Department of Entomology, National Museum, Cirkusova 1740, Praha-Horni Pocernice, Czech Republic</mods:affiliation>
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<mods:namePart id="69DBC77E043F40CDC0AF93E2DBB2BA81">Prokop, Jakub</mods:namePart>
<mods:nameIdentifier id="D99FD46046E04C339C96C6CAB3DD37C4" type="ORCID">https://orcid.org/0000-0001-6996-7832</mods:nameIdentifier>
<mods:affiliation id="106F653621D181E656BF399CD5B4F90D">Charles University, Faculty of Science, Department of Zoology, Vinicna 7, 12800 Prague 2, Czech Republic</mods:affiliation>
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<mods:namePart id="663AA2F909A480775FD60A2AC3BF60F8">Lackner, Tomas</mods:namePart>
<mods:nameIdentifier id="7F08DF8B57C1BE538638F04320B8E9CE" type="ORCID">https://orcid.org/0000-0002-0108-5785</mods:nameIdentifier>
<mods:affiliation id="8C0932EA1E70676B7FE8E4340C31F08F">Bavarian State Collection of Zoology, Muenchhausenstrasse 21, 81247 Munich, Germany</mods:affiliation>
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<mods:title id="080400483BA5990D6731E74E7DEC21FF">Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:part id="B17C70FB5D40A8ABFDA5F60B475EE5E6">
<mods:date id="36D3974AD7AB8E6990E5854A767FBCD2">2023</mods:date>
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<mods:number id="2002C93F31A53F2363897083E5EDF160">2023-04-28</mods:number>
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<mods:number id="D954EB8C84647FF6CDCE24391175B7AF">81</mods:number>
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<mods:start id="8C93C28A094DE02A7B73C04470FFB720">439</mods:start>
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<mods:classification id="16B66DB8522A524A3877F828C58B31FF">journal article</mods:classification>
<mods:identifier id="57C5F3ABD62BF6CEA76230844863D1B3" type="DOI">http://dx.doi.org/10.3897/asp.81.e102404</mods:identifier>
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<treatment id="4E1C441CB8C25040AF7AC8A471E329FB" ID-GBIF-Taxon="209579172" LSID="urn:lsid:zoobank.org:act:1E124B42-BF71-4DA4-9171-60B147356678" httpUri="http://treatment.plazi.org/id/4E1C441CB8C25040AF7AC8A471E329FB" lastPageNumber="439" pageId="0" pageNumber="439" scope_family="Histeridae" scope_order="Coleoptera" scope_subFamily="Histerinae">
<subSubSection id="D750405CD7F58C80264FA8ACC2E6B5E3" pageId="0" pageNumber="439" type="nomenclature">
<paragraph id="0D0DEDC33B9464DBCA22C1FE8E8A9E79" pageId="0" pageNumber="439">
<taxonomicName id="AE7AB0F10B2B196A032E5849B20B1CDA" LSID="https://zoobank.org/1E124B42-BF71-4DA4-9171-60B147356678" authority="Simon Pražák &amp; Lackner" authorityName="Simon Prazak &amp; Lackner" authorityYear="2023" class="Insecta" family="Histeridae" genus="Cretanapleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cretanapleus seideli" order="Coleoptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="seideli" status="sp. nov.">
Cretanapleus seideli Simon
<normalizedToken id="1FCFFAA18B1E895F43F050EA13506D02" originalValue="Pražák">Prazak</normalizedToken>
&amp; Lackner
</taxonomicName>
<taxonomicNameLabel id="3C30796AA2560332063B5E621D9386DE" pageId="0" pageNumber="439">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="23894E71A76FF3BEC0D3CC7816F8294F" pageId="0" pageNumber="439" type="type material">
<paragraph id="90DF30CEA4451E865614CADCA1DA49AA" pageId="0" pageNumber="439">Type material.</paragraph>
<paragraph id="EF9AD7A4349EA3B4BCE426F295B28B95" pageId="0" pageNumber="439">
Holotype specimen (1409/E), unknown sex, Northern Myanmar, inclusion in burmite (ca. 99 Ma), amber piece clear, 15
<normalizedToken id="10BCB49460301225C35E9CBDBC2BD936" originalValue="×6×">x6x</normalizedToken>
2 mm, with small pieces of debris.
</paragraph>
</subSubSection>
<subSubSection id="DA9C0C642DCBB117B6741CE6A34567DE" pageId="0" pageNumber="439" type="description">
<paragraph id="4320E5B7535A3D6A5C623CE5F7B5C79B" pageId="0" pageNumber="439">Description.</paragraph>
<paragraph id="BB2C9570C4946F9DE5AC48A4B4CB8689" pageId="0" pageNumber="439">
<emphasis id="64AF836120AF7070F19B46AC076CD7B5" bold="true" italics="true" pageId="0" pageNumber="439">Measurements.</emphasis>
Head width: 0.14 mm, width between anterior pronotal angles: 0.175 mm, width between posterior pronotal angles: 0.4 mm, pronotal length: 0.21 mm, elytral length: 0.38 mm, elytral width (across widest point): 0.46 mm. -
<emphasis id="9E73E4858E6F34CC1890EB3D90260EF2" bold="true" italics="true" pageId="0" pageNumber="439">Body</emphasis>
regularly round, convex, cuticle brown to black, covered with white debris coating. Body margins dorsally explanate. Elytra with punctures (diameter 10 microns) separated by their diameter. Pronotum with irregular weak longitudinal rugae. -
<emphasis id="0D6825C76837C7048B573B8D6E2E44D0" bold="true" italics="true" pageId="0" pageNumber="439">Head.</emphasis>
Frontoclypeal area laterally depressed, almost constricted. Frontal stria carinate, widely interrupted medially, slightly continuous to clypeus. Frontal surface laterad of frontal stria forming a depression fit to receive antennal scape in repose, pressed against outer margin of frontal stria. Clypeus anteriorly elevated with large punctures. Labrum dorsally with depressions, with a pair of long prominent setae. Right mandible bidentate with a prominent acute subapical tooth, inner margin of left mandible not visible. Terminal maxillary palpomere elongate, apically acute, slightly flattened. Antennal scape long and robust. Shape of the scape coincident with frontal carina. Pedicel as long as antennomeres 3-6 combined. Antennomeres 3-8 short. Antennal club round, with two apparent straight annuli. Eyes not visible. -
<emphasis id="E5AD4462CC2206A419E897920ADA96A2" bold="true" italics="true" pageId="0" pageNumber="439">Thorax.</emphasis>
Pronotal margins narrowing anteriorly; anterior angles acute. Lateral and anterior margins explanate. Median part of frontal disc punctuate, convex with irregular longitudinal rugae. Posterior margin subungulate. Scutellum minute, triangular. Elytra apically truncate. Lateral and basal elytral margins distinctly explanate. Elytral suture elevated. Elytra punctuate at least medially (punctures separated by their diameter); rest of elytral surface covered with debris. Dorsal striation not discernible. Elytral epipleuron unusually wide with two protuberant costae on basal half. Epipleuron largely conceals metepisternum. Prosternal process rectangular, medially depressed. Prosternal lobe short with distinct margin and with antennal fissures laterally. Apex of prosternal lobe keel-like, elevated. Antennal cavity located near procoxa, deep, closed anteriorly. Mesoventrite transverse, 4x wider than long. Meso-metaventral suture invisible. Meso-metaventral stria carinate, anteriorly arcuate. Anterolateral corners with distinct round depressions. Metaventrite with delicate sparse punctures. Lateral metaventral stria complete, carinate. Metaventral disc convex. Metepisternum largely concealed by elytral epipleuron, with rugose punctuation. -
<emphasis id="BD060B5F9055BC36664409CE4DB093B0" bold="true" italics="true" pageId="0" pageNumber="439">Abdomen.</emphasis>
First visible abdominal sternite with large punctures separated by their diameter. Pygidium with similarly patterned punctures. Other abdominal segments inflexed and covered with debris, unobservable. -
<emphasis id="AC77F31E4E418BAC460DE6734B80E903" bold="true" italics="true" pageId="0" pageNumber="439">Legs.</emphasis>
Profemur with a weakly developed groove for protibial reception. Protibia moderately broad, with a costa along inner margin and slightly thickened along the outer margin, creating a tarsal groove. Outer protibial margin with a dense row of short setae diminishing in size basally, a row of short slender setae also present on the inner margin. Protibial spur tiny. Terminal protarsomere as long as tarsomeres 2-4 combined, claws 1/3 of terminal tarsomere length. Mesotibia slenderer, with a row of short slender setae both on outer and inner margin. Mesotibia with a shallow groove for mesofemoral reception. Mesotarsus similar to protarsus. Metafemur with a groove for metatibial reception, otherwise hind leg identical to middle leg.
</paragraph>
</subSubSection>
<subSubSection id="5A0B302C0CAE9840EA77BB1E3C86E1EF" pageId="0" pageNumber="439" type="taxonomic assignment">
<paragraph id="573059A56E6DCD1097713DDD630671B5" pageId="0" pageNumber="439">Taxonomic assignment.</paragraph>
<paragraph id="3BBC6E73C18B1EC62FA2DC1F82995244" pageId="0" pageNumber="439">
<taxonomicName id="A47CF5C1247E8B76B4561C5DC84EBF42" authorityName="Simon Prazak &amp; Lackner" authorityYear="2023" class="Insecta" family="Histeridae" genus="Cretanapleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cretanapleus seideli" order="Coleoptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="seideli">
<emphasis id="DED0F981577221325BBF7AB2CCCD36A9" italics="true" pageId="0" pageNumber="439">Cretanapleus seideli</emphasis>
</taxonomicName>
can be clearly placed in the
<taxonomicName id="2957DAF0353CC441568A33B2CDF95FD3" authorityName="Reitter" authorityYear="1909" lsidName="" pageId="0" pageNumber="439" rank="subFamily" subFamily="Dendrophilinae">Dendrophilinae</taxonomicName>
based on the prosternum with a short prosternal lobe with antennal fissures laterally. We further place this species in the monophyletic tribe
<taxonomicName id="05903BFA04930CE66002410EF6A3A6CE" authorityName="Olexa" authorityYear="1982" lsidName="" pageId="0" pageNumber="439" rank="tribe" tribe="Anapleini">Anapleini</taxonomicName>
(
<bibRefCitation id="3819C4B7332BB1C2C1C7B6A45845FC9B" DOI="https://doi.org/10.1111/cla.12418" author="Zhou, YL" journalOrPublisher="Cladistics" pageId="0" pageNumber="439" pagination="521 - 539" refId="B34" refString="Zhou, YL, Caterino, MS, Ren, D, Ślipinski, A, 2020. Phylogeny and evolution of Mesozoic and extant lineages of Histeridae (Coleoptera), with discovery of a new subfamily Antigracilinae from the Lower Cretaceous. Cladistics 36 (5): 521 - 539, DOI: https://doi.org/10.1111/cla.12418" title="Phylogeny and evolution of Mesozoic and extant lineages of Histeridae (Coleoptera), with discovery of a new subfamily Antigracilinae from the Lower Cretaceous." url="https://doi.org/10.1111/cla.12418" volume="36" year="2020">Zhou et al. 2020</bibRefCitation>
), as defined by
<bibRefCitation id="A5D23D18DAD6083EF16FA779A0909B34" author="Olexa, A" journalOrPublisher="Acta entomologica bohemoslovaca" pageId="0" pageNumber="439" pagination="113 - 131" refId="B26" refString="Olexa, A, 1984. Revision der Arten der Gattung Pholioxenus Reichardt (Coleoptera, Histeridae). Acta entomologica bohemoslovaca 81 (2): 113 - 131" title="Revision der Arten der Gattung Pholioxenus Reichardt (Coleoptera, Histeridae)." volume="81" year="1984">Olexa (1984)</bibRefCitation>
, based on protibial characters: protarsal groove with distinctly elevated margins, outer protibial margin simple, devoid of teeth or denticles, with a simple row of setae (Fig.
<figureCitation id="C10C4EC2CE9BF0FC771F2B58E78CC585" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cretanapleus seideli Simon Prazak &amp; Lackner, gen. &amp; sp. nov. (A, B, C, E, G) and Anapleus sp. (Turkey; D, F, H; SEM micrographs). A dorsal view; B amber specimen; C - D protibia; E - F ventral view; G - H frontal view." figureDoi="10.3897/asp.81.e102404.figure3" httpUri="https://binary.pensoft.net/fig/843422" pageId="0" pageNumber="439">3C, D</figureCitation>
). However,
<taxonomicName id="03B2F497CCBBC59142B02976242A4F16" lsidName="C. seideli" pageId="0" pageNumber="439" rank="species" species="seideli">
<emphasis id="45641357F9CC17014F9493B78DA24F23" italics="true" pageId="0" pageNumber="439">C. seideli</emphasis>
</taxonomicName>
differs from the only extant genus of the tribe,
<taxonomicName id="7CE064D01B6A886FEBE6E14636D26712" authorityName="Horn" authorityYear="1873" class="Insecta" family="Histeridae" genus="Anapleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anapleus" order="Coleoptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
<emphasis id="9E72C64A6BC739ED24C7BA0AED6B4FF7" italics="true" pageId="0" pageNumber="439">Anapleus</emphasis>
</taxonomicName>
Horn, 1873, in several significant characters: presence of frontal stria (absent in
<taxonomicName id="EEF7ACF84094874DCD3B2EDDADE1D3B1" authorityName="Horn" authorityYear="1873" class="Insecta" family="Histeridae" genus="Anapleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anapleus" order="Coleoptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
<emphasis id="EF28BFCDCF84C57B88232B939D57582F" italics="true" pageId="0" pageNumber="439">Anapleus</emphasis>
</taxonomicName>
, Fig.
<figureCitation id="72FE35DFC10E6FC44BA5EF0BC1C83157" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cretanapleus seideli Simon Prazak &amp; Lackner, gen. &amp; sp. nov. (A, B, C, E, G) and Anapleus sp. (Turkey; D, F, H; SEM micrographs). A dorsal view; B amber specimen; C - D protibia; E - F ventral view; G - H frontal view." figureDoi="10.3897/asp.81.e102404.figure3" httpUri="https://binary.pensoft.net/fig/843422" pageId="0" pageNumber="439">3G, H</figureCitation>
), deep and closed antennal cavity (shallow and widely open in
<taxonomicName id="EC5200213682505BBB02686522128011" authorityName="Horn" authorityYear="1873" class="Insecta" family="Histeridae" genus="Anapleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anapleus" order="Coleoptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
<emphasis id="3B0EC9F1B7C1E3D22D5FF4B8454B3641" italics="true" pageId="0" pageNumber="439">Anapleus</emphasis>
</taxonomicName>
), and strongly emarginated lateral body margins (Fig.
<figureCitation id="937F46609999228F78CFBE69F05FEF21" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cretanapleus seideli Simon Prazak &amp; Lackner, gen. &amp; sp. nov. (A, B, C, E, G) and Anapleus sp. (Turkey; D, F, H; SEM micrographs). A dorsal view; B amber specimen; C - D protibia; E - F ventral view; G - H frontal view." figureDoi="10.3897/asp.81.e102404.figure3" httpUri="https://binary.pensoft.net/fig/843422" pageId="0" pageNumber="439">3G</figureCitation>
, rounded in
<taxonomicName id="B974159F1ACD6F9B3C5AA7230FB95032" authorityName="Horn" authorityYear="1873" class="Insecta" family="Histeridae" genus="Anapleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anapleus" order="Coleoptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
<emphasis id="613C9AFE9FF0885F1A642F256893E808" italics="true" pageId="0" pageNumber="439">Anapleus</emphasis>
</taxonomicName>
). Our phylogenetic analyses confirmed the placement of
<taxonomicName id="3469F6CB418023469E6F4C994040B212" lsidName="C. seideli" pageId="0" pageNumber="439" rank="species" species="seideli">
<emphasis id="5B88B95C00BBB3962CF54C2752EC0B1B" italics="true" pageId="0" pageNumber="439">C. seideli</emphasis>
</taxonomicName>
in
<taxonomicName id="707D90A908F393F00CAE84D2BE72BC34" authorityName="Olexa" authorityYear="1982" lsidName="" pageId="0" pageNumber="439" rank="tribe" tribe="Anapleini">Anapleini</taxonomicName>
(Fig.
<figureCitation id="162CDB2EBA709B3E4FD7220B130168B0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Maximum parsimony analysis of the position of known fossil species of the Histeridae. A the complete tree showing the position of the fossils and the current subfamily classification of the lineages; B Dendrophilinae part of the tree with mapped characters; C Histerinae part of the tree with mapped characters. Bold taxa = fossils, red taxa = fossils described in this paper. For characters see Zhou et al. (2020)." figureDoi="10.3897/asp.81.e102404.figure5" httpUri="https://binary.pensoft.net/fig/843424" pageId="0" pageNumber="439">5</figureCitation>
).
</paragraph>
<caption id="1E984B9B02FD0FC7014D84AE31C59410" doi="10.3897/asp.81.e102404.figure3" httpUri="https://binary.pensoft.net/fig/843422" pageId="0" pageNumber="439" start="Figure 3" startId="F3">
<paragraph id="14CF70F38C3684358A83EC55B93F65AC" pageId="0" pageNumber="439">
<emphasis id="A1CF2F5379348A2E41F9719C008EA98F" bold="true" pageId="0" pageNumber="439">Figure 3.</emphasis>
<taxonomicName id="AB8AACE3A50EE056735EC0B1E455979A" authorityName="Simon Prazak &amp; Lackner" authorityYear="2023" class="Insecta" family="Histeridae" genus="Cretanapleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cretanapleus seideli" order="Coleoptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="seideli">
<emphasis id="A445D1EB99020BA29B3CC8ADAFD4A028" italics="true" pageId="0" pageNumber="439">Cretanapleus seideli</emphasis>
</taxonomicName>
Simon
<normalizedToken id="2BA3B0F5369EA947E40941047032217C" originalValue="Pražák">Prazak</normalizedToken>
&amp; Lackner, gen. &amp; sp. nov. (A, B, C, E, G) and
<taxonomicName id="BB748007A4B86216930FC9679AE49307" authorityName="Horn" authorityYear="1873" class="Insecta" family="Histeridae" genus="Anapleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anapleus" order="Coleoptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
<emphasis id="5FADC88A6A84E78E517748EAD0E2282C" italics="true" pageId="0" pageNumber="439">Anapleus</emphasis>
</taxonomicName>
sp. (Turkey; D, F, H; SEM micrographs).
<emphasis id="29F5977774A9BF6F59902C964803034B" bold="true" pageId="0" pageNumber="439">A</emphasis>
dorsal view;
<emphasis id="4585C077BF60A07A0BD8CB172CCCE079" bold="true" pageId="0" pageNumber="439">B</emphasis>
amber specimen;
<emphasis id="5EA6DF2A2A54D4200C21AE4CAD9876E8" bold="true" pageId="0" pageNumber="439">C</emphasis>
-
<emphasis id="A4E6E2F77C1ADF1C42124346B2E1294F" bold="true" pageId="0" pageNumber="439">D</emphasis>
protibia;
<emphasis id="42944EB3BB8F253C3A298209C7B121D6" bold="true" pageId="0" pageNumber="439">E</emphasis>
-
<emphasis id="E204E8D9715519256E23402CE20CE9E1" bold="true" pageId="0" pageNumber="439">F</emphasis>
ventral view;
<emphasis id="686EDC3C5765F7BBEEB9C7B4C24E1B43" bold="true" pageId="0" pageNumber="439">G</emphasis>
-
<emphasis id="A0784808D59D1A7F86E68495238042DF" bold="true" pageId="0" pageNumber="439">H</emphasis>
frontal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="0CAA7F19F24B2FA69303B0AA82D44405" pageId="0" pageNumber="439" type="etymology">
<paragraph id="204460266E1011C3F075630518AF6B18" pageId="0" pageNumber="439">Etymology.</paragraph>
<paragraph id="AB4F41319AA78C329CB44CE6BA07E53A" pageId="0" pageNumber="439">We dedicate the species to Dr. Matthias Seidel (Naturhistorisches Museum, Vienna, Austria) who introduced the first author of this paper to the study of fossil beetle inclusions.</paragraph>
<paragraph id="5CCDE23C1C6714B2D4DDBBC7BCEB821A" pageId="0" pageNumber="439">
<emphasis id="DC65BC3D2E9165D3C9332D3AA7887C1E" bold="true" pageId="0" pageNumber="439">Without tribal placement</emphasis>
</paragraph>
<paragraph id="22FCCAC232B8F1214226DE936500827E" pageId="0" pageNumber="439">
<emphasis id="68049FD54BDD5340E6C7479BF7D76816" bold="true" pageId="0" pageNumber="439">
Genus
<taxonomicName id="C30E1D5993675A6C6C9ACA8765D76FEE" authorityName="Caterino" authorityYear="2021" class="Insecta" family="Histeridae" genus="Yethiha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Yethiha" order="Coleoptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="genus">
<emphasis id="2F832AF886044F542ED05CC225DEA8F2" italics="true" pageId="0" pageNumber="439">Yethiha</emphasis>
</taxonomicName>
Caterino, 2021
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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