404 lines
36 KiB
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404 lines
36 KiB
XML
<document ID-CLB-Dataset="55415" ID-DOI="http://dx.doi.org/10.3897/zookeys.1176.106224" ID-GBIF-Dataset="7bb917e2-6ad1-4f3f-a1ae-df3feeb99862" ID-PMC="PMC10468690" ID-Pensoft-Pub="1313-2970-1176-165" ID-Pensoft-UUID="C538BFA63A3455D089E0A07BC22A97C6" ID-PubMed="37664866" ID-ZooBank="91E50336528C43DE826642D12D0C7C1C" ModsDocID="1313-2970-1176-165" checkinTime="1692810056931" checkinUser="pensoft" docAuthor="Khan, Eid Muhammad, Kamran, Muhammad, Mirza, Jawwad Hassan & Alatawi, Fahad Jaber" docDate="2023" docId="4E0A4286D8715265AAD699D360ED0DEF" docLanguage="en" docName="ZooKeys 1176: 165-180" docOrigin="ZooKeys 1176" docPubDate="2023-08-23" docSource="http://dx.doi.org/10.3897/zookeys.1176.106224" docTitle="Raphignathus (Diraphignathus) neohecmatanaensis sp. nov." docType="treatment" docVersion="4" id="C538BFA63A3455D089E0A07BC22A97C6" lastPageNumber="165" masterDocId="C538BFA63A3455D089E0A07BC22A97C6" masterDocTitle="New subgenera and a new species of the genus Raphignathus Duges (Prostigmata, Raphignathidae), with taxonomic notes on the genus Neoraphignathus Smiley & Moser" masterLastPageNumber="180" masterPageNumber="165" pageNumber="165" updateTime="1732480068003" updateUser="ExternalLinkService">
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<mods:mods id="FC5E696042307DF836A79A797583409B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A20A8CCC482A9E34090B759F32A9F7BB">
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<mods:title id="175644203169857B2838C54ECF696B19">New subgenera and a new species of the genus Raphignathus Duges (Prostigmata, Raphignathidae), with taxonomic notes on the genus Neoraphignathus Smiley & Moser</mods:title>
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<mods:name id="F670F1BC52022F51E703CEB50A448000" type="personal">
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<mods:roleTerm id="94E5A2386B80DBBA1E439451DB1B0B3F">Author</mods:roleTerm>
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<mods:namePart id="3B502A6554CB83B055762940E75F9EDF">Khan, Eid Muhammad</mods:namePart>
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<mods:nameIdentifier id="B32EA993DD8B47BFAE2AD5C0C0988A84" type="ORCID">https://orcid.org/0000-0001-5008-2524</mods:nameIdentifier>
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<mods:affiliation id="DF53BCA3BA06F1CA1D222F5803A574DE">Acarology Research Laboratory, Department of Plant Protection, College of Food and Agriculture Sciences, King Saud University, Riyadh, Saudi Arabia</mods:affiliation>
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<mods:namePart id="19070C5E02EB36FAE94301254724C5F1">Kamran, Muhammad</mods:namePart>
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<mods:nameIdentifier id="6966FA9EB40074D30CBB3E12B99E8B88" type="ORCID">https://orcid.org/0000-0001-6084-203X</mods:nameIdentifier>
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<mods:affiliation id="5F7459BDDE1D464186CE45FDA5F2875B">Acarology Research Laboratory, Department of Plant Protection, College of Food and Agriculture Sciences, King Saud University, Riyadh, Saudi Arabia</mods:affiliation>
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<mods:roleTerm id="435C7FC516F557117CA87B0931038184">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="03A5C8F0E1B3487FB56CC70F642873B7">Mirza, Jawwad Hassan</mods:namePart>
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<mods:nameIdentifier id="BFFB2213BFC7F047AC269ED4E514EE23" type="ORCID">https://orcid.org/0000-0002-1726-4331</mods:nameIdentifier>
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<mods:affiliation id="36194B52E32C0406F06A446DB5ED4B1E">Acarology Research Laboratory, Department of Plant Protection, College of Food and Agriculture Sciences, King Saud University, Riyadh, Saudi Arabia</mods:affiliation>
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<mods:namePart id="E6B65850B0A34F94EF22058A0C2947BB">Alatawi, Fahad Jaber</mods:namePart>
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<mods:nameIdentifier id="D7F3ABA67F488B09B904E839EDA18896" type="ORCID">https://orcid.org/0000-0002-6824-2650</mods:nameIdentifier>
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<mods:affiliation id="3A16C71E64714C5BCD2D993FBAC7BBC2">Acarology Research Laboratory, Department of Plant Protection, College of Food and Agriculture Sciences, King Saud University, Riyadh, Saudi Arabia</mods:affiliation>
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<mods:nameIdentifier id="06B76433C2F36100A5DE1D95A8588F37" type="email">falatawi@ksu.edu.sa</mods:nameIdentifier>
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<mods:title id="7569BFC9B1B26FC205129B9DB76D6B42">ZooKeys</mods:title>
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<mods:part id="ABF1F8D8CED77DDBC48BD56F1E2804A5">
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<mods:date id="381087EEA6E9E8388DE3323142ACB9E3">2023</mods:date>
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<mods:number id="A7A7E32BECC75F42203D4B511D0FC1F2">2023-08-23</mods:number>
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<mods:number id="A98430AAC4D8966CDA04FFC11DD3E30F">1176</mods:number>
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<mods:start id="5F3903B4CB47F4CE155A9641C058C4BF">165</mods:start>
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<mods:end id="4537BAA0C303332C3819EA348D9FBA75">180</mods:end>
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<mods:url id="631B27F659E16BF537274F42C6314A2B">http://dx.doi.org/10.3897/zookeys.1176.106224</mods:url>
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<mods:classification id="BC14F810C55AC6D8E2407F279399860E">journal article</mods:classification>
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<mods:identifier id="D21A54B14033F12CA2110FB15C09B698" type="DOI">http://dx.doi.org/10.3897/zookeys.1176.106224</mods:identifier>
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<mods:identifier id="EF0F7AE0E585A30967E527F712790FD6" type="Pensoft-Pub">1313-2970-1176-165</mods:identifier>
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</mods:mods>
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<treatment id="4E0A4286D8715265AAD699D360ED0DEF" ID-GBIF-Taxon="212453373" LSID="urn:lsid:zoobank.org:act:12E13CEE-478F-4A3F-A47E-B3C7F34C73FA" httpUri="http://treatment.plazi.org/id/4E0A4286D8715265AAD699D360ED0DEF" lastPageNumber="165" pageId="0" pageNumber="165" scope_family="Raphignathidae" scope_subGenus="Raphignathus" scope_subOrder="Prostigmata">
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<subSubSection id="8CF8D0C57A6E27E5CA5C9B034A8FDA6E" pageId="0" pageNumber="165" type="nomenclature">
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<paragraph id="4922FD89CE8A0924410A159D05E3AF33" pageId="0" pageNumber="165">
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<taxonomicName id="C3CFF774450E2C6530C5D05A1D53B9B9" LSID="https://zoobank.org/12E13CEE-478F-4A3F-A47E-B3C7F34C73FA" authority="sp. nov." authorityName="sp. nov." family="Raphignathidae" genus="Raphignathus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Raphignathus (Diraphignathus) neohecmatanaensis" order="Prostigmata" pageId="0" pageNumber="165" rank="species" species="neohecmatanaensis" subGenus="Diraphignathus">Raphignathus (Diraphignathus) neohecmatanaensis sp. nov.</taxonomicName>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="2BB2961A89646967D0D8DCB132D1507E" pageId="0" pageNumber="165" type="description">
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<paragraph id="7B6077DF8D0AD3F408BF764FEBCC930D" pageId="0" pageNumber="165">
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<figureCitation id="D17D1D727BBE4A6DA0FCF3989E113C1D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), dorsum. Scale bar: 100 µm." figureDoi="10.3897/zookeys.1176.106224.figure1" httpUri="https://binary.pensoft.net/fig/895996" pageId="0" pageNumber="165">Figs 1</figureCitation>
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||
<figureCitation id="B80DE894D2387CB16AFE67A3F979CD90" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), venter. Scale bar: 100 µm." figureDoi="10.3897/zookeys.1176.106224.figure2" httpUri="https://binary.pensoft.net/fig/895997" pageId="0" pageNumber="165">, 2</figureCitation>
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<figureCitation id="5478F47D6EEFE884045A7CD9317B82B4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), gnathosoma. Scale bar: 50 µm." figureDoi="10.3897/zookeys.1176.106224.figure3" httpUri="https://binary.pensoft.net/fig/895998" pageId="0" pageNumber="165">, 3</figureCitation>
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||
<figureCitation id="F07A3F508491F12A2C6847CA1DF83A2C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), palp. Scale bar: 20 µm." figureDoi="10.3897/zookeys.1176.106224.figure4" httpUri="https://binary.pensoft.net/fig/895999" pageId="0" pageNumber="165">, 4</figureCitation>
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||
<figureCitation id="A533C337FB02E9B01F869756CD2D858C" captionStart="Figures 5–8" captionStartId="F5" captionText="Figures 5 - 8. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), Legs I-IV. Scale bar: 20 µm." figureDoi="10.3897/zookeys.1176.106224.figures5-8" httpUri="https://binary.pensoft.net/fig/896000" pageId="0" pageNumber="165">, 5-8</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="00BC80EA5C78C315A01B101751325AE4" pageId="0" pageNumber="165" type="diagnosis">
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||
<paragraph id="0314D918205F73BE2803145CA87F7A42" pageId="0" pageNumber="165">Diagnosis.</paragraph>
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<paragraph id="6D2F4560C18FEB5EE75CB8F0363022F9" pageId="0" pageNumber="165">
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<emphasis id="33213AA86DAEE1B52FC0143162E8A3AB" bold="true" pageId="0" pageNumber="165">Female</emphasis>
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: endopodal shield absent between coxae I-IV; two small shields present posterolateral to median prodorsal shield; palp femora with two setae; femora 6-5-3-2; genua 5(+κ)-5(+κ)-4-4; tibiae 5(+φρ)-5(+φρ)-4(+φρ)-4 (+φρ); tarsi 21(1ω) -20(1ω)-15-14.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="40A79FFB43CC71D9ED60AA1C6B98E764" pageId="0" pageNumber="165" type="description">
|
||
<paragraph id="A597CE30088B3EEFD9979567178F615A" pageId="0" pageNumber="165">Description.</paragraph>
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||
<paragraph id="ADF27E13EE6CD27924DC52BE0B91A00C" pageId="0" pageNumber="165">
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||
<emphasis id="5F9A63C89102FFAD7033C0C3B3DB5FD3" bold="true" pageId="0" pageNumber="165">Female</emphasis>
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(
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||
<emphasis id="55FB77A38A345A78107EFC684B462C35" italics="true" pageId="0" pageNumber="165">n</emphasis>
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= 4). Idiosoma oval, length of body (including gnathosoma) 533 (525-545); width of body 345 (338-353).
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||
</paragraph>
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||
<paragraph id="FA0E15B418CC3C17C0563EBB5A0F7E71" pageId="0" pageNumber="165">
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||
<emphasis id="A3982EB227EF62761746C64322D6CF24" bold="true" italics="true" pageId="0" pageNumber="165">Dorsum</emphasis>
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(Fig.
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||
<figureCitation id="EDF99A8F8C328A8765CBF54326D559F7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), dorsum. Scale bar: 100 µm." figureDoi="10.3897/zookeys.1176.106224.figure1" httpUri="https://binary.pensoft.net/fig/895996" pageId="0" pageNumber="165">1</figureCitation>
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). Propodosoma with one medial and two lateral shields each containing three setae; medial sclerite with setae
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<emphasis id="6BA17EFE03731755083A3EB18126DB23" italics="true" pageId="0" pageNumber="165">vi</emphasis>
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,
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<emphasis id="52B2DBE155665D3DDD55067CFC4EB7AB" italics="true" pageId="0" pageNumber="165">si</emphasis>
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and
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<emphasis id="88DA99B2BC137374D40CDB7AE25D3F2E" italics="true" pageId="0" pageNumber="165">c1</emphasis>
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; paired ovoid lateral shields each with an eye, one cupule (
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<emphasis id="787D7690F516FE9E51EC3080104AD05A" italics="true" pageId="0" pageNumber="165">ia</emphasis>
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) and
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||
<emphasis id="345F126FA8629D0B9DD93E77376EA307" italics="true" pageId="0" pageNumber="165">sci</emphasis>
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,
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<emphasis id="AB890E78D57CCC11983F40D47BD3F8B5" italics="true" pageId="0" pageNumber="165">sce</emphasis>
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and seta
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||
<emphasis id="626ADD6358F688D543D96E078C4C727C" italics="true" pageId="0" pageNumber="165">c2</emphasis>
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seta; opisthosomal setal pairs
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||
<emphasis id="047E03FA445712422474027C3D8B62A1" italics="true" pageId="0" pageNumber="165">d1</emphasis>
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,
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<emphasis id="87F79207329C7D0E03481246E5A4AF6E" italics="true" pageId="0" pageNumber="165">e1</emphasis>
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and the pair of cupuli (
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||
<emphasis id="8B218D5DFC063DF63AD99A9731B54415" italics="true" pageId="0" pageNumber="165">im</emphasis>
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); posterior opisthosomal shield large, rectangular, bearing four pairs of setae (
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<emphasis id="099CE1C72BF469A1466C7990F3941B9D" italics="true" pageId="0" pageNumber="165">f1</emphasis>
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||
,
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||
<emphasis id="859229FF63E12CD2F382881271D0A92C" italics="true" pageId="0" pageNumber="165">h1-3</emphasis>
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) and one pair of the cupule (
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<emphasis id="477F4F0266D3EACC3DFB0EA0185EC037" italics="true" pageId="0" pageNumber="165">ip</emphasis>
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); all dorsal shields finely punctate; dorsal body setae setiform, smooth and acute; pseudanal setae
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<emphasis id="653F8C60E0DB831D91B9336F04B02660" italics="true" pageId="0" pageNumber="165">ps1</emphasis>
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dorsally located. Lengths of dorsal setae:
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<emphasis id="D773E15E1E627B35E6EDCD3FCC28A80F" italics="true" pageId="0" pageNumber="165">vi</emphasis>
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25 (23-27),
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<emphasis id="026D5802CF682B6DF0693B9B0817E007" italics="true" pageId="0" pageNumber="165">ve</emphasis>
|
||
28 (26-30),
|
||
<emphasis id="BE37D46DA3029A775DA88288A6B9E192" italics="true" pageId="0" pageNumber="165">sci</emphasis>
|
||
27 (26-29),
|
||
<emphasis id="17695013FFA2654F0458ECDFBCDD9BF9" italics="true" pageId="0" pageNumber="165">sce</emphasis>
|
||
28 (26-29),
|
||
<emphasis id="B8DA7DAB5D00ED239F9AE5BF983C1D67" italics="true" pageId="0" pageNumber="165">c1</emphasis>
|
||
23 (17-19),
|
||
<emphasis id="2C63D3F9CCBA07DEB5528EE11E92D270" italics="true" pageId="0" pageNumber="165">c2</emphasis>
|
||
25 (19-23),
|
||
<emphasis id="E745E6C70EF89B301C2BD6BC1A0C7ECC" italics="true" pageId="0" pageNumber="165">d1</emphasis>
|
||
21 (20-23),
|
||
<emphasis id="64E4C6A428A5DD548F3276BAEFA7F31A" italics="true" pageId="0" pageNumber="165">e1</emphasis>
|
||
22 (21-23),
|
||
<emphasis id="68E6D94BD11171E5B6234EF06079729F" italics="true" pageId="0" pageNumber="165">f1</emphasis>
|
||
21 (19-22),
|
||
<emphasis id="DFEE3DC235123C860BD60A1F631C5361" italics="true" pageId="0" pageNumber="165">h1</emphasis>
|
||
24 (23-25),
|
||
<emphasis id="BC7E58937DED8E301665D1F0D69EA21C" italics="true" pageId="0" pageNumber="165">h2</emphasis>
|
||
25 (23-25),
|
||
<emphasis id="9BBC95B942A2455E76EC20A1CCB67AE1" italics="true" pageId="0" pageNumber="165">h3</emphasis>
|
||
22 (21-24),
|
||
<emphasis id="F41AC7128A68947D71136AA76D2E565A" italics="true" pageId="0" pageNumber="165">ps1</emphasis>
|
||
23 (21-25); distances between dorsal 195 setae:
|
||
<emphasis id="0E2CC7804F676544ACAC6FFA00ACC31B" italics="true" pageId="0" pageNumber="165">vi-vi</emphasis>
|
||
27 (29-31),
|
||
<emphasis id="0E2533F98113BAF11E1D82078552E169" italics="true" pageId="0" pageNumber="165">sci-sci</emphasis>
|
||
123 (118-128),
|
||
<emphasis id="E2C31D465DA108783472F0794BE87F56" italics="true" pageId="0" pageNumber="165">vi- sci</emphasis>
|
||
70 (66-72),
|
||
<emphasis id="04A88E229A7B2B28A62A1C64807FE235" italics="true" pageId="0" pageNumber="165">sce-c2</emphasis>
|
||
68 (65-72),
|
||
<emphasis id="8F62A2F13BC39D64783F66782DC121B2" italics="true" pageId="0" pageNumber="165">c1-c1</emphasis>
|
||
30 (28-32),
|
||
<emphasis id="63AAB95C2B2A805B24E84A4D3E5C7814" italics="true" pageId="0" pageNumber="165">d1-d1</emphasis>
|
||
99 (92-101),
|
||
<emphasis id="B093EF6D96BE4EE61A1A7DE93C3F1278" italics="true" pageId="0" pageNumber="165">c1-d1</emphasis>
|
||
53 (49-55),
|
||
<emphasis id="77B2C305E076670321C6DA0D119EBB0B" italics="true" pageId="0" pageNumber="165">d1-e1</emphasis>
|
||
22 (21-24),
|
||
<emphasis id="5E8023CE6F8E1A83A85924FBDBDE2E93" italics="true" pageId="0" pageNumber="165">f1-f1</emphasis>
|
||
88 (83-90),
|
||
<emphasis id="6572B2F6050EA2EB400696B1F4F40DC6" italics="true" pageId="0" pageNumber="165">e1-f1</emphasis>
|
||
65 (61-68),
|
||
<emphasis id="EB903D514D05279931C912CFB3D72246" italics="true" pageId="0" pageNumber="165">h1-h1</emphasis>
|
||
45 (42-48),
|
||
<emphasis id="BB8EB5C9464D00B66740A1AA08F41E43" italics="true" pageId="0" pageNumber="165">h1-h2</emphasis>
|
||
43 (41-46),
|
||
<emphasis id="BDA11720C2E7847D7AC0A73F07F11510" italics="true" pageId="0" pageNumber="165">h2-h2</emphasis>
|
||
77 (72-80),
|
||
<emphasis id="4F949D6FB34FF7DE4542B401A407AA35" italics="true" pageId="0" pageNumber="165">h3-h3</emphasis>
|
||
101 (98-106).
|
||
</paragraph>
|
||
<caption id="7706F519EECD021F4A52541C7D5D5C17" doi="10.3897/zookeys.1176.106224.figure1" httpUri="https://binary.pensoft.net/fig/895996" pageId="0" pageNumber="165" start="Figure 1" startId="F1">
|
||
<paragraph id="32EBB8F7273DAE93D41090BF60D05BC9" pageId="0" pageNumber="165">
|
||
<emphasis id="C38BCF998EF48A55D4DCE54D2BCD25EB" bold="true" pageId="0" pageNumber="165">Figure 1.</emphasis>
|
||
<taxonomicName id="ED9D257BCCBCED5C80DB4D20BBE655F4" family="Raphignathidae" genus="Raphignathus" kingdom="Animalia" lsidName="Raphignathus (Diraphignathus) neohecmatanaensis" order="Prostigmata" pageId="0" pageNumber="165" rank="species" species="neohecmatanaensis" subGenus="Diraphignathus">Raphignathus (Diraphignathus) neohecmatanaensis</taxonomicName>
|
||
sp. nov. (female), dorsum. Scale bar: 100
|
||
<normalizedToken id="E1B69706D66917DD077D9FC1CF3F5672" originalValue="µm">µm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9F537A7161FA6FA204610D4350FFE834" pageId="0" pageNumber="165">
|
||
<emphasis id="D48A040823374A3D3DE888681853A87C" bold="true" italics="true" pageId="0" pageNumber="165">Venter</emphasis>
|
||
(Fig.
|
||
<figureCitation id="764BB44EA7BF71EEA0B3D56742437928" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), venter. Scale bar: 100 µm." figureDoi="10.3897/zookeys.1176.106224.figure2" httpUri="https://binary.pensoft.net/fig/895997" pageId="0" pageNumber="165">2</figureCitation>
|
||
). Venter entirely striated, without punctations; coxisternal shields absent (Fig.
|
||
<figureCitation id="E676F5E7B616AB2A0E98E72D8B86819A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), venter. Scale bar: 100 µm." figureDoi="10.3897/zookeys.1176.106224.figure2" httpUri="https://binary.pensoft.net/fig/895997" pageId="0" pageNumber="165">2</figureCitation>
|
||
); ventral setae lengths:
|
||
<emphasis id="D83F127E9B9B0809AE3893159DD2024B" italics="true" pageId="0" pageNumber="165">1a</emphasis>
|
||
42 (41-44),
|
||
<emphasis id="FE40C9594C7C5050F0F47D490704660F" italics="true" pageId="0" pageNumber="165">1b</emphasis>
|
||
38 (35-39),
|
||
<emphasis id="2FA577762AC667B6820CA2E663EF5410" italics="true" pageId="0" pageNumber="165">1c</emphasis>
|
||
34 (33-36),
|
||
<emphasis id="FD7F984AA8132BD50C598163D83033FD" italics="true" pageId="0" pageNumber="165">2b</emphasis>
|
||
36 (30-34),
|
||
<emphasis id="EC39C37CA7D5FE2837E5D6A71785741C" italics="true" pageId="0" pageNumber="165">2c</emphasis>
|
||
32 (30-34),
|
||
<emphasis id="5B36365DD116959B4778DC570A993958" italics="true" pageId="0" pageNumber="165">3a</emphasis>
|
||
24 (23-26),
|
||
<emphasis id="A2D7395346B5F63FE096F60CA56D587A" italics="true" pageId="0" pageNumber="165">3b</emphasis>
|
||
18 (17-20),
|
||
<emphasis id="F769DA7B4D80D918B278476A59DD5737" italics="true" pageId="0" pageNumber="165">3c</emphasis>
|
||
36 (34-38),
|
||
<emphasis id="2DC688253FAC612EE2E78C5BACEBBAC1" italics="true" pageId="0" pageNumber="165">4a</emphasis>
|
||
22 (23-27),
|
||
<emphasis id="949B0F07CD8FB64A622BDADAD9C218F3" italics="true" pageId="0" pageNumber="165">4c</emphasis>
|
||
38 (35-40); two pairs of aggenital setae (
|
||
<emphasis id="B3A9613BF39D01078955AB1E19A93867" italics="true" pageId="0" pageNumber="165">ag1-2</emphasis>
|
||
) with one cupule (
|
||
<emphasis id="E86D54643E084F25B09FA0021C520AF0" italics="true" pageId="0" pageNumber="165">ih</emphasis>
|
||
) on each side of the genital shield; anal opening and genital shields separate; genital shield prominent with a few punctations, bearing three pairs of genital setae (
|
||
<emphasis id="860B3695EA8F1A41BDB40EFC783A754F" italics="true" pageId="0" pageNumber="165">g1-3</emphasis>
|
||
); anal opening terminal, with three pairs of setae (
|
||
<emphasis id="5461B2D23947866D040D34135F05A629" italics="true" pageId="0" pageNumber="165">ps1</emphasis>
|
||
-3),
|
||
<emphasis id="089CFF49C57EF13333B7538EAE1D3585" italics="true" pageId="0" pageNumber="165">ps1</emphasis>
|
||
dorsal
|
||
<emphasis id="97C4B898100515117D38C85C87E46563" italics="true" pageId="0" pageNumber="165">ps3</emphasis>
|
||
and
|
||
<emphasis id="12071EBE7C68D87E835C2361FE4CA32E" italics="true" pageId="0" pageNumber="165">ps2</emphasis>
|
||
ventral; ventral setal lengths:
|
||
<emphasis id="0BBBA18DE2F5DAED2E270D32FBB5D91B" italics="true" pageId="0" pageNumber="165">ag1</emphasis>
|
||
29 (27-32);
|
||
<emphasis id="776FCBBA81A1F573F43890F5834335B5" italics="true" pageId="0" pageNumber="165">ag2</emphasis>
|
||
27 (25-28);
|
||
<emphasis id="3A6FA4E986113005BA48D459962CCF2A" italics="true" pageId="0" pageNumber="165">g1</emphasis>
|
||
31 (28-33);
|
||
<emphasis id="B3969441A36E1F65FAAE05E48798EF1E" italics="true" pageId="0" pageNumber="165">g2</emphasis>
|
||
25 (23-28);
|
||
<emphasis id="E264A67426BC3B3980F578144D1FE311" italics="true" pageId="0" pageNumber="165">g3</emphasis>
|
||
20 (19-22);
|
||
<emphasis id="DA9DDB48C4E07EF1A803750C6609E499" italics="true" pageId="0" pageNumber="165">ps2</emphasis>
|
||
22 (21-23);
|
||
<emphasis id="B6F1C802ED999362B35853BE47BC0C09" italics="true" pageId="0" pageNumber="165">ps3</emphasis>
|
||
21 (20-22). Distances between ventral setae:
|
||
<emphasis id="4B726D45C6F7D2418E5ABCDCDA1D4742" italics="true" pageId="0" pageNumber="165">1a-1a</emphasis>
|
||
55 (53-58),
|
||
<emphasis id="6EDB98491537AFEFF6C1DAF972C4FDBD" italics="true" pageId="0" pageNumber="165">3a-3a</emphasis>
|
||
115 (97-109),
|
||
<emphasis id="CFDF88823B027B37D99566A7EA3E621A" italics="true" pageId="0" pageNumber="165">4a-4a</emphasis>
|
||
70 (68-73),
|
||
<emphasis id="F216DC042E6EEDED74501C74DACE70B9" italics="true" pageId="0" pageNumber="165">ag2-ag2</emphasis>
|
||
42 (41-43),
|
||
<emphasis id="1C37F655B74ABC969E143019D45A422E" italics="true" pageId="0" pageNumber="165">g1-g1</emphasis>
|
||
38 (35-40),
|
||
<emphasis id="B7AA2E718B902E87770BBD415585A78F" italics="true" pageId="0" pageNumber="165">g2-g2</emphasis>
|
||
52 (48-55),
|
||
<emphasis id="DFB21224E703CB1B9E5F0BB1A60BFDF6" italics="true" pageId="0" pageNumber="165">g3-g3</emphasis>
|
||
75 (74-75),
|
||
<emphasis id="A4B7F61D0B5753FAF95CB9BF42F5C5AF" italics="true" pageId="0" pageNumber="165">2b-2c</emphasis>
|
||
30 (25-29),
|
||
<emphasis id="D662DB3AAE73978808B2855C693CD0EB" italics="true" pageId="0" pageNumber="165">1a-3a</emphasis>
|
||
50 (49-53),
|
||
<emphasis id="FDA56B09BC9321FBC2E779206FD5CCF6" italics="true" pageId="0" pageNumber="165">3a-4a</emphasis>
|
||
72 (68-75),
|
||
<emphasis id="8DD56E78FC6E76A9ED68B81F0E5E8047" italics="true" pageId="0" pageNumber="165">4a-ag1</emphasis>
|
||
85 (82-88),
|
||
<emphasis id="3A73AB2213A54695F7D0007761D456E8" italics="true" pageId="0" pageNumber="165">ag1-ag2</emphasis>
|
||
65 (63-68),
|
||
<emphasis id="CD5484D2220E0C466F5241D37D11BE05" italics="true" pageId="0" pageNumber="165">ag2-g1</emphasis>
|
||
63 (60-65),
|
||
<emphasis id="CF2C1A07AAB990743A5F526296F05EAD" italics="true" pageId="0" pageNumber="165">g1-g2</emphasis>
|
||
15 (14-17),
|
||
<emphasis id="D2CCD3A8C9AC1E0EEEEDE1F574617F0D" italics="true" pageId="0" pageNumber="165">g2-g3</emphasis>
|
||
22 (21-25),
|
||
<emphasis id="397D892DC5C297ACC007484580CBB0B2" italics="true" pageId="0" pageNumber="165">ag-g1</emphasis>
|
||
42 (40-45),
|
||
<emphasis id="D5556BEBE6243373B17CB090237B7995" italics="true" pageId="0" pageNumber="165">g3-ps3</emphasis>
|
||
16 (15-18),
|
||
<emphasis id="E470E54E13415DEFFAD7721B4508D508" italics="true" pageId="0" pageNumber="165">ps2- ps3</emphasis>
|
||
16 (14-18).
|
||
</paragraph>
|
||
<caption id="B353663262C7AB3AB847C9D98BAA8975" doi="10.3897/zookeys.1176.106224.figure2" httpUri="https://binary.pensoft.net/fig/895997" pageId="0" pageNumber="165" start="Figure 2" startId="F2">
|
||
<paragraph id="7E62751D3356277EC5071A35A3EB40BA" pageId="0" pageNumber="165">
|
||
<emphasis id="4A95E12020F3119122400E07BE60B249" bold="true" pageId="0" pageNumber="165">Figure 2.</emphasis>
|
||
<taxonomicName id="44B27DC94A412BEFD6704767E4782039" family="Raphignathidae" genus="Raphignathus" kingdom="Animalia" lsidName="Raphignathus (Diraphignathus) neohecmatanaensis" order="Prostigmata" pageId="0" pageNumber="165" rank="species" species="neohecmatanaensis" subGenus="Diraphignathus">Raphignathus (Diraphignathus) neohecmatanaensis</taxonomicName>
|
||
sp. nov. (female), venter. Scale bar: 100
|
||
<normalizedToken id="C3A8AB46282211C29A6CBA8AC8B43674" originalValue="µm">µm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3485598787B3561DA464CD239BF9859F" pageId="0" pageNumber="165">
|
||
<emphasis id="2B110FCC382916035B63F9ED11F2C302" bold="true" italics="true" pageId="0" pageNumber="165">Gnathosoma</emphasis>
|
||
(Figs
|
||
<figureCitation id="DE2BD1519CC2757D6B5EA5A8C13DBFFD" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), gnathosoma. Scale bar: 50 µm." figureDoi="10.3897/zookeys.1176.106224.figure3" httpUri="https://binary.pensoft.net/fig/895998" pageId="0" pageNumber="165">3</figureCitation>
|
||
,
|
||
<figureCitation id="6D1186D6258D8BFB6A7034A41BD0EA0A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), palp. Scale bar: 20 µm." figureDoi="10.3897/zookeys.1176.106224.figure4" httpUri="https://binary.pensoft.net/fig/895999" pageId="0" pageNumber="165">4</figureCitation>
|
||
). Ventral infracapitular with two pairs of very long setae (
|
||
<emphasis id="34B8F035CF5598EDECC65145AD7A2F90" italics="true" pageId="0" pageNumber="165">m</emphasis>
|
||
and
|
||
<emphasis id="163C472C653C6F9CD933D67CDB18BEED" italics="true" pageId="0" pageNumber="165">n</emphasis>
|
||
),
|
||
<emphasis id="D97E911FC5BFD56327C15CE0D8E1C2A1" italics="true" pageId="0" pageNumber="165">m</emphasis>
|
||
40 (39-42),
|
||
<emphasis id="1C5714584ED08789DF317577EC03090D" italics="true" pageId="0" pageNumber="165">n</emphasis>
|
||
52 (49-54) and two pairs of pilose adoral setae (
|
||
<emphasis id="DA20C8F7E8964061D545ABBA708456FD" italics="true" pageId="0" pageNumber="165">or1-2</emphasis>
|
||
),
|
||
<emphasis id="6F096FE313351E7F8B7D04C4BAE6CC7E" italics="true" pageId="0" pageNumber="165">or1</emphasis>
|
||
23 (21-24),
|
||
<emphasis id="290D2B5FD415843D5417B6E4A9376AC5" italics="true" pageId="0" pageNumber="165">or2</emphasis>
|
||
20 (19-22) (Fig.
|
||
<figureCitation id="2E6E8E229F9A60EFC5854213BB87193F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), gnathosoma. Scale bar: 50 µm." figureDoi="10.3897/zookeys.1176.106224.figure3" httpUri="https://binary.pensoft.net/fig/895998" pageId="0" pageNumber="165">3</figureCitation>
|
||
); stylophore conical and striated; palp chaetotaxy (femur-tarsus) as follows: 3-2-4+1 claw 4+1ω+4 eupathidia (ζ) (Fig.
|
||
<figureCitation id="C01816F56444D87B64B6E704D7F07F07" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), palp. Scale bar: 20 µm." figureDoi="10.3897/zookeys.1176.106224.figure4" httpUri="https://binary.pensoft.net/fig/895999" pageId="0" pageNumber="165">4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="D4B1DC67656D031CED23222AAEB19E99" doi="10.3897/zookeys.1176.106224.figure3" httpUri="https://binary.pensoft.net/fig/895998" pageId="0" pageNumber="165" start="Figure 3" startId="F3">
|
||
<paragraph id="A459F84CAC9B693158C9B7C372A3A553" pageId="0" pageNumber="165">
|
||
<emphasis id="5FC90FD43638FF2AA0F3E19B30795D8C" bold="true" pageId="0" pageNumber="165">Figure 3.</emphasis>
|
||
<taxonomicName id="CEC31D6A81CF87DD8593BABAC6F77E18" family="Raphignathidae" genus="Raphignathus" kingdom="Animalia" lsidName="Raphignathus (Diraphignathus) neohecmatanaensis" order="Prostigmata" pageId="0" pageNumber="165" rank="species" species="neohecmatanaensis" subGenus="Diraphignathus">Raphignathus (Diraphignathus) neohecmatanaensis</taxonomicName>
|
||
sp. nov. (female), gnathosoma. Scale bar: 50
|
||
<normalizedToken id="8D5D38FDEF4D23382FADB42137D94092" originalValue="µm">µm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="763F214419CA5B0D4BAF7337CF7315ED" doi="10.3897/zookeys.1176.106224.figure4" httpUri="https://binary.pensoft.net/fig/895999" pageId="0" pageNumber="165" start="Figure 4" startId="F4">
|
||
<paragraph id="F3181648521A2898D4A1544C8C9E78CF" pageId="0" pageNumber="165">
|
||
<emphasis id="D0620E781959D161F94311060171B342" bold="true" pageId="0" pageNumber="165">Figure 4.</emphasis>
|
||
<taxonomicName id="106CE6BFB6869F2CB1B4FA9ADEEEAE7C" family="Raphignathidae" genus="Raphignathus" kingdom="Animalia" lsidName="Raphignathus (Diraphignathus) neohecmatanaensis" order="Prostigmata" pageId="0" pageNumber="165" rank="species" species="neohecmatanaensis" subGenus="Diraphignathus">Raphignathus (Diraphignathus) neohecmatanaensis</taxonomicName>
|
||
sp. nov. (female), palp. Scale bar: 20
|
||
<normalizedToken id="2F6DB6DFB742B35FCC07CB20DEB0F6D2" originalValue="µm">µm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BCB309D92F600505E6BC1727981C902D" pageId="0" pageNumber="165">
|
||
<emphasis id="B0D8AD988A80E809488CA6D7FA794702" bold="true" italics="true" pageId="0" pageNumber="165">Legs</emphasis>
|
||
(Figs
|
||
<figureCitation id="818DDE4CA37DCC48FE6FF5740977EA44" captionStart="Figures 5–8" captionStartId="F5" captionText="Figures 5 - 8. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), Legs I-IV. Scale bar: 20 µm." figureDoi="10.3897/zookeys.1176.106224.figures5-8" httpUri="https://binary.pensoft.net/fig/896000" pageId="0" pageNumber="165">5-8</figureCitation>
|
||
). Length of legs I-IV (without coxae): 340 (328-348); 270 (276-286); 325 (317-328); 375 (367-384), respectively. Chaetotaxy on legs I-IV (solenidia in parentheses and not included in setal counts): coxa 2-2-2-1; trochanter 1-1-2-1; femora 6-5-3-2; genua 5(+κ)-5(+κ)-4-4; tibiae 5(+φρ)-5(+φρ)-4(+φρ)-4(+φρ); tarsi 21(1ω) -20(1ω)-15-14.
|
||
</paragraph>
|
||
<caption id="C7EE2A971E65B24A9509C9B65AAA4AAC" doi="10.3897/zookeys.1176.106224.figures5-8" httpUri="https://binary.pensoft.net/fig/896000" pageId="0" pageNumber="165" start="Figures 5–8" startId="F5">
|
||
<paragraph id="85C107D04589A42CE80D43411D6FEDA6" pageId="0" pageNumber="165">
|
||
<emphasis id="BC056304AD26DCFB15A3C38450C1BD0E" bold="true" pageId="0" pageNumber="165">Figures 5-8.</emphasis>
|
||
<taxonomicName id="89F74B7C4FA2012F0D0D98EEFD346F8C" family="Raphignathidae" genus="Raphignathus" kingdom="Animalia" lsidName="Raphignathus (Diraphignathus) neohecmatanaensis" order="Prostigmata" pageId="0" pageNumber="165" rank="species" species="neohecmatanaensis" subGenus="Diraphignathus">Raphignathus (Diraphignathus) neohecmatanaensis</taxonomicName>
|
||
sp. nov. (female), Legs I-IV. Scale bar: 20
|
||
<normalizedToken id="58D35E83F99ECEA27947F9DAA39CFD3D" originalValue="µm">µm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="A4D9A2266785B173E2BE5E1223902448" pageId="0" pageNumber="165" type="male and immature stages">
|
||
<paragraph id="957428EDE88C6A7CA56C3F6538C7AC79" pageId="0" pageNumber="165">Male and immature stages.</paragraph>
|
||
<paragraph id="05EECDE8A5E31E8C26BD3A93032A760B" pageId="0" pageNumber="165">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8CF52E08A3678DC18A34701A3449C41F" pageId="0" pageNumber="165" type="type material">
|
||
<paragraph id="7BE510CDE2101453F96EACFC636C4338" pageId="0" pageNumber="165">Type materials.</paragraph>
|
||
<paragraph id="5440C05D5EDD4CE263A912D80D649F03" pageId="0" pageNumber="165">
|
||
<emphasis id="C53C2579D97F5A652457A9E80A3989C6" bold="true" italics="true" pageId="0" pageNumber="165">Holotype</emphasis>
|
||
female and three paratype females, Faifa, Jazan,
|
||
<geoCoordinate id="26822D3D5C45BA90E0577579CBE404D8" degrees="24" direction="north" minutes="30.412" orientation="latitude" precision="1" value="24.506866">24°30.412'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="05C0502F8D3DA4120C1999652B045DFC" degrees="39" direction="east" minutes="36.578" orientation="longitude" precision="1" value="39.609634">39°36.578'E</geoCoordinate>
|
||
, 8 Oct., 2020, collected from
|
||
<taxonomicName id="3CF8582E92EF550F1C7648E5B68A02D6" class="Dicotyledoneae" family="Rhamnaceae" genus="Ziziphus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Ziziphus spina-christi" order="Rhamnales" pageId="0" pageNumber="165" phylum="Angiospermae" rank="species" species="spina-christi">
|
||
<emphasis id="0BA2FA7006917362924F1B1F5ABC354B" italics="true" pageId="0" pageNumber="165">Ziziphus spina-christi</emphasis>
|
||
</taxonomicName>
|
||
Mill. (
|
||
<taxonomicName id="0DD9B63BE6CA09470CEC37F7B4F6EC25" class="Dicotyledoneae" family="Rhamnaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Rhamnales" pageId="0" pageNumber="165" phylum="Angiospermae" rank="family">Rhamnaceae</taxonomicName>
|
||
) by Eid M. Khan, Jawwad H. Mirza & Hafiz S. Mushtaq.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6DD3E53261C5EADC1C8FEC22476E063E" pageId="0" pageNumber="165" type="etymology">
|
||
<paragraph id="BC20FEE951DD1852DD91FEE60CE85E76" pageId="0" pageNumber="165">Etymology.</paragraph>
|
||
<paragraph id="63141A627A90AA539A87920D8050E49A" pageId="0" pageNumber="165">
|
||
The specific epithet is in reference to the similarity of the new species to
|
||
<taxonomicName id="24CE9AB489FCCD60AF5CF5EE8E353D06" class="Arachnida" family="Raphignathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="R. (D.) hecmatanaensis" order="Prostigmata" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="hecmatanaensis" subGenus="D.">R. (D.) hecmatanaensis</taxonomicName>
|
||
;
|
||
<emphasis id="4A7685E1AD8CC8DC7343909A1922AC64" italics="true" pageId="0" pageNumber="165">neo</emphasis>
|
||
= new.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8D66110CB16D78D1F225D32A7800C020" pageId="0" pageNumber="165" type="remarks">
|
||
<paragraph id="C1E48608C433D44219785AAB2C42EE2F" pageId="0" pageNumber="165">Remarks.</paragraph>
|
||
<paragraph id="BD8447BEE5BCEA3D29791693B003C24B" pageId="0" pageNumber="165">
|
||
<taxonomicName id="33ACBC9DEC7AB1A7A032951E485BE9A4" family="Raphignathidae" genus="Raphignathus" kingdom="Animalia" lsidName="Raphignathus (D.) neohecmatanaensis" order="Prostigmata" pageId="0" pageNumber="165" rank="species" species="neohecmatanaensis" subGenus="D.">Raphignathus (D.) neohecmatanaensis</taxonomicName>
|
||
sp. nov., belongs to the subgenus
|
||
<taxonomicName id="C904A0B0A9CB1E5CEDA3D9408D198008" genus="Diraphignathus" lsidName="Diraphignathus" pageId="0" pageNumber="165" rank="genus">Diraphignathus</taxonomicName>
|
||
subgen. nov. The new species resembles to
|
||
<taxonomicName id="FDD73418DCFB696BA7C3C03B9A654286" class="Arachnida" family="Raphignathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="R. (D.) hecmatanaensis" order="Prostigmata" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="hecmatanaensis" subGenus="D.">R. (D.) hecmatanaensis</taxonomicName>
|
||
Khanjani & Ueckermann in having two pairs of setae (
|
||
<emphasis id="DBDB7D8499F68B3C91D55FD3E3BA0BBF" italics="true" pageId="0" pageNumber="165">d1</emphasis>
|
||
and
|
||
<emphasis id="F28D44F2A9B77CBBEAA38A8CEBA06AAC" italics="true" pageId="0" pageNumber="165">e1</emphasis>
|
||
) on the interscutal membrane, two setae on the palp femora, and two small plates present posterolateral to the median prodorsal shield. However, the new species differs from
|
||
<taxonomicName id="F31CD9ACE0F30C6E88232424136C1ADE" class="Arachnida" family="Raphignathidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="R. (D.) hecmatanaensis" order="Prostigmata" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="hecmatanaensis" subGenus="D.">R. (D.) hecmatanaensis</taxonomicName>
|
||
in the absence of an endopodal shield (vs present), femur IV with two setae (vs three), and leg tarsus I with one solenidion (vs two solenidia).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |