treatments-xml/data/4E/04/87/4E0487A2FFE07C50D7A9C702A0887377.xml

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<mods:title id="78D186A43548ABC57FA3ED90D1A19B1F">A new species of Alacran (Scorpiones: Typhlochactidae) from a cave in Oaxaca, Mexico</mods:title>
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<mods:namePart id="493A7E382121BC29E0441BE9CDEA9F12">Francke, Oscar F.</mods:namePart>
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<mods:date id="8E94CDEAC645C107E2CDB8356C328E0D">2009</mods:date>
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<paragraph id="C61236B4FFE07C58D7A9C702A60B7386" blockId="1.[151,525,1399,1460]" box="[151,525,1399,1425]" pageId="1" pageNumber="47">
<heading id="9D5A81D8FFE07C58D7A9C702A60B7386" bold="true" box="[151,525,1399,1425]" fontSize="11" level="1" pageId="1" pageNumber="47" reason="1">
<emphasis id="F4D9EAA6FFE07C58D7A9C702A60B7386" bold="true" box="[151,525,1399,1425]" pageId="1" pageNumber="47">
<taxonomicName id="01AD4D37FFE07C58D7A9C702A5697386" box="[151,367,1399,1425]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="47" phylum="Arthropoda" rank="species" species="chamuco" status="sp. nov.">
<emphasis id="F4D9EAA6FFE07C58D7A9C702A5697386" bold="true" box="[151,367,1399,1425]" italics="true" pageId="1" pageNumber="47">Alacran chamuco</emphasis>
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, 
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(
<figureCitation id="5E962A31FFE07C58D7A1C7EFA54D73A3" box="[159,331,1434,1460]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 – 4" captionStart-2="FIGURES 5 – 6" captionStart-3="FIGURES 7 – 8" captionStart-4="FIGURES 9 – 11" captionStart-5="FIGURES 12 – 13" captionStart-6="FIGURES 14 – 15" captionStart-7="FIGURES 16 – 19" captionStartId-0="2.[151,255,1882,1906]" captionStartId-1="3.[151,274,566,590]" captionStartId-2="3.[151,269,1976,2000]" captionStartId-3="5.[151,269,1499,1523]" captionStartId-4="6.[151,269,1747,1771]" captionStartId-5="7.[151,273,948,972]" captionStartId-6="7.[151,271,1973,1997]" captionStartId-7="8.[151,271,1252,1276]" captionTargetBox-0="[222,1352,563,1882]" captionTargetBox-1="[155,1430,197,539]" captionTargetBox-2="[157,1423,699,1929]" captionTargetBox-3="[179,1398,214,1458]" captionTargetBox-4="[158,1432,597,1704]" captionTargetBox-5="[157,1433,202,922]" captionTargetBox-6="[162,1410,1085,1935]" captionTargetBox-7="[173,1410,206,1214]" captionTargetId-0="figure@2.[222,1365,547,1882]" captionTargetId-1="figure@3.[151,1436,197,541]" captionTargetId-2="figure@3.[151,1436,689,1954]" captionTargetId-3="figure@5.[157,1430,194,1475]" captionTargetId-4="figure@6.[151,1436,581,1723]" captionTargetId-5="figure@7.[151,1436,194,931]" captionTargetId-6="figure@7.[156,1430,1060,1948]" captionTargetId-7="figure@8.[156,1430,194,1228]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="7" captionTargetPageId-7="8" captionText-0="FIGURE 1. Map of the state of Oaxaca, Mexico, indicating the known distributions for the troglobitic Alacran chamuco, sp. nov. (circle), and Alacran tartarus Francke (triangle)." captionText-1="FIGURES 2 – 4. Holotype female of Alacran chamuco, sp. nov. 2. Dorsal aspect of prosoma. 3. Ventral aspect of prosoma + metasoma. 4. Detail of right pectinal tooth (scales = 1 mm)." captionText-2="FIGURES 5 – 6. Metasomal segment V and telson. 5. Holotype female of Alacran chamuco, sp. nov. 6. Adult female of Alacran tartarus Francke (scale = 1 mm)." captionText-3="FIGURES 7 – 8. Holotype female of Alacran chamuco, sp. nov. 7. Dorsal view of right pedipalp femur. 8. Dorsal view of right pedipalp patella (scale = 1 mm)." captionText-4="FIGURES 9 – 11. Trichobothrial pattern on external aspect of pedipalp patella. 9. Holotype female of Alacran chamuco, sp. nov., left patella with 21 external trichobothria; trichobothrium not present on the right patella is highlighted in black; relative positions of the 3 ventral trichobothria also indicated. 10. Holotype female of Alacran chamuco, sp. nov., right patella with 20 external trichobothria. 11. Composite illustration, prepared directly from the camera lucida (by drawing one pattern directly over the other), showing the trichobothrial patterns on external aspect of the pedipalp patella of the holotype female of Alacran chamuco sp. nov. (circles), and an adult female Alacran tartarus Francke (crosses) (scale = 1 mm)." captionText-5="FIGURES 12 – 13. Dorsal view of left pedipalp chela. 12. Holotype female of Alacran chamuco, sp. nov. 13. Adult female Alacran tartarus Francke." captionText-6="FIGURES 14 – 15. External view of left pedipalp chela. 14. Holotype female of Alacran chamuco, sp. nov. 15. Adult female Alacran tartarus Francke (scale = 1 mm)." captionText-7="FIGURES 16 – 19. Dentition on pedipalp chela fingers. 16, 18. Holotype female of Alacran chamuco, sp. nov. 17, 19. Adult female Alacran tartarus Francke. 16, 17. Fixed finger. 18, 19. Movable finger (scale = 1 mm)." httpUri-0="https://zenodo.org/record/190120/files/figure.png" httpUri-1="https://zenodo.org/record/190121/files/figure.png" httpUri-2="https://zenodo.org/record/190122/files/figure.png" httpUri-3="https://zenodo.org/record/190123/files/figure.png" httpUri-4="https://zenodo.org/record/190124/files/figure.png" httpUri-5="https://zenodo.org/record/190125/files/figure.png" httpUri-6="https://zenodo.org/record/190126/files/figure.png" httpUri-7="https://zenodo.org/record/190127/files/figure.png" pageId="1" pageNumber="47">Figs. 1–16, 18</figureCitation>
, 
<figureCitation id="5E962A31FFE07C58D669C7EFA57073A3" box="[343,374,1434,1460]" captionStart="FIGURE 20" captionStartId="8.[151,255,1948,1972]" captionTargetBox="[166,1435,1376,1915]" captionTargetId="figure@8.[151,1436,1364,1924]" captionTargetPageId="8" captionText="FIGURE 20. Telotarsus III of holotype female of Alacran chamuco, sp. nov. (scale = 1 mm)." httpUri="https://zenodo.org/record/190128/files/figure.png" pageId="1" pageNumber="47">20</figureCitation>
)
</paragraph>
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<taxonomicName id="01AD4D37FFE07C58D7A9C792A4EB73E9" box="[151,237,1511,1534]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="F4D9EAA6FFE07C58D7A9C792A4EB73E9" box="[151,237,1511,1534]" italics="true" pageId="1" pageNumber="47">Alacran</emphasis>
</taxonomicName>
sp.: Vignoli &amp; Prendini, 2009: 4 (
<figureCitation id="5E962A31FFE07C58D567C792A69373E9" box="[601,661,1511,1534]" captionStart="FIGURE 1" captionStartId="2.[151,255,1882,1906]" captionTargetBox="[222,1352,563,1882]" captionTargetId="figure@2.[222,1365,547,1882]" captionTargetPageId="2" captionText="FIGURE 1. Map of the state of Oaxaca, Mexico, indicating the known distributions for the troglobitic Alacran chamuco, sp. nov. (circle), and Alacran tartarus Francke (triangle)." httpUri="https://zenodo.org/record/190120/files/figure.png" pageId="1" pageNumber="47">Fig. 1</figureCitation>
A), 8 (
<tableCitation id="8B2F030FFFE07C58D5E2C792A72973E9" box="[732,815,1511,1534]" captionStart="TABLE 2" captionStartId="4.[151,243,1354,1378]" captionTargetBox="[151,1437,1551,1821]" captionTargetPageId="4" captionText="TABLE 2. Morphometric comparisons between the holotype female of Alacran chamuco, sp. nov., and five adult females of Alacran tartarus Francke, including the holotype. Abbreviations are L = length, W = width and D = depth. Measurements in the first two columns are from Francke (1982); the second two are the WDS specimens measured by Vignoli &amp; Prendini (2009, Table 6, pp. 36 – 37)." httpUri="http://table.plazi.org/id/92D2663CFFE57C5DD7A9C73FA68373DE" pageId="1" pageNumber="47" tableUuid="92D2663CFFE57C5DD7A9C73FA68373DE">Table 2</tableCitation>
), 35.
</paragraph>
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<paragraph id="C61236B4FFE07C58D7A9C458A72B70F5" blockId="1.[151,1437,1581,2033]" pageId="1" pageNumber="47">
<emphasis id="F4D9EAA6FFE07C58D7A9C458A51C7050" bold="true" box="[151,282,1581,1607]" pageId="1" pageNumber="47">
<typeStatus id="19168816FFE07C58D7A9C458A4D27050" box="[151,212,1581,1607]" pageId="1" pageNumber="47">Type</typeStatus>
data.
</emphasis>
<typeStatus id="19168816FFE07C58D61FC45BA588705F" box="[289,398,1582,1608]" pageId="1" pageNumber="47" type="holotype">Holotype</typeStatus>
female, and only known specimen, from Te Cimutaa (NAD 27 777899 1987404) [
<geoCoordinate id="A3995073FFE07C58D26AC45BA50F7079" direction="north" orientation="latitude" pageId="1" pageNumber="47" precision="1" value="17.904028">N 17° 54’ 14.5”</geoCoordinate>
<geoCoordinate id="A3995073FFE07C58D62FC421A5DA7079" box="[273,476,1620,1646]" direction="west" orientation="longitude" pageId="1" pageNumber="47" precision="1" value="-96.37711">W 96° 22’ 37.6”</geoCoordinate>
, elev 
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<quantity id="01559B51FFE07C58D51AC421A6717079" box="[548,631,1620,1646]" metricMagnitude="2" metricUnit="m" metricValue="9.44" pageId="1" pageNumber="47" unit="m" value="944.0">944 m</quantity>
.
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], Municipio de Valle Nacional, Oaxaca, 
<collectingCountry id="BEBA7624FFE07C58D356C421A0C57079" box="[1128,1219,1620,1646]" name="Mexico" pageId="1" pageNumber="47">Mexico</collectingCountry>
(
<figureCitation id="5E962A31FFE07C58D3EAC421A11D7079" box="[1236,1307,1620,1646]" captionStart="FIGURE 1" captionStartId="2.[151,255,1882,1906]" captionTargetBox="[222,1352,563,1882]" captionTargetId="figure@2.[222,1365,547,1882]" captionTargetPageId="2" captionText="FIGURE 1. Map of the state of Oaxaca, Mexico, indicating the known distributions for the troglobitic Alacran chamuco, sp. nov. (circle), and Alacran tartarus Francke (triangle)." httpUri="https://zenodo.org/record/190120/files/figure.png" pageId="1" pageNumber="47">Fig. 1</figureCitation>
); 
<date id="B2131074FFE07C58D209C421A4D17082" pageId="1" pageNumber="47" value="2008-04-25">25 April 2008</date>
, Paul Bryant; approximately 
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<quantity id="01559B51FFE07C58D500C40EA6867082" box="[574,640,1659,1685]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="47" unit="m" value="50.0">50 m</quantity>
.
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from the cave entrance. Deposited in the Colección Nacional de Arácnidos (
<collectionCode id="A0BCAE71FFE07C58D61BC4D7A57B70AB" box="[293,381,1698,1724]" pageId="1" pageNumber="47">CNAN</collectionCode>
T-0401), Instituto de Biología, 
<collectionCode id="A0BCAE71FFE07C58D439C4D7A76E70AB" box="[775,872,1698,1724]" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="1" pageNumber="47">UNAM</collectionCode>
. “Te” means “cave” in Chinanteco, the native language of the inhabitants of the Valle Nacional region.
</paragraph>
</subSubSection>
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<emphasis id="F4D9EAA6FFE07C58D7F8C49AA561711E" bold="true" box="[198,359,1775,1801]" pageId="1" pageNumber="47">Distribution.</emphasis>
Known only from the 
<typeStatus id="19168816FFE07C58D54CC49AA6DB711E" box="[626,733,1775,1801]" pageId="1" pageNumber="47" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="8EB7653FFFE07C58D7F8C560A7AC7141" pageId="1" pageNumber="47" type="etymology">
<paragraph id="C61236B4FFE07C58D7F8C560A7AC7141" blockId="1.[151,1437,1581,2033]" pageId="1" pageNumber="47">
<emphasis id="F4D9EAA6FFE07C58D7F8C560A5557138" bold="true" box="[198,339,1813,1839]" pageId="1" pageNumber="47">Etymology.</emphasis>
The specific epithet is a noun in apposition, a name given in parts of 
<collectingCountry id="BEBA7624FFE07C58D3ABC563A0E87127" box="[1173,1262,1814,1840]" name="Mexico" pageId="1" pageNumber="47">Mexico</collectingCountry>
(including the Chinanteca region) to the devil—which inhabits in the underworld.
</paragraph>
</subSubSection>
<subSubSection id="8EB7653FFFE07C5BD7F8C517A0B876E8" lastPageId="2" lastPageNumber="48" pageId="1" pageNumber="47" type="diagnosis">
<paragraph id="C61236B4FFE07C5BD7F8C517A0B876E8" blockId="1.[151,1437,1581,2033]" lastBlockId="2.[151,1436,152,487]" lastPageId="2" lastPageNumber="48" pageId="1" pageNumber="47">
<emphasis id="F4D9EAA6FFE07C58D7F8C517A540716B" bold="true" box="[198,326,1890,1916]" pageId="1" pageNumber="47">Diagnosis.</emphasis>
The most conspicuous differences between 
<taxonomicName id="01AD4D37FFE07C58D472C516A7D1716B" box="[844,983,1891,1916]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="47" phylum="Arthropoda" rank="species" species="chamuco">
<emphasis id="F4D9EAA6FFE07C58D472C516A7D1716B" box="[844,983,1891,1916]" italics="true" pageId="1" pageNumber="47">A. chamuco</emphasis>
</taxonomicName>
and 
<taxonomicName id="01AD4D37FFE07C58D32EC516A088716B" box="[1040,1166,1891,1916]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="47" phylum="Arthropoda" rank="species" species="tartarus">
<emphasis id="F4D9EAA6FFE07C58D32EC516A088716B" box="[1040,1166,1891,1916]" italics="true" pageId="1" pageNumber="47">A. tartarus</emphasis>
</taxonomicName>
are: (1) the former has five inner denticles on the fixed finger of the pedipalp chela, and six inner denticles on the movable finger, whereas the latter has six and seven inner denticles, respectively; (2) in 
<taxonomicName id="01AD4D37FFE07C58D367C5C4A0F471DD" box="[1113,1266,1969,1994]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="47" phylum="Arthropoda" rank="species" species="chamuco">
<emphasis id="F4D9EAA6FFE07C58D367C5C4A0F471DD" box="[1113,1266,1969,1994]" italics="true" pageId="1" pageNumber="47">A. chamuco</emphasis>
</taxonomicName>
the telson is proportionately smaller: the ratio telson length/carapace length in 
<taxonomicName id="01AD4D37FFE07C58D49AC5A2A03671E7" box="[932,1072,2007,2032]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="47" phylum="Arthropoda" rank="species" species="chamuco">
<emphasis id="F4D9EAA6FFE07C58D49AC5A2A03671E7" box="[932,1072,2007,2032]" italics="true" pageId="1" pageNumber="47">A. chamuco</emphasis>
</taxonomicName>
is 1.08, and in 
<taxonomicName id="01AD4D37FFE07C58D3D6C5A2A16E71E7" box="[1256,1384,2007,2032]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="47" phylum="Arthropoda" rank="species" species="tartarus">
<emphasis id="F4D9EAA6FFE07C58D3D6C5A2A16E71E7" box="[1256,1384,2007,2032]" italics="true" pageId="1" pageNumber="47">A. tartarus</emphasis>
</taxonomicName>
it is 1.25 (n=5, range 1.24–1.27) (16% shorter); the ratio telson width/carapace length in 
<taxonomicName id="01AD4D37FFE37C5BD3BEC2EDA10D76A6" box="[1152,1291,152,177]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="chamuco">
<emphasis id="F4D9EAA6FFE37C5BD3BEC2EDA10D76A6" box="[1152,1291,152,177]" italics="true" pageId="2" pageNumber="48">A. chamuco</emphasis>
</taxonomicName>
is 0.47, and in 
<taxonomicName id="01AD4D37FFE37C5BD788C2CAA53076CF" box="[182,310,191,216]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="tartarus">
<emphasis id="F4D9EAA6FFE37C5BD788C2CAA53076CF" box="[182,310,191,216]" italics="true" pageId="2" pageNumber="48">A. tartarus</emphasis>
</taxonomicName>
it is 0.58 (n=5, range 0.55–0.62) (23% narrower); and the ratio of telson depth/carapace length in 
<taxonomicName id="01AD4D37FFE37C5BD78BC290A54676E9" box="[181,320,229,254]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="chamuco">
<emphasis id="F4D9EAA6FFE37C5BD78BC290A54676E9" box="[181,320,229,254]" italics="true" pageId="2" pageNumber="48">A. chamuco</emphasis>
</taxonomicName>
is 0.43, and in 
<taxonomicName id="01AD4D37FFE37C5BD6CDC290A67776E9" box="[499,625,229,254]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="tartarus">
<emphasis id="F4D9EAA6FFE37C5BD6CDC290A67776E9" box="[499,625,229,254]" italics="true" pageId="2" pageNumber="48">A. tartarus</emphasis>
</taxonomicName>
it is 0.57 (n=5, range 0.55–0.62) (32% less deep).
</paragraph>
</subSubSection>
<subSubSection id="8EB7653FFFE37C50D7F8C37EA0887377" lastPageId="9" lastPageNumber="55" pageId="2" pageNumber="48" type="description">
<paragraph id="C61236B4FFE37C5BD7F8C37EA7847731" blockId="2.[151,1436,152,487]" box="[198,898,267,294]" pageId="2" pageNumber="48">
<emphasis id="F4D9EAA6FFE37C5BD7F8C37EA5587732" bold="true" box="[198,350,267,293]" pageId="2" pageNumber="48">Description.</emphasis>
Based on 
<typeStatus id="19168816FFE37C5BD6E7C379A6397731" box="[473,575,268,294]" pageId="2" pageNumber="48" type="holotype">holotype</typeStatus>
and only known specimen.
</paragraph>
<paragraph id="C61236B4FFE37C5BD7FBC347A6DA7764" blockId="2.[151,1436,152,487]" pageId="2" pageNumber="48">
<emphasis id="F4D9EAA6FFE37C5BD7FBC347A51F775B" bold="true" box="[197,281,306,332]" pageId="2" pageNumber="48">Color:</emphasis>
Dorsum pale yellow-brown, venter and legs pale yellow; pedipalps orange-brown; granules on metasomal and pedipalpal carinae reddish brown.
</paragraph>
<paragraph id="C61236B4FFE37C5BD7FBC3F5A74D77F0" blockId="2.[151,1436,152,487]" pageId="2" pageNumber="48">
Prosoma: Carapace subquadrangular, anterior margin straight; without distinct furrows or ridges; lacking median and lateral ocelli; with scattered, moderately dense small granulation (
<figureCitation id="5E962A31FFE37C5BD378C3D3A09677D7" box="[1094,1168,422,448]" captionStart="FIGURES 2 – 4" captionStartId="3.[151,274,566,590]" captionTargetBox="[155,1430,197,539]" captionTargetId="figure@3.[151,1436,197,541]" captionTargetPageId="3" captionText="FIGURES 2 – 4. Holotype female of Alacran chamuco, sp. nov. 2. Dorsal aspect of prosoma. 3. Ventral aspect of prosoma + metasoma. 4. Detail of right pectinal tooth (scales = 1 mm)." httpUri="https://zenodo.org/record/190121/files/figure.png" pageId="2" pageNumber="48">Fig. 2</figureCitation>
). Sternum longer than wide; with a shallow, broad posteromedian furrow (
<figureCitation id="5E962A31FFE37C5BD5CAC3B8A73D77F0" box="[756,827,461,487]" captionStart="FIGURES 2 – 4" captionStartId="3.[151,274,566,590]" captionTargetBox="[155,1430,197,539]" captionTargetId="figure@3.[151,1436,197,541]" captionTargetPageId="3" captionText="FIGURES 2 – 4. Holotype female of Alacran chamuco, sp. nov. 2. Dorsal aspect of prosoma. 3. Ventral aspect of prosoma + metasoma. 4. Detail of right pectinal tooth (scales = 1 mm)." httpUri="https://zenodo.org/record/190121/files/figure.png" pageId="2" pageNumber="48">Fig. 3</figureCitation>
).
</paragraph>
<caption id="92D2663CFFE37C5BD7A9C52FA6FC7183" httpUri="https://zenodo.org/record/190120/files/figure.png" pageId="2" pageNumber="48" targetBox="[222,1352,563,1882]" targetPageId="2">
<paragraph id="C61236B4FFE37C5BD7A9C52FA6FC7183" blockId="2.[151,1436,1882,1941]" pageId="2" pageNumber="48">
<emphasis id="F4D9EAA6FFE37C5BD7A9C52FA51E7165" bold="true" box="[151,280,1882,1906]" pageId="2" pageNumber="48">FIGURE 1.</emphasis>
Map of the state of Oaxaca, Mexico, indicating the known distributions for the troglobitic 
<emphasis id="F4D9EAA6FFE37C5BD3E7C52EA19A7165" box="[1241,1436,1883,1906]" italics="true" pageId="2" pageNumber="48">
<taxonomicName id="01AD4D37FFE37C5BD3E7C52EA19E7165" box="[1241,1432,1883,1906]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="chamuco" status="sp. nov.">Alacran chamuco</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="EFEA57DDFFE37C5BD7A9C508A4EA7182" box="[151,236,1917,1941]" pageId="2" pageNumber="48" rank="species">
<emphasis id="F4D9EAA6FFE37C5BD7A9C508A4EA7182" bold="true" box="[151,236,1917,1941]" pageId="2" pageNumber="48">sp. nov.</emphasis>
</taxonomicNameLabel>
(circle), and 
<taxonomicName id="01AD4D37FFE37C5BD644C50BA68F7183" authority="Francke" authorityName="Francke" box="[378,649,1917,1941]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="tartarus">
<emphasis id="F4D9EAA6FFE37C5BD644C50BA62B7182" box="[378,557,1918,1941]" italics="true" pageId="2" pageNumber="48">Alacran tartarus</emphasis>
Francke
</taxonomicName>
(triangle).
</paragraph>
</caption>
<caption id="92D2663CFFE27C5AD7A9C043A7837467" httpUri="https://zenodo.org/record/190121/files/figure.png" pageId="3" pageNumber="49" targetBox="[155,1430,197,539]" targetPageId="3">
<paragraph id="C61236B4FFE27C5AD7A9C043A7837467" blockId="3.[151,1436,566,624]" pageId="3" pageNumber="49">
<emphasis id="F4D9EAA6FFE27C5AD7A9C043A54D7459" bold="true" box="[151,331,566,590]" pageId="3" pageNumber="49">FIGURES 2–4.</emphasis>
Holotype female of 
<taxonomicName id="01AD4D37FFE27C5AD506C042A6F97459" box="[568,767,567,590]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="chamuco" status="sp. nov.">
<emphasis id="F4D9EAA6FFE27C5AD506C042A6F97459" box="[568,767,567,590]" italics="true" pageId="3" pageNumber="49">Alacran chamuco</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="EFEA57DDFFE27C5AD430C043A76F7459" box="[782,873,566,590]" pageId="3" pageNumber="49" rank="species">
<emphasis id="F4D9EAA6FFE27C5AD430C043A76F7459" bold="true" box="[782,873,566,590]" pageId="3" pageNumber="49">sp. nov.</emphasis>
</taxonomicNameLabel>
2. Dorsal aspect of prosoma. 3. Ventral aspect of prosoma + metasoma. 4. Detail of right pectinal tooth (scales = 1mm).
</paragraph>
</caption>
<caption id="92D2663CFFE27C5AD7A9C5CDA64871E5" httpUri="https://zenodo.org/record/190122/files/figure.png" pageId="3" pageNumber="49" targetBox="[157,1423,699,1929]" targetPageId="3">
<paragraph id="C61236B4FFE27C5AD7A9C5CDA64871E5" blockId="3.[151,1436,1976,2034]" pageId="3" pageNumber="49">
<emphasis id="F4D9EAA6FFE27C5AD7A9C5CDA54471C7" bold="true" box="[151,322,1976,2000]" pageId="3" pageNumber="49">FIGURES 5–6.</emphasis>
Metasomal segment V and telson. 5. Holotype female of 
<taxonomicName id="01AD4D37FFE27C5AD491C5CCA06A71C7" box="[943,1132,1977,2000]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="chamuco" status="sp. nov.">
<emphasis id="F4D9EAA6FFE27C5AD491C5CCA06A71C7" box="[943,1132,1977,2000]" italics="true" pageId="3" pageNumber="49">Alacran chamuco</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="EFEA57DDFFE27C5AD344C5CDA0D671C7" box="[1146,1232,1976,2000]" pageId="3" pageNumber="49" rank="species">
<emphasis id="F4D9EAA6FFE27C5AD344C5CDA0D671C7" bold="true" box="[1146,1232,1976,2000]" pageId="3" pageNumber="49">sp. nov.</emphasis>
</taxonomicNameLabel>
6. Adult female of 
<taxonomicName id="01AD4D37FFE27C5AD7A9C5AEA5A071E5" authority="Francke" authorityName="Francke" box="[151,422,2011,2034]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="tartarus">
<emphasis id="F4D9EAA6FFE27C5AD7A9C5AEA54F71E5" box="[151,329,2011,2034]" italics="true" pageId="3" pageNumber="49">Alacran tartarus</emphasis>
Francke
</taxonomicName>
(scale = 1mm).
</paragraph>
</caption>
<caption id="92D2663CFFE57C5DD7A9C2E2A70176C6" ID-Table-UUID="92D2663CFFE57C5DD7A9C2E2A70176C6" httpUri="http://table.plazi.org/id/92D2663CFFE57C5DD7A9C2E2A70176C6" pageId="4" pageNumber="50" targetBox="[151,1437,237,1241]" targetIsTable="true" targetPageId="4">
<paragraph id="C61236B4FFE57C5DD7A9C2E2A70176C6" blockId="4.[151,1437,151,209]" pageId="4" pageNumber="50">
<emphasis id="F4D9EAA6FFE57C5DD7A9C2E2A50E76B8" bold="true" box="[151,264,151,175]" pageId="4" pageNumber="50">TABLE 1</emphasis>
. Measurements (in mm.) of the holotype female of 
<taxonomicName id="01AD4D37FFE57C5DD46AC2EDA01C76B8" box="[852,1050,152,175]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="chamuco" status="sp. nov.">
<emphasis id="F4D9EAA6FFE57C5DD46AC2EDA01C76B8" box="[852,1050,152,175]" italics="true" pageId="4" pageNumber="50">Alacran chamuco</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="EFEA57DDFFE57C5DD317C2E2A08276B8" box="[1065,1156,151,175]" pageId="4" pageNumber="50" rank="species">
<emphasis id="F4D9EAA6FFE57C5DD317C2E2A08276B8" bold="true" box="[1065,1156,151,175]" pageId="4" pageNumber="50">sp. nov.</emphasis>
</taxonomicNameLabel>
, from Te Cimutaa, Valle Nacional, Oaxaca, Mexico. L=length, W=width, D=depth.
</paragraph>
</caption>
<paragraph id="C61236B4FFE57C5DD7A1C298A01F72CE" pageId="4" pageNumber="50">
<table id="B4ADC414FFE583A6D7A9C298A19B72CE" box="[151,1437,237,1241]" gridcols="3" gridrows="16" pageId="4" pageNumber="50">
<tr id="789D34F6FFE583A6D7A9C298A19B7713" box="[151,1437,237,260]" gridrow="0" pageId="4" pageNumber="50">
<th id="3B4C5D8AFFE583A6D7A9C298A5427713" box="[151,324,237,260]" gridcol="0" gridrow="0" pageId="4" pageNumber="50">Total</th>
<th id="3B4C5D8AFFE583A6D5B7C298A6D07713" box="[649,726,237,260]" gridcol="1" gridrow="0" pageId="4" pageNumber="50">L</th>
<th id="3B4C5D8AFFE583A6D4C6C298A19B7713" box="[1016,1437,237,260]" gridcol="2" gridrow="0" pageId="4" pageNumber="50">57.5</th>
</tr>
<tr id="789D34F6FFE583A6D7A9C36DA19B774E" box="[151,1437,280,345]" gridrow="1" pageId="4" pageNumber="50">
<th id="3B4C5D8AFFE583A6D7A9C36DA542774E" box="[151,324,280,345]" gridcol="0" gridrow="1" pageId="4" pageNumber="50">Carapace Mesosoma</th>
<td id="3B4C5D8AFFE583A6D5B7C36DA6D0774E" box="[649,726,280,345]" gridcol="1" gridrow="1" pageId="4" pageNumber="50">L/W L</td>
<td id="3B4C5D8AFFE583A6D4C6C36DA19B774E" box="[1016,1437,280,345]" gridcol="2" gridrow="1" pageId="4" pageNumber="50">5.3/5.5 15</td>
</tr>
<tr id="789D34F6FFE583A6D7A9C318A19B7793" box="[151,1437,365,388]" gridrow="2" pageId="4" pageNumber="50">
<th id="3B4C5D8AFFE583A6D7A9C318A5427793" box="[151,324,365,388]" gridcol="0" gridrow="2" pageId="4" pageNumber="50">Metasoma</th>
<td id="3B4C5D8AFFE583A6D5B7C318A6D07793" box="[649,726,365,388]" gridcol="1" gridrow="2" pageId="4" pageNumber="50">Total L</td>
<td id="3B4C5D8AFFE583A6D4C6C318A19B7793" box="[1016,1437,365,388]" gridcol="2" gridrow="2" pageId="4" pageNumber="50">37.2</td>
</tr>
<tr id="789D34F6FFE583A6D7A9C3EDA19B77CE" box="[151,1437,408,473]" gridrow="3" pageId="4" pageNumber="50">
<th id="3B4C5D8AFFE583A6D7A9C3EDA54277CE" box="[151,324,408,473]" gridcol="0" gridrow="3" pageId="4" pageNumber="50">I II</th>
<td id="3B4C5D8AFFE583A6D5B7C3EDA6D077CE" box="[649,726,408,473]" gridcol="1" gridrow="3" pageId="4" pageNumber="50">L/W L/W</td>
<td id="3B4C5D8AFFE583A6D4C6C3EDA19B77CE" box="[1016,1437,408,473]" gridcol="2" gridrow="3" pageId="4" pageNumber="50">4.5/2.5 5.3/2.2</td>
</tr>
<tr id="789D34F6FFE583A6D7A9C398A19B7413" box="[151,1437,493,516]" gridrow="4" pageId="4" pageNumber="50">
<th id="3B4C5D8AFFE583A6D7A9C398A5427413" box="[151,324,493,516]" gridcol="0" gridrow="4" pageId="4" pageNumber="50">III</th>
<td id="3B4C5D8AFFE583A6D5B7C398A6D07413" box="[649,726,493,516]" gridcol="1" gridrow="4" pageId="4" pageNumber="50">L/W</td>
<td id="3B4C5D8AFFE583A6D4C6C398A19B7413" box="[1016,1437,493,516]" gridcol="2" gridrow="4" pageId="4" pageNumber="50">6.2/2.0</td>
</tr>
<tr id="789D34F6FFE583A6D7A9C06DA19B744E" box="[151,1437,536,601]" gridrow="5" pageId="4" pageNumber="50">
<th id="3B4C5D8AFFE583A6D7A9C06DA542744E" box="[151,324,536,601]" gridcol="0" gridrow="5" pageId="4" pageNumber="50">IV V</th>
<td id="3B4C5D8AFFE583A6D5B7C06DA6D0744E" box="[649,726,536,601]" gridcol="1" gridrow="5" pageId="4" pageNumber="50">L/W L/W</td>
<td id="3B4C5D8AFFE583A6D4C6C06DA19B744E" box="[1016,1437,536,601]" gridcol="2" gridrow="5" pageId="4" pageNumber="50">8.6/1.6 12.6/1.3</td>
</tr>
<tr id="789D34F6FFE583A6D7A9C018A19B7493" box="[151,1437,621,644]" gridrow="6" pageId="4" pageNumber="50">
<th id="3B4C5D8AFFE583A6D7A9C018A5427493" box="[151,324,621,644]" gridcol="0" gridrow="6" pageId="4" pageNumber="50">Telson</th>
<td id="3B4C5D8AFFE583A6D5B7C018A6D07493" box="[649,726,621,644]" gridcol="1" gridrow="6" pageId="4" pageNumber="50">L/W/D</td>
<td id="3B4C5D8AFFE583A6D4C6C018A19B7493" box="[1016,1437,621,644]" gridcol="2" gridrow="6" pageId="4" pageNumber="50">5.7/2.5/2.3</td>
</tr>
<tr id="789D34F6FFE583A6D7A9C0EDA19B74CE" box="[151,1437,664,729]" gridrow="7" pageId="4" pageNumber="50">
<th id="3B4C5D8AFFE583A6D7A9C0EDA54274CE" box="[151,324,664,729]" gridcol="0" gridrow="7" pageId="4" pageNumber="50">Pedipalp Femur</th>
<td id="3B4C5D8AFFE583A6D5B7C0EDA6D074CE" box="[649,726,664,729]" gridcol="1" gridrow="7" pageId="4" pageNumber="50">Total L L/W/D</td>
<td id="3B4C5D8AFFE583A6D4C6C0EDA19B74CE" box="[1016,1437,664,729]" gridcol="2" gridrow="7" pageId="4" pageNumber="50">28.2 7.9/1.8/1.0</td>
</tr>
<tr id="789D34F6FFE583A6D7A9C098A19B7513" box="[151,1437,749,772]" gridrow="8" pageId="4" pageNumber="50">
<th id="3B4C5D8AFFE583A6D7A9C098A5427513" box="[151,324,749,772]" gridcol="0" gridrow="8" pageId="4" pageNumber="50">Patella</th>
<td id="3B4C5D8AFFE583A6D5B7C098A6D07513" box="[649,726,749,772]" gridcol="1" gridrow="8" pageId="4" pageNumber="50">L/W/D</td>
<td id="3B4C5D8AFFE583A6D4C6C098A19B7513" box="[1016,1437,749,772]" gridcol="2" gridrow="8" pageId="4" pageNumber="50">7.2/1.9/1.8</td>
</tr>
<tr id="789D34F6FFE583A6D7A9C16DA19B754E" box="[151,1437,792,857]" gridrow="9" pageId="4" pageNumber="50">
<th id="3B4C5D8AFFE583A6D7A9C16DA542754E" box="[151,324,792,857]" gridcol="0" gridrow="9" pageId="4" pageNumber="50">Chela Movable finger</th>
<td id="3B4C5D8AFFE583A6D5B7C16DA6D0754E" box="[649,726,792,857]" gridcol="1" gridrow="9" pageId="4" pageNumber="50">L/W/D L</td>
<td id="3B4C5D8AFFE583A6D4C6C16DA19B754E" box="[1016,1437,792,857]" gridcol="2" gridrow="9" pageId="4" pageNumber="50">13.1/2.8/3.2 8.1</td>
</tr>
<tr id="789D34F6FFE583A6D7A9C118A19B7593" box="[151,1437,877,900]" gridrow="10" pageId="4" pageNumber="50">
<th id="3B4C5D8AFFE583A6D7A9C118A5427593" box="[151,324,877,900]" gridcol="0" gridrow="10" pageId="4" pageNumber="50">Fixed finger</th>
<td id="3B4C5D8AFFE583A6D5B7C118A6D07593" box="[649,726,877,900]" gridcol="1" gridrow="10" pageId="4" pageNumber="50">L</td>
<td id="3B4C5D8AFFE583A6D4C6C118A19B7593" box="[1016,1437,877,900]" gridcol="2" gridrow="10" pageId="4" pageNumber="50">6.8</td>
</tr>
<tr id="789D34F6FFE583A6D7A9C1EDA19B75CE" box="[151,1437,920,985]" gridrow="11" pageId="4" pageNumber="50">
<th id="3B4C5D8AFFE583A6D7A9C1EDA54275CE" box="[151,324,920,985]" gridcol="0" gridrow="11" pageId="4" pageNumber="50">Femur I Femur II</th>
<td id="3B4C5D8AFFE583A6D5B7C1EDA6D075CE" box="[649,726,920,985]" gridcol="1" gridrow="11" pageId="4" pageNumber="50">L L</td>
<td id="3B4C5D8AFFE583A6D4C6C1EDA19B75CE" box="[1016,1437,920,985]" gridcol="2" gridrow="11" pageId="4" pageNumber="50">5.1 6.1</td>
</tr>
<tr id="789D34F6FFE583A6D7A9C198A19B7213" box="[151,1437,1005,1028]" gridrow="12" pageId="4" pageNumber="50">
<th id="3B4C5D8AFFE583A6D7A9C198A5427213" box="[151,324,1005,1028]" gridcol="0" gridrow="12" pageId="4" pageNumber="50">Femur III</th>
<td id="3B4C5D8AFFE583A6D5B7C198A6D07213" box="[649,726,1005,1028]" gridcol="1" gridrow="12" pageId="4" pageNumber="50">L</td>
<td id="3B4C5D8AFFE583A6D4C6C198A19B7213" box="[1016,1437,1005,1028]" gridcol="2" gridrow="12" pageId="4" pageNumber="50">7.6</td>
</tr>
<tr id="789D34F6FFE583A6D7A9C66DA19B724E" box="[151,1437,1048,1113]" gridrow="13" pageId="4" pageNumber="50">
<th id="3B4C5D8AFFE583A6D7A9C66DA542724E" box="[151,324,1048,1113]" gridcol="0" gridrow="13" pageId="4" pageNumber="50">Femur IV Patella I</th>
<td id="3B4C5D8AFFE583A6D5B7C66DA6D0724E" box="[649,726,1048,1113]" gridcol="1" gridrow="13" pageId="4" pageNumber="50">L L</td>
<td id="3B4C5D8AFFE583A6D4C6C66DA19B724E" box="[1016,1437,1048,1113]" gridcol="2" gridrow="13" pageId="4" pageNumber="50">8.5 4.5</td>
</tr>
<tr id="789D34F6FFE583A6D7A9C618A19B7293" box="[151,1437,1133,1156]" gridrow="14" pageId="4" pageNumber="50">
<th id="3B4C5D8AFFE583A6D7A9C618A5427293" box="[151,324,1133,1156]" gridcol="0" gridrow="14" pageId="4" pageNumber="50">Patella II</th>
<td id="3B4C5D8AFFE583A6D5B7C618A6D07293" box="[649,726,1133,1156]" gridcol="1" gridrow="14" pageId="4" pageNumber="50">L</td>
<td id="3B4C5D8AFFE583A6D4C6C618A19B7293" box="[1016,1437,1133,1156]" gridcol="2" gridrow="14" pageId="4" pageNumber="50">5.1</td>
</tr>
<tr id="789D34F6FFE583A6D7A9C6EDA19B72CE" box="[151,1437,1176,1241]" gridrow="15" pageId="4" pageNumber="50">
<th id="3B4C5D8AFFE583A6D7A9C6EDA54272CE" box="[151,324,1176,1241]" gridcol="0" gridrow="15" pageId="4" pageNumber="50">Patella III Patella IV</th>
<td id="3B4C5D8AFFE583A6D5B7C6EDA6D072CE" box="[649,726,1176,1241]" gridcol="1" gridrow="15" pageId="4" pageNumber="50">L L</td>
<td id="3B4C5D8AFFE583A6D4C6C6EDA19B72CE" box="[1016,1437,1176,1241]" gridcol="2" gridrow="15" pageId="4" pageNumber="50">5.9 6.2</td>
</tr>
</table>
</paragraph>
<caption id="92D2663CFFE57C5DD7A9C73FA68373DE" ID-Table-UUID="92D2663CFFE57C5DD7A9C73FA68373DE" httpUri="http://table.plazi.org/id/92D2663CFFE57C5DD7A9C73FA68373DE" pageId="4" pageNumber="50" targetBox="[151,1437,1551,1821]" targetIsTable="true" targetPageId="4">
<paragraph id="C61236B4FFE57C5DD7A9C73FA68373DE" blockId="4.[151,1437,1354,1530]" pageId="4" pageNumber="50">
<emphasis id="F4D9EAA6FFE57C5DD7A9C73FA50E7375" bold="true" box="[151,264,1354,1378]" pageId="4" pageNumber="50">TABLE 2</emphasis>
. Morphometric comparisons between the holotype female of 
<taxonomicName id="01AD4D37FFE57C5DD4FFC73FA0817376" box="[961,1159,1354,1377]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="chamuco" status="sp. nov.">
<emphasis id="F4D9EAA6FFE57C5DD4FFC73FA0817376" box="[961,1159,1354,1377]" italics="true" pageId="4" pageNumber="50">Alacran chamuco</emphasis>
</taxonomicName>
, 
<emphasis id="F4D9EAA6FFE57C5DD3A9C73FA0FF7375" bold="true" box="[1175,1273,1354,1378]" pageId="4" pageNumber="50">
<taxonomicNameLabel id="EFEA57DDFFE57C5DD3A9C73FA0F57375" box="[1175,1267,1354,1378]" pageId="4" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
and five adult females of 
<taxonomicName id="01AD4D37FFE57C5DD62AC718A6337393" authority="Francke" authorityName="Francke" box="[276,565,1389,1412]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="tartarus">
<emphasis id="F4D9EAA6FFE57C5DD62AC718A5D77393" box="[276,465,1389,1412]" italics="true" pageId="4" pageNumber="50">Alacran tartarus</emphasis>
Francke
</taxonomicName>
, including the holotype. Abbreviations are L=length, W=width and D=depth. Measurements in the first two columns are from Francke (1982); the second two are the WDS specimens measured by Vignoli &amp; Prendini (2009, Table 6, pp.36–37).
</paragraph>
</caption>
<paragraph id="C61236B4FFE57C5DD69CC790A13A7167" blockId="4.[151,1437,1354,1530]" lastBlockId="4.[272,1340,1842,1904]" pageId="4" pageNumber="50">
<emphasis id="F4D9EAA6FFE57C5DD69CC790A07973ED" box="[418,1151,1509,1530]" italics="true" pageId="4" pageNumber="50">
<taxonomicName id="01AD4D37FFE57C5DD69CC790A00773ED" box="[418,1025,1509,1530]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="tartarus">A. tartarus A. tartarus A. tartarus A. tartarus A. tartarus</taxonomicName>
<taxonomicName id="01AD4D37FFE57C5DD328C790A07973ED" authority="Standard" authorityName="Standard" box="[1046,1151,1509,1530]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="tartarus">A. tartarus</taxonomicName>
</emphasis>
Standard 
<taxonomicName id="01AD4D37FFE57C5DD22CC790A54C7150" authority="Tel W" authorityName="Tel W" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="chamuco">
<emphasis id="F4D9EAA6FFE57C5DD22CC790A18473ED" box="[1298,1410,1509,1530]" italics="true" pageId="4" pageNumber="50">A. chamuco</emphasis>
Tel W
</taxonomicName>
/ 
<collectingCountry id="BEBA7624FFE57C5DD66CC547A5727150" box="[338,372,1842,1863]" name="Central African Republic" pageId="4" pageNumber="50">Car</collectingCountry>
L 0.57 0.55 0.59 0.62 0.59 0.584 0.026076 0.47 Tel D/ 
<collectingCountry id="BEBA7624FFE57C5DD672C52EA5697167" box="[332,367,1883,1904]" name="Central African Republic" pageId="4" pageNumber="50">Car</collectingCountry>
L 0.55 0.55 0.56 0.62 0.55 0.566 0.030495 0.43
</paragraph>
<paragraph id="C61236B4FFE57C5DD69CC47AA13A710A" pageId="4" pageNumber="50">
<table id="B4ADC414FFE583A6D7A9C47AA19B710A" box="[151,1437,1551,1821]" gridcols="7" gridrows="7" pageId="4" pageNumber="50">
<tr id="789D34F6FFE583A6D7A9C47AA19B7033" box="[151,1437,1551,1572]" gridrow="0" pageId="4" pageNumber="50" rowspan-0="1">
<th id="3B4C5D8AFFE583A6D69CC47AA5F07033" box="[418,502,1551,1572]" gridcol="1" gridrow="0" pageId="4" pageNumber="50">holotype</th>
<th id="3B4C5D8AFFE583A6D521C47AA6497033" box="[543,591,1551,1572]" gridcol="2" gridrow="0" pageId="4" pageNumber="50">adult</th>
<th id="3B4C5D8AFFE583A6D5A3C47AA7CF7033" box="[669,969,1551,1572]" gridcol="3" gridrow="0" pageId="4" pageNumber="50">adult WDS adult WDS adult</th>
<th id="3B4C5D8AFFE583A6D328C47AA0677033" box="[1046,1121,1551,1572]" gridcol="4" gridrow="0" pageId="4" pageNumber="50">average</th>
<th id="3B4C5D8AFFE583A6D3AAC47AA0F57033" box="[1172,1267,1551,1572]" gridcol="5" gridrow="0" pageId="4" pageNumber="50">Deviation</th>
<th id="3B4C5D8AFFE583A6D22CC47AA19B7033" box="[1298,1437,1551,1572]" gridcol="6" gridrow="0" pageId="4" pageNumber="50">holotype</th>
</tr>
<tr id="789D34F6FFE583A6D7A9C44DA19B705A" box="[151,1437,1592,1613]" gridrow="1" pageId="4" pageNumber="50" rowspan-5="1">
<th id="3B4C5D8AFFE583A6D7A9C44DA584705A" box="[151,386,1592,1613]" gridcol="0" gridrow="1" pageId="4" pageNumber="50">Carapace L</th>
<td id="3B4C5D8AFFE583A6D69CC44DA5F0705A" box="[418,502,1592,1613]" gridcol="1" gridrow="1" pageId="4" pageNumber="50">5</td>
<td id="3B4C5D8AFFE583A6D521C44DA649705A" box="[543,591,1592,1613]" gridcol="2" gridrow="1" pageId="4" pageNumber="50">5.3</td>
<td id="3B4C5D8AFFE583A6D5A3C44DA7CF705A" box="[669,969,1592,1613]" gridcol="3" gridrow="1" pageId="4" pageNumber="50">6.3 5.5 5.6</td>
<td id="3B4C5D8AFFE583A6D328C44DA067705A" box="[1046,1121,1592,1613]" gridcol="4" gridrow="1" pageId="4" pageNumber="50">5.54</td>
<td id="3B4C5D8AFFE583A6D22CC44DA19B705A" box="[1298,1437,1592,1613]" gridcol="6" gridrow="1" pageId="4" pageNumber="50">5.3</td>
</tr>
<tr id="789D34F6FFE583A6D7A9C417A19B7060" box="[151,1437,1634,1655]" gridrow="2" pageId="4" pageNumber="50" rowspan-5="1">
<th id="3B4C5D8AFFE583A6D7A9C417A5847060" box="[151,386,1634,1655]" gridcol="0" gridrow="2" pageId="4" pageNumber="50">Femur L</th>
<td id="3B4C5D8AFFE583A6D69CC417A5F07060" box="[418,502,1634,1655]" gridcol="1" gridrow="2" pageId="4" pageNumber="50">7</td>
<td id="3B4C5D8AFFE583A6D521C417A6497060" box="[543,591,1634,1655]" gridcol="2" gridrow="2" pageId="4" pageNumber="50">7.6</td>
<td id="3B4C5D8AFFE583A6D5A3C417A7CF7060" box="[669,969,1634,1655]" gridcol="3" gridrow="2" pageId="4" pageNumber="50">7.4 8.3 8</td>
<td id="3B4C5D8AFFE583A6D328C417A0677060" box="[1046,1121,1634,1655]" gridcol="4" gridrow="2" pageId="4" pageNumber="50">7.66</td>
<td id="3B4C5D8AFFE583A6D22CC417A19B7060" box="[1298,1437,1634,1655]" gridcol="6" gridrow="2" pageId="4" pageNumber="50">7.8</td>
</tr>
<tr id="789D34F6FFE583A6D7A9C4FEA19B70B7" box="[151,1437,1675,1696]" gridrow="3" pageId="4" pageNumber="50" rowspan-5="1">
<th id="3B4C5D8AFFE583A6D7A9C4FEA58470B7" box="[151,386,1675,1696]" gridcol="0" gridrow="3" pageId="4" pageNumber="50">Telson L</th>
<td id="3B4C5D8AFFE583A6D69CC4FEA5F070B7" box="[418,502,1675,1696]" gridcol="1" gridrow="3" pageId="4" pageNumber="50">6.2</td>
<td id="3B4C5D8AFFE583A6D521C4FEA64970B7" box="[543,591,1675,1696]" gridcol="2" gridrow="3" pageId="4" pageNumber="50">6.6</td>
<td id="3B4C5D8AFFE583A6D5A3C4FEA7CF70B7" box="[669,969,1675,1696]" gridcol="3" gridrow="3" pageId="4" pageNumber="50">7.8 7 7</td>
<td id="3B4C5D8AFFE583A6D328C4FEA06770B7" box="[1046,1121,1675,1696]" gridcol="4" gridrow="3" pageId="4" pageNumber="50">6.92</td>
<td id="3B4C5D8AFFE583A6D22CC4FEA19B70B7" box="[1298,1437,1675,1696]" gridcol="6" gridrow="3" pageId="4" pageNumber="50">5.7</td>
</tr>
<tr id="789D34F6FFE583A6D7A9C4C0A19B70DD" box="[151,1437,1717,1738]" gridrow="4" pageId="4" pageNumber="50" rowspan-5="1">
<th id="3B4C5D8AFFE583A6D7A9C4C0A58470DD" box="[151,386,1717,1738]" gridcol="0" gridrow="4" pageId="4" pageNumber="50">W</th>
<td id="3B4C5D8AFFE583A6D69CC4C0A5F070DD" box="[418,502,1717,1738]" gridcol="1" gridrow="4" pageId="4" pageNumber="50">2.85</td>
<td id="3B4C5D8AFFE583A6D521C4C0A64970DD" box="[543,591,1717,1738]" gridcol="2" gridrow="4" pageId="4" pageNumber="50">2.9</td>
<td id="3B4C5D8AFFE583A6D5A3C4C0A7CF70DD" box="[669,969,1717,1738]" gridcol="3" gridrow="4" pageId="4" pageNumber="50">3.7 3.4 3.3</td>
<td id="3B4C5D8AFFE583A6D328C4C0A06770DD" box="[1046,1121,1717,1738]" gridcol="4" gridrow="4" pageId="4" pageNumber="50">3.23</td>
<td id="3B4C5D8AFFE583A6D22CC4C0A19B70DD" box="[1298,1437,1717,1738]" gridcol="6" gridrow="4" pageId="4" pageNumber="50">2.5</td>
</tr>
<tr id="789D34F6FFE583A6D7A9C4AAA19B70E3" box="[151,1437,1759,1780]" gridrow="5" pageId="4" pageNumber="50" rowspan-5="1">
<th id="3B4C5D8AFFE583A6D7A9C4AAA58470E3" box="[151,386,1759,1780]" gridcol="0" gridrow="5" pageId="4" pageNumber="50">D</th>
<td id="3B4C5D8AFFE583A6D69CC4AAA5F070E3" box="[418,502,1759,1780]" gridcol="1" gridrow="5" pageId="4" pageNumber="50">2.75</td>
<td id="3B4C5D8AFFE583A6D521C4AAA64970E3" box="[543,591,1759,1780]" gridcol="2" gridrow="5" pageId="4" pageNumber="50">2.9</td>
<td id="3B4C5D8AFFE583A6D5A3C4AAA7CF70E3" box="[669,969,1759,1780]" gridcol="3" gridrow="5" pageId="4" pageNumber="50">3.5 3.4 3.1</td>
<td id="3B4C5D8AFFE583A6D328C4AAA06770E3" box="[1046,1121,1759,1780]" gridcol="4" gridrow="5" pageId="4" pageNumber="50">3.13</td>
<td id="3B4C5D8AFFE583A6D22CC4AAA19B70E3" box="[1298,1437,1759,1780]" gridcol="6" gridrow="5" pageId="4" pageNumber="50">2.3</td>
</tr>
<tr id="789D34F6FFE583A6D7A9C57DA19B710A" box="[151,1437,1800,1821]" gridrow="6" pageId="4" pageNumber="50">
<th id="3B4C5D8AFFE583A6D7A9C57DA584710A" box="[151,386,1800,1821]" gridcol="0" gridrow="6" pageId="4" pageNumber="50">RATIO: Tel L/Car L</th>
<td id="3B4C5D8AFFE583A6D69CC57DA5F0710A" box="[418,502,1800,1821]" gridcol="1" gridrow="6" pageId="4" pageNumber="50">1.24</td>
<td id="3B4C5D8AFFE583A6D521C57DA649710A" box="[543,591,1800,1821]" gridcol="2" gridrow="6" pageId="4" pageNumber="50">1.25</td>
<td id="3B4C5D8AFFE583A6D5A3C57DA7CF710A" box="[669,969,1800,1821]" gridcol="3" gridrow="6" pageId="4" pageNumber="50">1.24 1.27 1.25</td>
<td id="3B4C5D8AFFE583A6D328C57DA067710A" box="[1046,1121,1800,1821]" gridcol="4" gridrow="6" pageId="4" pageNumber="50">1.25</td>
<td id="3B4C5D8AFFE583A6D3AAC57DA0F5710A" box="[1172,1267,1800,1821]" gridcol="5" gridrow="6" pageId="4" pageNumber="50">0.012247</td>
<td id="3B4C5D8AFFE583A6D22CC57DA19B710A" box="[1298,1437,1800,1821]" gridcol="6" gridrow="6" pageId="4" pageNumber="50">1.08</td>
</tr>
</table>
</paragraph>
<caption id="92D2663CFFE47C5CD7A9C7AEA6277003" httpUri="https://zenodo.org/record/190123/files/figure.png" pageId="5" pageNumber="51" targetBox="[179,1398,214,1458]" targetPageId="5">
<paragraph id="C61236B4FFE47C5CD7A9C7AEA6277003" blockId="5.[151,1436,1499,1556]" pageId="5" pageNumber="51">
<emphasis id="F4D9EAA6FFE47C5CD7A9C7AEA54573E4" bold="true" box="[151,323,1499,1523]" pageId="5" pageNumber="51">FIGURES 7–8.</emphasis>
Holotype female of 
<taxonomicName id="01AD4D37FFE47C5CD523C7AEA6DF73E5" box="[541,729,1499,1522]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="chamuco" status="sp. nov.">
<emphasis id="F4D9EAA6FFE47C5CD523C7AEA6DF73E5" box="[541,729,1499,1522]" italics="true" pageId="5" pageNumber="51">Alacran chamuco</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="EFEA57DDFFE47C5CD5D8C7AEA73D73E4" box="[742,827,1499,1523]" pageId="5" pageNumber="51" rank="species">
<emphasis id="F4D9EAA6FFE47C5CD5D8C7AEA73D73E4" bold="true" box="[742,827,1499,1523]" pageId="5" pageNumber="51">sp. nov.</emphasis>
</taxonomicNameLabel>
7. Dorsal view of right pedipalp femur. 8. Dorsal view of right pedipalp patella (scale = 1mm).
</paragraph>
</caption>
<paragraph id="C61236B4FFE47C5CD7F8C432A6C57135" blockId="5.[151,1437,1607,2020]" pageId="5" pageNumber="51">
Mesosoma: Tergites I–VI smooth, lustrous, acarinate. Tergite VII with scattered, medium-sized granules posterolaterally; without distinct carinae; lateral margins finely serrate. Genital operculum obovate, with a complete median longitudinal membranous connection. Pectines each with three distinct marginal lamellae, five teeth and no fulcra; teeth finger-like, 3–4 times longer than wide, with sensory pegs on distal 1/3 to ¼ only (
<figureCitation id="5E962A31FFE47C5CD7E5C497A52570EB" box="[219,291,1762,1788]" captionStart="FIGURES 2 – 4" captionStartId="3.[151,274,566,590]" captionTargetBox="[155,1430,197,539]" captionTargetId="figure@3.[151,1436,197,541]" captionTargetPageId="3" captionText="FIGURES 2 – 4. Holotype female of Alacran chamuco, sp. nov. 2. Dorsal aspect of prosoma. 3. Ventral aspect of prosoma + metasoma. 4. Detail of right pectinal tooth (scales = 1 mm)." httpUri="https://zenodo.org/record/190121/files/figure.png" pageId="5" pageNumber="51">Fig. 4</figureCitation>
). Sternites III–VI smooth, lustrous, with few scattered macrosetae; stigmata small, round. Sternite VII with very few scattered granules, acarinate.
</paragraph>
<paragraph id="C61236B4FFE47C5CD7FBC55AA11971AA" blockId="5.[151,1437,1607,2020]" pageId="5" pageNumber="51">
Metasoma: All segments longer than wide. I and II with three pairs of carinae each: Dorsolateral, ventrolateral and lateral supramedian keels moderately strong, granulose; III–V with two pairs of carinae each, dorsolateral and ventrolateral, all moderately strong, granulose. Intercarinal spaces smooth, lustrous. Telson globose, smooth, lustrous; aculeus short, strongly curved (
<figureCitation id="5E962A31FFE47C5CD4A8C5D6A7D871AA" box="[918,990,1955,1981]" captionStart="FIGURES 5 – 6" captionStartId="3.[151,269,1976,2000]" captionTargetBox="[157,1423,699,1929]" captionTargetId="figure@3.[151,1436,689,1954]" captionTargetPageId="3" captionText="FIGURES 5 – 6. Metasomal segment V and telson. 5. Holotype female of Alacran chamuco, sp. nov. 6. Adult female of Alacran tartarus Francke (scale = 1 mm)." httpUri="https://zenodo.org/record/190122/files/figure.png" pageId="5" pageNumber="51">Fig. 6</figureCitation>
), without lateral serrations.
</paragraph>
<paragraph id="C61236B4FFE47C5FD7FBC5BFA7F876CF" blockId="5.[151,1437,1607,2020]" lastBlockId="6.[151,1438,152,526]" lastPageId="6" lastPageNumber="52" pageId="5" pageNumber="51">Chelicera: Fixed finger with four teeth; basal and middle teeth not forming a bicusp. Movable finger dorsal margin with five teeth; ventral margin distinctly ridged, with distal tooth twice as long as dorsal counterpart. Both fingers densely covered with long, white hairs ventrally.</paragraph>
<paragraph id="C61236B4FFE77C5FD7FBC290A17E775B" blockId="6.[151,1438,152,526]" pageId="6" pageNumber="52">
Pedipalp femur: Strongly compressed dorsoventrally, about 3–4 times wider than deep; with five distinct, granular carinae (prodorsal, proventral, retrodorsal, retromedian and retroventral), and with three trichobothria (
<figureCitation id="5E962A31FFE77C5FD600C347A580775B" box="[318,390,306,332]" captionStart="FIGURES 7 – 8" captionStartId="5.[151,269,1499,1523]" captionTargetBox="[179,1398,214,1458]" captionTargetId="figure@5.[157,1430,194,1475]" captionTargetPageId="5" captionText="FIGURES 7 – 8. Holotype female of Alacran chamuco, sp. nov. 7. Dorsal view of right pedipalp femur. 8. Dorsal view of right pedipalp patella (scale = 1 mm)." httpUri="https://zenodo.org/record/190123/files/figure.png" pageId="6" pageNumber="52">Fig. 7</figureCitation>
). Dorsal face with few scattered granules; ventral face with very few granules basally.
</paragraph>
<paragraph id="C61236B4FFE77C5FD7F8C32CA64E7419" blockId="6.[151,1438,152,526]" pageId="6" pageNumber="52">
Pedipalp patella: Subquadrangular in cross-section, slightly wider than deep. Four distinct, granular carinae: prodorsal, proventral, retrodorsal and retroventral. Anterior face smooth, lustrous. Dorsal face with three trichobothria, each surrounded by scattered granules (
<figureCitation id="5E962A31FFE77C5FD45CC3D3A7AA77D7" box="[866,940,422,448]" captionStart="FIGURES 7 – 8" captionStartId="5.[151,269,1499,1523]" captionTargetBox="[179,1398,214,1458]" captionTargetId="figure@5.[157,1430,194,1475]" captionTargetPageId="5" captionText="FIGURES 7 – 8. Holotype female of Alacran chamuco, sp. nov. 7. Dorsal view of right pedipalp femur. 8. Dorsal view of right pedipalp patella (scale = 1 mm)." httpUri="https://zenodo.org/record/190123/files/figure.png" pageId="6" pageNumber="52">Fig. 8</figureCitation>
). Posterior face with 19–20 trichobothria (
<figureCitation id="5E962A31FFE77C5FD79EC3B8A53477F0" box="[160,306,461,487]" captionStart="FIGURES 9 – 11" captionStartId="6.[151,269,1747,1771]" captionTargetBox="[158,1432,597,1704]" captionTargetId="figure@6.[151,1436,581,1723]" captionTargetPageId="6" captionText="FIGURES 9 – 11. Trichobothrial pattern on external aspect of pedipalp patella. 9. Holotype female of Alacran chamuco, sp. nov., left patella with 21 external trichobothria; trichobothrium not present on the right patella is highlighted in black; relative positions of the 3 ventral trichobothria also indicated. 10. Holotype female of Alacran chamuco, sp. nov., right patella with 20 external trichobothria. 11. Composite illustration, prepared directly from the camera lucida (by drawing one pattern directly over the other), showing the trichobothrial patterns on external aspect of the pedipalp patella of the holotype female of Alacran chamuco sp. nov. (circles), and an adult female Alacran tartarus Francke (crosses) (scale = 1 mm)." httpUri="https://zenodo.org/record/190124/files/figure.png" pageId="6" pageNumber="52">Figs. 9, 10</figureCitation>
), each surrounded by granules. Ventral face with sparse scattered granules; with three trichobothria along posterior margin.
</paragraph>
<caption id="92D2663CFFE77C5FD7A9C4A6A4D871AD" httpUri="https://zenodo.org/record/190124/files/figure.png" pageId="6" pageNumber="52" targetBox="[158,1432,597,1704]" targetPageId="6">
<paragraph id="C61236B4FFE77C5FD7A9C4A6A4D871AD" blockId="6.[151,1436,1747,1978]" pageId="6" pageNumber="52">
<emphasis id="F4D9EAA6FFE77C5FD7A9C4A6A54970FC" bold="true" box="[151,335,1747,1771]" pageId="6" pageNumber="52">FIGURES 9–11.</emphasis>
Trichobothrial pattern on external aspect of pedipalp patella. 9. Holotype female of 
<taxonomicName id="01AD4D37FFE77C5FD3E6C4A6A19370FD" box="[1240,1429,1747,1770]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="chamuco" status="sp. nov.">
<emphasis id="F4D9EAA6FFE77C5FD3E6C4A6A19370FD" box="[1240,1429,1747,1770]" italics="true" pageId="6" pageNumber="52">Alacran chamuco</emphasis>
</taxonomicName>
, 
<emphasis id="F4D9EAA6FFE77C5FD7A9C480A4F4711A" bold="true" box="[151,242,1781,1805]" pageId="6" pageNumber="52">
<taxonomicNameLabel id="EFEA57DDFFE77C5FD7A9C480A4EB711A" box="[151,237,1781,1805]" pageId="6" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
left patella with 21 external trichobothria; trichobothrium not present on the right patella is highlighted in black; relative positions of the 3 ventral trichobothria also indicated. 10. Holotype female of 
<taxonomicName id="01AD4D37FFE77C5FD307C56CA0F17127" box="[1081,1271,1817,1840]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="chamuco" status="sp. nov.">
<emphasis id="F4D9EAA6FFE77C5FD307C56CA0F17127" box="[1081,1271,1817,1840]" italics="true" pageId="6" pageNumber="52">Alacran chamuco</emphasis>
</taxonomicName>
, 
<emphasis id="F4D9EAA6FFE77C5FD23BC56DA1647127" bold="true" box="[1285,1378,1816,1840]" pageId="6" pageNumber="52">
<taxonomicNameLabel id="EFEA57DDFFE77C5FD23BC56DA15B7127" box="[1285,1373,1816,1840]" pageId="6" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
right patella with 20 external trichobothria. 11. Composite illustration, prepared directly from the camera lucida (by drawing one pattern directly over the other), showing the trichobothrial patterns on external aspect of the pedipalp patella of the holotype female of 
<taxonomicName id="01AD4D37FFE77C5FD657C5F4A620718F" box="[361,550,1921,1944]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="chamuco" status="sp. nov.">
<emphasis id="F4D9EAA6FFE77C5FD657C5F4A620718F" box="[361,550,1921,1944]" italics="true" pageId="6" pageNumber="52">Alacran chamuco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EFEA57DDFFE77C5FD513C5F5A682718F" box="[557,644,1920,1944]" pageId="6" pageNumber="52" rank="species">
<emphasis id="F4D9EAA6FFE77C5FD513C5F5A682718F" bold="true" box="[557,644,1920,1944]" pageId="6" pageNumber="52">sp. nov.</emphasis>
</taxonomicNameLabel>
(circles), and an adult female 
<taxonomicName id="01AD4D37FFE77C5FD4F4C5F4A0DC7180" authority="Francke" authorityName="Francke" box="[970,1242,1920,1944]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="tartarus">
<emphasis id="F4D9EAA6FFE77C5FD4F4C5F4A07B718F" box="[970,1149,1921,1944]" italics="true" pageId="6" pageNumber="52">Alacran tartarus</emphasis>
Francke
</taxonomicName>
(crosses) (scale = 1mm).
</paragraph>
</caption>
<caption id="92D2663CFFE67C5ED7A9C1C1A5FA75F9" httpUri="https://zenodo.org/record/190125/files/figure.png" pageId="7" pageNumber="53" targetBox="[157,1433,202,922]" targetPageId="7">
<paragraph id="C61236B4FFE67C5ED7A9C1C1A5FA75F9" blockId="7.[151,1436,948,1007]" pageId="7" pageNumber="53">
<emphasis id="F4D9EAA6FFE67C5ED7A9C1C1A56575DB" bold="true" box="[151,355,948,972]" pageId="7" pageNumber="53">FIGURES 12–13.</emphasis>
Dorsal view of left pedipalp chela. 12. Holotype female of 
<taxonomicName id="01AD4D37FFE67C5ED33EC1C0A0C575DB" box="[1024,1219,949,972]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="chamuco" status="sp. nov.">
<emphasis id="F4D9EAA6FFE67C5ED33EC1C0A0C575DB" box="[1024,1219,949,972]" italics="true" pageId="7" pageNumber="53">Alacran chamuco</emphasis>
</taxonomicName>
, 
<emphasis id="F4D9EAA6FFE67C5ED3ECC1C1A12A75DB" bold="true" box="[1234,1324,948,972]" pageId="7" pageNumber="53">
<taxonomicNameLabel id="EFEA57DDFFE67C5ED3ECC1C1A12A75DB" box="[1234,1324,948,972]" pageId="7" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
13. Adult female 
<taxonomicName id="01AD4D37FFE67C5ED7D8C1ADA5FA75F9" authority="Francke." authorityName="Francke." box="[230,508,983,1007]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="tartarus">
<emphasis id="F4D9EAA6FFE67C5ED7D8C1ADA59F75F8" box="[230,409,984,1007]" italics="true" pageId="7" pageNumber="53">Alacran tartarus</emphasis>
Francke.
</taxonomicName>
</paragraph>
</caption>
<caption id="92D2663CFFE67C5ED7A9C5C0A69B71F8" httpUri="https://zenodo.org/record/190126/files/figure.png" pageId="7" pageNumber="53" targetBox="[162,1410,1085,1935]" targetPageId="7">
<paragraph id="C61236B4FFE67C5ED7A9C5C0A69B71F8" blockId="7.[151,1436,1973,2031]" pageId="7" pageNumber="53">
<emphasis id="F4D9EAA6FFE67C5ED7A9C5C0A56771DA" bold="true" box="[151,353,1973,1997]" pageId="7" pageNumber="53">FIGURES 14–15.</emphasis>
External view of left pedipalp chela. 14. Holotype female of 
<taxonomicName id="01AD4D37FFE67C5ED338C5C0A0C171DB" box="[1030,1223,1973,1996]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="chamuco" status="sp. nov.">
<emphasis id="F4D9EAA6FFE67C5ED338C5C0A0C171DB" box="[1030,1223,1973,1996]" italics="true" pageId="7" pageNumber="53">Alacran chamuco</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="EFEA57DDFFE67C5ED3EBC5C0A12B71DA" box="[1237,1325,1973,1997]" pageId="7" pageNumber="53" rank="species">
<emphasis id="F4D9EAA6FFE67C5ED3EBC5C0A12B71DA" bold="true" box="[1237,1325,1973,1997]" pageId="7" pageNumber="53">sp. nov.</emphasis>
</taxonomicNameLabel>
15. Adult female 
<taxonomicName id="01AD4D37FFE67C5ED7D8C5ADA5F371F8" authority="Francke" authorityName="Francke" box="[230,501,2008,2031]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="tartarus">
<emphasis id="F4D9EAA6FFE67C5ED7D8C5ADA59F71F8" box="[230,409,2008,2031]" italics="true" pageId="7" pageNumber="53">Alacran tartarus</emphasis>
Francke
</taxonomicName>
(scale = 1mm).
</paragraph>
</caption>
<caption id="92D2663CFFE97C51D7A9C691A0C47309" httpUri="https://zenodo.org/record/190127/files/figure.png" pageId="8" pageNumber="54" targetBox="[173,1410,206,1214]" targetPageId="8">
<paragraph id="C61236B4FFE97C51D7A9C691A0C47309" blockId="8.[151,1436,1252,1310]" pageId="8" pageNumber="54">
<emphasis id="F4D9EAA6FFE97C51D7A9C691A56672EB" bold="true" box="[151,352,1252,1276]" pageId="8" pageNumber="54">FIGURES 16–19.</emphasis>
Dentition on pedipalp chela fingers. 16, 18. Holotype female of 
<taxonomicName id="01AD4D37FFE97C51D31AC691A0E272EC" box="[1060,1252,1252,1275]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="54" phylum="Arthropoda" rank="species" species="chamuco" status="sp. nov.">
<emphasis id="F4D9EAA6FFE97C51D31AC691A0E272EC" box="[1060,1252,1252,1275]" italics="true" pageId="8" pageNumber="54">Alacran chamuco</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="EFEA57DDFFE97C51D3CCC691A14C72EB" box="[1266,1354,1252,1276]" pageId="8" pageNumber="54" rank="species">
<emphasis id="F4D9EAA6FFE97C51D3CCC691A14C72EB" bold="true" box="[1266,1354,1252,1276]" pageId="8" pageNumber="54">sp. nov.</emphasis>
</taxonomicNameLabel>
17, 19. Adult female 
<taxonomicName id="01AD4D37FFE97C51D617C772A6467309" authority="Francke." authorityName="Francke." box="[297,576,1287,1310]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="54" phylum="Arthropoda" rank="species" species="tartarus">
<emphasis id="F4D9EAA6FFE97C51D617C772A5DA7309" box="[297,476,1287,1310]" italics="true" pageId="8" pageNumber="54">Alacran tartarus</emphasis>
Francke.
</taxonomicName>
16, 17. Fixed finger. 18, 19. Movable finger (scale = 1mm).
</paragraph>
</caption>
<caption id="92D2663CFFE97C51D7A9C5E9A07071A4" box="[151,1142,1948,1972]" httpUri="https://zenodo.org/record/190128/files/figure.png" pageId="8" pageNumber="54" targetBox="[166,1435,1376,1915]" targetPageId="8">
<paragraph id="C61236B4FFE97C51D7A9C5E9A07071A4" blockId="8.[151,1142,1948,1972]" box="[151,1142,1948,1972]" pageId="8" pageNumber="54">
<emphasis id="F4D9EAA6FFE97C51D7A9C5E9A52071A3" bold="true" box="[151,294,1948,1972]" pageId="8" pageNumber="54">FIGURE 20.</emphasis>
Telotarsus III of holotype female of 
<taxonomicName id="01AD4D37FFE97C51D591C5E9A76A71A4" box="[687,876,1948,1971]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="54" phylum="Arthropoda" rank="species" species="chamuco" status="sp. nov.">
<emphasis id="F4D9EAA6FFE97C51D591C5E9A76A71A4" box="[687,876,1948,1971]" italics="true" pageId="8" pageNumber="54">Alacran chamuco</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="EFEA57DDFFE97C51D447C5E9A7C871A3" box="[889,974,1948,1972]" pageId="8" pageNumber="54" rank="species">
<emphasis id="F4D9EAA6FFE97C51D447C5E9A7C871A3" bold="true" box="[889,974,1948,1972]" pageId="8" pageNumber="54">sp. nov.</emphasis>
</taxonomicNameLabel>
(scale = 1mm).
</paragraph>
</caption>
<paragraph id="C61236B4FFE87C50D7F8C2EDA4C377D7" blockId="9.[151,1437,152,1376]" pageId="9" pageNumber="55">
Pedipalp chela: Digital carina with fairly distinct, single row of medium-sized granules. Dorsal secondary carina with somewhat scattered medium-sized granules. Dorsal marginal carina with widely scattered medium-sized granules. Retrolateral secondary carina present only on basal one-half, with somewhat scattered medium-sized granules. Proventral and retroventral carinae with somewhat scattered, medium-sized granules. Prodorsal and proventral carinae weak, with few scattered granules (
<figureCitation id="5E962A31FFE87C50D378C347A0AD775B" box="[1094,1195,306,332]" captionStart="FIGURES 12 – 13" captionStartId="7.[151,273,948,972]" captionTargetBox="[157,1433,202,922]" captionTargetId="figure@7.[151,1436,194,931]" captionTargetPageId="7" captionText="FIGURES 12 – 13. Dorsal view of left pedipalp chela. 12. Holotype female of Alacran chamuco, sp. nov. 13. Adult female Alacran tartarus Francke." httpUri="https://zenodo.org/record/190125/files/figure.png" pageId="9" pageNumber="55">Figs. 12</figureCitation>
, 
<figureCitation id="5E962A31FFE87C50D387C347A0DC775B" box="[1209,1242,306,332]" captionStart="FIGURES 14 – 15" captionStartId="7.[151,271,1973,1997]" captionTargetBox="[162,1410,1085,1935]" captionTargetId="figure@7.[156,1430,1060,1948]" captionTargetPageId="7" captionText="FIGURES 14 – 15. External view of left pedipalp chela. 14. Holotype female of Alacran chamuco, sp. nov. 15. Adult female Alacran tartarus Francke (scale = 1 mm)." httpUri="https://zenodo.org/record/190126/files/figure.png" pageId="9" pageNumber="55">14</figureCitation>
). Trichobothrial pattern as in 
<emphasis id="F4D9EAA6FFE87C50D604C32CA5CD7765" box="[314,459,345,370]" italics="true" pageId="9" pageNumber="55">
<taxonomicName id="01AD4D37FFE87C50D604C32CA5C17765" box="[314,455,345,370]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="55" phylum="Arthropoda" rank="species" species="tartarus">A. tartarus</taxonomicName>
.
</emphasis>
Fixed finger with six distinct rows of granules, with five outer and five inner denticles (
<figureCitation id="5E962A31FFE87C50D62FC3F5A56D778D" box="[273,363,384,410]" captionStart="FIGURES 16 – 19" captionStartId="8.[151,271,1252,1276]" captionTargetBox="[173,1410,206,1214]" captionTargetId="figure@8.[156,1430,194,1228]" captionTargetPageId="8" captionText="FIGURES 16 – 19. Dentition on pedipalp chela fingers. 16, 18. Holotype female of Alacran chamuco, sp. nov. 17, 19. Adult female Alacran tartarus Francke. 16, 17. Fixed finger. 18, 19. Movable finger (scale = 1 mm)." httpUri="https://zenodo.org/record/190127/files/figure.png" pageId="9" pageNumber="55">Fig. 16</figureCitation>
). Movable finger with seven rows of granules, with six outer and six inner denticles (
<figureCitation id="5E962A31FFE87C50D252C3F5A4B177D7" captionStart="FIGURES 16 – 19" captionStartId="8.[151,271,1252,1276]" captionTargetBox="[173,1410,206,1214]" captionTargetId="figure@8.[156,1430,194,1228]" captionTargetPageId="8" captionText="FIGURES 16 – 19. Dentition on pedipalp chela fingers. 16, 18. Holotype female of Alacran chamuco, sp. nov. 17, 19. Adult female Alacran tartarus Francke. 16, 17. Fixed finger. 18, 19. Movable finger (scale = 1 mm)." httpUri="https://zenodo.org/record/190127/files/figure.png" pageId="9" pageNumber="55">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="C61236B4FFE87C50D7FBC3B8A5B07423" blockId="9.[151,1437,152,1376]" pageId="9" pageNumber="55">
Legs. Long and slender (selected measurements in 
<tableCitation id="8B2F030FFFE87C50D411C3B8A78E77F0" box="[815,904,461,487]" captionStart="TABLE 1" captionStartId="4.[151,243,151,175]" captionTargetBox="[151,1437,237,1241]" captionTargetPageId="4" captionText="TABLE 1. Measurements (in mm.) of the holotype female of Alacran chamuco, sp. nov., from Te Cimutaa, Valle Nacional, Oaxaca, Mexico. L = length, W = width, D = depth." httpUri="http://table.plazi.org/id/92D2663CFFE57C5DD7A9C2E2A70176C6" pageId="9" pageNumber="55" tableUuid="92D2663CFFE57C5DD7A9C2E2A70176C6">Table 1</tableCitation>
). Tibial spurs absent. Prolateral pedal spurs present on all legs. Retrolateral pedal spurs absent. Telotarsus without ventromedian spinules (
<figureCitation id="5E962A31FFE87C50D20EC381A1887419" box="[1328,1422,500,526]" captionStart="FIGURE 20" captionStartId="8.[151,255,1948,1972]" captionTargetBox="[166,1435,1376,1915]" captionTargetId="figure@8.[151,1436,1364,1924]" captionTargetPageId="8" captionText="FIGURE 20. Telotarsus III of holotype female of Alacran chamuco, sp. nov. (scale = 1 mm)." httpUri="https://zenodo.org/record/190128/files/figure.png" pageId="9" pageNumber="55">Fig. 20</figureCitation>
); ungues long and curved.
</paragraph>
<paragraph id="C61236B4FFE87C50D7F8C035A622750B" blockId="9.[151,1437,152,1376]" pageId="9" pageNumber="55">
<emphasis id="F4D9EAA6FFE87C50D7F8C035A575744D" bold="true" box="[198,371,576,602]" pageId="9" pageNumber="55">Comparisons.</emphasis>
<taxonomicName id="01AD4D37FFE87C50D644C034A602744D" box="[378,516,577,602]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="55" phylum="Arthropoda" rank="species" species="chamuco">
<emphasis id="F4D9EAA6FFE87C50D644C034A602744D" box="[378,516,577,602]" italics="true" pageId="9" pageNumber="55">A. chamuco</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="01AD4D37FFE87C50D5A2C034A71D744D" box="[668,795,577,602]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="55" phylum="Arthropoda" rank="species" species="tartarus">
<emphasis id="F4D9EAA6FFE87C50D5A2C034A71D744D" box="[668,795,577,602]" italics="true" pageId="9" pageNumber="55">A. tartarus</emphasis>
</taxonomicName>
, the only other known species in the genus, as follows: (1) on 
<taxonomicName id="01AD4D37FFE87C50D7DBC01DA5657496" box="[229,355,616,641]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="55" phylum="Arthropoda" rank="species" species="tartarus">
<emphasis id="F4D9EAA6FFE87C50D7DBC01DA5657496" box="[229,355,616,641]" italics="true" pageId="9" pageNumber="55">A. tartarus</emphasis>
</taxonomicName>
the fixed finger of the pedipalps has seven rows of granules (
<figureCitation id="5E962A31FFE87C50D30AC01DA0847495" box="[1076,1154,616,642]" captionStart="FIGURES 16 – 19" captionStartId="8.[151,271,1252,1276]" captionTargetBox="[173,1410,206,1214]" captionTargetId="figure@8.[156,1430,194,1228]" captionTargetPageId="8" captionText="FIGURES 16 – 19. Dentition on pedipalp chela fingers. 16, 18. Holotype female of Alacran chamuco, sp. nov. 17, 19. Adult female Alacran tartarus Francke. 16, 17. Fixed finger. 18, 19. Movable finger (scale = 1 mm)." httpUri="https://zenodo.org/record/190127/files/figure.png" pageId="9" pageNumber="55">Fig.17</figureCitation>
) instead of six (
<figureCitation id="5E962A31FFE87C50D200C01DA1957495" box="[1342,1427,616,642]" captionStart="FIGURES 16 – 19" captionStartId="8.[151,271,1252,1276]" captionTargetBox="[173,1410,206,1214]" captionTargetId="figure@8.[156,1430,194,1228]" captionTargetPageId="8" captionText="FIGURES 16 – 19. Dentition on pedipalp chela fingers. 16, 18. Holotype female of Alacran chamuco, sp. nov. 17, 19. Adult female Alacran tartarus Francke. 16, 17. Fixed finger. 18, 19. Movable finger (scale = 1 mm)." httpUri="https://zenodo.org/record/190127/files/figure.png" pageId="9" pageNumber="55">Fig. 16</figureCitation>
) clearly separated by six outer denticles instead of five; (2) the movable finger has seven inner denticles (
<figureCitation id="5E962A31FFE87C50D252C0FBA4B074D8" captionStart="FIGURES 16 – 19" captionStartId="8.[151,271,1252,1276]" captionTargetBox="[173,1410,206,1214]" captionTargetId="figure@8.[156,1430,194,1228]" captionTargetPageId="8" captionText="FIGURES 16 – 19. Dentition on pedipalp chela fingers. 16, 18. Holotype female of Alacran chamuco, sp. nov. 17, 19. Adult female Alacran tartarus Francke. 16, 17. Fixed finger. 18, 19. Movable finger (scale = 1 mm)." httpUri="https://zenodo.org/record/190127/files/figure.png" pageId="9" pageNumber="55">Fig. 19</figureCitation>
) instead of six (
<figureCitation id="5E962A31FFE87C50D6BFC0C0A5D874D8" box="[385,478,693,719]" captionStart="FIGURES 16 – 19" captionStartId="8.[151,271,1252,1276]" captionTargetBox="[173,1410,206,1214]" captionTargetId="figure@8.[156,1430,194,1228]" captionTargetPageId="8" captionText="FIGURES 16 – 19. Dentition on pedipalp chela fingers. 16, 18. Holotype female of Alacran chamuco, sp. nov. 17, 19. Adult female Alacran tartarus Francke. 16, 17. Fixed finger. 18, 19. Movable finger (scale = 1 mm)." httpUri="https://zenodo.org/record/190127/files/figure.png" pageId="9" pageNumber="55">Fig. 18</figureCitation>
); (3) the telson is considerably larger (
<tableCitation id="8B2F030FFFE87C50D4FEC0C0A02774D8" box="[960,1057,693,719]" captionStart="TABLE 2" captionStartId="4.[151,243,1354,1378]" captionTargetBox="[151,1437,1551,1821]" captionTargetPageId="4" captionText="TABLE 2. Morphometric comparisons between the holotype female of Alacran chamuco, sp. nov., and five adult females of Alacran tartarus Francke, including the holotype. Abbreviations are L = length, W = width and D = depth. Measurements in the first two columns are from Francke (1982); the second two are the WDS specimens measured by Vignoli &amp; Prendini (2009, Table 6, pp. 36 – 37)." httpUri="http://table.plazi.org/id/92D2663CFFE57C5DD7A9C73FA68373DE" pageId="9" pageNumber="55" tableUuid="92D2663CFFE57C5DD7A9C73FA68373DE">Table 2</tableCitation>
; 
<figureCitation id="5E962A31FFE87C50D30EC0C0A0AE74D8" box="[1072,1192,693,719]" captionStart="FIGURES 5 – 6" captionStartId="3.[151,269,1976,2000]" captionTargetBox="[157,1423,699,1929]" captionTargetId="figure@3.[151,1436,689,1954]" captionTargetPageId="3" captionText="FIGURES 5 – 6. Metasomal segment V and telson. 5. Holotype female of Alacran chamuco, sp. nov. 6. Adult female of Alacran tartarus Francke (scale = 1 mm)." httpUri="https://zenodo.org/record/190122/files/figure.png" pageId="9" pageNumber="55">Figs. 5, 6</figureCitation>
); (4) the carinae on metasomal segment V are more finely granulose (
<figureCitation id="5E962A31FFE87C50D5E1C0A9A75674E1" box="[735,848,732,758]" captionStart="FIGURES 5 – 6" captionStartId="3.[151,269,1976,2000]" captionTargetBox="[157,1423,699,1929]" captionTargetId="figure@3.[151,1436,689,1954]" captionTargetPageId="3" captionText="FIGURES 5 – 6. Metasomal segment V and telson. 5. Holotype female of Alacran chamuco, sp. nov. 6. Adult female of Alacran tartarus Francke (scale = 1 mm)." httpUri="https://zenodo.org/record/190122/files/figure.png" pageId="9" pageNumber="55">Figs. 5, 6</figureCitation>
); (5) the pedipalp chela is rounder and the carinae are more granulose (
<figureCitation id="5E962A31FFE87C50D6B9C177A611750B" box="[391,535,770,796]" captionStart-0="FIGURES 12 – 13" captionStart-1="FIGURES 14 – 15" captionStartId-0="7.[151,273,948,972]" captionStartId-1="7.[151,271,1973,1997]" captionTargetBox-0="[157,1433,202,922]" captionTargetBox-1="[162,1410,1085,1935]" captionTargetId-0="figure@7.[151,1436,194,931]" captionTargetId-1="figure@7.[156,1430,1060,1948]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionText-0="FIGURES 12 – 13. Dorsal view of left pedipalp chela. 12. Holotype female of Alacran chamuco, sp. nov. 13. Adult female Alacran tartarus Francke." captionText-1="FIGURES 14 – 15. External view of left pedipalp chela. 14. Holotype female of Alacran chamuco, sp. nov. 15. Adult female Alacran tartarus Francke (scale = 1 mm)." httpUri-0="https://zenodo.org/record/190125/files/figure.png" httpUri-1="https://zenodo.org/record/190126/files/figure.png" pageId="9" pageNumber="55">Figs. 12–15</figureCitation>
).
</paragraph>
<paragraph id="C61236B4FFE87C50D7F8C15CA515723C" blockId="9.[151,1437,152,1376]" pageId="9" pageNumber="55">
Although both species posses enlarged vesicles, a phenomenon known in several other troglobitic scorpions (Volschenk &amp; Prendini, 2008), in the new species it is not as large as in 
<taxonomicName id="01AD4D37FFE87C50D3ABC125A11D757E" box="[1173,1307,848,873]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="55" phylum="Arthropoda" rank="species" species="tartarus">
<emphasis id="F4D9EAA6FFE87C50D3ABC125A11D757E" box="[1173,1307,848,873]" italics="true" pageId="9" pageNumber="55">A. tartarus</emphasis>
</taxonomicName>
(
<tableCitation id="8B2F030FFFE87C50D212C125A188757D" box="[1324,1422,848,874]" captionStart="TABLE 2" captionStartId="4.[151,243,1354,1378]" captionTargetBox="[151,1437,1551,1821]" captionTargetPageId="4" captionText="TABLE 2. Morphometric comparisons between the holotype female of Alacran chamuco, sp. nov., and five adult females of Alacran tartarus Francke, including the holotype. Abbreviations are L = length, W = width and D = depth. Measurements in the first two columns are from Francke (1982); the second two are the WDS specimens measured by Vignoli &amp; Prendini (2009, Table 6, pp. 36 – 37)." httpUri="http://table.plazi.org/id/92D2663CFFE57C5DD7A9C73FA68373DE" pageId="9" pageNumber="55" tableUuid="92D2663CFFE57C5DD7A9C73FA68373DE">Table 2</tableCitation>
). Considering the five adult females of 
<taxonomicName id="01AD4D37FFE87C50D569C102A6D17587" box="[599,727,887,912]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="55" phylum="Arthropoda" rank="species" species="tartarus">
<emphasis id="F4D9EAA6FFE87C50D569C102A6D17587" box="[599,727,887,912]" italics="true" pageId="9" pageNumber="55">A. tartarus</emphasis>
</taxonomicName>
(mean and standard deviation) and comparing them against the single 
<taxonomicName id="01AD4D37FFE87C50D62CC1E8A59875A1" box="[274,414,925,950]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="55" phylum="Arthropoda" rank="species" species="chamuco">
<emphasis id="F4D9EAA6FFE87C50D62CC1E8A59875A1" box="[274,414,925,950]" italics="true" pageId="9" pageNumber="55">A. chamuco</emphasis>
</taxonomicName>
available we find statistically highly significant differences between them: (a) for the ratio telson L/carapace L C = (1.25–1.08)/0.0122=13.93 (p&lt;0.001); (b) for the ratio telson W/carapace L C = (0.584–0.47)/0.0261=4.37 (p&lt;0.001); and (c) for the ratio telson D/carapace L C = (0.566–0.43)/0.0305=4.46 (p&lt;0.001).
</paragraph>
<paragraph id="C61236B4FFE87C50D7F8C642A0887377" blockId="9.[151,1437,152,1376]" pageId="9" pageNumber="55">
<emphasis id="F4D9EAA6FFE87C50D7F8C642A65E7246" bold="true" box="[198,600,1079,1105]" pageId="9" pageNumber="55">Biogeographical considerations.</emphasis>
The caves in which the two species of 
<taxonomicName id="01AD4D37FFE87C50D300C64DA09A7246" box="[1086,1180,1080,1105]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="F4D9EAA6FFE87C50D300C64DA09A7246" box="[1086,1180,1080,1105]" italics="true" pageId="9" pageNumber="55">Alacran</emphasis>
</taxonomicName>
have been found are only about 
<quantity id="01559B51FFE87C50D625C62BA560726F" box="[283,358,1118,1144]" metricMagnitude="4" metricUnit="m" metricValue="7.5" pageId="9" pageNumber="55" unit="km" value="75.0">75 km</quantity>
apart on a straight line (
<figureCitation id="5E962A31FFE87C50D5BBC62BA6CD726F" box="[645,715,1118,1144]" captionStart="FIGURE 1" captionStartId="2.[151,255,1882,1906]" captionTargetBox="[222,1352,563,1882]" captionTargetId="figure@2.[222,1365,547,1882]" captionTargetPageId="2" captionText="FIGURE 1. Map of the state of Oaxaca, Mexico, indicating the known distributions for the troglobitic Alacran chamuco, sp. nov. (circle), and Alacran tartarus Francke (triangle)." httpUri="https://zenodo.org/record/190120/files/figure.png" pageId="9" pageNumber="55">Fig. 1</figureCitation>
). However, those on the Huautla Plateau are located at about 
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in elevation, whereas Te Cimutaa is at 
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.
</specimenCount>
Furthermore two deep canyons separate the Mazateca and the Chinateca regions: in a straight-line intersect from Huautla to Valle Nacional the contour first climbs to about 
<quantity id="01559B51FFE87C50D63FC6A6A55172FA" box="[257,343,1235,1261]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="55" unit="m" value="2300.0">2300m</quantity>
; then, the canyon of the Rio Santo 
<collectingCountry id="BEBA7624FFE87C50D43FC6A6A77772FA" box="[769,881,1235,1261]" name="Dominican Republic" pageId="9" pageNumber="55">Domingo</collectingCountry>
has an elevation of approximately 
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; the contour raises again to 
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in elevation before dropping once more, at the Rio Perfume, down to 
<quantity id="01559B51FFE87C50D20AC68CA1787304" box="[1332,1406,1273,1299]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="55" unit="m" value="100.0">100 m</quantity>
in elevation; finally, is the climb to Te Cimutaa (topographical contour lines shown in 
<figureCitation id="5E962A31FFE87C50D3BAC755A0CF732D" box="[1156,1225,1312,1338]" captionStart="FIGURE 1" captionStartId="2.[151,255,1882,1906]" captionTargetBox="[222,1352,563,1882]" captionTargetId="figure@2.[222,1365,547,1882]" captionTargetPageId="2" captionText="FIGURE 1. Map of the state of Oaxaca, Mexico, indicating the known distributions for the troglobitic Alacran chamuco, sp. nov. (circle), and Alacran tartarus Francke (triangle)." httpUri="https://zenodo.org/record/190120/files/figure.png" pageId="9" pageNumber="55">Fig. 1</figureCitation>
). The probability that the two cave systems are interconnected beneath the Sierra Madre is infinitesimal.
</paragraph>
</subSubSection>
</treatment>
</document>