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<document id="3383B2B726C72509415FEB46CD354ABF" ID-DOI="10.5281/zenodo.275907" ID-GBIF-Dataset="d5d27825-82a5-4aaf-b47d-35d24b33aaaa" ID-ISSN="1175-5326" ID-Zenodo-Dep="275907" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460189977646" checkinUser="plazi" docAuthor="Lott, Derek A." docDate="2010" docId="4D54A843FFA41A7C0694FAB5EF59FB92" docLanguage="en" docName="zt02402p051.pdf" docOrigin="Zootaxa 2402" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Acylophorus trigonocephalus Cameron" docType="treatment" docVersion="5" lastPageNumber="30" masterDocId="B16DD03BFFB81A610603FFAFEE40FFEE" masterDocTitle="The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa" masterLastPageNumber="51" masterPageNumber="1" pageNumber="29" updateTime="1698236417163" updateUser="plazi">
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<mods:title id="FD996C52CEE8662D445AC9B5761989BE">The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa</mods:title>
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<mods:namePart id="203DD3E34CC7FBE1874D38B6E9C9D796">Lott, Derek A.</mods:namePart>
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<mods:date id="B57268467D9DA08253726D3584A31228">2010</mods:date>
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<mods:identifier id="ECC8F5736DF7B433B10CE5A2F548866D" type="DOI">10.5281/zenodo.275907</mods:identifier>
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<treatment id="4D54A843FFA41A7C0694FAB5EF59FB92" ID-DOI="http://doi.org/10.5281/zenodo.6196054" ID-GBIF-Taxon="127655338" ID-Zenodo-Dep="6196054" LSID="urn:lsid:plazi:treatment:4D54A843FFA41A7C0694FAB5EF59FB92" httpUri="http://treatment.plazi.org/id/4D54A843FFA41A7C0694FAB5EF59FB92" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<subSubSection id="8DE74ADEFFA41A7D0694FAB5EFD4FAB9" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="C5421955FFA41A7D0694FAB5EC36FADA" blockId="28.[151,630,1306,1367]" box="[151,630,1306,1332]" pageId="28" pageNumber="29">
<heading id="9E0AAE39FFA41A7D0694FAB5EC36FADA" bold="true" box="[151,630,1306,1332]" fontSize="11" level="1" pageId="28" pageNumber="29" reason="1">
<taxonomicName id="02FD62D6FFA41A7D0694FAB5EC36FADA" ID-CoL="64R8X" authority="Cameron" authorityName="Cameron" box="[151,630,1306,1332]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="trigonocephalus">
<emphasis id="F789C547FFA41A7D0694FAB5EC36FADA" bold="true" box="[151,630,1306,1332]" pageId="28" pageNumber="29">
<emphasis id="F789C547FFA41A7D0694FAB5EFBAFADA" bold="true" box="[151,506,1306,1332]" italics="true" pageId="28" pageNumber="29">Acylophorus trigonocephalus</emphasis>
<collectingCountry id="BDEA59C5FFA41A7D0401FAB5EC36FADA" box="[514,630,1306,1332]" name="Cameroon" pageId="28" pageNumber="29">Cameron</collectingCountry>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="C5421955FFA41A7D0694FA92EFD4FAB9" blockId="28.[151,630,1306,1367]" box="[151,404,1341,1367]" pageId="28" pageNumber="29">(Figs 21, 70, 97, 135)</paragraph>
</subSubSection>
<subSubSection id="8DE74ADEFFA41A7D0694FA27EA10F9BC" pageId="28" pageNumber="29" type="reference_group">
<paragraph id="C5421955FFA41A7D0694FA27EDE4FA71" blockId="28.[151,1026,1416,1471]" box="[151,932,1416,1439]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="E5ED3E7BFFA41A7D0694FA27EDE4FA71" box="[151,932,1416,1439]" pageId="28" pageNumber="29">
<taxonomicName id="02FD62D6FFA41A7D0694FA27ECF6FA71" ID-CoL="64R8X" authority="Cameron, 1949: 322" authorityName="Cameron" authorityPageNumber="322" authorityYear="1949" box="[151,694,1416,1439]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="trigonocephalus">
<emphasis id="F789C547FFA41A7D0694FA27EF91FA71" box="[151,465,1416,1439]" italics="true" pageId="28" pageNumber="29">Acylophorus trigonocephalus</emphasis>
<treatmentCitation id="445C3F44FFA41A7D07DBFA27ECF6FA71" author="Cameron" box="[472,694,1416,1439]" page="322" pageId="28" pageNumber="29" year="1949">
<bibRefCitation id="A16C64A4FFA41A7D07DBFA27ECC0FA71" author="Cameron" box="[472,640,1416,1439]" pageId="28" pageNumber="50" refString="Cameron, M. (1949). New species of African Staphylinidae. Bulletin de l'Institut Fondamental d'Afrique Noire, 11, 313 - 326." type="journal article" year="1949">
<collectingCountry id="BDEA59C5FFA41A7D07DBFA27EC7DFA71" box="[472,573,1416,1439]" name="Cameroon" pageId="28" pageNumber="29">Cameron</collectingCountry>
, 1949
</bibRefCitation>
: 322
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="445C3F44FFA41A7D04C2FA27EDE0FA71" author="Herman" box="[705,928,1416,1439]" page="3037" pageId="28" pageNumber="29" year="2001">
<bibRefCitation id="A16C64A4FFA41A7D04C2FA27ED1DFA71" author="Herman" box="[705,861,1416,1439]" pageId="28" pageNumber="50" refString="Herman, L. H. (2001). Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group (part 3). Bulletin of the American Museum of Natural History, 265, 3021 - 3840." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 3037
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C5421955FFA41A7D0694FA07EA42FA51" blockId="28.[151,1026,1416,1471]" box="[151,1026,1447,1471]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="E5ED3E7BFFA41A7D0694FA07EA42FA51" box="[151,1026,1447,1471]" pageId="28" pageNumber="29">
<taxonomicName id="02FD62D6FFA41A7D0694FA07EC21FA51" ID-CoL="9W85" authority="Bordoni, 1994: 312" authorityName="Bordoni" authorityPageNumber="312" authorityYear="1994" box="[151,609,1448,1471]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="leonensis">
<emphasis id="F789C547FFA41A7D0694FA07EFC7FA51" box="[151,391,1448,1471]" italics="true" pageId="28" pageNumber="29">Acylophorus leonensis</emphasis>
<treatmentCitation id="445C3F44FFA41A7D078DFA07EC21FA51" author="Bordoni" box="[398,609,1448,1471]" page="312" pageId="28" pageNumber="29" year="1994">
<bibRefCitation id="A16C64A4FFA41A7D078DFA07EC6CFA51" author="Bordoni" box="[398,556,1448,1471]" pageId="28" pageNumber="50" refString="Bordoni, A. (1994). Nuove specie di Acylophorus Nordmann della Sierra Leone. Problemi attuali di Scienzia e di Cultura, 267, 307 - 314." type="journal article" year="1994">Bordoni, 1994</bibRefCitation>
: 312
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="445C3F44FFA41A7D046FFA07ED08FA51" author="Herman" box="[620,840,1448,1471]" page="3033" pageId="28" pageNumber="29" year="2001">
<bibRefCitation id="A16C64A4FFA41A7D046FFA07ED48FA51" author="Herman" box="[620,776,1448,1471]" pageId="28" pageNumber="50" refString="Herman, L. H. (2001). Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group (part 3). Bulletin of the American Museum of Natural History, 265, 3021 - 3840." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 3033
</treatmentCitation>
<emphasis id="F789C547FFA41A7D054CFA08EDBBFA51" bold="true" box="[847,1019,1447,1471]" pageId="28" pageNumber="29">(new synonym)</emphasis>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C5421955FFA41A7D0694FA42EA10F9BC" blockId="28.[151,1437,1517,2028]" pageId="28" pageNumber="29">
<emphasis id="F789C547FFA41A7D0694FA42EF10F9E9" bold="true" box="[151,336,1517,1543]" pageId="28" pageNumber="29">Redescription.</emphasis>
Length 6.5mm. Body black with iridescent abdomen, pronotum sometimes slightly paler. I have also seen one specimen that is completely pale brown. Antennae brown to dark brown, first segment often paler. Legs pale, sometimes with darkened femora and tibiae. Palpi all pale.
</paragraph>
</subSubSection>
<subSubSection id="8DE74ADEFFA41A7C06C7F9F2EC8BFE82" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="description">
<paragraph id="C5421955FFA41A7D06C7F9F2EAC8F8B9" blockId="28.[151,1437,1517,2028]" pageId="28" pageNumber="29">
Head small in comparison with pronotum (pronotum 1.9x wider than head) more or less as long as wide with well developed temples behind large eyes (Fig. 21). Head pigmented in front of antennal insertion. Micro-punctures very sparse and scarcely detectable at
<date id="B1433F95FFA41A7D0539F907ED28F92C" box="[826,872,1704,1730]" pageId="28" pageNumber="29" value="1940-10">40x</date>
magnification. Short pubescence behind eyes. Three pairs of interocular setae and a line of five postocular setae just visible from above on each side. Maxillary palpi with last two segments pubescent, terminal segment elongate, rounded on both sides, but slightly asymmetric, longer than penultimate segment which is also elongate (
<figureCitation id="5DC605D0FFA41A7D025DF8B7EAFCF8DC" box="[1118,1212,1816,1842]" captionStart="FIGURES 51 – 76" captionStartId="36.[151,271,1818,1842]" captionTargetBox="[176,1425,305,1773]" captionTargetId="figure@36.[155,1440,293,1797]" captionTargetPageId="36" captionText="FIGURES 51 – 76. Terminal segments of the maxilliary palpi of Acylophorus species. 51, A. orientalis Fauvel; 52, A. nitens new species; 53, A. rossii Bordoni, 54, A. bumbunae Bordoni; 55, A. capensis Cameron; 56, A. dankalensis Bordoni; 57, A. tshuapensis new species; 58, A. makhoreae new species; 59, A. densipennis Bernhauer; 60, A. antennalis Cameron; 61, A. collarti Cameron; 62. A. micans new species; 63, A. allardi Levasseur; 64, A. congoensis Cameron; 65, A. ziloensis Levasseur; 66, A. mareei Bernhauer; 67, A. salifi new species; 68, A. lomaensis Levasseur; 69. A. minor new species; 70, A. trigonocephalus Cameron; 71, A. setiger new species; 72, A. usambarae Bernhauer; 73, A. grandis Bernhauer; 74, A. bafutensis Levasseur; 75, A. kambuiensis Bordoni, 76. A. lualabaensis Levasseur." httpUri="https://zenodo.org/record/275914/files/figure.png" pageId="28" pageNumber="29">Fig. 70</figureCitation>
). First segment of antenna longer than next five. Segments I to IV elongate, IX to X transverse (
<figureCitation id="5DC605D0FFA41A7D0220F892EA3BF8B9" box="[1059,1147,1853,1879]" captionStart="FIGURES 97 – 103" captionStartId="38.[151,270,1806,1830]" captionTargetBox="[178,1406,194,1766]" captionTargetId="figure@38.[161,1418,178,1785]" captionTargetPageId="38" captionText="FIGURES 97 – 103. Antennae of Acylophorus species. 97, A. trigonocephalus Cameron; 98, A. setiger new species; 99, A. usambarae Bernhauer; 100, A. grandis Bernhauer; 101, A. bafutensis Levasseur; 102, A. kambuiensis Bordoni, 103. A. lualabaensis Levasseur. FIGURES 104 – 107. Details of Acylophorus delphinus Fauvel. 104, forebody; 105, terminal segments of maxilliary palpi; 106, antenna; 107, aedeagus. FIGURE 108. Lateral view of head of Acylophorus salifi new species." httpUri="https://zenodo.org/record/275916/files/figure.png" pageId="28" pageNumber="29">Fig. 97</figureCitation>
).
</paragraph>
<paragraph id="C5421955FFA41A7C06C7F8CDEC85FF12" blockId="28.[151,1437,1517,2028]" lastBlockId="29.[151,1437,152,1148]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
Pronotum transverse (1.3x wider than long) with rounded sides, widest in basal half. Shining with no micro-punctures. Marginal setae very long and more numerous than in
<taxonomicName id="02FD62D6FFA41A7D05EFF827EA3FF84F" box="[1004,1151,1928,1953]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="F789C547FFA41A7D05EFF827EA3FF84F" box="[1004,1151,1928,1953]" italics="true" pageId="28" pageNumber="29">A. orientalis</emphasis>
</taxonomicName>
. One pair of dorsal and one pair of lateral setae. Elytra transverse (1.65x wider than long) with long but sparse pubescence and coarse asperate punctures which are stronger than in
<taxonomicName id="02FD62D6FFA41A7D04B6F87CED07F802" box="[693,839,2003,2028]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="F789C547FFA41A7D04B6F87CED07F802" box="[693,839,2003,2028]" italics="true" pageId="28" pageNumber="29">A. orientalis</emphasis>
</taxonomicName>
. Apical bristles much longer than hairs on the rest of the elytra. Abdominal tergites also with long, sparse pubescence. Punctures much sparser on apical half of each tergite than on basal half. Apical fringe of bristles on each tergite of two distinct lengths, the shorter bristles more numerous than the longer bristles.
</paragraph>
<paragraph id="C5421955FFA51A7C06C7FEA7EC8BFE82" blockId="29.[151,1437,152,1148]" pageId="29" pageNumber="30">
<collectingCountry id="BDEA59C5FFA51A7C06C7FEA7EF42FECC" box="[196,258,264,290]" name="Malta" pageId="29" pageNumber="30">Male</collectingCountry>
sternite IX with apex simple. Aedeagus with bilobed paramere, each lobe narrow, parallel, pegs confusedly arranged toward apex, which has numerous short marginal hairs (
<figureCitation id="5DC605D0FFA51A7C0214FE82EA3DFEA9" box="[1047,1149,301,327]" captionStart="FIGURES 134 – 137" captionStartId="42.[151,269,1913,1937]" captionTargetBox="[314,1249,189,1879]" captionTargetId="figure@42.[294,1296,170,1909]" captionTargetPageId="42" captionText="FIGURES 134 – 137. Aedeagi of species in the Acylophorus trigonocephalus group. 134, A. grandis Bernhauer; 135, A. trigonocephalus Cameron; 136, A. setiger new species; 137, A. bafutensis Levasseur (reproduced from Levasseur, 1968 and not drawn to scale). FIGURES 138 – 139. Aedeagi of species in the Acylophorus kambuiensis group. 138, A. kambuiensis Bordoni, 139. A. lualabaensis Levasseur." httpUri="https://zenodo.org/record/275920/files/figure.png" pageId="29" pageNumber="30">Fig. 135</figureCitation>
). Median lobe with apex barely expanded, only just surpassing paramere.
</paragraph>
</subSubSection>
<subSubSection id="8DE74ADEFFA51A7C06C7FED8EC3FFCBC" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="C5421955FFA51A7C06C7FED8EA22FDC9" blockId="29.[151,1437,152,1148]" pageId="29" pageNumber="30">
<emphasis id="F789C547FFA51A7C06C7FED8EF3FFE7F" bold="true" box="[196,383,375,401]" pageId="29" pageNumber="30">
<typeStatus id="1A46A7F7FFA51A7C06C7FED8EF41FE7F" box="[196,257,375,401]" pageId="29" pageNumber="30">Type</typeStatus>
material.
</emphasis>
<collectingCountry id="BDEA59C5FFA51A7C0789FED7EFB9FE7C" box="[394,505,376,402]" name="Cameroon" pageId="29" pageNumber="30">Cameron</collectingCountry>
described
<taxonomicName id="02FD62D6FFA51A7C047DFED7ED27FE7F" box="[638,871,376,401]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="trigonocephalus">
<emphasis id="F789C547FFA51A7C047DFED7ED27FE7F" box="[638,871,376,401]" italics="true" pageId="29" pageNumber="30">A. trigonocephalus</emphasis>
</taxonomicName>
from material collected in
<collectingCountry id="BDEA59C5FFA51A7C02B5FED7EB1EFE7C" box="[1206,1374,376,402]" name="Ivory Coast" pageId="29" pageNumber="30">Côte d’Ivoire</collectingCountry>
. One
<typeStatus id="1A46A7F7FFA51A7C0694FE32EEB5FE59" box="[151,245,413,439]" pageId="29" pageNumber="30" type="syntype">syntype</typeStatus>
has been located and is here designated as the
<typeStatus id="1A46A7F7FFA51A7C0530FE32EDE2FE59" box="[819,930,413,439]" pageId="29" pageNumber="30" type="lectotype">lectotype</typeStatus>
in order to fix the identity of the species.
<emphasis id="F789C547FFA51A7C0694FE6DEF5DFE32" bold="true" box="[151,285,450,476]" pageId="29" pageNumber="30">
<typeStatus id="1A46A7F7FFA51A7C0694FE6DEF5DFE32" box="[151,285,450,476]" pageId="29" pageNumber="30" type="lectotype">Lectotype</typeStatus>
</emphasis>
Ƥ: “IFAN-1946 TONKUI C.I.
<quantity id="0205B4B0FFA51A7C04B3FE6DED06FE32" box="[688,838,450,476]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="29" pageNumber="30" unit="m" value="1050.0" valueMax="1200.0" valueMin="900.0">900–1200m</quantity>
A. VILLIERS /
<collectionCode id="A3EC8190FFA51A7C021DFE6DEA9BFE32" box="[1054,1243,450,476]" pageId="29" pageNumber="30">FORET PRIM</collectionCode>
;
<date id="B1433F95FFA51A7C02EBFE6DEB1EFE32" box="[1256,1374,450,476]" bridgedPair="-" pageId="29" pageNumber="30" value="1920-09-30">20-30-IX</date>
/ M.
<collectingCountry id="BDEA59C5FFA51A7C0694FE47EF43FDEC" box="[151,259,488,514]" name="Cameroon" pageId="29" pageNumber="30">Cameron</collectingCountry>
Bequest B.M. 1955-147 / Dr. M.
<collectingCountry id="BDEA59C5FFA51A7C048EFE47ECB8FDEC" box="[653,760,488,514]" name="Cameroon" pageId="29" pageNumber="30">Cameron</collectingCountry>
det., 1948;
<taxonomicName id="02FD62D6FFA51A7C0580FE47EB63FDEC" authority="Cam." authorityName="Cam." box="[899,1315,488,514]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="trigonocephalus">
<emphasis id="F789C547FFA51A7C0580FE47EA9DFDEF" box="[899,1245,488,513]" italics="true" pageId="29" pageNumber="30">Acylophorus trigonocephalus</emphasis>
Cam.
</taxonomicName>
COTYPE /
<typeStatus id="1A46A7F7FFA51A7C06A5FDA2EF0DFDC9" box="[166,333,525,551]" pageId="29" pageNumber="30" type="lectotype">LECTOTYPE</typeStatus>
<taxonomicName id="02FD62D6FFA51A7C0756FDA2ED7FFDC9" authority="Cameron P" authorityName="Cameron P" box="[341,831,525,551]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="trigonocephalus">
<emphasis id="F789C547FFA51A7C0756FDA2ECEFFDC8" box="[341,687,525,550]" italics="true" pageId="29" pageNumber="30">Acylophorus trigonocephalus</emphasis>
<collectingCountry id="BDEA59C5FFA51A7C04B5FDA2ED62FDC9" box="[694,802,525,551]" name="Cameroon" pageId="29" pageNumber="30">Cameron</collectingCountry>
Ƥ
</taxonomicName>
det. DA Lott” (
<collectionCode id="A3EC8190FFA51A7C05F4FDA2EA15FDC9" box="[1015,1109,525,551]" pageId="29" pageNumber="30">BMNH</collectionCode>
).
</paragraph>
<paragraph id="C5421955FFA51A7C06C7FD9DEABBFD55" blockId="29.[151,1437,152,1148]" pageId="29" pageNumber="30">
<taxonomicName id="02FD62D6FFA51A7C06C7FD9DEC22FDA2" authority="Bordoni." authorityName="Bordoni." box="[196,610,562,588]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="leonensis">
<emphasis id="F789C547FFA51A7C06C7FD9DEFA7FDA5" box="[196,487,562,587]" italics="true" pageId="29" pageNumber="30">Acylophorus leonensis</emphasis>
Bordoni.
</taxonomicName>
<emphasis id="F789C547FFA51A7C046FFD9DECA8FDA2" bold="true" box="[620,744,562,588]" pageId="29" pageNumber="30">
<typeStatus id="1A46A7F7FFA51A7C046FFD9DECA8FDA2" box="[620,744,562,588]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
</emphasis>
3: “S. Leone W. Area Sugar Loaf Mt. Rossi
<date id="B1433F95FFA51A7C0339FD9DEBCBFDA2" box="[1338,1419,562,588]" pageId="29" pageNumber="30" value="92-03-07">7.3.92</date>
/
<typeStatus id="1A46A7F7FFA51A7C0694FDF7EF0AFD9C" box="[151,330,600,626]" pageId="29" pageNumber="30" type="holotype">HOLOTYPUS</typeStatus>
/
<typeStatus id="1A46A7F7FFA51A7C0760FDF7EC56FD9C" box="[355,534,600,626]" pageId="29" pageNumber="30" type="holotype">HOLOTYPUS</typeStatus>
<taxonomicName id="02FD62D6FFA51A7C041CFDF7ED72FD9F" box="[543,818,600,625]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="leonensis" status="sp. nov.">
<emphasis id="F789C547FFA51A7C041CFDF7ED72FD9F" box="[543,818,600,625]" italics="true" pageId="29" pageNumber="30">Acylophorus leonensis</emphasis>
</taxonomicName>
<emphasis id="F789C547FFA51A7C0538FDF8EDC1FD9F" bold="true" box="[827,897,599,625]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="ECBA783CFFA51A7C0538FDF8EDC1FD9F" box="[827,897,599,625]" pageId="29" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Bordoni det. 1992” (cBord). In the original description Bordoni compared his new species to
<emphasis id="F789C547FFA51A7C04FAFDD2ED26FD78" box="[761,870,637,662]" italics="true" pageId="29" pageNumber="30">
<taxonomicName id="02FD62D6FFA51A7C04FAFDD2ED21FD78" box="[761,865,637,662]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="rossii">A. rossii</taxonomicName>
,
</emphasis>
but not to
<taxonomicName id="02FD62D6FFA51A7C05EEFDD2EA95FD78" box="[1005,1237,637,662]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="trigonocephalus">
<emphasis id="F789C547FFA51A7C05EEFDD2EA95FD78" box="[1005,1237,637,662]" italics="true" pageId="29" pageNumber="30">A. trigonocephalus</emphasis>
</taxonomicName>
with which it is similar in all respects. Consequently,
<taxonomicName id="02FD62D6FFA51A7C044EFD0DEC9BFD55" box="[589,731,674,699]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="leonensis">
<emphasis id="F789C547FFA51A7C044EFD0DEC9BFD55" box="[589,731,674,699]" italics="true" pageId="29" pageNumber="30">A. leonensis</emphasis>
</taxonomicName>
is here synonymised with
<emphasis id="F789C547FFA51A7C0217FD0DEABBFD55" box="[1044,1275,674,699]" italics="true" pageId="29" pageNumber="30">
<taxonomicName id="02FD62D6FFA51A7C0217FD0DEAB7FD55" box="[1044,1271,674,699]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="trigonocephalus">A. trigonocephalus</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="C5421955FFA51A7C06C7FD68EC3FFCBC" blockId="29.[151,1437,152,1148]" pageId="29" pageNumber="30">
<emphasis id="F789C547FFA51A7C06C7FD68ED41FD0F" bold="true" box="[196,769,711,737]" pageId="29" pageNumber="30">
Further material examined. R.D.
<collectingCountry id="BDEA59C5FFA51A7C048EFD68ED41FD0F" box="[653,769,711,737]" name="Republic of the Congo" pageId="29" pageNumber="30">CONGO</collectingCountry>
</emphasis>
:
<emphasis id="F789C547FFA51A7C0517FD68ED21FD0F" bold="true" box="[788,865,711,737]" pageId="29" pageNumber="30">Kivu:</emphasis>
Kabari, SE Kahuzi, 2080/
<quantity id="0205B4B0FFA51A7C02B5FD67EB4FFD0C" box="[1206,1295,712,738]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="29" pageNumber="30" unit="m" value="2200.0">2200m</quantity>
, N Leleup,
<date id="B1433F95FFA51A7C0694FD42EF64FCE9" box="[151,292,749,775]" pageId="29" pageNumber="30" value="1951-08-01">viii.1951, 1</date>
Ƥ (
<collectionCode id="A3EC8190FFA51A7C074EFD42EFEDFCE9" box="[333,429,749,775]" pageId="29" pageNumber="30">ISRNB</collectionCode>
); Mwenga, Luiko,
<quantity id="0205B4B0FFA51A7C0495FD42ECAEFCE9" box="[662,750,749,775]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="29" pageNumber="30" unit="m" value="2050.0">2050m</quantity>
, N Leleup, 21.
<date id="B1433F95FFA51A7C05A0FD42EA52FCE9" box="[931,1042,749,775]" pageId="29" pageNumber="30" value="1952-01-01">i.1952, 1</date>
Ƥ (
<collectionCode id="A3EC8190FFA51A7C0239FD42EADAFCE9" box="[1082,1178,749,775]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="29" pageNumber="30">IRSNB</collectionCode>
).
<emphasis id="F789C547FFA51A7C02B3FD42EBD3FCE9" bold="true" box="[1200,1427,749,775]" pageId="29" pageNumber="30">
<collectingCountry id="BDEA59C5FFA51A7C02B3FD42EBD3FCE9" box="[1200,1427,749,775]" name="Sierra Leone" pageId="29" pageNumber="30">SIERRA LEONE</collectingCountry>
</emphasis>
:
<emphasis id="F789C547FFA51A7C0694FCBDEFD0FCC2" bold="true" box="[151,400,786,812]" pageId="29" pageNumber="30">Northern Province</emphasis>
: Sinikoro, W Rossi, 31.
<date id="B1433F95FFA51A7C04BDFCBDED1FFCC5" box="[702,863,785,812]" pageId="29" pageNumber="30" value="1982-12-13">xii.1982, 13</date>
(cBord).
<emphasis id="F789C547FFA51A7C05D8FCBDEAC9FCC2" bold="true" box="[987,1161,786,812]" pageId="29" pageNumber="30">
<collectingCountry id="BDEA59C5FFA51A7C05D8FCBDEAC9FCC2" box="[987,1161,786,812]" name="Zimbabwe" pageId="29" pageNumber="30">ZIMBABWE</collectingCountry>
</emphasis>
: Kutsaga near Harare Airport, W Rossi, 18.
<date id="B1433F95FFA51A7C0793FC97EC5EFCBE" box="[400,542,822,850]" pageId="29" pageNumber="30" value="1997-06-13">vi.1997, 13</date>
(cLott).
</paragraph>
</subSubSection>
<subSubSection id="8DE74ADEFFA51A7C06C7FCF2EF59FB92" pageId="29" pageNumber="30" type="distribution">
<paragraph id="C5421955FFA51A7C06C7FCF2EFC3FC2C" blockId="29.[151,1437,152,1148]" pageId="29" pageNumber="30">
<emphasis id="F789C547FFA51A7C06C7FCF2EC62FC99" bold="true" box="[196,546,861,887]" pageId="29" pageNumber="30">Distribution and bionomics.</emphasis>
Recorded from
<collectingCountry id="BDEA59C5FFA51A7C04E0FCF2ED3CFC99" box="[739,892,861,887]" name="Sierra Leone" pageId="29" pageNumber="30">Sierra Leone</collectingCountry>
,
<collectingCountry id="BDEA59C5FFA51A7C058AFCF2EA69FC99" box="[905,1065,861,887]" name="Ivory Coast" pageId="29" pageNumber="30">Côte d’Ivoire</collectingCountry>
,
<collectingCountry id="BDEA59C5FFA51A7C0235FCF2EAC4FC99" box="[1078,1156,861,887]" name="Republic of the Congo" pageId="29" pageNumber="30">Congo</collectingCountry>
and
<collectingCountry id="BDEA59C5FFA51A7C02BDFCF2EB7EFC99" box="[1214,1342,861,887]" name="Zimbabwe" pageId="29" pageNumber="30">Zimbabwe</collectingCountry>
, always in small numbers (
<figureCitation id="5DC605D0FFA51A7C0775FC2DEF9FFC72" box="[374,479,898,924]" captionStart="FIGURE 147" captionStartId="47.[151,255,1866,1890]" captionTargetBox="[151,1436,1004,1838]" captionTargetId="figure@47.[151,1436,1004,1838]" captionTargetPageId="47" captionText="FIGURE 147. Distribution of species in the Acylophorus trigonocephalus group = A. trigonocephalus Cameron, = A. setiger new species, = A. usambarae Bernhauer, = A. grandis Bernhauer; = A. bafutensis Levasseur)." httpUri="https://zenodo.org/record/275928/files/figure.png" pageId="29" pageNumber="30">Fig. 147</figureCitation>
). Recorded habitats include primary forest, montane forest and the margins of a fast-flowing stream.
</paragraph>
<paragraph id="C5421955FFA51A7C06C7FC62EF59FB92" blockId="29.[151,1437,152,1148]" pageId="29" pageNumber="30">
<emphasis id="F789C547FFA51A7C06C7FC62EF80FC09" bold="true" box="[196,448,973,999]" pageId="29" pageNumber="30">Comparative notes.</emphasis>
Easily distinguished from
<taxonomicName id="02FD62D6FFA51A7C050FFC62EDE4FC08" box="[780,932,973,998]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="F789C547FFA51A7C050FFC62EDE4FC08" box="[780,932,973,998]" italics="true" pageId="29" pageNumber="30">A. orientalis</emphasis>
</taxonomicName>
by the characters common to the species group, two pubescent terminal segments on the maxillary palpi, the three pairs of interocular punctures and the larger number of marginal setae. Within the group it can be recognised by a combination of the long setae on the head and pronotum, the large eyes and the moderately transverse elytra. The aedeagus is also distinctive.
</paragraph>
</subSubSection>
</treatment>
</document>
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