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<document id="F2D9016FE5F7096C109F22BCB9649783" ID-CLB-Dataset="20928" ID-DOI="10.11646/zootaxa.5093.4.4" ID-GBIF-Dataset="d6297960-52dd-4b0a-a51b-05bda0e7d69f" ID-ISSN="1175-5326" ID-Zenodo-Dep="5912781" ID-ZooBank="4FE93B11-0714-400F-B9A6-41A54E2FCDC9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643354449005" checkinUser="plazi" docAuthor="Spinelli, Gustavo R., Ronderos, Maria M. &amp; Grogan, William L." docDate="2022" docId="4D4A9349FFEA39311AD2FCE2FF3CF94F" docLanguage="en" docName="zootaxa.5093.4.4.pdf" docOrigin="Zootaxa 5093 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrurohelea ventanensis Spinelli &amp; Ronderos &amp; Grogan 2022, n. sp." docType="treatment" docVersion="7" lastPageNumber="462" masterDocId="B173EB31FFE739201A45FFD5FFFBFF95" masterDocTitle="Five new species in the predaceous midge genus Macrurohelea Ingram &amp; Macfie from Argentina, and descriptions of the previously unknown males of M. kuscheli Wirth and M. monotheca Spinelli &amp; Grogan (Diptera: Ceratopogonidae)" masterLastPageNumber="464" masterPageNumber="445" pageNumber="458" updateTime="1699304617883" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="2F93BE6216929F6FCA0A793AE57DB20E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="29074653B38D7D242047D86946F32ADD">
<mods:title id="78AD2FD31BB9CCEEDB8E1C0144E34117">Five new species in the predaceous midge genus Macrurohelea Ingram &amp; Macfie from Argentina, and descriptions of the previously unknown males of M. kuscheli Wirth and M. monotheca Spinelli &amp; Grogan (Diptera: Ceratopogonidae)</mods:title>
</mods:titleInfo>
<mods:name id="9F2435F063EDA8B0C8D8931A226D5A98" type="personal">
<mods:role id="3B799F9D246CD1FABFE5EFF9DAB9A3B5">
<mods:roleTerm id="3484BFC80B17CB4B258B60D5AD256E80">Author</mods:roleTerm>
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<mods:namePart id="B3494E0FBE3386E577617514908D2278">Spinelli, Gustavo R.</mods:namePart>
<mods:affiliation id="EBB1B91C83DF42D83042A7C0F0E2E502">División Entomología, Museo de La Plata, Paseo del Bosque s / n, 1900 La Plata, Argentina. &amp; Instituto de Limnología “ Dr. Raúl A. Ringuelet ” (Consejo Nacional de Investigaciones Científicas y Técnicas de la Argentina - Universidad Nacional de La Plata), Boulevard 120 y 62 s / n, 1900 La Plata, Buenos Aires, Argentina.</mods:affiliation>
</mods:name>
<mods:name id="5714DD06D284B5291F2F05238E762230" type="personal">
<mods:role id="3258B95C66D55E461C8866CB12C2C497">
<mods:roleTerm id="C7A1DBA2BC2FB32DE3AD2E6FF28E3A51">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="49E2346CCC6FD3EDB9773D6F819F5C5E">Ronderos, Maria M.</mods:namePart>
<mods:nameIdentifier id="CEC6F65CCC8AAE76722123E5B082ED08" type="ORCID">0000-0001-8322-1471</mods:nameIdentifier>
<mods:affiliation id="41AEEFBB337CC0D65B852789C8F7A8F4">División Entomología, Museo de La Plata, Paseo del Bosque s / n, 1900 La Plata, Argentina. &amp; Centro de Estudios Parasitológicos y de Vectores (Consejo Nacional de Investigaciones Científicas y Técnicas de la Argentina - Universidad Nacional de La Plata), Boulevard 120 y 62 s / n, 1900 La Plata, Buenos Aires, Argentina. &amp; ronderos @ fcnym. unlp. edu. ar; https: // orcid. org / 0000 - 0001 - 8322 - 1471</mods:affiliation>
<mods:nameIdentifier id="00DB8863B96629508AD9CFD8A219D60C" type="email">ronderos@fcnym.unlp.edu.ar</mods:nameIdentifier>
</mods:name>
<mods:name id="33ED2F3324DA56000F351A433159D274" type="personal">
<mods:role id="D1498B55ACC470E794A9565211C7651C">
<mods:roleTerm id="B62636F76D68989515A3EF0C975C78B8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2B3B224CEF5210D852D413FC483C7931">Grogan, William L.</mods:namePart>
<mods:nameIdentifier id="AE5CFEF9074BDA12EC0249B2220FC2C3" type="ORCID">0000-0002-6475-8692</mods:nameIdentifier>
<mods:affiliation id="DF793D3966A589331E50669CE5FF716C">Florida State Collection of Arthropods, Florida Department of Agriculture and Consumer Services, Gainesville, Florida 32614 - 7100 USA. &amp; William. Grogan @ fdacs. gov; https: // orcid. org / 0000 - 0002 - 6475 - 8692</mods:affiliation>
<mods:nameIdentifier id="FDD635620B69CD655EF66683C1F26664" type="email">rogan@fdacs.gov</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="786311996041CBF34377CC21120888AD">text</mods:typeOfResource>
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<mods:title id="B04C936A3A99ADD3E9B0A7CFF00D3ADA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="AC95E9DDFB667F2405E628E5C3CD631D">
<mods:date id="02470FBAD681DF5F509673D6B0875CDA">2022</mods:date>
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<mods:number id="46922070EB2E9087FC4095FA74476371">2022-01-28</mods:number>
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<mods:number id="E4155BEF49262B82CF0161FE2BBC2E0D">5093</mods:number>
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<mods:classification id="591F57E1E4CF13560B93E8C478980BBD">journal article</mods:classification>
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<mods:identifier id="DAAB629A2149721D5A96F4897C98DC0A" type="DOI">10.11646/zootaxa.5093.4.4</mods:identifier>
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<treatment id="4D4A9349FFEA39311AD2FCE2FF3CF94F" ID-DOI="http://doi.org/10.5281/zenodo.5914103" ID-GBIF-Taxon="192700744" ID-Zenodo-Dep="5914103" LSID="urn:lsid:plazi:treatment:4D4A9349FFEA39311AD2FCE2FF3CF94F" httpUri="http://treatment.plazi.org/id/4D4A9349FFEA39311AD2FCE2FF3CF94F" lastPageId="17" lastPageNumber="462" pageId="13" pageNumber="458">
<subSubSection id="8DF971D4FFEA392D1AD2FCE2FDD8FCC7" box="[151,547,823,850]" pageId="13" pageNumber="458" type="nomenclature">
<paragraph id="C55C225FFFEA392D1AD2FCE2FDD8FCC7" blockId="13.[151,1065,823,923]" box="[151,547,823,850]" pageId="13" pageNumber="458">
<heading id="9E149533FFEA392D1AD2FCE2FDD8FCC7" bold="true" box="[151,547,823,850]" fontSize="11" level="1" pageId="13" pageNumber="458" reason="1">
<emphasis id="F797FE4DFFEA392D1AD2FCE2FDD8FCC7" bold="true" box="[151,547,823,850]" pageId="13" pageNumber="458">
<taxonomicName id="02E359DCFFEA392D1AD2FCE2FE22FCC7" ID-CoL="9HXXG" authority="Spinelli &amp; Ronderos &amp; Grogan, 2022" authorityName="Spinelli &amp; Ronderos &amp; Grogan" authorityYear="2022" box="[151,473,823,850]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="458" phylum="Arthropoda" rank="species" species="ventanensis" status="sp. nov.">
<emphasis id="F797FE4DFFEA392D1AD2FCE2FE22FCC7" bold="true" box="[151,473,823,850]" italics="true" pageId="13" pageNumber="458">Macrurohelea ventanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECA44336FFEA392D1BA5FCEDFDD8FCC7" box="[480,547,824,850]" pageId="13" pageNumber="458" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8DF971D4FFEA392D1AD2FC89FBD2FC0E" pageId="13" pageNumber="458" type="description">
<paragraph id="C55C225FFFEA392D1AD2FC89FE79FCE2" blockId="13.[151,1065,823,923]" box="[151,386,860,887]" pageId="13" pageNumber="458">
(
<figureCitation id="5DD83EDAFFEA392D1ADAFC89FEDCFCE3" box="[159,295,860,886]" captionStart="FIGURES 33–36" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[240,1349,181,1917]" captionTargetId="figure-19@15.[237,1350,181,1920]" captionTargetPageId="15" captionText="FIGURES 33–36. Macrurohelea ventanensis n. sp. Holotype male. 33 head. 34 thorax. 35 wing. 36 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912793" httpUri="https://zenodo.org/record/5912793/files/figure.png" pageId="13" pageNumber="458">Figs. 33–36</figureCitation>
,
<figureCitation id="5DD83EDAFFEA392D1B76FC89FE81FCE2" box="[307,378,860,887]" captionStart="FIGURES 47–50" captionStartId="18.[151,264,1227,1251]" captionTargetBox="[250,1333,183,1189]" captionTargetId="figure-23@18.[242,1346,181,1203]" captionTargetPageId="18" captionText="FIGURES 47–50. Macrurohelea sirii n. sp. Holotype male, 47–48. Macrurohelea ventanensis n. sp. Holotype male, 49–50. 47, 49 parameres. 48, 50 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912801" httpUri="https://zenodo.org/record/5912801/files/figure.png" pageId="13" pageNumber="458">49–50</figureCitation>
)
</paragraph>
<paragraph id="C55C225FFFEA392D1AD2FC55FBD2FC0E" blockId="13.[151,1065,823,923]" box="[151,1065,896,923]" pageId="13" pageNumber="458">
Zoobank
<uri id="B1722E5DFFEA392D1B47FC55FBD2FC0E" box="[258,1065,896,923]" pageId="13" pageNumber="458">
urn:lsid:zoobank.org:act:
<uuid id="B145188AFFEA392D1851FC55FBD2FC0E" box="[532,1065,896,923]" pageId="13" pageNumber="458">9A500E88-CAE8-4F9E-976F-6F39BF21C962</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="8DF971D4FFEA392D1AD2FC1DFE59F81F" pageId="13" pageNumber="458" type="diagnosis">
<paragraph id="C55C225FFFEA392D1AD2FC1DFCAEFBE7" blockId="13.[151,1437,968,2039]" pageId="13" pageNumber="458">
<emphasis id="F797FE4DFFEA392D1AD2FC1DFEE9FC77" bold="true" box="[151,274,968,994]" pageId="13" pageNumber="458">Diagnosis.</emphasis>
Males: the only species with a pale brown halter; tergite 9 abruptly tapered on basal 1/5, with rounded apex; tergite 10 with straight margins, apex straight with small, rounded apicolateral process with single divergent seta; parameres stout, nearly straight, widely separated, subparallel, with slender, slightly hooked, curved, fingerlike apical process; aedeagus basal arch deeply concave, heavily sclerotized, distal portion more lightly sclerotized, apex with a short hyaline quadrate process. Females unknown.
</paragraph>
<paragraph id="C55C225FFFEA392D1A82FBA9FAB3F8F2" blockId="13.[151,1437,968,2039]" pageId="13" pageNumber="458">
<emphasis id="F797FE4DFFEA392D1A82FBA9FEFFFB03" bold="true" box="[199,260,1148,1174]" pageId="13" pageNumber="458">Male</emphasis>
. Head (
<figureCitation id="5DD83EDAFFEA392D1B2DFBA9FE3BFB03" box="[360,448,1148,1174]" captionStart="FIGURES 33–36" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[240,1349,181,1917]" captionTargetId="figure-19@15.[237,1350,181,1920]" captionTargetPageId="15" captionText="FIGURES 33–36. Macrurohelea ventanensis n. sp. Holotype male. 33 head. 34 thorax. 35 wing. 36 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912793" httpUri="https://zenodo.org/record/5912793/files/figure.png" pageId="13" pageNumber="458">Fig. 33</figureCitation>
) brown. Eyes separated medially by width of 2.5 ommatidia with numerous short interommatidial spicules. Antennal flagellomeres distinctly separated; flagellomeres 1–8 broadly moniliform, 9–10 narrow, slightly elongate, 11–13 more elongate, 13 longest. Palpus brown; segment 3 with small sensory pit slightly distad of mid-length; segment 4 0.66 length of segment 3, segment 5 slightly longer than segment 3, with bulbous apex; palpal ratio 2.00–2.50 (2.25, n =2). Thorax (
<figureCitation id="5DD83EDAFFEA392D1880FAD9FCEDFAB3" box="[709,790,1292,1318]" captionStart="FIGURES 33–36" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[240,1349,181,1917]" captionTargetId="figure-19@15.[237,1350,181,1920]" captionTargetPageId="15" captionText="FIGURES 33–36. Macrurohelea ventanensis n. sp. Holotype male. 33 head. 34 thorax. 35 wing. 36 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912793" httpUri="https://zenodo.org/record/5912793/files/figure.png" pageId="13" pageNumber="458">Fig. 34</figureCitation>
) uniformly dark brown; scutum with 3 stout prealar setae, one postalar seta; scutellum with 4 stout, long setae. Legs medium brown; femora, tibiae very slender, greatly elongate; ventral palisade setae on tarsomere 1 of fore, hind legs; tarsomeres 4 cordiform; tarsal claws small, mostly straight, slightly curved on extreme bifid tips. Wing (
<figureCitation id="5DD83EDAFFEA392D194FFAADFCA4FA06" box="[778,863,1400,1427]" captionStart="FIGURES 33–36" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[240,1349,181,1917]" captionTargetId="figure-19@15.[237,1350,181,1920]" captionTargetPageId="15" captionText="FIGURES 33–36. Macrurohelea ventanensis n. sp. Holotype male. 33 head. 34 thorax. 35 wing. 36 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912793" httpUri="https://zenodo.org/record/5912793/files/figure.png" pageId="13" pageNumber="458">Fig. 35</figureCitation>
) membrane pale, slightly infuscated with minute microtrichia; anterior veins yellowish brown, posterior veins paler; 2
<superScript id="32968F17FFEA392D19E7FA49FC49FA3F" attach="left" box="[930,946,1436,1450]" fontSize="6" pageId="13" pageNumber="458">nd</superScript>
radial cell twice as long as 1st; cell r
<subScript id="5967201AFFEA392D1F1FFA7BFA99FA29" attach="left" box="[1370,1378,1454,1468]" fontSize="6" pageId="13" pageNumber="458">3</subScript>
with poorly developed intercalary vein; r-m crossvein slightly shorter than petiole of M; medium-size macrotrichiae on costa; wing length 1.31–1.35 (1.33, n =2) mm, width 0.45–0.47 (0.46, n = 2) mm; costal ratio 0.59–0.63 (0.61, n =2); halter pale brown. Abdomen brown. Genitalia (
<figureCitation id="5DD83EDAFFEA392D1898F9DDFCC0F9B7" box="[733,827,1544,1570]" captionStart="FIGURES 33–36" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[240,1349,181,1917]" captionTargetId="figure-19@15.[237,1350,181,1920]" captionTargetPageId="15" captionText="FIGURES 33–36. Macrurohelea ventanensis n. sp. Holotype male. 33 head. 34 thorax. 35 wing. 36 genitalia. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5912793" httpUri="https://zenodo.org/record/5912793/files/figure.png" pageId="13" pageNumber="458">Figs. 36</figureCitation>
,
<figureCitation id="5DD83EDAFFEA392D1903F9DDFC69F9B6" box="[838,914,1544,1571]" captionStart="FIGURES 47–50" captionStartId="18.[151,264,1227,1251]" captionTargetBox="[250,1333,183,1189]" captionTargetId="figure-23@18.[242,1346,181,1203]" captionTargetPageId="18" captionText="FIGURES 47–50. Macrurohelea sirii n. sp. Holotype male, 47–48. Macrurohelea ventanensis n. sp. Holotype male, 49–50. 47, 49 parameres. 48, 50 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912801" httpUri="https://zenodo.org/record/5912801/files/figure.png" pageId="13" pageNumber="458">49–50</figureCitation>
). Tergite 9 abruptly tapered on basal 1/5, distal portion broad, elongate with rounded apex, extending 0.75 length of gonocoxites. Tergite 10 with straight margins, apex straight with small rounded divergent apicolateral process with single medium-size seta; cercus slender, fingerlike. Sternite 9 2.4x broader than long, posteromedian margin with shallow excavation. Gonocoxite stout, twice as long as wide, with rounded sub-basal mesal extension; gonostylus 0.77 length of gonocoxite, proximal ¼ broadest, distal ¾ greatly curved with pointed apex. Parameres (
<figureCitation id="5DD83EDAFFEA392D18A9F969FCC7F943" box="[748,828,1724,1750]" captionStart="FIGURES 47–50" captionStartId="18.[151,264,1227,1251]" captionTargetBox="[250,1333,183,1189]" captionTargetId="figure-23@18.[242,1346,181,1203]" captionTargetPageId="18" captionText="FIGURES 47–50. Macrurohelea sirii n. sp. Holotype male, 47–48. Macrurohelea ventanensis n. sp. Holotype male, 49–50. 47, 49 parameres. 48, 50 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912801" httpUri="https://zenodo.org/record/5912801/files/figure.png" pageId="13" pageNumber="458">Fig. 49</figureCitation>
) broadly separated, heavily sclerotized; basal apodeme stout, laterally directed, apex blunt; distal halves stout, nearly straight, sub-parallel, with slender, slightly hooked, apical, curved finger-like process.Aedeagus (
<figureCitation id="5DD83EDAFFEA392D18D5F8D1FD1BF88A" box="[656,736,1796,1823]" captionStart="FIGURES 47–50" captionStartId="18.[151,264,1227,1251]" captionTargetBox="[250,1333,183,1189]" captionTargetId="figure-23@18.[242,1346,181,1203]" captionTargetPageId="18" captionText="FIGURES 47–50. Macrurohelea sirii n. sp. Holotype male, 47–48. Macrurohelea ventanensis n. sp. Holotype male, 49–50. 47, 49 parameres. 48, 50 aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5912801" httpUri="https://zenodo.org/record/5912801/files/figure.png" pageId="13" pageNumber="458">Fig. 50</figureCitation>
) broadly triangular, 0.80 length of basal width; basal arm stout, laterally directed with curved truncate tip; basal arch deeply concave, heavily sclerotized, extending nearly 2/3 of total length; distal portion lightly sclerotized, tapered distally, with a short, apical hyaline quadrate process.
</paragraph>
<paragraph id="C55C225FFFEA392D1A82F8A5FE59F81F" blockId="13.[151,1437,968,2039]" box="[199,418,1904,1930]" pageId="13" pageNumber="458">
<emphasis id="F797FE4DFFEA392D1A82F8A5FEDEF81F" bold="true" box="[199,293,1904,1930]" pageId="13" pageNumber="458">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="8DF971D4FFEA392D1A82F841FC82F862" pageId="13" pageNumber="458" type="materials_examined">
<paragraph id="C55C225FFFEA392D1A82F841FC82F862" blockId="13.[151,1437,968,2039]" pageId="13" pageNumber="458">
<emphasis id="F797FE4DFFEA392D1A82F841FE82F83B" bold="true" box="[199,377,1940,1966]" pageId="13" pageNumber="458">Type material.</emphasis>
<materialsCitation id="758B2802FFEA392D1BC0F841FC82F862" ID-GBIF-Occurrence="3458732310" collectingDate="2012-11" collectorName="A. Siri &amp; M. Donato" country="Argentina" elevation="574" latitude="-38.058804" location="Ventana" longLatPrecision="1" longitude="-62.019943" municipality="Reserva Parque Prov." pageId="13" pageNumber="458" specimenCount="2" stateProvince="Ciudad Autonoma de Buenos Aires" typeStatus="holotype">
<typeStatus id="1A589CFDFFEA392D1BC0F841FE16F83B" box="[389,493,1940,1966]" pageId="13" pageNumber="458" type="holotype">Holotype</typeStatus>
male,
<typeStatus id="1A589CFDFFEA392D1804F840FD5AF83A" box="[577,673,1941,1967]" pageId="13" pageNumber="458" type="paratype">paratype</typeStatus>
male, labeled “
<taxonomicName id="02E359DCFFEA392D191EF841FF0BF847" authority="Spinelli, Ronderos and Grogan" authorityName="Spinelli, Ronderos and Grogan" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="458" phylum="Arthropoda" rank="species" species="ventanensis">Macrurohelea ventanensis Spinelli, Ronderos and Grogan</taxonomicName>
”, “
<collectingCountry id="BDF462CFFFEA392D1B50F86DFE71F847" box="[277,394,1976,2002]" name="Argentina" pageId="13" pageNumber="458">Argentina</collectingCountry>
,
<collectingRegion id="0727ECBDFFEA392D1BD9F86CFDC1F847" box="[412,570,1976,2003]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="13" pageNumber="458">Buenos Aires</collectingRegion>
,
<collectingMunicipality id="2538B825FFEA392D1809F86CFCB2F846" box="[588,841,1977,2003]" pageId="13" pageNumber="458">Reserva Parque Prov.</collectingMunicipality>
Ernesto Torquinst, arroyo
<location id="C03C7484FFEA392D1EC9F86CFB12F846" LSID="urn:lsid:plazi:treatment:4D4A9349FFEA39311AD2FCE2FF3CF94F:C03C7484FFEA392D1EC9F86CFB12F846" box="[1164,1257,1977,2003]" country="Argentina" latitude="-38.058804" longLatPrecision="1" longitude="-62.019943" municipality="Reserva Parque Prov." name="Ventana" pageId="13" pageNumber="458" stateProvince="Ciudad Autonoma de Buenos Aires">Ventana</location>
,
<geoCoordinate id="A0D74498FFEA392D1EBFF86DFA6CF847" box="[1274,1431,1976,2002]" degrees="38" direction="south" minutes="03" orientation="latitude" pageId="13" pageNumber="458" precision="1" seconds="31.7" value="-38.058804">38º03’31.7’’S</geoCoordinate>
,
<geoCoordinate id="A0D74498FFEA392D1AD2F809FEC7F863" box="[151,316,2012,2038]" degrees="62" direction="west" minutes="01" orientation="longitude" pageId="13" pageNumber="458" precision="1" seconds="11.8" value="-62.019943">62º01’11.8’’W</geoCoordinate>
,
<quantity id="021B8FBAFFEA392D1B0DF809FE6AF862" box="[328,401,2012,2039]" metricMagnitude="2" metricUnit="m" metricValue="5.74" pageId="13" pageNumber="458" unit="m" value="574.0">
<elevation id="4ECEC56CFFEA392D1B0DF809FE6AF862" box="[328,401,2012,2039]" metricMagnitude="2" metricUnit="m" metricValue="5.74" pageId="13" pageNumber="458" unit="m" value="574.0">574 m</elevation>
</quantity>
,
<date id="B15D049FFFEA392D1BD9F809FE07F863" box="[412,508,2012,2038]" bridgedPair="/" pageId="13" pageNumber="458" value="2012-11">
<collectingDate id="A119FD77FFEA392D1BD9F809FE07F863" box="[412,508,2012,2038]" bridgedPair="/" pageId="13" pageNumber="458" value="2012-11">XI/2012</collectingDate>
</date>
,
<collectorName id="68164789FFEA392D1843F809FDA9F863" box="[518,594,2012,2038]" pageId="13" pageNumber="458">A. Siri</collectorName>
–
<collectorName id="68164789FFEA392D1824F809FD20F862" box="[609,731,2012,2039]" pageId="13" pageNumber="458">M. Donato</collectorName>
,
<collectingMethod id="1CA25A48FFEA392D18A3F809FC8EF862" box="[742,885,2012,2039]" pageId="13" pageNumber="458">Malaise trap</collectingMethod>
.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="919C72D7FFE9392E1AD2F842FCFFF846" ID-DOI="http://doi.org/10.5281/zenodo.5912791" ID-Zenodo-Dep="5912791" httpUri="https://zenodo.org/record/5912791/files/figure.png" pageId="14" pageNumber="459" startId="14.[151,264,1943,1967]" targetBox="[211,1381,185,1920]" targetPageId="14">
<paragraph id="C55C225FFFE9392E1AD2F842FCFFF846" blockId="14.[151,1436,1943,2004]" pageId="14" pageNumber="459">
<emphasis id="F797FE4DFFE9392E1AD2F842FEADF83A" bold="true" box="[151,342,1943,1967]" pageId="14" pageNumber="459">FIGURES 26–32.</emphasis>
<taxonomicName id="02E359DCFFE9392E1B1BF84DFDDBF83A" authority="Spinelli &amp; Ronderos &amp; Grogan, 2022" authorityName="Spinelli &amp; Ronderos &amp; Grogan" authorityYear="2022" box="[350,544,1944,1968]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="459" phylum="Arthropoda" rank="species" species="sirii" status="sp. nov.">
<emphasis id="F797FE4DFFE9392E1B1BF84DFDDBF83A" box="[350,544,1944,1968]" italics="true" pageId="14" pageNumber="459">Macrurohelea sirii</emphasis>
</taxonomicName>
<emphasis id="F797FE4DFFE9392E186DF84DFD98F825" bold="true" box="[552,611,1944,1968]" pageId="14" pageNumber="459">
<taxonomicNameLabel id="ECA44336FFE9392E186DF84DFD98F825" box="[552,611,1944,1968]" pageId="14" pageNumber="459" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Allotype female, 26–29. Holotype male, 30–32. 26, 30 head. 27 thorax. 28, 31 wing. 29 tip of abdomen. 32 genitalia. Scale bars = 0.05 mm.
</paragraph>
</caption>
<caption id="919C72D7FFE8392F1AD2F842FF3DF846" ID-DOI="http://doi.org/10.5281/zenodo.5912793" ID-Zenodo-Dep="5912793" httpUri="https://zenodo.org/record/5912793/files/figure.png" pageId="15" pageNumber="460" startId="15.[151,264,1943,1967]" targetBox="[240,1349,181,1917]" targetPageId="15">
<paragraph id="C55C225FFFE8392F1AD2F842FF3DF846" blockId="15.[151,1436,1943,2003]" pageId="15" pageNumber="460">
<emphasis id="F797FE4DFFE8392F1AD2F842FEAFF83A" bold="true" box="[151,340,1943,1967]" pageId="15" pageNumber="460">FIGURES 33–36.</emphasis>
<taxonomicName id="02E359DCFFE8392F1B1CF84DFD9FF83A" authority="Spinelli &amp; Ronderos &amp; Grogan, 2022" authorityName="Spinelli &amp; Ronderos &amp; Grogan" authorityYear="2022" box="[345,612,1944,1968]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="460" phylum="Arthropoda" rank="species" species="ventanensis" status="sp. nov.">
<emphasis id="F797FE4DFFE8392F1B1CF84DFD9FF83A" box="[345,612,1944,1968]" italics="true" pageId="15" pageNumber="460">Macrurohelea ventanensis</emphasis>
</taxonomicName>
<emphasis id="F797FE4DFFE8392F182FF84DFD58F825" bold="true" box="[618,675,1944,1968]" pageId="15" pageNumber="460">
<taxonomicNameLabel id="ECA44336FFE8392F182FF84DFD58F825" box="[618,675,1944,1968]" pageId="15" pageNumber="460" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Holotype male. 33 head. 34 thorax. 35 wing. 36 genitalia. Scale bars = 0.05 mm.
</paragraph>
</caption>
<caption id="919C72D7FFF739301AD2F8ACFBF3F820" ID-DOI="http://doi.org/10.5281/zenodo.5912797" ID-Zenodo-Dep="5912797" httpUri="https://zenodo.org/record/5912797/files/figure.png" pageId="16" pageNumber="461" startId="16.[151,264,1913,1937]" targetBox="[214,1370,182,1882]" targetPageId="16">
<paragraph id="C55C225FFFF739301AD2F8ACFBF3F820" blockId="16.[151,1437,1913,1973]" pageId="16" pageNumber="461">
<emphasis id="F797FE4DFFF739301AD2F8ACFEACF804" bold="true" box="[151,343,1913,1937]" pageId="16" pageNumber="461">FIGURES 37–42.</emphasis>
<taxonomicName id="02E359DCFFF739301B1AF8ACFDC2F804" authority="Spinelli &amp; Ronderos &amp; Grogan, 2022" authorityName="Spinelli &amp; Ronderos &amp; Grogan" authorityYear="2022" box="[351,569,1913,1937]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="461" phylum="Arthropoda" rank="species" species="bassoi" status="sp. nov.">
<emphasis id="F797FE4DFFF739301B1AF8ACFDC2F804" box="[351,569,1913,1937]" italics="true" pageId="16" pageNumber="461">Macrurohelea bassoi</emphasis>
</taxonomicName>
<emphasis id="F797FE4DFFF739301804F8ACFD86F804" bold="true" box="[577,637,1913,1937]" pageId="16" pageNumber="461">
<taxonomicNameLabel id="ECA44336FFF739301804F8ACFD86F804" box="[577,637,1913,1937]" pageId="16" pageNumber="461" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Holotype male, 37–38.
<taxonomicName id="02E359DCFFF739301939F8ACFB99F804" authority="Spinelli &amp; Ronderos &amp; Grogan, 2022" authorityName="Spinelli &amp; Ronderos &amp; Grogan" authorityYear="2022" box="[892,1122,1913,1937]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="461" phylum="Arthropoda" rank="species" species="donatoi" status="sp. nov.">
<emphasis id="F797FE4DFFF739301939F8ACFB99F804" box="[892,1122,1913,1937]" italics="true" pageId="16" pageNumber="461">Macrurohelea donatoi</emphasis>
</taxonomicName>
<emphasis id="F797FE4DFFF739301E2EF8ACFB5DF804" bold="true" box="[1131,1190,1913,1937]" pageId="16" pageNumber="461">
<taxonomicNameLabel id="ECA44336FFF739301E2EF8ACFB5DF804" box="[1131,1190,1913,1937]" pageId="16" pageNumber="461" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Holotype male, 39–40.
<taxonomicName id="02E359DCFFF739301AD2F848FE3FF820" authority="Wirth" authorityName="Wirth" authorityYear="1965" box="[151,452,1949,1973]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="461" phylum="Arthropoda" rank="species" species="kuscheli">
<emphasis id="F797FE4DFFF739301AD2F848FE7AF820" box="[151,385,1949,1973]" italics="true" pageId="16" pageNumber="461">Macrurohelea kuscheli</emphasis>
Wirth
</taxonomicName>
, male, 41–42. 37, 39, 41 parameres. 38, 40, 42 aedeagus.
</paragraph>
</caption>
<caption id="919C72D7FFF639311AD2FA77FDBCFA4B" ID-DOI="http://doi.org/10.5281/zenodo.5912799" ID-Zenodo-Dep="5912799" httpUri="https://zenodo.org/record/5912799/files/figure.png" pageId="17" pageNumber="462" startId="17.[151,264,1442,1466]" targetBox="[166,1437,186,1402]" targetPageId="17">
<paragraph id="C55C225FFFF639311AD2FA77FDBCFA4B" blockId="17.[151,1436,1442,1502]" pageId="17" pageNumber="462">
<emphasis id="F797FE4DFFF639311AD2FA77FEADFA2F" bold="true" box="[151,342,1442,1466]" pageId="17" pageNumber="462">FIGURES 43–46.</emphasis>
<taxonomicName id="02E359DCFFF639311B1BFA77FCD4FA2F" authority="Spinelli &amp; Grogan" authorityName="Spinelli &amp; Grogan" authorityYear="1984" box="[350,815,1442,1466]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="462" phylum="Arthropoda" rank="species" species="monotheca">
<emphasis id="F797FE4DFFF639311B1BFA77FD9EFA2F" box="[350,613,1442,1466]" italics="true" pageId="17" pageNumber="462">Macrurohelea monotheca</emphasis>
Spinelli &amp; Grogan
</taxonomicName>
, male, 43–44.
<taxonomicName id="02E359DCFFF63931198DFA77FB48FA2F" authority="Spinelli &amp; Ronderos &amp; Grogan, 2022" authorityName="Spinelli &amp; Ronderos &amp; Grogan" authorityYear="2022" box="[968,1203,1442,1466]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="462" phylum="Arthropoda" rank="species" species="morenoi" status="sp. nov.">
<emphasis id="F797FE4DFFF63931198DFA77FB48FA2F" box="[968,1203,1442,1466]" italics="true" pageId="17" pageNumber="462">Macrurohelea morenoi</emphasis>
</taxonomicName>
<emphasis id="F797FE4DFFF639311EFEFA77FB0DFA2F" bold="true" box="[1211,1270,1442,1466]" pageId="17" pageNumber="462">
<taxonomicNameLabel id="ECA44336FFF639311EFEFA77FB0DFA2F" box="[1211,1270,1442,1466]" pageId="17" pageNumber="462" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Holotype male, 45–46. 43, 45 parameres. 44, 46 aedeagus.
</paragraph>
</caption>
<subSubSection id="8DF971D4FFF639311A82F9D9FECCF9FB" pageId="17" pageNumber="462" type="distribution">
<paragraph id="C55C225FFFF639311A82F9D9FB0EF9B3" blockId="17.[151,1437,1548,1754]" box="[199,1269,1548,1575]" pageId="17" pageNumber="462">
<emphasis id="F797FE4DFFF639311A82F9D9FE9AF9B3" bold="true" box="[199,353,1548,1574]" pageId="17" pageNumber="462">Distribution.</emphasis>
<collectingCountry id="BDF462CFFFF639311B23F9D9FE23F9B3" box="[358,472,1548,1574]" name="Argentina" pageId="17" pageNumber="462">Argentina</collectingCountry>
, known only from the type-locality in southern
<collectingRegion id="0727ECBDFFF6393119AAF9D8FB09F9B3" box="[1007,1266,1548,1575]" country="Argentina" name="Buenos Aires" pageId="17" pageNumber="462">Buenos Aires province</collectingRegion>
.
</paragraph>
<paragraph id="C55C225FFFF639311A82F9E5FECCF9FB" blockId="17.[151,1437,1548,1754]" pageId="17" pageNumber="462">
<emphasis id="F797FE4DFFF639311A82F9E5FDD9F9DF" bold="true" box="[199,546,1584,1610]" pageId="17" pageNumber="462">Derivation of specific epithet.</emphasis>
The specific epithet is in reference to Sierra de la Ventana, where the type-series was collected.
</paragraph>
</subSubSection>
<subSubSection id="8DF971D4FFF639311A82F9ADFF3CF94F" pageId="17" pageNumber="462" type="discussion">
<paragraph id="C55C225FFFF639311A82F9ADFF3CF94F" blockId="17.[151,1437,1548,1754]" pageId="17" pageNumber="462">
<emphasis id="F797FE4DFFF639311A82F9ADFEB7F907" bold="true" box="[199,332,1656,1682]" pageId="17" pageNumber="462">Discussion.</emphasis>
Males of this new species are similar to those of
<taxonomicName id="02E359DCFFF639311920F9ACFC1EF907" authorityName="Wirth" authorityYear="1965" box="[869,997,1656,1683]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="462" phylum="Arthropoda" rank="species" species="kuscheli">
<emphasis id="F797FE4DFFF639311920F9ACFC87F907" box="[869,892,1657,1682]" italics="true" pageId="17" pageNumber="462">M</emphasis>
.
<emphasis id="F797FE4DFFF6393119CCF9ADFC1EF907" box="[905,997,1656,1682]" italics="true" pageId="17" pageNumber="462">kuscheli</emphasis>
</taxonomicName>
. However, in
<taxonomicName id="02E359DCFFF639311E3AF9ACFB04F907" authorityName="Wirth" authorityYear="1965" box="[1151,1279,1656,1683]" class="Insecta" family="Ceratopogonidae" genus="Macrurohelea" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="462" phylum="Arthropoda" rank="species" species="kuscheli">
<emphasis id="F797FE4DFFF639311E3AF9ACFB6DF907" box="[1151,1174,1657,1682]" italics="true" pageId="17" pageNumber="462">M</emphasis>
.
<emphasis id="F797FE4DFFF639311EE6F9ADFB04F907" box="[1187,1279,1656,1682]" italics="true" pageId="17" pageNumber="462">kuscheli</emphasis>
</taxonomicName>
flagellomeres 4–9 are fused, and the aedeagus is slightly triangular with the distal portion slender, bifid, with short, separated tips.
</paragraph>
</subSubSection>
</treatment>
</document>
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