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<document id="EA5E6F820159209DA8854E61ADB94DBB" ID-DOI="10.5281/zenodo.12586261" ID-ISSN="2064-2474" ID-Zenodo-Dep="12586261" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="admin" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719595820211" checkinUser="felipe" docAuthor="Zwick, P &amp; Surenkhorloo, P" docDate="2005" docId="4D1DFF14BF487264FDB7101DFDF570F6" docLanguage="en" docName="ActaZoolAcadSciH.51.3.253-276.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 51 (3)" docStyle="DocumentStyle:4FD749C6DF7B87D8940F4D8C7B0A0514.3:ActaZoolAcadSciH.2002-2014.journal_article" docStyleId="4FD749C6DF7B87D8940F4D8C7B0A0514" docStyleName="ActaZoolAcadSciH.2002-2014.journal_article" docStyleVersion="3" docTitle="Isoperla altaica SAMAL 1939" docType="treatment" docVersion="4" lastPageNumber="262" masterDocId="B124876CBF4E726DFFDF1343FFD0721D" masterDocTitle="The Mongolian Species Of Isoperlinae (Plecoptera: Perlodidae)" masterLastPageNumber="276" masterPageNumber="253" pageNumber="259" updateTime="1719713003434" updateUser="admin" zenodo-license-document="CC-BY-NC-4.0">
<mods:mods id="617C7D089D06C3A62F63E5C7DC6CF0BB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3F6B4632AC945B5DCAA287038B3E2A58">
<mods:title id="366E516269070AA9C583ED9898AF2559">The Mongolian Species Of Isoperlinae (Plecoptera: Perlodidae)</mods:title>
</mods:titleInfo>
<mods:name id="5C0B0F702A49EE45D7BF13659135A605" type="personal">
<mods:role id="29A05FDC84522A8CFAAC8D386E44DB46">
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<mods:namePart id="CA9003D6140EE0151C840533CF1955D7">Zwick, P</mods:namePart>
</mods:name>
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<mods:role id="8CFE96B6A89335B66F4AA57BEEB1AEEF">
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<mods:namePart id="D6748468130DCD740A7E52AFFFEFEA0C">Surenkhorloo, P</mods:namePart>
</mods:name>
<mods:typeOfResource id="C19CEAE2A50889D7FB2E5EAF1E0992A2">text</mods:typeOfResource>
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<mods:title id="76E78A26568483E37378EF8B296173C2">Acta Zoologica Academiae Scientiarum Hungaricae</mods:title>
</mods:titleInfo>
<mods:part id="22BB76BFAE570D900B924086B4380620">
<mods:date id="CFFF787AD0B3E808447747E1720B62E1">2005</mods:date>
<mods:detail id="1B648DC54378779BBEB0D2E8BD6E848E" type="pubDate">
<mods:number id="0F9271E21887BC23B49AD374A7A9001A">2005-10-31</mods:number>
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<mods:detail id="82666FF0D1C75E077BCD65BB867CEF85" type="volume">
<mods:number id="2DE16DDC77E8DE7A8A66B73E90FEFB84">51</mods:number>
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<mods:number id="4EA9898E0176F5C4F87333C8AB5D57E3">3</mods:number>
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<mods:start id="CAD102FB8C31B62B8D0140E763478E9B">253</mods:start>
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<mods:classification id="4D94B4A431BDB75B7846CC19E4D0D0E3">journal article</mods:classification>
<mods:identifier id="A345DC5AB02C7339E1FD52D090686CC6" type="DOI">10.5281/zenodo.12586261</mods:identifier>
<mods:identifier id="CA1DF2871C3324923E6E8A8820E63F3C" type="ISSN">2064-2474</mods:identifier>
<mods:identifier id="A8A24600164808D37E37279A8D245D62" type="Zenodo-Dep">12586261</mods:identifier>
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<treatment id="4D1DFF14BF487264FDB7101DFDF570F6" ID-DOI="http://doi.org/10.5281/zenodo.12586344" ID-Zenodo-Dep="12586344" LSID="urn:lsid:plazi:treatment:4D1DFF14BF487264FDB7101DFDF570F6" httpUri="http://treatment.plazi.org/id/4D1DFF14BF487264FDB7101DFDF570F6" lastPageId="9" lastPageNumber="262" pageId="6" pageNumber="259">
<subSubSection id="8DAE1D89BF48726BFDB7101DFC1B7165" box="[616,971,862,888]" pageId="6" pageNumber="259" type="nomenclature">
<paragraph id="C50B4E02BF48726BFDB7101DFC1B7165" blockId="6.[616,971,862,888]" box="[616,971,862,888]" pageId="6" pageNumber="259">
<heading id="9E43F96EBF48726BFDB7101DFC1B7165" box="[616,971,862,888]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="259" reason="6">
<taxonomicName id="02B43581BF48726BFDB7101DFC1B7165" ID-CoL="6MZVM" authority="SAMAL, 1939" authorityName="SAMAL" authorityYear="1939" box="[616,971,862,888]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="altaica">
<emphasis id="F7C09210BF48726BFDB7101DFCF27165" box="[616,802,862,888]" italics="true" pageId="6" pageNumber="259">Isoperla altaica</emphasis>
<bibRefCitation id="A12533F3BF48726BFCF6101DFC1B7165" author="SAMAL, J." box="[809,971,862,888]" pageId="6" pageNumber="259" pagination="419 - 426" refId="ref9697" refString="SAMAL, J. (1939) Contribution a l'etude de la faune des Plecopteres d'Altai. Vest. Ceskosl. spol. zool. 6 - 7: 419 - 426." type="journal article" year="1939">
<smallCapsWord id="C3EDD8DEBF48726BFCF6101DFC52716A" baselines="881,881" box="[809,898,862,888]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Samal" pageId="6" pageNumber="259">ŠÁMAL</smallCapsWord>
, 1939
</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8DAE1D89BF487264FD7A10C2FDF570F6" lastPageId="9" lastPageNumber="262" pageId="6" pageNumber="259" type="description">
<paragraph id="C50B4E02BF48726BFD7A10C2FC5E7186" blockId="6.[677,910,897,923]" box="[677,910,897,923]" pageId="6" pageNumber="259">
<heading id="9E43F96EBF48726BFD7A10C2FC5E7186" box="[677,910,897,923]" centered="true" fontSize="11" level="3" pageId="6" pageNumber="259" reason="3">
(
<figureCitation id="5D8F5287BF48726BFD7210C2FD297186" box="[685,761,897,923]" captionStart="Fig" captionStartId="7.[317,352,1705,1726]" captionTargetBox="[319,1267,408,1673]" captionTargetId="figure-14@7.[319,1267,404,1677]" captionTargetPageId="7" captionText="Fig. 3. Isoperla altaica Šámal. a = head and prothorax, b, c = male abdominal tip in ventral and dorsal views, respectively; d, e = penis sclerite in lateral and caudal views, respectively, f = spicules on penial membrane next to large sclerite, g = male with everted penis, lateral view, h = female abdomi-" figureDoi="http://doi.org/10.5281/zenodo.12586267" httpUri="https://zenodo.org/record/12586267/files/figure.png" pageId="6" pageNumber="259">Figs 3</figureCitation>
,
<figureCitation id="5D8F5287BF48726BFCD910C2FCC77186" box="[774,791,897,923]" captionStart="Fig" captionStartId="8.[376,411,1791,1812]" captionTargetBox="[328,1260,1255,1738]" captionTargetId="figure-382@8.[319,1267,1234,1763]" captionTargetPageId="8" captionText="Fig. 4. Isoperla altaica ŠÁMAL, egg and details of collar; specimen from Khonin Nuga" figureDoi="http://doi.org/10.5281/zenodo.12586269" httpUri="https://zenodo.org/record/12586269/files/figure.png" pageId="6" pageNumber="259">4</figureCitation>
,
<figureCitation id="5D8F5287BF48726BFCFC10C2FCE07186" box="[803,816,897,923]" captionStart="Fig" captionStartId="9.[317,352,1772,1793]" captionTargetBox="[363,1234,925,1735]" captionTargetId="figure-127@9.[350,1238,909,1744]" captionTargetPageId="9" captionText="Fig. 5. Larvae of Isoperla spp. a =I.lunigera (KLAPÁLEK), b–d = I.altaica ŠÁMAL, with details of labium and labial segment" figureDoi="http://doi.org/10.5281/zenodo.12586271" httpUri="https://zenodo.org/record/12586271/files/figure.png" pageId="6" pageNumber="259">5</figureCitation>
b-d, 6a)
</heading>
</paragraph>
<paragraph id="C50B4E02BF48726BFEE21086FD7971C4" blockId="6.[317,886,964,1014]" box="[317,681,964,986]" pageId="6" pageNumber="259">
<taxonomicName id="02B43581BF48726BFEE21086FDCC71C4" authority="Samal" authorityName="Samal" box="[317,540,964,986]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="altaica">
<emphasis id="F7C09210BF48726BFEE21086FE0671C7" box="[317,470,965,986]" italics="true" pageId="6" pageNumber="259">Isoperla altaica</emphasis>
Šámal
</taxonomicName>
: 421, figs 4, 5.
</paragraph>
<paragraph id="C50B4E02BF48726BFEE210A2FCA671EB" blockId="6.[317,886,964,1014]" box="[317,886,993,1014]" pageId="6" pageNumber="259">
<taxonomicName id="02B43581BF48726BFEE210A2FE0671EB" box="[317,470,993,1014]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="altaica">
<emphasis id="F7C09210BF48726BFEE210A2FE0671EB" box="[317,470,993,1014]" italics="true" pageId="6" pageNumber="259">Isoperla altaica</emphasis>
</taxonomicName>
-
<smallCapsWord id="C3EDD8DEBF48726BFE3B10A2FD9A71EB" baselines="1009,1009" box="[484,586,993,1014]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Zapekina" pageId="6" pageNumber="259">ZAPEKINA</smallCapsWord>
<smallCapsWord id="C3EDD8DEBF48726BFD8F10A2FD7971E8" baselines="1009,1009" box="[592,681,993,1014]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Dulkeit" pageId="6" pageNumber="259">DULKEIT</smallCapsWord>
, 1955. 169, figs 5–9.
</paragraph>
<paragraph id="C50B4E02BF48726BFEA11758FB3176C0" blockId="6.[317,1270,1051,1446]" pageId="6" pageNumber="259">
<materialsCitation id="75DC445FBF48726BFEA11758FB0A76C6" collectingDate="1966-07-08" collectingDateMax="2003-08-04" collectingDateMin="1966-07-08" collectionCode="V, ZWICK" collectorName="River" country="Mongolia" county="Material" elevation="1750" latitude="49.966667" location="Gebirge" longLatPrecision="1100" longitude="82.6" municipality="Mongol" pageId="6" pageNumber="259" specimenCount="1" stateProvince="Altay">
<collectingCounty id="2C6A368EBF48726BFEA11758FE1F762D" box="[382,463,1051,1072]" pageId="6" pageNumber="259">Material</collectingCounty>
studied.
<collectingCountry id="BDA30E92BF48726BFDF01758FD66762D" box="[559,694,1051,1072]" name="Mongolia" pageId="6" pageNumber="259">MONGOLIA</collectingCountry>
: Chovd aimak, ca
<locationDeviation id="D4B95720BF48726BFCA81758FBB1762D" box="[887,1121,1051,1072]" location="Somon Uenc" pageId="6" pageNumber="259">
<quantity id="024CE3E7BF48726BFCA81758FC69762D" box="[887,953,1051,1072]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="259" unit="km" value="35.0">35 km</quantity>
N Somon Uenč
</locationDeviation>
,
<collectingMunicipality id="256FD478BF48726BFBB01758FB6B762D" box="[1135,1211,1051,1072]" pageId="6" pageNumber="259">Mongol</collectingMunicipality>
<collectingRegion id="077080E0BF48726BFB1A1758FB25762D" box="[1221,1269,1051,1072]" country="Russia" name="Altay" pageId="6" pageNumber="259">Altaj</collectingRegion>
<location id="C06B18D9BF48726BFEE2177BFE5D7650" LSID="urn:lsid:plazi:treatment:4D1DFF14BF487264FDB7101DFDF570F6:C06B18D9BF48726BFEE2177BFE5D7650" box="[317,397,1080,1101]" country="Mongolia" county="Material" latitude="49.966667" longLatPrecision="1100" longitude="82.6" municipality="Mongol" name="Gebirge" pageId="6" pageNumber="259" stateProvince="Altay">Gebirge</location>
,
<quantity id="024CE3E7BF48726BFE46177BFE0E7650" box="[409,478,1080,1101]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="6" pageNumber="259" unit="m" value="1750.0">
<elevation id="4E99A931BF48726BFE46177BFE0E7650" box="[409,478,1080,1101]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="6" pageNumber="259" unit="m" value="1750.0">1750m</elevation>
</quantity>
, kl.
<location id="C06B18D9BF48726BFDD4177BFD0A7650" LSID="urn:lsid:plazi:treatment:4D1DFF14BF487264FDB7101DFDF570F6:C06B18D9BF48726BFDD4177BFD0A7650" box="[523,730,1080,1101]" country="Mongolia" county="Material" latitude="49.966667" longLatPrecision="1100" longitude="82.6" municipality="Mongol" name="Nebenbach des Uenc" pageId="6" pageNumber="259" stateProvince="Altay">Nebenbach des Uenč</location>
gol,
<date id="B10A68C2BF48726BFCD0177BFCAB7650" box="[783,891,1080,1101]" pageId="6" pageNumber="259" value="1966-07-08">
<collectingDate id="A14E912ABF48726BFCD0177BFCAB7650" box="[783,891,1080,1101]" pageId="6" pageNumber="259" value="1966-07-08">8.VII.1966</collectingDate>
</date>
,
<quantity id="024CE3E7BF48726BFC58177BFC6F7650" box="[903,959,1079,1101]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="259" unit="mm" value="1.0">1 mm</quantity>
[
<smallCapsWord id="C3EDD8DEBF48726BFC12177BFBCE7656" baselines="1096,1095" box="[973,1054,1080,1101]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Kaszab" pageId="6" pageNumber="259">KASZAB</smallCapsWord>
646]. Selenge aimak,
<location id="C06B18D9BF48726BFEE21717FE567674" LSID="urn:lsid:plazi:treatment:4D1DFF14BF487264FDB7101DFDF570F6:C06B18D9BF48726BFEE21717FE567674" box="[317,390,1108,1129]" country="Mongolia" county="Material" latitude="49.966667" longLatPrecision="1100" longitude="82.6" municipality="Mongol" name="Mandal" pageId="6" pageNumber="259" stateProvince="Altay">Mandal</location>
sum,
<location id="C06B18D9BF48726BFE611717FDD67674" LSID="urn:lsid:plazi:treatment:4D1DFF14BF487264FDB7101DFDF570F6:C06B18D9BF48726BFE611717FDD67674" box="[446,518,1108,1129]" country="Mongolia" county="Material" latitude="49.966667" longLatPrecision="1100" longitude="82.6" municipality="Mongol" name="Khonin" pageId="6" pageNumber="259" stateProvince="Altay">Khonin</location>
nuga:
<location id="C06B18D9BF48726BFD9B1717FD267674" LSID="urn:lsid:plazi:treatment:4D1DFF14BF487264FDB7101DFDF570F6:C06B18D9BF48726BFD9B1717FD267674" box="[580,758,1108,1129]" country="Mongolia" county="Material" latitude="49.966667" longLatPrecision="1100" longitude="82.6" municipality="Mongol" name="Bar-Chuluut River" pageId="6" pageNumber="259" stateProvince="Altay">Bar-Chuluut River</location>
[48°58’638N,
<geoCoordinate id="A08028C5BF48726BFC5B1717FBD97674" box="[900,1033,1108,1129]" degrees="106" direction="east" minutes="57" orientation="longitude" pageId="6" pageNumber="259" precision="15" seconds="013" value="106.953606">106°57’013E</geoCoordinate>
], tributary of
<location id="C06B18D9BF48726BFB581717FB227674" LSID="urn:lsid:plazi:treatment:4D1DFF14BF487264FDB7101DFDF570F6:C06B18D9BF48726BFB581717FB227674" box="[1159,1266,1108,1129]" country="Mongolia" county="Material" latitude="49.966667" longLatPrecision="1100" longitude="82.6" municipality="Mongol" name="Eroo River" pageId="6" pageNumber="259" stateProvince="Altay">Eroo River</location>
, 1 f,
<date id="B10A68C2BF48726BFEAC1732FE33769B" box="[371,483,1137,1158]" pageId="6" pageNumber="259" value="1999-09-07">
<collectingDate id="A14E912ABF48726BFEAC1732FE33769B" box="[371,483,1137,1158]" pageId="6" pageNumber="259" value="1999-09-07">07.IX.1999</collectingDate>
</date>
; 3 ff,
<date id="B10A68C2BF48726BFDE71732FD68769B" box="[568,696,1137,1158]" pageId="6" pageNumber="259" value="2001-08-21">
<collectingDate id="A14E912ABF48726BFDE71732FD68769B" box="[568,696,1137,1158]" pageId="6" pageNumber="259" value="2001-08-21">21.VIII.2001</collectingDate>
</date>
;
<quantity id="024CE3E7BF48726BFD1A1732FD2E769A" box="[709,766,1137,1159]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="259" unit="mm" value="2.0">2 mm</quantity>
, 4 ff,
<date id="B10A68C2BF48726BFC8B1732FC01769B" box="[852,977,1137,1158]" pageId="6" pageNumber="259" value="2003-08-04">
<collectingDate id="A14E912ABF48726BFC8B1732FC01769B" box="[852,977,1137,1158]" pageId="6" pageNumber="259" value="2003-08-04">04.VIII.2003</collectingDate>
</date>
(all leg. P.
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). KAZACHSTAN, surroundings of
<location id="C06B18D9BF48726BFD5217CDFCEC76BE" LSID="urn:lsid:plazi:treatment:4D1DFF14BF487264FDB7101DFDF570F6:C06B18D9BF48726BFD5217CDFCEC76BE" box="[653,828,1166,1187]" country="Mongolia" county="Material" latitude="49.966667" longLatPrecision="1100" longitude="82.6" municipality="Mongol" name="Ust Kamenogorsk" pageId="6" pageNumber="259" stateProvince="Altay">Ust Kamenogorsk</location>
[
<geoCoordinate id="A08028C5BF48726BFC9617CDFC7076BE" box="[841,928,1166,1187]" degrees="49" direction="north" minutes="58" orientation="latitude" pageId="6" pageNumber="259" precision="925" value="49.966667">49°58’N</geoCoordinate>
,
<geoCoordinate id="A08028C5BF48726BFC7317CDFC2F76BE" box="[940,1023,1166,1187]" degrees="82" direction="east" minutes="36" orientation="longitude" pageId="6" pageNumber="259" precision="925" value="82.6">82°36’E</geoCoordinate>
], basin of
<location id="C06B18D9BF48726BFBB917CDFE5D76DD" LSID="urn:lsid:plazi:treatment:4D1DFF14BF487264FDB7101DFDF570F6:C06B18D9BF48726BFBB917CDFE5D76DD" country="Mongolia" county="Material" latitude="49.966667" longLatPrecision="1100" longitude="82.6" municipality="Mongol" name="River Gornaya Ulbinka" pageId="6" pageNumber="259" stateProvince="Altay">River Gornaya Ulbinka</location>
,
<date id="B10A68C2BF48726BFE4817E8FE2976DD" box="[407,505,1195,1216]" pageId="6" pageNumber="259" value="2000-11-06">
<collectingDate id="A14E912ABF48726BFE4817E8FE2976DD" box="[407,505,1195,1216]" pageId="6" pageNumber="259" value="2000-11-06">11.6.2000</collectingDate>
</date>
, leg. &amp; det.
<collectionCode id="A3A5D6C7BF48726BFDB017E8FD5376DD" box="[623,643,1195,1216]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="259" type="Museum">V</collectionCode>
.
<smallCapsWord id="C3EDD8DEBF48726BFD5217E8FCD676A3" baselines="1211,1210" box="[653,774,1195,1216]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Devyatkov" pageId="6" pageNumber="259">DEVYATKOV</smallCapsWord>
,
<quantity id="024CE3E7BF48726BFCCD17E8FC9876DD" box="[786,840,1194,1216]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="259" unit="mm" value="8.0">8 mm</quantity>
, 9 ff (in coll.
<smallCapsWord id="C3EDD8DEBF48726BFC3D17E8FBF376A3" baselines="1211,1210" box="[994,1059,1195,1216]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Zwick" pageId="6" pageNumber="259">
<collectionCode id="A3A5D6C7BF48726BFC3D17E8FBF376A3" box="[994,1059,1195,1216]" pageId="6" pageNumber="259">ZWICK</collectionCode>
</smallCapsWord>
).
<collectingCountry id="BDA30E92BF48726BFBE817E8FB5F76DD" box="[1079,1167,1195,1216]" name="Russia" pageId="6" pageNumber="259">RUSSIA</collectingCountry>
,
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,
<location id="C06B18D9BF48726BFEA5178BFE2676C0" LSID="urn:lsid:plazi:treatment:4D1DFF14BF487264FDB7101DFDF570F6:C06B18D9BF48726BFEA5178BFE2676C0" box="[378,502,1224,1245]" country="Mongolia" county="Material" latitude="49.966667" longLatPrecision="1100" longitude="82.6" municipality="Mongol" name="Iturup Island" pageId="6" pageNumber="259" stateProvince="Altay">Iturup Island</location>
,
<quantity id="024CE3E7BF48726BFE21178BFDF076C0" box="[510,544,1223,1245]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="259" unit="m" value="1.0">
<elevation id="4E99A931BF48726BFE21178BFDF076C0" box="[510,544,1223,1245]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="259" unit="m" value="1.0">1 m</elevation>
</quantity>
, 1 f,
<date id="B10A68C2BF48726BFD89178BFD7C76C0" box="[598,684,1224,1245]" pageId="6" pageNumber="259" value="1998-08-01">
<collectingDate id="A14E912ABF48726BFD89178BFD7C76C0" box="[598,684,1224,1245]" pageId="6" pageNumber="259" value="1998-08-01">1.8.1998</collectingDate>
</date>
,
<location id="C06B18D9BF48726BFD6A178BFCF676C0" LSID="urn:lsid:plazi:treatment:4D1DFF14BF487264FDB7101DFDF570F6:C06B18D9BF48726BFD6A178BFCF676C0" box="[693,806,1224,1245]" country="Mongolia" county="Material" latitude="49.966667" longLatPrecision="1100" longitude="82.6" municipality="Mongol" name="Lake Iturup" pageId="6" pageNumber="259" stateProvince="Altay">Lake Iturup</location>
,
<location id="C06B18D9BF48726BFCF1178BFC3D76C0" LSID="urn:lsid:plazi:treatment:4D1DFF14BF487264FDB7101DFDF570F6:C06B18D9BF48726BFCF1178BFC3D76C0" box="[814,1005,1224,1245]" country="Mongolia" county="Material" latitude="49.966667" longLatPrecision="1100" longitude="82.6" municipality="Mongol" name="River Blagodatnaya" pageId="6" pageNumber="259" stateProvince="Altay">
<collectorName id="68412BD4BF48726BFCF1178BFCB376C0" box="[814,867,1224,1245]" pageId="6" pageNumber="259">River</collectorName>
Blagodatnaya
</location>
, leg. &amp; det.
<collectionCode id="A3A5D6C7BF48726BFB84178BFBBF76C0" box="[1115,1135,1224,1245]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="259" type="Museum">V</collectionCode>
.
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</materialsCitation>
.
</paragraph>
<paragraph id="C50B4E02BF48726BFEA117A7FDFC77BB" blockId="6.[317,1270,1051,1446]" pageId="6" pageNumber="259">
<materialsCitation id="75DC445FBF48726BFEA117A7FDF977BB" collectionCode="SAMAL" county="The" location="Lake Kara-Koli" municipality="River Karas" pageId="6" pageNumber="259" specimenCount="2" stateProvince="Leningrad">
<collectingCounty id="2C6A368EBF48726BFEA117A7FE7376E4" box="[382,419,1252,1273]" pageId="6" pageNumber="259">The</collectingCounty>
male
<typeStatus id="1A0FF0A0BF48726BFE0217A7FDD676E4" box="[477,518,1252,1273]" pageId="6" pageNumber="259">type</typeStatus>
specimen was collected on the
<collectingMunicipality id="256FD478BF48726BFCED17A7FC7676E4" box="[818,934,1252,1273]" pageId="6" pageNumber="259">River Karas</collectingMunicipality>
, outflow of
<location id="C06B18D9BF48726BFBC617A7FB6276E4" LSID="urn:lsid:plazi:treatment:4D1DFF14BF487264FDB7101DFDF570F6:C06B18D9BF48726BFBC617A7FB6276E4" box="[1049,1202,1252,1273]" county="The" municipality="River Karas" name="Lake Kara-Koli" pageId="6" pageNumber="259" stateProvince="Leningrad">Lake Kara-Koli</location>
, by the staff of the
<location id="C06B18D9BF48726BFE721642FD07770B" LSID="urn:lsid:plazi:treatment:4D1DFF14BF487264FDB7101DFDF570F6:C06B18D9BF48726BFE721642FD07770B" box="[429,727,1281,1302]" county="The" municipality="River Karas" name="Hydrobiological State Institute" pageId="6" pageNumber="259" stateProvince="Leningrad">Hydrobiological State Institute</location>
in
<collectingRegion id="077080E0BF48726BFD281642FC8A770B" box="[759,858,1281,1302]" country="Russia" name="Leningrad" pageId="6" pageNumber="259">Leningrad</collectingRegion>
and was given to J.
<smallCapsWord id="C3EDD8DEBF48726BFBFD1642FBB67708" baselines="1297,1297" box="[1058,1126,1281,1302]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Samal" pageId="6" pageNumber="259">
<collectionCode id="A3A5D6C7BF48726BFBFD1642FBB67708" box="[1058,1126,1281,1302]" pageId="6" pageNumber="259">ŠÁMAL</collectionCode>
</smallCapsWord>
for study. The
<typeStatus id="1A0FF0A0BF48726BFEE2165DFEB6772E" box="[317,358,1310,1331]" pageId="6" pageNumber="259">type</typeStatus>
is not in the collection of the
<location id="C06B18D9BF48726BFD55165DFC98772E" LSID="urn:lsid:plazi:treatment:4D1DFF14BF487264FDB7101DFDF570F6:C06B18D9BF48726BFD55165DFC98772E" box="[650,840,1310,1331]" county="The" municipality="River Karas" name="Zoological Institute" pageId="6" pageNumber="259" stateProvince="Leningrad">Zoological Institute</location>
<collectingRegion id="077080E0BF48726BFC91165DFC06772E" box="[846,982,1310,1331]" country="Russia" name="Saint Petersburg" pageId="6" pageNumber="259">St. Petersburg</collectingRegion>
today (
<bibRefCitation id="A12533F3BF48726BFBC0165DFB15772E" author="ZHILTZOVA, L. A." box="[1055,1221,1310,1331]" pageId="6" pageNumber="259" refId="ref10213" refString="ZHILTZOVA, L. A. (1995) Catalogue of type specimens in the collection of the Zoological Institute, Russian Academy of Sciences. Stoneflies (Plecoptera). Zoological Institute St. Petersburg, 38 pp. [in Russian]" type="book" year="1995">
<smallCapsWord id="C3EDD8DEBF48726BFBC0165DFB5D772F" baselines="1326,1326" box="[1055,1165,1310,1331]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Zhiltzova" pageId="6" pageNumber="259">ZHILTZOVA</smallCapsWord>
1995
</bibRefCitation>
) and may be lost.
<smallCapsWord id="C3EDD8DEBF48726BFE691678FE2A7753" baselines="1355,1354" box="[438,506,1339,1360]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Samal" pageId="6" pageNumber="259">
<collectionCode id="A3A5D6C7BF48726BFE691678FE2A7753" box="[438,506,1339,1360]" pageId="6" pageNumber="259">ŠÁMAL</collectionCode>
</smallCapsWord>
emphasized its slight brachyptery. His illustration shows a large ventral lobe on sternite 8. Tergite 10 was described and illustrated as “partly open” medially. However, this condition was probably an artifact, perhaps through drying because there is really only a weakly sclerotized median strip
</materialsCitation>
.
</paragraph>
<paragraph id="C50B4E02BF48726BFEA11688FCC3746D" blockId="6.[317,1270,1483,1821]" pageId="6" pageNumber="259">
The original description of body pattern is not detailed. We accept the interpretation of the first revising author (
<bibRefCitation id="A12533F3BF48726BFE0716ABFD0577E0" author="ZAPEKINA-DULKEIT, J. I." box="[472,725,1512,1533]" pageId="6" pageNumber="259" pagination="167 - 177" refId="ref9819" refString="ZAPEKINA-DULKEIT, J. I. (1955) [On the Plecoptera-fauna of the Telezkoe basin]. Entomol. Obozr. 34: 167 - 177. [in Russian]" type="journal article" year="1955">
<smallCapsWord id="C3EDD8DEBF48726BFE0716ABFD4677E6" baselines="1528,1527,1528,1528,1527" box="[472,662,1512,1533]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Zapekina-Dulkeit" pageId="6" pageNumber="259">ZAPEKINA-DULKEIT</smallCapsWord>
, 1955
</bibRefCitation>
;
<bibRefCitation id="A12533F3BF48726BFD0216ABFCDE77E0" author="ZAPEKINA-DULKEIT, J. I." box="[733,782,1512,1533]" pageId="6" pageNumber="259" pagination="199 - 217" refId="ref9852" refString="ZAPEKINA-DULKEIT, J. I. (1975) [To the fauna of stoneflies (Plecoptera, Insecta) of rivers of the Lake Baikal basin]. Trudi Gosud. Zapovedn. &quot; Stolbi &quot; 10: 199 - 217. [in Russian] [quoted from ZHILTZOVA 2003]." type="journal article" year="1975">1975</bibRefCitation>
) who distinguished
<taxonomicName id="02B43581BF48726BFC1216ABFBF377E0" box="[973,1059,1512,1533]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="altaica">
<emphasis id="F7C09210BF48726BFC1216ABFBF377E0" box="[973,1059,1512,1533]" italics="true" pageId="6" pageNumber="259">I. altaica</emphasis>
</taxonomicName>
from a new species,
<taxonomicName id="02B43581BF48726BFB3716ABFEAD7407" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="eximia">
<emphasis id="F7C09210BF48726BFB3716ABFEAD7407" italics="true" pageId="6" pageNumber="259">I. eximia</emphasis>
</taxonomicName>
; see also there. The central dark mark on the head is entire and extends to the front edge of the pronotum. In front of it, the head is pale to its front border. A less dark band-like transverse mark extends from the centre of head to the compound eyes. The median part of the meso- and methathoracic scuta is darker than the lateral portion (
<figureCitation id="5D8F5287BF48726BFD601518FCD7746D" box="[703,775,1627,1648]" captionStart="Fig" captionStartId="7.[317,352,1705,1726]" captionTargetBox="[319,1267,408,1673]" captionTargetId="figure-14@7.[319,1267,404,1677]" captionTargetPageId="7" captionText="Fig. 3. Isoperla altaica Šámal. a = head and prothorax, b, c = male abdominal tip in ventral and dorsal views, respectively; d, e = penis sclerite in lateral and caudal views, respectively, f = spicules on penial membrane next to large sclerite, g = male with everted penis, lateral view, h = female abdomi-" figureDoi="http://doi.org/10.5281/zenodo.12586267" httpUri="https://zenodo.org/record/12586267/files/figure.png" pageId="6" pageNumber="259">Fig. 3a</figureCitation>
).
</paragraph>
<paragraph id="C50B4E02BF48726BFEA1153BFC7F74B4" blockId="6.[317,1270,1483,1821]" pageId="6" pageNumber="259">
Size. Fore wing length, mm
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, ff
<quantity id="024CE3E7BF48726BFCE2153BFC417490" box="[829,913,1656,1677]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="6" pageNumber="259" unit="mm" value="10.5">10.5 mm</quantity>
(specimens from Khonin nuga); mm 8.0–
<quantity id="024CE3E7BF48726BFEB815D7FE6574B4" box="[359,437,1684,1705]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="6" pageNumber="259" unit="mm" value="8.2">8.2 mm</quantity>
, ff
<quantity id="024CE3E7BF48726BFE2D15D7FDAF74B4" box="[498,639,1684,1705]" metricMagnitude="-2" metricUnit="m" metricValue="1.075" metricValueMax="1.1199999999999999" metricValueMin="1.03" pageId="6" pageNumber="259" unit="mm" value="10.75" valueMax="11.2" valueMin="10.3">10.3–11.2 mm</quantity>
(specimens from
<collectingCountry id="BDA30E92BF48726BFCEF15D7FC7574B4" box="[816,933,1684,1705]" name="Kazakhstan" pageId="6" pageNumber="259">Kazakhstan</collectingCountry>
).
</paragraph>
<paragraph id="C50B4E02BF487265FEA115F2FB00703D" blockId="6.[317,1270,1483,1821]" lastBlockId="8.[317,1270,436,1120]" lastPageId="8" lastPageNumber="261" pageId="6" pageNumber="259">
Male. Specimens from
<collectingCountry id="BDA30E92BF48726BFDBD15F2FD0274DB" box="[610,722,1713,1734]" name="Kazakhstan" pageId="6" pageNumber="259">Kazakhstan</collectingCountry>
are slightly brachypterous, i.e., the wings extend barely beyond the abdominal tip while those from Khonin Nuga (
<collectingCountry id="BDA30E92BF48726BFCAD158DFC0374FE" box="[882,979,1742,1763]" name="Mongolia" pageId="6" pageNumber="259">Mongolia</collectingCountry>
, upper reaches of Eroo River) have normal wings. Abdomen dark brown both dorsally and ventrally, except the large ventral lobe on sternite 8 which is strikingly pale yellow, especially the rounded distal edge. Tergite 10, nal segments 6–9, ventral view paraprocts and general shape of subgenital plate normal (
<figureCitation id="5D8F5287BF467265FC8412F7FC6C73D4" box="[859,956,436,457]" captionStart="Fig" captionStartId="7.[317,352,1705,1726]" captionTargetBox="[319,1267,408,1673]" captionTargetId="figure-14@7.[319,1267,404,1677]" captionTargetPageId="7" captionText="Fig. 3. Isoperla altaica Šámal. a = head and prothorax, b, c = male abdominal tip in ventral and dorsal views, respectively; d, e = penis sclerite in lateral and caudal views, respectively, f = spicules on penial membrane next to large sclerite, g = male with everted penis, lateral view, h = female abdomi-" figureDoi="http://doi.org/10.5281/zenodo.12586267" httpUri="https://zenodo.org/record/12586267/files/figure.png" pageId="8" pageNumber="261">Figs 3b, c</figureCitation>
). Pilosity of dark brown cerci resembling those of
<taxonomicName id="02B43581BF467265FE361292FD9773FB" box="[489,583,465,486]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="261" phylum="Arthropoda" rank="species" species="asiatica">
<emphasis id="F7C09210BF467265FE361292FD9773FB" box="[489,583,465,486]" italics="true" pageId="8" pageNumber="261">I. asiatica</emphasis>
</taxonomicName>
; see there. Penis membraneous, armed with a large asymmetrical sclerite (
<figureCitation id="5D8F5287BF467265FE9B12ADFE4E701E" box="[324,414,494,515]" captionStart="Fig" captionStartId="7.[317,352,1705,1726]" captionTargetBox="[319,1267,408,1673]" captionTargetId="figure-14@7.[319,1267,404,1677]" captionTargetPageId="7" captionText="Fig. 3. Isoperla altaica Šámal. a = head and prothorax, b, c = male abdominal tip in ventral and dorsal views, respectively; d, e = penis sclerite in lateral and caudal views, respectively, f = spicules on penial membrane next to large sclerite, g = male with everted penis, lateral view, h = female abdomi-" figureDoi="http://doi.org/10.5281/zenodo.12586267" httpUri="https://zenodo.org/record/12586267/files/figure.png" pageId="8" pageNumber="261">Figs 3e, f</figureCitation>
) which stands on the rear face of the everted organ, its slender, slightly twisted apical section points downward (
<figureCitation id="5D8F5287BF467265FDC91148FD8C703D" box="[534,604,523,544]" captionStart="Fig" captionStartId="7.[317,352,1705,1726]" captionTargetBox="[319,1267,408,1673]" captionTargetId="figure-14@7.[319,1267,404,1677]" captionTargetPageId="7" captionText="Fig. 3. Isoperla altaica Šámal. a = head and prothorax, b, c = male abdominal tip in ventral and dorsal views, respectively; d, e = penis sclerite in lateral and caudal views, respectively, f = spicules on penial membrane next to large sclerite, g = male with everted penis, lateral view, h = female abdomi-" figureDoi="http://doi.org/10.5281/zenodo.12586267" httpUri="https://zenodo.org/record/12586267/files/figure.png" pageId="8" pageNumber="261">
Fig.
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</figureCitation>
). Minute scales (
<figureCitation id="5D8F5287BF467265FD251148FCE9703D" box="[762,825,523,544]" captionStart="Fig" captionStartId="7.[317,352,1705,1726]" captionTargetBox="[319,1267,408,1673]" captionTargetId="figure-14@7.[319,1267,404,1677]" captionTargetPageId="7" captionText="Fig. 3. Isoperla altaica Šámal. a = head and prothorax, b, c = male abdominal tip in ventral and dorsal views, respectively; d, e = penis sclerite in lateral and caudal views, respectively, f = spicules on penial membrane next to large sclerite, g = male with everted penis, lateral view, h = female abdomi-" figureDoi="http://doi.org/10.5281/zenodo.12586267" httpUri="https://zenodo.org/record/12586267/files/figure.png" pageId="8" pageNumber="261">Fig. 3f</figureCitation>
) cover most of the surrounding membrane.
</paragraph>
<caption id="91CB1E8ABF49726AFEE215EAFB2674E5" ID-DOI="http://doi.org/10.5281/zenodo.12586267" ID-Zenodo-Dep="12586267" httpUri="https://zenodo.org/record/12586267/files/figure.png" pageId="7" pageNumber="260" startId="7.[317,352,1705,1726]" targetBox="[319,1267,408,1673]" targetPageId="7" targetType="figure">
<paragraph id="C50B4E02BF49726AFEE215EAFB2674E5" blockId="7.[317,1270,1705,1813]" pageId="7" pageNumber="260">
<emphasis id="F7C09210BF49726AFEE215EAFEAB74A3" bold="true" box="[317,379,1705,1726]" pageId="7" pageNumber="260">Fig. 3.</emphasis>
<taxonomicName id="02B43581BF49726AFEA015E9FD8D74A2" authority="Samal." authorityName="Samal." box="[383,605,1706,1727]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="altaica">
<emphasis id="F7C09210BF49726AFEA015E9FDC674A2" box="[383,534,1706,1727]" italics="true" pageId="7" pageNumber="260">Isoperla altaica</emphasis>
Šámal.
</taxonomicName>
a = head and prothorax, b, c = male abdominal tip in ventral and dorsal views, respectively; d, e = penis sclerite in lateral and caudal views, respectively, f = spicules on penial membrane next to large sclerite, g = male with everted penis, lateral view, h = female abdomi-
</paragraph>
</caption>
<paragraph id="C50B4E02BF467265FEA1116BFEB270D0" blockId="8.[317,1270,436,1120]" pageId="8" pageNumber="261">
The appearance of the asymmetrical penis sclerite differs depending on slight differences in the positioning of specimens. In side view, the basal part of the sclerite is broad, more than 3 times wider than the apical part. However, when viewed from the apex of the sclerite one notices that only that side of the sclerite is enlarged which lies on top when the curved apex points to the right hand side. The upper edge of this side forms a blunt crest, the opposite side is low and rounded (
<figureCitation id="5D8F5287BF467265FB4D11D8FB2170AD" box="[1170,1265,667,688]" captionStart="Fig" captionStartId="7.[317,352,1705,1726]" captionTargetBox="[319,1267,408,1673]" captionTargetId="figure-14@7.[319,1267,404,1677]" captionTargetPageId="7" captionText="Fig. 3. Isoperla altaica Šámal. a = head and prothorax, b, c = male abdominal tip in ventral and dorsal views, respectively; d, e = penis sclerite in lateral and caudal views, respectively, f = spicules on penial membrane next to large sclerite, g = male with everted penis, lateral view, h = female abdomi-" figureDoi="http://doi.org/10.5281/zenodo.12586267" httpUri="https://zenodo.org/record/12586267/files/figure.png" pageId="8" pageNumber="261">Figs 3d, e</figureCitation>
,
<figureCitation id="5D8F5287BF467265FEE211FBFE8670D0" box="[317,342,696,717]" captionStart="Fig" captionStartId="10.[317,352,1753,1774]" captionTargetBox="[357,1232,1226,1717]" captionTargetId="figure-238@10.[347,1240,1219,1725]" captionTargetPageId="10" captionText="Fig. 6. Isoperla altaica ŠÁMAL (a) and I. eximia ZAPEKINA-DULKEIT (b, c). Penis sclerites (a, b; to same scale) and fore body (c; not to scale)" figureDoi="http://doi.org/10.5281/zenodo.12586273" httpUri="https://zenodo.org/record/12586273/files/figure.png" pageId="8" pageNumber="261">6a</figureCitation>
).
</paragraph>
<paragraph id="C50B4E02BF467265FEA11197FBF6711B" blockId="8.[317,1270,436,1120]" pageId="8" pageNumber="261">
Female (
<figureCitation id="5D8F5287BF467265FE0C1197FDCB70F4" box="[467,539,724,745]" captionStart="Fig" captionStartId="7.[317,352,1705,1726]" captionTargetBox="[319,1267,408,1673]" captionTargetId="figure-14@7.[319,1267,404,1677]" captionTargetPageId="7" captionText="Fig. 3. Isoperla altaica Šámal. a = head and prothorax, b, c = male abdominal tip in ventral and dorsal views, respectively; d, e = penis sclerite in lateral and caudal views, respectively, f = spicules on penial membrane next to large sclerite, g = male with everted penis, lateral view, h = female abdomi-" figureDoi="http://doi.org/10.5281/zenodo.12586267" httpUri="https://zenodo.org/record/12586267/files/figure.png" pageId="8" pageNumber="261">Fig. 3h</figureCitation>
). Subgenital plate short, projecting only a little, distal margin shallowly indented in the middle, shiny, dark, finely pilose, like the rest of the segment.
</paragraph>
<paragraph id="C50B4E02BF467265FEA1104DFCE37164" blockId="8.[317,1270,436,1120]" pageId="8" pageNumber="261">
Egg. Plump, regular oval with blunt poles. There is a prominent egg shoulder because a raised ring surrounds the plateau from which the collar rises (
<figureCitation id="5D8F5287BF467265FC901068FC5A715D" box="[847,906,811,832]" captionStart="Fig" captionStartId="8.[376,411,1791,1812]" captionTargetBox="[328,1260,1255,1738]" captionTargetId="figure-382@8.[319,1267,1234,1763]" captionTargetPageId="8" captionText="Fig. 4. Isoperla altaica ŠÁMAL, egg and details of collar; specimen from Khonin Nuga" figureDoi="http://doi.org/10.5281/zenodo.12586269" httpUri="https://zenodo.org/record/12586269/files/figure.png" pageId="8" pageNumber="261">Fig. 4</figureCitation>
). The collar lacks distinct meshes but has a finely crenulate margin. The base of the fungiform anchor in the deeply embedded center of the collar is straight and flat. Micropyles not observed.
</paragraph>
<paragraph id="C50B4E02BF467265FEA110C2FD877614" blockId="8.[317,1270,436,1120]" pageId="8" pageNumber="261">Eggs from Khonin Nuga measure 360×250 µm and have a thin, structureless chorion of uniform width (7–8 µm). In contrast, eggs from near Ust Kamenogorsk are smaller (295×235 µm), their chorion is finely punctured, in optical section a fine radial striation extends across the chorion which is almost 13 µm thick near the equator of the egg, but only about 9 µm near the opercular pole or next to the ring around the collar.</paragraph>
<paragraph id="C50B4E02BF467265FEA11752FD84767D" blockId="8.[317,1270,436,1120]" pageId="8" pageNumber="261">
Larva (
<figureCitation id="5D8F5287BF467265FE1C1752FDDA763B" box="[451,522,1041,1062]" captionStart="Fig" captionStartId="9.[317,352,1772,1793]" captionTargetBox="[363,1234,925,1735]" captionTargetId="figure-127@9.[350,1238,909,1744]" captionTargetPageId="9" captionText="Fig. 5. Larvae of Isoperla spp. a =I.lunigera (KLAPÁLEK), b–d = I.altaica ŠÁMAL, with details of labium and labial segment" figureDoi="http://doi.org/10.5281/zenodo.12586271" httpUri="https://zenodo.org/record/12586271/files/figure.png" pageId="8" pageNumber="261">Fig. 5b</figureCitation>
). We illustrate a pharate male in its larval skin which had the penis sclerite already developed. It differs from the larva previously assigned to
<taxonomicName id="02B43581BF467265FC17176DFEA8767D" authority="(ZAPEKINA-DULKEIT, 1955)" baseAuthorityName="ZAPEKINA-DULKEIT" baseAuthorityYear="1955" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="261" phylum="Arthropoda" rank="species" species="altaica">
<emphasis id="F7C09210BF467265FC17176DFBF2765E" box="[968,1058,1070,1091]" italics="true" pageId="8" pageNumber="261">I. altaica</emphasis>
(
<bibRefCitation id="A12533F3BF467265FBEE176DFEA1767D" author="ZAPEKINA-DULKEIT, J. I." pageId="8" pageNumber="261" pagination="167 - 177" refId="ref9819" refString="ZAPEKINA-DULKEIT, J. I. (1955) [On the Plecoptera-fauna of the Telezkoe basin]. Entomol. Obozr. 34: 167 - 177. [in Russian]" type="journal article" year="1955">
<smallCapsWord id="C3EDD8DEBF467265FBEE176DFB3F765F" baselines="1086,1086,1086,1086,1086" box="[1073,1263,1070,1091]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Zapekina-Dulkeit" pageId="8" pageNumber="261">ZAPEKINA-DULKEIT</smallCapsWord>
, 1955
</bibRefCitation>
)
</taxonomicName>
; see under
<taxonomicName id="02B43581BF467265FE371708FD9E767D" baseAuthorityName="KLAPALEK" baseAuthorityYear="1923" box="[488,590,1099,1120]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="261" phylum="Arthropoda" rank="species" species="lunigera">
<emphasis id="F7C09210BF467265FE371708FD9E767D" box="[488,590,1099,1120]" italics="true" pageId="8" pageNumber="261">I. lunigera</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="91CB1E8ABF467265FEA715BCFB6B7508" ID-DOI="http://doi.org/10.5281/zenodo.12586269" ID-Zenodo-Dep="12586269" box="[376,1211,1791,1813]" httpUri="https://zenodo.org/record/12586269/files/figure.png" pageId="8" pageNumber="261" startId="8.[376,411,1791,1812]" targetBox="[328,1260,1255,1738]" targetPageId="8" targetType="figure">
<paragraph id="C50B4E02BF467265FEA715BCFB6B7508" blockId="8.[376,1211,1791,1813]" box="[376,1211,1791,1813]" pageId="8" pageNumber="261">
<emphasis id="F7C09210BF467265FEA715BCFE677509" bold="true" box="[376,439,1791,1812]" pageId="8" pageNumber="261">Fig. 4.</emphasis>
<taxonomicName id="02B43581BF467265FE621443FD747509" authority="SAMAL" authorityName="SAMAL" box="[445,676,1792,1813]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="261" phylum="Arthropoda" rank="species" species="altaica">
<emphasis id="F7C09210BF467265FE621443FD867508" box="[445,598,1792,1813]" italics="true" pageId="8" pageNumber="261">Isoperla altaica</emphasis>
<smallCapsWord id="C3EDD8DEBF467265FD831443FD747509" baselines="1808,1807" box="[604,676,1792,1813]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Samal" pageId="8" pageNumber="261">ŠÁMAL</smallCapsWord>
</taxonomicName>
, egg and details of collar; specimen from Khonin Nuga
</paragraph>
</caption>
<paragraph id="C50B4E02BF477264FEA112FBFB567005" blockId="9.[317,1270,439,747]" pageId="9" pageNumber="262">
<emphasis id="F7C09210BF477264FEA112FBFDBB73CF" box="[382,619,440,466]" italics="true" pageId="9" pageNumber="262">Notes and affinities.</emphasis>
The penial sclerites of all specimens are similar. However, there are differences in wing length, egg size and egg chorion thickness between populations. Future studies based on more material are desirable.
</paragraph>
<paragraph id="C50B4E02BF477264FEA11162FDF570F6" blockId="9.[317,1270,439,747]" pageId="9" pageNumber="262">
<emphasis id="F7C09210BF477264FEA11162FDBD7026" box="[382,621,545,571]" italics="true" pageId="9" pageNumber="262">
<taxonomicName id="02B43581BF477264FEA11162FE2B7026" baseAuthorityName="KLAPALEK" baseAuthorityYear="1923" box="[382,507,545,571]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="species" species="lunigera">I. lunigera</taxonomicName>
,
<taxonomicName id="02B43581BF477264FDDA1162FDBD7026" box="[517,621,545,571]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="species" species="altaica">I. altaica</taxonomicName>
</emphasis>
and
<taxonomicName id="02B43581BF477264FD7B1162FCD97026" box="[676,777,545,571]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="species" species="eximia">
<emphasis id="F7C09210BF477264FD7B1162FCD97026" box="[676,777,545,571]" italics="true" pageId="9" pageNumber="262">I. eximia</emphasis>
</taxonomicName>
belong to the
<taxonomicName id="02B43581BF477264FC751162FBCB7026" authorityName="Banks" authorityYear="1906" box="[938,1051,545,571]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="species" species="sordida">
<emphasis id="F7C09210BF477264FC751162FBCB7026" box="[938,1051,545,571]" italics="true" pageId="9" pageNumber="262">I. sordida</emphasis>
</taxonomicName>
species-group distinguished by a large unpaired penis sclerite freely projecting from the membrane. At this time, the present three are the only known Palaearctic members of the group which includes a number of species in western North America (
<bibRefCitation id="A12533F3BF477264FB8C11C8FE2870D5" author="SZCZYTKO, S. W. &amp; STEWART, K. W." pageId="9" pageNumber="262" pagination="1 - 120" refId="ref9768" refString="SZCZYTKO, S. W. and STEWART, K. W. (1979) The genus Isoperla (Plecoptera) of western North America; holomorphology and systematics, and a new stonefly genus Cascadoperla. Mems Amer. Ent. Soc. 32: 1 - 120." type="journal article" year="1979">
<smallCapsWord id="C3EDD8DEBF477264FB8C11C8FB0370BE" baselines="670,670" box="[1107,1235,651,677]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Szczytko" pageId="9" pageNumber="262">SZCZYTKO</smallCapsWord>
&amp;
<smallCapsWord id="C3EDD8DEBF477264FEE211EDFE6170DB" baselines="705,705" box="[317,433,686,712]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Stewart" pageId="9" pageNumber="262">STEWART</smallCapsWord>
1979
</bibRefCitation>
). The asymmetry of the sclerite suggests that
<taxonomicName id="02B43581BF477264FBD011EDFBA870D5" box="[1039,1144,686,712]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="species" species="altaica">
<emphasis id="F7C09210BF477264FBD011EDFBA870D5" box="[1039,1144,686,712]" italics="true" pageId="9" pageNumber="262">I. altaica</emphasis>
</taxonomicName>
and
<taxonomicName id="02B43581BF477264FB6E11EDFEB970F6" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="species" species="eximia">
<emphasis id="F7C09210BF477264FB6E11EDFEB970F6" italics="true" pageId="9" pageNumber="262">I. eximia</emphasis>
</taxonomicName>
are close allies.
</paragraph>
</subSubSection>
</treatment>
</document>
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