301 lines
31 KiB
XML
301 lines
31 KiB
XML
<document id="D46C608D186C0433BA7B93461D9A5DF9" ID-DOI="10.1649/0010-065X(2001)055[0433:NSOHKC]2.0.CO;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="4900383" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622828115451" checkinUser="carolina" docAuthor="Ratcliffe, Brett C." docDate="2001" docId="4C4A87F74454E03BFE389212FE20FE7F" docLanguage="en" docName="TheColeopBull.55.4.433-443.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (4)" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Hemiphileurus kahni Dupuis and Dechambre" docType="treatment" docVersion="3" lastPageNumber="436" masterDocId="B073FF8F4455E038FF98931DFFFEFFA1" masterDocTitle="New Species Of Hemiphileurus Kolbe (Coleoptera: Scarabaeidae: Dynastinae) From Mexico, Guatemala, Colombia, And Brazil" masterLastPageNumber="443" masterPageNumber="433" pageNumber="434" updateTime="1698955217201" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="C75989DF2A987E4C282358646152CA44" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="DA4A394E23372192BDDAE8E89618D8DB">
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<mods:title id="0C327C1B13E86C45C5617064C7787FEC">New Species Of Hemiphileurus Kolbe (Coleoptera: Scarabaeidae: Dynastinae) From Mexico, Guatemala, Colombia, And Brazil</mods:title>
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</mods:titleInfo>
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<mods:name id="F7B5EADD304CFD3AAA5C264E9148EDB6" type="personal">
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<mods:role id="3CE46AE9A9EB66F27CF1B75F9DBFFB1C">
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<mods:roleTerm id="C111893DC2A3BF9F6E491B94B1C9EA5B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4D298DA87A491F9F8E1DE19A1BC1EA9B">Ratcliffe, Brett C.</mods:namePart>
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<mods:affiliation id="DA76BB277546203A0BCD72D9E61E7D47">Systematics Research Collections W 436 Nebraska Hall University of Nebraska Lincoln, NE 68588 0514, U. S. A.</mods:affiliation>
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<mods:nameIdentifier id="4CC910CFDB5114C06090557224B6C663" type="email">bratcliffe1@unl.edu</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="705244B4F6A9285813B7FBDF10A70551">text</mods:typeOfResource>
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<mods:relatedItem id="D0BBA0904D55C3043D2E440C6C12EFB3" type="host">
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<mods:titleInfo id="6646DE3AC75798BF85A06212D4256C62">
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<mods:title id="24FA201C35E08DB726DF94F23D3638A5">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="3133A83C38EA2B045A96ADF1D1D85626">
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<mods:date id="DC0BFE8D11C0B75EB84A2585D339BC45">2001</mods:date>
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<mods:detail id="917DCD8C6A6527B33F3232D8E62DABCE" type="pubDate">
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<mods:number id="9139DA777AE0039F2B841D656DBB3032">2001-12-01</mods:number>
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</mods:detail>
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<mods:detail id="A28124883ED70A30421ECA353104CDCA" type="volume">
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<mods:number id="F28B65169EFEDB68F0BB58B08FDC18BE">55</mods:number>
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</mods:detail>
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<mods:detail id="F441E07D8B562B6A7C1B7D1BF684F493" type="issue">
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<mods:number id="6760B629042B0C5B78CC292816358892">4</mods:number>
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</mods:detail>
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<mods:extent id="A43FA433F951AC41F672FD3AA58B896C" unit="page">
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<mods:start id="E8CE8166B8247C7F18CFEF3F10F454DB">433</mods:start>
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<mods:end id="AA9286965A2D18BB54D98E3F90AE802F">443</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="E0ABDE9F47826B382B9C4C098F03E6F4">journal article</mods:classification>
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<mods:identifier id="D02CDA8669417D0007D0409426C7FB23" type="DOI">10.1649/0010-065X(2001)055[0433:NSOHKC]2.0.CO;2</mods:identifier>
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<mods:identifier id="244F957E19B91877D63DED09C745149A" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="0352CACE3920E50D0C729E83B824C1C4" type="Zenodo-Dep">4900383</mods:identifier>
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</mods:mods>
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<treatment id="4C4A87F74454E03BFE389212FE20FE7F" LSID="urn:lsid:plazi:treatment:4C4A87F74454E03BFE389212FE20FE7F" httpUri="http://treatment.plazi.org/id/4C4A87F74454E03BFE389212FE20FE7F" lastPageId="3" lastPageNumber="436" pageId="1" pageNumber="434">
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<subSubSection id="8CF9656A4454E039FE389212FC7CFE85" box="[416,898,271,292]" pageId="1" pageNumber="434" type="nomenclature">
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<paragraph id="C45C36E14454E039FE389212FC7CFE85" blockId="1.[416,898,271,319]" box="[416,898,271,292]" pageId="1" pageNumber="434">
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<heading id="9F14818D4454E039FE389212FC7CFE85" box="[416,898,271,292]" centered="true" fontSize="9" level="2" pageId="1" pageNumber="434" reason="2">
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<taxonomicName id="03E34D624454E039FE389212FC7CFE85" authority="Dupuis and Dechambre" authorityName="Dupuis and Dechambre" box="[416,898,271,292]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="434" phylum="Arthropoda" rank="species" species="kahni">
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<emphasis id="F697EAF34454E039FE389212FD80FE85" box="[416,638,271,292]" italics="true" pageId="1" pageNumber="434">Hemiphileurus kahni</emphasis>
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Dupuis and Dechambre
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="8CF9656A4454E039FDC39237FD2FFE9E" box="[603,721,298,319]" pageId="1" pageNumber="434" type="description">
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<paragraph id="C45C36E14454E039FDC39237FD2FFE9E" blockId="1.[416,898,271,319]" box="[603,721,298,319]" pageId="1" pageNumber="434">
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(
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<figureCitation id="5CD82A644454E039FDFB9237FD54FE9E" box="[611,682,298,319]" captionStart="Figs" captionStartId="2.[272,315,1403,1422]" captionTargetBox="[249,1081,265,1382]" captionTargetId="figure-11@2.[249,1081,265,1382]" captionTargetPageId="2" captionText="Figs. 1–2. Parameres of H. kahni Dupuis and Dechambre." figureDoi="http://doi.org/10.5281/zenodo.4900385" httpUri="https://zenodo.org/record/4900385/files/figure.png" pageId="1" pageNumber="434">Figs. 1</figureCitation>
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–3)
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</paragraph>
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</subSubSection>
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<subSubSection id="8CF9656A4454E039FF6E9244FBC9FE05" pageId="1" pageNumber="434" type="reference_group">
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<paragraph id="C45C36E14454E039FF6E9244FCC0FECF" blockId="1.[246,1079,345,797]" box="[246,830,345,366]" pageId="1" pageNumber="434">
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<taxonomicName id="03E34D624454E039FF6E9244FCC4FECF" authority="Dupuis and Dechambre 2000: 26" authorityName="Dupuis and Dechambre" authorityPageNumber="26" authorityYear="2000" box="[246,826,345,366]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="434" phylum="Arthropoda" rank="species" species="kahni">
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<emphasis id="F697EAF34454E039FF6E9244FE2AFECF" box="[246,468,345,366]" italics="true" pageId="1" pageNumber="434">Hemiphileurus kahni</emphasis>
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Dupuis and Dechambre 2000:26
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="C45C36E14454E039FE889269FBC9FE05" blockId="1.[246,1079,345,797]" pageId="1" pageNumber="434">
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This species was described from a single male specimen from Genaro Herrera,
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<collectingRegion id="0627F8034454E039FEB69292FE88FE05" box="[302,374,399,420]" country="Peru" name="Loreto" pageId="1" pageNumber="434">Loreto</collectingRegion>
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,
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<collectingCountry id="BCF476714454E039FE1C9292FE49FE05" box="[388,439,399,420]" name="Peru" pageId="1" pageNumber="434">Peru</collectingCountry>
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. The specimens listed below represent a NEW COUNTRY
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</paragraph>
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</subSubSection>
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<subSubSection id="8CF9656A4454E039FF6E92B4FE96FE1F" box="[246,360,425,446]" pageId="1" pageNumber="434" type="description">
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<paragraph id="C45C36E14454E039FF6E92B4FE96FE1F" blockId="1.[246,1079,345,797]" box="[246,360,425,446]" pageId="1" pageNumber="434">RECORD:</paragraph>
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</subSubSection>
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<subSubSection id="8CF9656A4454E039FE8892D9FE14FCBC" pageId="1" pageNumber="434" type="materials_examined">
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<paragraph id="C45C36E14454E039FE8892D9FE37FD8B" blockId="1.[246,1079,345,797]" pageId="1" pageNumber="434">
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<materialsCitation id="748B3CBC4454E039FE8892D9FE3BFD8B" collectingDate="1976-09-02" collectingDateMax="1976-11-10" collectingDateMin="1976-09-02" collectionCode="Record" collectorName="J. Adis & B. C. Ratcliffe" country="Brazil" county="Instituto Nacional de Pesquisas da Amazonia" location="Amazonas" municipality="Rio Taruma Mirim - Igapo" pageId="1" pageNumber="434" specimenCount="3">
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<collectingCountry id="BCF476714454E039FE8892D9FE8DFE78" box="[272,371,452,473]" name="Brazil" pageId="1" pageNumber="434">BRAZIL</collectingCountry>
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:
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<location id="C13C603A4454E039FE1B92D9FE0BFE78" LSID="urn:lsid:plazi:treatment:4C4A87F74454E03BFE389212FE20FE7F:C13C603A4454E039FE1B92D9FE0BFE78" box="[387,501,452,473]" country="Brazil" county="Instituto Nacional de Pesquisas da Amazonia" municipality="Rio Taruma Mirim - Igapo" name="Amazonas" pageId="1" pageNumber="434">Amazonas</location>
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;
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<collectingMunicipality id="2438AC9B4454E039FD9F92D9FCCDFE78" box="[519,819,452,473]" pageId="1" pageNumber="434">Rio Taruma Mirim—Igapo</collectingMunicipality>
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;
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<collectorName id="691653374454E039FCDD92D9FC69FE78" box="[837,919,452,473]" pageId="1" pageNumber="434">J. Adis</collectorName>
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,
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<date id="B05D10214454E039FC3192D9FBEDFE78" box="[937,1043,452,473]" pageId="1" pageNumber="434" value="1976-09-02">
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<collectingDate id="A019E9C94454E039FC3192D9FBEDFE78" box="[937,1043,452,473]" pageId="1" pageNumber="434" value="1976-09-02">2.IX.1976</collectingDate>
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</date>
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(
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<specimenCount id="D2E5FD684454E039FBB092D9FE9BFE55" pageId="1" pageNumber="434" type="generic">2 specimens</specimenCount>
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) and
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<date id="B05D10214454E039FE3B92C2FDE4FE55" box="[419,538,479,500]" pageId="1" pageNumber="434" value="1976-11-10">
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<collectingDate id="A019E9C94454E039FE3B92C2FDE4FE55" box="[419,538,479,500]" pageId="1" pageNumber="434" value="1976-11-10">10.XI.1976</collectingDate>
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</date>
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(
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<specimenCount id="D2E5FD684454E039FDB592C2FD57FE55" box="[557,681,479,500]" pageId="1" pageNumber="434" type="generic">1 specimen</specimenCount>
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). Specimens are deposited at the
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<collectingCounty id="2D3D4E6D4454E039FB9592C2FD40FDAE" pageId="1" pageNumber="434">Instituto Naçional de Pesquisas da Amazonia</collectingCounty>
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(Manaus, Brazil) and in the
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<collectorName id="691653374454E039FC6492E7FEAFFD8B" pageId="1" pageNumber="434">B. C. Ratcliffe</collectorName>
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collection
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="C45C36E14454E039FE88912DFC75FD6D" blockId="1.[246,1079,345,797]" pageId="1" pageNumber="434">
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The Peruvian specimen was collected to the SSW of Iquitos where the El Capite river empties into the Ucayali which, in turn, almost immediately empties into the Amazon. The Brazilian specimens were taken near Manaus just above where the Rio Negro joins the Amazon. These two localites are approximately
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<quantity id="031B9B044454E039FEE59181FE1CFD10" box="[381,482,668,689]" metricMagnitude="6" metricUnit="m" metricValue="1.54" pageId="1" pageNumber="434" unit="km" value="1540.0">1,540 km</quantity>
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apart and are both characterized by lowland rainforest. The
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<collectingCountry id="BCF476714454E039FEB091AAFE96FD6D" box="[296,360,695,716]" name="Brazil" pageId="1" pageNumber="434">Brazil</collectingCountry>
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specimens represent a substantial range extension.
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</paragraph>
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<paragraph id="C45C36E14454E039FE8891CFFE14FCBC" blockId="1.[246,1079,345,797]" pageId="1" pageNumber="434">
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Inasmuch as the description by Dupuis and Dechambre did not include a female, is based on only
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<specimenCount id="D2E5FD684454E039FD9C91F0FD66FCA3" box="[516,664,749,770]" pageId="1" pageNumber="434" type="generic">one specimen</specimenCount>
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, and is overly brief, I provide below a detailed redescription.
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</paragraph>
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</subSubSection>
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<subSubSection id="8CF9656A4454E039FE959028FCD3FA14" pageId="1" pageNumber="434" type="description">
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<paragraph id="C45C36E14454E039FE959028FCC4FA91" blockId="1.[246,1080,821,1329]" pageId="1" pageNumber="434">
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<emphasis id="F697EAF34454E039FE959028FEC1FCE9" bold="true" box="[269,319,821,840]" pageId="1" pageNumber="434">Male</emphasis>
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. Length
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<quantity id="031B9B044454E039FE05902BFE06FCE8" box="[413,504,822,841]" metricMagnitude="-2" metricUnit="m" metricValue="2.2399999999999998" pageId="1" pageNumber="434" unit="mm" value="22.4">22.4 mm</quantity>
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; width
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<quantity id="031B9B044454E039FDDF902BFD5CFCE8" box="[583,674,822,841]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="1" pageNumber="434" unit="mm" value="10.7">10.7 mm</quantity>
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. Color black.
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<emphasis id="F697EAF34454E039FCAC902BFC98FCE8" box="[820,870,822,841]" italics="true" pageId="1" pageNumber="434">Head</emphasis>
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: Surface completely, transversely rugose. Frons with large, laterally compressed tubercle mesad of anterior border of each eye; surface between tubercles foveate, moderately rugose. Clypeus triangular, apex acute and strongly reflexed, a distinct carina extending from apex to base of each tubercle. Interocular width equals 4.5 transverse eye diameters. Antenna with 10 segments, club slightly longer than segments 2–7. Mandibles arcuate on lateral edge, apices acute.
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<emphasis id="F697EAF34454E039FEEC90DAFE2FFC7B" box="[372,465,967,986]" italics="true" pageId="1" pageNumber="434">Pronotum</emphasis>
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: Surface finely shagreened, moderately punctate; punctures moderately large, each with a minute seta, punctures becoming slightly denser on sides. Median, longitudinal furrow shallow (nearly obsolete), becoming wider apically, not reaching base or apex. Base with complete marginal bead.
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<emphasis id="F697EAF34454E039FC81970DFCADFB82" box="[793,851,1040,1059]" italics="true" pageId="1" pageNumber="434">Elytra</emphasis>
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: Surface with 5 distinct, punctate striae between suture and humeral umbone; punctures large, ocellate, separated from one another in each row by 1–2 puncture diameters. Intervals convex, with sparse micropunctures.
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<taxonomicName id="03E34D624454E039FE0C9744FDBCFBCD" authority=": Surface" authorityName="Surface" box="[404,578,1113,1132]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="434" phylum="Chordata" rank="genus">
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<emphasis id="F697EAF34454E039FE0C9744FE13FBCD" box="[404,493,1113,1132]" italics="true" pageId="1" pageNumber="434">Pygidium</emphasis>
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: Surface
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</taxonomicName>
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completely, densely punctate; punctures large, deep, setigerous; setae tawny in color, short, a few longer at center apex. In lateral view, surface strongly convex.
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<emphasis id="F697EAF34454E039FE0F9797FE3DFB3C" box="[407,451,1162,1181]" italics="true" pageId="1" pageNumber="434">Legs</emphasis>
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: Foretibia tridentate, teeth subequally separated. Median transverse carina on meso and metatibiae ending in a long, acute spine on upper surface. Apex of posterior tibia with strong, acute spine on upper angle and with 8 short, broad spinules below spine. Apex of first tarsomere on posterior tarsus with strong, acute spine dorsally.
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<emphasis id="F697EAF34454E039FF6E97F1FECAFB5E" box="[246,308,1260,1279]" italics="true" pageId="1" pageNumber="434">Venter</emphasis>
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: Prosternal process broad, moderate in length, apex flattened into a concave, suboval plate (Fig. 3). Last sternite with small, dense punctures in lateral angles, elsewhere with small, sparse punctures.
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<emphasis id="F697EAF34454E039FDC89603FD48FA90" box="[592,694,1310,1329]" italics="true" pageId="1" pageNumber="434">Parameres</emphasis>
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:
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<figureCitation id="5CD82A644454E039FD5C9600FCC8FA91" box="[708,822,1309,1328]" captionStart="Figs" captionStartId="2.[272,315,1403,1422]" captionTargetBox="[249,1081,265,1382]" captionTargetId="figure-11@2.[249,1081,265,1382]" captionTargetPageId="2" captionText="Figs. 1–2. Parameres of H. kahni Dupuis and Dechambre." figureDoi="http://doi.org/10.5281/zenodo.4900385" httpUri="https://zenodo.org/record/4900385/files/figure.png" pageId="1" pageNumber="434">Figures 1–2</figureCitation>
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.
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</paragraph>
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<paragraph id="C45C36E14454E039FE889653FCD3FA14" blockId="1.[246,1079,1358,1652]" pageId="1" pageNumber="434">
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<emphasis id="F697EAF34454E039FE889653FE96FAC2" bold="true" box="[272,360,1358,1379]" pageId="1" pageNumber="434">Female.</emphasis>
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Length 23.0–
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<quantity id="031B9B044454E039FD9C9652FD97FAC5" box="[516,617,1359,1380]" metricMagnitude="-2" metricUnit="m" metricValue="2.34" pageId="1" pageNumber="434" unit="mm" value="23.4">23.4 mm</quantity>
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; width
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<quantity id="031B9B044454E039FD269652FCA0FAC5" box="[702,862,1359,1380]" metricMagnitude="-2" metricUnit="m" metricValue="1.125" metricValueMax="1.13" metricValueMin="1.1199999999999999" pageId="1" pageNumber="434" unit="mm" value="11.25" valueMax="11.3" valueMin="11.2">11.2–11.3 mm</quantity>
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. As male except in the following respects:
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<emphasis id="F697EAF34454E039FE6F9677FD9EFADE" box="[503,608,1386,1407]" italics="true" pageId="1" pageNumber="434">Pronotum</emphasis>
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: Punctures slightly larger.
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<emphasis id="F697EAF34454E039FC1F9677FC36FADE" box="[903,968,1386,1407]" italics="true" pageId="1" pageNumber="434">Elytra</emphasis>
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: Intervals slightly wider.
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<taxonomicName id="03E34D624454E039FE0D9698FDBFFA3B" authority=": Setae" authorityName="Setae" box="[405,577,1413,1434]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="434" phylum="Chordata" rank="genus">
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<emphasis id="F697EAF34454E039FE0D9698FE07FA3B" box="[405,505,1413,1434]" italics="true" pageId="1" pageNumber="434">Pygidium</emphasis>
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: Setae
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</taxonomicName>
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shorter.
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<emphasis id="F697EAF34454E039FD069698FD1AFA3B" box="[670,740,1413,1434]" italics="true" pageId="1" pageNumber="434">Venter</emphasis>
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: Last sternite completely, moderately densely punctate, punctures moderate in size.
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</paragraph>
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</subSubSection>
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<subSubSection id="8CF9656A4454E03BFE8896A6FE20FE7F" lastPageId="3" lastPageNumber="436" pageId="1" pageNumber="434" type="discussion">
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<paragraph id="C45C36E14454E039FE8896A6FBC8F9D5" blockId="1.[246,1079,1358,1652]" pageId="1" pageNumber="434">
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<emphasis id="F697EAF34454E039FE8896A6FE83FA71" bold="true" box="[272,381,1467,1488]" pageId="1" pageNumber="434">Remarks.</emphasis>
|
||
<taxonomicName id="03E34D624454E039FE1196A1FD92FA70" box="[393,620,1468,1489]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="434" phylum="Arthropoda" rank="species" species="kahni">
|
||
<emphasis id="F697EAF34454E039FE1196A1FD92FA70" box="[393,620,1468,1489]" italics="true" pageId="1" pageNumber="434">Hemiphileurus kahni</emphasis>
|
||
</taxonomicName>
|
||
will key only to couplet
|
||
<quantity id="031B9B044454E039FC0196A1FC2AFA70" box="[921,980,1468,1489]" metricMagnitude="0" metricUnit="m" metricValue="1.0413999999999999" pageId="1" pageNumber="434" unit="in" value="41.0">41 in</quantity>
|
||
Endrödi (1985) where the following two choices are for Mesoamerican species [
|
||
<taxonomicName id="03E34D624454E039FB8596CAFE42F9A6" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1888" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="434" phylum="Arthropoda" rank="species" species="laevicauda">
|
||
<emphasis id="F697EAF34454E039FB8596CAFE97F9A6" italics="true" pageId="1" pageNumber="434">H. laevicauda</emphasis>
|
||
(Bates)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="03E34D624454E039FE6B96EFFCE8F9A6" authority="(Bates)" baseAuthorityName="Bates" box="[499,790,1522,1543]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="434" phylum="Arthropoda" rank="species" species="punctatostriatus">
|
||
<emphasis id="F697EAF34454E039FE6B96EFFD41F9A6" box="[499,703,1522,1543]" italics="true" pageId="1" pageNumber="434">H. punctatostriatus</emphasis>
|
||
(Bates)
|
||
</taxonomicName>
|
||
] that each have totally different parameres than
|
||
<emphasis id="F697EAF34454E039FE759510FDADF983" box="[493,595,1549,1570]" italics="true" pageId="1" pageNumber="434">
|
||
<taxonomicName id="03E34D624454E039FE759510FDAEF983" box="[493,592,1549,1570]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="434" phylum="Arthropoda" rank="species" species="kahni">H. kahni</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The parameres (
|
||
<figureCitation id="5CD82A644454E039FC889510FC89F983" box="[784,887,1549,1570]" captionStart="Figs" captionStartId="2.[272,315,1403,1422]" captionTargetBox="[249,1081,265,1382]" captionTargetId="figure-11@2.[249,1081,265,1382]" captionTargetPageId="2" captionText="Figs. 1–2. Parameres of H. kahni Dupuis and Dechambre." figureDoi="http://doi.org/10.5281/zenodo.4900385" httpUri="https://zenodo.org/record/4900385/files/figure.png" pageId="1" pageNumber="434">Figs. 1–2</figureCitation>
|
||
) of
|
||
<taxonomicName id="03E34D624454E039FC349510FBF5F983" box="[940,1035,1549,1570]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="434" phylum="Arthropoda" rank="species" species="kahni">
|
||
<emphasis id="F697EAF34454E039FC349510FBF5F983" box="[940,1035,1549,1570]" italics="true" pageId="1" pageNumber="434">H. kahni</emphasis>
|
||
</taxonomicName>
|
||
are unique among species of
|
||
<taxonomicName id="03E34D624454E039FD8C9534FD51F99F" authorityName="Kolbe" authorityYear="1910" box="[532,687,1577,1598]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="434" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F697EAF34454E039FD8C9534FD51F99F" box="[532,687,1577,1598]" italics="true" pageId="1" pageNumber="434">Hemiphileurus</emphasis>
|
||
</taxonomicName>
|
||
and are diagnostic for the species. The distinct microsetae in the pronotal punctures are also unusual for a species of
|
||
<emphasis id="F697EAF34454E039FE8E9542FE46F9D5" box="[278,440,1631,1652]" italics="true" pageId="1" pageNumber="434">
|
||
<taxonomicName id="03E34D624454E039FE8E9542FE4AF9D5" authorityName="Kolbe" authorityYear="1910" box="[278,436,1631,1652]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="434" phylum="Arthropoda" rank="genus">Hemiphileurus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
but this may simply reflect a pristine specimen where the
|
||
</paragraph>
|
||
<caption id="909C66694457E03AFE889666FCB4FA2F" ID-DOI="http://doi.org/10.5281/zenodo.4900385" ID-Zenodo-Dep="4900385" box="[272,842,1403,1422]" httpUri="https://zenodo.org/record/4900385/files/figure.png" pageId="2" pageNumber="435" startId="2.[272,315,1403,1422]" targetBox="[249,1081,265,1382]" targetPageId="2">
|
||
<paragraph id="C45C36E14457E03AFE889666FCB4FA2F" blockId="2.[272,1082,1403,1614]" box="[272,842,1403,1422]" pageId="2" pageNumber="435">
|
||
<emphasis id="F697EAF34457E03AFE889666FE8EFA2F" bold="true" box="[272,368,1403,1422]" pageId="2" pageNumber="435">Figs. 1–2.</emphasis>
|
||
Parameres of
|
||
<taxonomicName id="03E34D624457E03AFD929666FCB4FA2F" authority="Dupuis and Dechambre." authorityName="Dupuis and Dechambre." box="[522,842,1403,1422]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="species" species="kahni">
|
||
<emphasis id="F697EAF34457E03AFD929666FDA2FA2F" box="[522,604,1403,1422]" italics="true" pageId="2" pageNumber="435">H. kahni</emphasis>
|
||
Dupuis and Dechambre.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C45C36E14457E03AFE88968EFC8BFA07" blockId="2.[272,1082,1403,1614]" box="[272,885,1427,1446]" pageId="2" pageNumber="435">
|
||
<emphasis id="F697EAF34457E03AFE88968EFEAEFA07" bold="true" box="[272,336,1427,1446]" pageId="2" pageNumber="435">Fig. 3.</emphasis>
|
||
Prosternal process of
|
||
<taxonomicName id="03E34D624457E03AFDAD968EFC8BFA07" authority="Dupuis and Dechambre." authorityName="Dupuis and Dechambre." box="[565,885,1427,1446]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="species" species="kahni">
|
||
<emphasis id="F697EAF34457E03AFDAD968EFD79FA07" box="[565,647,1427,1446]" italics="true" pageId="2" pageNumber="435">H. kahni</emphasis>
|
||
Dupuis and Dechambre.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="C45C36E14457E03AFE8896B6FC67FA1F" blockId="2.[272,1082,1403,1614]" box="[272,921,1451,1470]" pageId="2" pageNumber="435">
|
||
<emphasis id="F697EAF34457E03AFE8896B6FE91FA1F" bold="true" box="[272,367,1451,1470]" pageId="2" pageNumber="435">Figs. 4–5.</emphasis>
|
||
Parameres of
|
||
<typeStatus id="1B5888434457E03AFD9296B6FDA5FA1F" box="[522,603,1451,1470]" pageId="2" pageNumber="435" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="03E34D624457E03AFDE696B6FCA4FA1F" authority="Ratcliffe" authorityName="Ratcliffe" authorityYear="2001" box="[638,858,1451,1470]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="species" species="bispinosus" status="sp. nov.">
|
||
<emphasis id="F697EAF34457E03AFDE696B6FD01FA1F" box="[638,767,1451,1470]" italics="true" pageId="2" pageNumber="435">H. bispinosus</emphasis>
|
||
Ratcliffe
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="EDA457884457E03AFCFD96B6FC67FA1F" box="[869,921,1451,1470]" pageId="2" pageNumber="435" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="C45C36E14457E03AFE8896DEFBEFFA77" blockId="2.[272,1082,1403,1614]" box="[272,1041,1475,1494]" pageId="2" pageNumber="435">
|
||
<emphasis id="F697EAF34457E03AFE8896DEFEAEFA77" bold="true" box="[272,336,1475,1494]" pageId="2" pageNumber="435">Fig. 6.</emphasis>
|
||
Prosternal process, lateral oblique view, of
|
||
<taxonomicName id="03E34D624457E03AFC9B96DEFC2CFA77" authority="Ratcliffe" authorityName="Ratcliffe" authorityYear="2001" box="[771,978,1475,1494]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="species" species="deslislesi" status="sp. nov.">
|
||
<emphasis id="F697EAF34457E03AFC9B96DEFC89FA77" box="[771,887,1475,1494]" italics="true" pageId="2" pageNumber="435">H. deslislesi</emphasis>
|
||
Ratcliffe
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="EDA457884457E03AFC4596DEFBEFFA77" box="[989,1041,1475,1494]" pageId="2" pageNumber="435" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="C45C36E14457E03AFE8896C6FC72FA4F" blockId="2.[272,1082,1403,1614]" box="[272,908,1499,1518]" pageId="2" pageNumber="435">
|
||
<emphasis id="F697EAF34457E03AFE8896C6FE91FA4F" bold="true" box="[272,367,1499,1518]" pageId="2" pageNumber="435">Figs. 7–8.</emphasis>
|
||
Parameres of
|
||
<typeStatus id="1B5888434457E03AFD9296C6FDA5FA4F" box="[522,603,1499,1518]" pageId="2" pageNumber="435" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="03E34D624457E03AFDE696C6FCB3FA4F" authority="Ratcliffe" authorityName="Ratcliffe" authorityYear="2001" box="[638,845,1499,1518]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="species" species="deslislesi" status="sp. nov.">
|
||
<emphasis id="F697EAF34457E03AFDE696C6FD0CFA4F" box="[638,754,1499,1518]" italics="true" pageId="2" pageNumber="435">H. deslislesi</emphasis>
|
||
Ratcliffe
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="EDA457884457E03AFCC096C6FC72FA4F" box="[856,908,1499,1518]" pageId="2" pageNumber="435" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="C45C36E14457E03AFE8896EEFD10F9A7" blockId="2.[272,1082,1403,1614]" box="[272,750,1523,1542]" pageId="2" pageNumber="435">
|
||
<emphasis id="F697EAF34457E03AFE8896EEFEAEF9A7" bold="true" box="[272,336,1523,1542]" pageId="2" pageNumber="435">Fig. 9.</emphasis>
|
||
Parameres of
|
||
<taxonomicName id="03E34D624457E03AFE7296EEFD14F9A7" authority="(Burmeister)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[490,746,1523,1542]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="species" species="variolosus">
|
||
<emphasis id="F697EAF34457E03AFE7296EEFD97F9A7" box="[490,617,1523,1542]" italics="true" pageId="2" pageNumber="435">H. variolosus</emphasis>
|
||
(Burmeister)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C45C36E14457E03AFE889516FBEDF9BF" blockId="2.[272,1082,1403,1614]" box="[272,1043,1547,1566]" pageId="2" pageNumber="435">
|
||
<emphasis id="F697EAF34457E03AFE889516FEA2F9BF" bold="true" box="[272,348,1547,1566]" pageId="2" pageNumber="435">Fig. 10.</emphasis>
|
||
Prosternal process, lateral oblique view, of
|
||
<taxonomicName id="03E34D624457E03AFC979516FBF1F9BF" authority="(Burmeister)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[783,1039,1547,1566]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="species" species="variolosus">
|
||
<emphasis id="F697EAF34457E03AFC979516FC70F9BF" box="[783,910,1547,1566]" italics="true" pageId="2" pageNumber="435">H. variolosus</emphasis>
|
||
(Burmeister)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C45C36E14457E03AFE88953EFE9DF9EF" blockId="2.[272,1082,1403,1614]" pageId="2" pageNumber="435">
|
||
<emphasis id="F697EAF34457E03AFE88953EFE78F997" bold="true" box="[272,390,1571,1590]" pageId="2" pageNumber="435">Figs. 11–13.</emphasis>
|
||
Parameres of
|
||
<taxonomicName id="03E34D624457E03AFDB8953EFD0CF997" authority="Endrodi." authorityName="Endrodi." box="[544,754,1571,1590]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="species" species="rugulosus">
|
||
<emphasis id="F697EAF34457E03AFDB8953EFD64F997" box="[544,666,1571,1590]" italics="true" pageId="2" pageNumber="435">H. rugulosus</emphasis>
|
||
Endrödi.
|
||
</taxonomicName>
|
||
<emphasis id="F697EAF34457E03AFD61953EFCC5F997" bold="true" box="[761,827,1571,1590]" pageId="2" pageNumber="435">11–12)</emphasis>
|
||
new drawing;
|
||
<emphasis id="F697EAF34457E03AFC51953EFC16F997" bold="true" box="[969,1000,1571,1590]" pageId="2" pageNumber="435">13)</emphasis>
|
||
Endrödi drawing.
|
||
</paragraph>
|
||
<paragraph id="C45C36E14456E03BFF6E9212FE57FE9F" blockId="3.[246,1078,271,478]" pageId="3" pageNumber="436">tiny setae have not been worn away; these setae are less noticeable in the two other specimens.</paragraph>
|
||
<paragraph id="C45C36E14456E03BFE889259FE20FE7F" blockId="3.[246,1078,271,478]" pageId="3" pageNumber="436">
|
||
The biology of this species is unknown. The specimens reported here were collected in arboreal photoeclectors mounted on tree trunks in an area of blackwater inundation forest (igapo) near where the Tarumã Mirim river empties into the Rio Negro about
|
||
<quantity id="031B9B044456E03BFD839289FD9DFE08" box="[539,611,404,425]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="436" unit="km" value="20.0">20 km</quantity>
|
||
upstream from Manaus (
|
||
<bibRefCitation id="A0724B104456E03BFCE79289FC08FE08" author="Adis" box="[895,1014,404,425]" pageId="3" pageNumber="436" pagination="87 - 173" refId="ref5439" refString="Adis, J. 1981. Comparative ecological studies of the terrestrial arthropod fauna in Central Amazonian inundation-forests. Amazoniana 7: 87 - 173." type="journal article" year="1981">Adis 1981</bibRefCitation>
|
||
). The photoeclector traps were modified to capture insects walking downward on the trunk of the trap tree.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |