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<document id="4A0868293B67AB0C31797E67B380A563" ID-DOI="10.11646/zootaxa.5492.4.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="13268886" ID-ZooBank="9EAFBF80-9862-480A-8CAA-9DB544AE614E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1723107730108" checkinUser="plazi" docAuthor="Sirenko, Boris I." docDate="2024" docId="4B6287D5B203FF8804DEF29CFE0FFEC6" docLanguage="en" docName="zootaxa.5492.4.2.pdf" docOrigin="Zootaxa 5492 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5492.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kaasichiton Sirenko, 2024, new genus" docType="treatment" docVersion="2" lastPageNumber="509" masterDocId="B75BFFADB202FF8C0449F04BFFE0FFDB" masterDocTitle="A new genus and two new species of brooding chitons (Mollusca: Polyplacophora) from the North-Western Pacific" masterLastPageNumber="529" masterPageNumber="505" pageNumber="506" updateTime="1723108186403" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="493E06B98D169998DCA87C8DF57255E7">A new genus and two new species of brooding chitons (Mollusca: Polyplacophora) from the North-Western Pacific</mods:title>
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<mods:date id="05494B87218329A54422702605714B5F">2024</mods:date>
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<paragraph id="C37436C3B203FF8D04DEF29CFDE9FD2A" blockId="1.[151,521,727,754]" box="[151,521,727,754]" pageId="1" pageNumber="506">
<heading id="983C81AFB203FF8D04DEF29CFDE9FD2A" bold="true" box="[151,521,727,754]" fontSize="11" level="1" pageId="1" pageNumber="506" reason="1">
<emphasis id="F1BFEAD1B203FF8D04DEF29CFDE9FD2A" bold="true" box="[151,521,727,754]" pageId="1" pageNumber="506">
Genus
<taxonomicName id="04CB4D40B203FF8D04A7F29CFE9FFD29" authorityName="Sirenko" authorityYear="2024" box="[238,383,727,754]" class="Polyplacophora" family="Leptochitonidae" genus="Kaasichiton" kingdom="Animalia" order="Lepidopleurida" pageId="1" pageNumber="506" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="F1BFEAD1B203FF8D04A7F29CFE9FFD29" bold="true" box="[238,383,727,754]" italics="true" pageId="1" pageNumber="506">Kaasichiton</emphasis>
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<taxonomicNameLabel id="EA8C57AAB203FF8D05CEF29CFDE9FD2A" box="[391,521,727,753]" pageId="1" pageNumber="506" rank="genus">new genus</taxonomicNameLabel>
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<emphasis id="F1BFEAD1B203FF8D04DEF354FED2FCE2" bold="true" box="[151,306,799,825]" pageId="1" pageNumber="506">
<typeStatus id="1C708861B203FF8D04DEF354FF32FCE2" box="[151,210,799,825]" pageId="1" pageNumber="506">Type</typeStatus>
species.
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<taxonomicName id="04CB4D40B203FF8D0570F354FD4DFCE2" ID-CoL="3TLYB" authority="Sirenko, 1990" authorityName="Sirenko" authorityYear="1990" box="[313,685,799,825]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="1" pageNumber="506" phylum="Mollusca" rank="species" species="lukini">
<emphasis id="F1BFEAD1B203FF8D0570F354FDE6FCE2" box="[313,518,799,825]" italics="true" pageId="1" pageNumber="506">Leptochiton lukini</emphasis>
<bibRefCitation id="A75A4B32B203FF8D0644F354FD4DFCE2" author="Sirenko, B. I." box="[525,685,799,825]" pageId="1" pageNumber="506" pagination="96 - 103" refId="ref8206" refString="Sirenko, B. I. (1990) New species of chitons of the genus Leptochiton (Mollusca, Polyplacophora) of the shelf and the slope of the Kurile Islands. Proceedings of the Zoological Institute RAS, 218, 96 - 103. [in Russian]" type="journal article" year="1990">Sirenko, 1990</bibRefCitation>
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.
</paragraph>
<paragraph id="C37436C3B203FF8D048EF308FF54FC59" blockId="1.[151,1437,799,1438]" pageId="1" pageNumber="506">
<emphasis id="F1BFEAD1B203FF8D048EF308FE4FFC86" bold="true" box="[199,431,835,861]" pageId="1" pageNumber="506">Genus distribution.</emphasis>
<materialsCitation id="73A33C9EB203FF8D05FFF308FF50FC59" county="Kurile Islands" elevation="887" location="Bering Sea" municipality="Sea of Okhotsk" pageId="1" pageNumber="506" specimenCount="1" stateProvince="Kamchatka">
<collectingCounty id="2A154E4FB203FF8D05FFF308FDB6FC86" box="[438,598,835,861]" pageId="1" pageNumber="506">Kurile Islands</collectingCounty>
, the
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and the
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, east
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,
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,
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.
</paragraph>
</subSubSection>
<subSubSection id="8BD16548B203FF8D048EF3C0FC72FBCA" pageId="1" pageNumber="506" type="etymology">
<paragraph id="C37436C3B203FF8D048EF3C0FC72FBCA" blockId="1.[151,1437,799,1438]" pageId="1" pageNumber="506">
<emphasis id="F1BFEAD1B203FF8D048EF3C0FEAEFC7E" bold="true" box="[199,334,907,933]" pageId="1" pageNumber="506">Etymology.</emphasis>
The genus name is a combination of the family name of Pieter Kaas (
<date id="B7751003B203FF8D0006F3C0FAD3FC7D" box="[1103,1331,907,934]" pageId="1" pageNumber="506" value="1915-11-14">
November 14
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1915
</date>
<date id="B7751003B203FF8D0101F3C7FF30FC12" pageId="1" pageNumber="506" value="1996-06-08">
June 8
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1996
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) and the Greek word χιτών (= chiton). Pieter, or more familar Piet,was a famous malacologist who described dozens of species and genera of chitons and was one of the authors of six volumes of the “Monograph of Living Chitons”, for which he, among others, compiled beautiful drawings.
</paragraph>
</subSubSection>
<subSubSection id="8BD16548B203FF8D048EF450FB7BFA46" pageId="1" pageNumber="506" type="diagnosis">
<paragraph id="C37436C3B203FF8D048EF450FB7BFA46" blockId="1.[151,1437,799,1438]" pageId="1" pageNumber="506">
<emphasis id="F1BFEAD1B203FF8D048EF450FEA2FBEE" bold="true" box="[199,322,1051,1077]" pageId="1" pageNumber="506">Diagnosis.</emphasis>
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Chitons of small size with
<collectionCode id="A5DAAE06B203FF8D0625F457FD6FFBED" box="[620,655,1052,1078]" pageId="1" pageNumber="506">BL</collectionCode>
up to
<quantity id="04339B26B203FF8D0698F450FCC8FBED" box="[721,808,1051,1078]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="1" pageNumber="506" unit="mm" value="6.8">6.8 mm</quantity>
, usually 3.04.0 mm, elongate, valves low to moderately elevated (dorsal elevation about 0.360.49 [
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]). Head valve wider than tail valve. Tail valve generally with a wide, flattened, posterior mucro shifted to the postmucronal area. Ratio of length of antemucronal area to length of postmucronal area 1.172.10. Tegmentum smooth to the naked eye or with very low, rare granules arranged in vague longitudinal rows in central areas of intermediate valves and in antemucronal areas of tail valve and in a random manner in other areas. Each granule or microsculpture element with one aesthete pore. Perinotum very narrow, 910 times narrower than valve V, covered with bluntly pointed, short or long scales with 35 distinct or vague longitudinal ribs. Radula long, about half of body length, with 60123 rows of mature teeth; central tooth narrow, first lateral teeth wing-shaped, major lateral teeth narrow, with uni-,bi- or tridentate cusps. Number of gills in adult specimens between 5 to 8, ranging from valves VIVII to anus. Living predominantly on shelf, bathymetrically ranging from
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. Feeding
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mainly detrivorous. All four species are brooding
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.
</paragraph>
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<emphasis id="F1BFEAD1B203FF8D04DEF5BAFEEAF9D2" bold="true" box="[151,266,1521,1545]" pageId="1" pageNumber="506">TABLE 1.</emphasis>
Comparison of some important features of shell, girdle and radula of four species of
<taxonomicName id="04CB4D40B203FF8D00D1F5BAFAF9F9D2" authorityName="Sirenko" authorityYear="2024" box="[1176,1305,1521,1545]" class="Polyplacophora" family="Leptochitonidae" genus="Kaasichiton" kingdom="Animalia" order="Lepidopleurida" pageId="1" pageNumber="506" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="F1BFEAD1B203FF8D00D1F5BAFAF9F9D2" box="[1176,1305,1521,1545]" italics="true" pageId="1" pageNumber="506">Kaasichiton</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EA8C57AAB203FF8D0156F5BAFA8AF9D3" box="[1311,1386,1521,1545]" pageId="1" pageNumber="506" rank="genus">n. gen.</taxonomicNameLabel>
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</caption>
<paragraph id="C37436C3B203FF8D04D0F656FAEAF8BB" pageId="1" pageNumber="506">
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<tr id="7DFB3481B203007304D0F656FA75F9EE" box="[153,1429,1565,1589]" gridrow="0" pageId="1" pageNumber="506">
<th id="3E2A5DFDB203007304D0F656FEA9F9EE" box="[153,329,1565,1589]" gridcol="0" gridrow="0" pageId="1" pageNumber="506">Species</th>
<th id="3E2A5DFDB203007305CCF656FE0BF9EE" box="[389,491,1565,1589]" gridcol="1" gridrow="0" pageId="1" pageNumber="506">Ratio of</th>
<th id="3E2A5DFDB20300730642F656FD40F9EE" box="[523,672,1565,1589]" gridcol="2" gridrow="0" pageId="1" pageNumber="506">Ratio of length</th>
<th id="3E2A5DFDB203007306F0F656FCB5F9EE" box="[697,853,1565,1589]" gridcol="3" gridrow="0" pageId="1" pageNumber="506">Ratio of width of</th>
<th id="3E2A5DFDB20300730731F656FC3DF9EE" box="[888,989,1565,1589]" gridcol="4" gridrow="0" pageId="1" pageNumber="506">Dorsal</th>
<th id="3E2A5DFDB203007307BAF656FBACF9EE" box="[1011,1100,1565,1589]" gridcol="5" gridrow="0" pageId="1" pageNumber="506">Length of</th>
<th id="3E2A5DFDB2030073003CF656FB38F9EE" box="[1141,1240,1565,1589]" gridcol="6" gridrow="0" pageId="1" pageNumber="506">Number of</th>
<th id="3E2A5DFDB203007300B4F656FA75F9EE" box="[1277,1429,1565,1589]" gridcol="7" gridrow="0" pageId="1" pageNumber="506">Number of</th>
</tr>
<tr id="7DFB3481B203007304D0F60AFA75F9A6" box="[153,1429,1601,1661]" gridrow="1" pageId="1" pageNumber="506" rowspan-0="1">
<td id="3E2A5DFDB203007305CCF60AFD40F9A6" box="[389,672,1601,1661]" colspan="2" colspanRight="1" gridcol="1" gridrow="1" pageId="1" pageNumber="506">width of head of antemucronal to width of area to length of</td>
<td id="3E2A5DFDB203007306F0F60AFA75F9A6" box="[697,1429,1601,1661]" colspan="5" colspanRight="4" gridcol="3" gridrow="1" pageId="1" pageNumber="506">jugal sinus to width elevation of dorsal scales rows of radula denticles in cusp of apophyses in valve V of girdle of major lateral</td>
</tr>
<tr id="7DFB3481B203007304D0F6C2FA75F97A" box="[153,1429,1673,1697]" gridrow="2" pageId="1" pageNumber="506" rowspan-0="1" rowspan-4="1" rowspan-6="1">
<td id="3E2A5DFDB203007305CCF6C2FE0BF97A" box="[389,491,1673,1697]" gridcol="1" gridrow="2" pageId="1" pageNumber="506">tail valve</td>
<td id="3E2A5DFDB20300730642F6C2FCB5F97A" box="[523,853,1673,1697]" colspan="2" colspanRight="1" gridcol="2" gridrow="2" pageId="1" pageNumber="506">postmucronal area valve V</td>
<td id="3E2A5DFDB203007307BAF6C2FBACF97A" box="[1011,1100,1673,1697]" gridcol="5" gridrow="2" pageId="1" pageNumber="506">(µm)</td>
<td id="3E2A5DFDB203007300B4F6C2FA75F97A" box="[1277,1429,1673,1697]" gridcol="7" gridrow="2" pageId="1" pageNumber="506">tooth of radula</td>
</tr>
<tr id="7DFB3481B203007304D0F6F2FA75F90A" box="[153,1429,1721,1745]" gridrow="3" pageId="1" pageNumber="506">
<th id="3E2A5DFDB203007304D0F6F2FEA9F90A" box="[153,329,1721,1745]" gridcol="0" gridrow="3" pageId="1" pageNumber="506">
<taxonomicName id="04CB4D40B203FF8D04D0F6F2FF10F90A" authorityName="Sirenko" authorityYear="1990" box="[153,240,1721,1745]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="1" pageNumber="506" phylum="Mollusca" rank="species" species="lukini">
<emphasis id="F1BFEAD1B203FF8D04D0F6F2FF10F90A" box="[153,240,1721,1745]" italics="true" pageId="1" pageNumber="506">K. lukini</emphasis>
</taxonomicName>
</th>
<td id="3E2A5DFDB203007305CCF6F2FE0BF90A" box="[389,491,1721,1745]" gridcol="1" gridrow="3" pageId="1" pageNumber="506">1.081.16</td>
<td id="3E2A5DFDB20300730642F6F2FD40F90A" box="[523,672,1721,1745]" gridcol="2" gridrow="3" pageId="1" pageNumber="506">1.32</td>
<td id="3E2A5DFDB203007306F0F6F2FCB5F90A" box="[697,853,1721,1745]" gridcol="3" gridrow="3" pageId="1" pageNumber="506">2.32.4</td>
<td id="3E2A5DFDB20300730731F6F2FC3DF90A" box="[888,989,1721,1745]" gridcol="4" gridrow="3" pageId="1" pageNumber="506">0.390.40</td>
<td id="3E2A5DFDB203007307BAF6F2FBACF90A" box="[1011,1100,1721,1745]" gridcol="5" gridrow="3" pageId="1" pageNumber="506">5590</td>
<td id="3E2A5DFDB2030073003CF6F2FB38F90A" box="[1141,1240,1721,1745]" gridcol="6" gridrow="3" pageId="1" pageNumber="506">90123</td>
<td id="3E2A5DFDB203007300B4F6F2FA75F90A" box="[1277,1429,1721,1745]" gridcol="7" gridrow="3" pageId="1" pageNumber="506">2</td>
</tr>
<tr id="7DFB3481B203007304D0F6A3FA75F8DA" box="[153,1429,1768,1793]" gridrow="4" pageId="1" pageNumber="506">
<th id="3E2A5DFDB203007304D0F6A3FEA9F8DA" box="[153,329,1768,1793]" gridcol="0" gridrow="4" pageId="1" pageNumber="506">
<taxonomicName id="04CB4D40B203FF8D04D0F6A2FEF9F8DA" authorityName="Sirenko" authorityYear="2024" baseAuthorityName="Sirenko" baseAuthorityYear="2017" box="[153,281,1769,1793]" class="Polyplacophora" family="Leptochitonidae" genus="Kaasichiton" kingdom="Animalia" order="Lepidopleurida" pageId="1" pageNumber="506" phylum="Mollusca" rank="species" species="incubatus">
<emphasis id="F1BFEAD1B203FF8D04D0F6A2FEF9F8DA" box="[153,281,1769,1793]" italics="true" pageId="1" pageNumber="506">K. incubatus</emphasis>
</taxonomicName>
</th>
<td id="3E2A5DFDB203007305CCF6A3FE0BF8DA" box="[389,491,1768,1793]" gridcol="1" gridrow="4" pageId="1" pageNumber="506">1.031.17</td>
<td id="3E2A5DFDB20300730642F6A3FD40F8DA" box="[523,672,1768,1793]" gridcol="2" gridrow="4" pageId="1" pageNumber="506">1.332.10</td>
<td id="3E2A5DFDB203007306F0F6A3FCB5F8DA" box="[697,853,1768,1793]" gridcol="3" gridrow="4" pageId="1" pageNumber="506">1.32</td>
<td id="3E2A5DFDB20300730731F6A3FC3DF8DA" box="[888,989,1768,1793]" gridcol="4" gridrow="4" pageId="1" pageNumber="506">0.360.49</td>
<td id="3E2A5DFDB203007307BAF6A3FBACF8DA" box="[1011,1100,1768,1793]" gridcol="5" gridrow="4" pageId="1" pageNumber="506">6167</td>
<td id="3E2A5DFDB2030073003CF6A3FB38F8DA" box="[1141,1240,1768,1793]" gridcol="6" gridrow="4" pageId="1" pageNumber="506">100</td>
<td id="3E2A5DFDB203007300B4F6A3FA75F8DA" box="[1277,1429,1768,1793]" gridcol="7" gridrow="4" pageId="1" pageNumber="506">1</td>
</tr>
<tr id="7DFB3481B203007304D0F753FA75F8EA" box="[153,1429,1816,1841]" gridrow="5" pageId="1" pageNumber="506">
<th id="3E2A5DFDB203007304D0F753FEA9F8EA" box="[153,329,1816,1841]" gridcol="0" gridrow="5" pageId="1" pageNumber="506">
<taxonomicName id="04CB4D40B203FF8D04D0F752FEE9F8EB" authorityName="Sirenko" authorityYear="2024" box="[153,265,1817,1840]" class="Polyplacophora" family="Leptochitonidae" genus="Kaasichiton" kingdom="Animalia" order="Lepidopleurida" pageId="1" pageNumber="506" phylum="Mollusca" rank="species" species="ionensis" status="sp. nov.">
<emphasis id="F1BFEAD1B203FF8D04D0F752FEE9F8EB" box="[153,265,1817,1840]" italics="true" pageId="1" pageNumber="506">K. ionensis</emphasis>
</taxonomicName>
<emphasis id="F1BFEAD1B203FF8D0559F752FEA9F8EA" bold="true" box="[272,329,1817,1841]" pageId="1" pageNumber="506">
<taxonomicNameLabel id="EA8C57AAB203FF8D0559F752FEA9F8EA" box="[272,329,1817,1841]" pageId="1" pageNumber="506" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</th>
<td id="3E2A5DFDB203007305CCF753FE0BF8EA" box="[389,491,1816,1841]" gridcol="1" gridrow="5" pageId="1" pageNumber="506">1.051.12</td>
<td id="3E2A5DFDB20300730642F753FD40F8EA" box="[523,672,1816,1841]" gridcol="2" gridrow="5" pageId="1" pageNumber="506">1.171.22</td>
<td id="3E2A5DFDB203007306F0F753FCB5F8EA" box="[697,853,1816,1841]" gridcol="3" gridrow="5" pageId="1" pageNumber="506">1.652.07</td>
<td id="3E2A5DFDB20300730731F753FC3DF8EA" box="[888,989,1816,1841]" gridcol="4" gridrow="5" pageId="1" pageNumber="506">0.380.45</td>
<td id="3E2A5DFDB203007307BAF753FBACF8EA" box="[1011,1100,1816,1841]" gridcol="5" gridrow="5" pageId="1" pageNumber="506">6570</td>
<td id="3E2A5DFDB2030073003CF753FB38F8EA" box="[1141,1240,1816,1841]" gridcol="6" gridrow="5" pageId="1" pageNumber="506">6064</td>
<td id="3E2A5DFDB203007300B4F753FA75F8EA" box="[1277,1429,1816,1841]" gridcol="7" gridrow="5" pageId="1" pageNumber="506">3</td>
</tr>
<tr id="7DFB3481B203007304D0F703FA75F8BB" box="[153,1429,1864,1888]" gridrow="6" pageId="1" pageNumber="506">
<th id="3E2A5DFDB203007304D0F703FE0BF8BB" box="[153,491,1864,1888]" colspan="2" colspanRight="1" gridcol="0" gridrow="6" pageId="1" pageNumber="506">
<taxonomicName id="04CB4D40B203FF8D04D0F702FEA3F8BB" authorityName="Sirenko" authorityYear="2024" box="[153,323,1864,1888]" class="Polyplacophora" family="Leptochitonidae" genus="Kaasichiton" kingdom="Animalia" order="Lepidopleurida" pageId="1" pageNumber="506" phylum="Mollusca" rank="species" species="kamchatkensis" status="sp. nov.">
<emphasis id="F1BFEAD1B203FF8D04D0F702FEA3F8BB" box="[153,323,1864,1888]" italics="true" pageId="1" pageNumber="506">K. kamchatkensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EA8C57AAB203FF8D0500F703FE60F8BB" box="[329,384,1864,1888]" pageId="1" pageNumber="506" rank="species">
<emphasis id="F1BFEAD1B203FF8D0500F703FE9AF8BB" bold="true" box="[329,378,1864,1888]" pageId="1" pageNumber="506">n. sp</emphasis>
.
</taxonomicNameLabel>
1.09
</th>
<td id="3E2A5DFDB20300730642F703FD40F8BB" box="[523,672,1864,1888]" gridcol="2" gridrow="6" pageId="1" pageNumber="506">1.19</td>
<td id="3E2A5DFDB203007306F0F703FCB5F8BB" box="[697,853,1864,1888]" gridcol="3" gridrow="6" pageId="1" pageNumber="506">1.6</td>
<td id="3E2A5DFDB20300730731F703FC3DF8BB" box="[888,989,1864,1888]" gridcol="4" gridrow="6" pageId="1" pageNumber="506">0.39</td>
<td id="3E2A5DFDB203007307BAF703FBACF8BB" box="[1011,1100,1864,1888]" gridcol="5" gridrow="6" pageId="1" pageNumber="506">66118</td>
<td id="3E2A5DFDB2030073003CF703FB38F8BB" box="[1141,1240,1864,1888]" gridcol="6" gridrow="6" pageId="1" pageNumber="506">6084</td>
<td id="3E2A5DFDB203007300B4F703FA75F8BB" box="[1277,1429,1864,1888]" gridcol="7" gridrow="6" pageId="1" pageNumber="506">2</td>
</tr>
</table>
</paragraph>
<caption id="97B4664BB200FF8E04DEF6EFFDD1F8F3" ID-DOI="http://doi.org/10.5281/zenodo.13268888" ID-Zenodo-Dep="13268888" httpUri="https://zenodo.org/record/13268888/files/figure.png" pageId="2" pageNumber="507" startId="2.[151,250,1700,1724]" targetBox="[151,1435,181,1676]" targetPageId="2" targetType="figure">
<paragraph id="C37436C3B200FF8E04DEF6EFFDD1F8F3" blockId="2.[151,1437,1700,1832]" pageId="2" pageNumber="507">
<emphasis id="F1BFEAD1B200FF8E04DEF6EFFEF1F967" bold="true" box="[151,273,1700,1724]" pageId="2" pageNumber="507">FIGURE 1.</emphasis>
<taxonomicName id="04CB4D40B200FF8E055FF6EFFD24F967" authority=", Kurile Islands, Chirpoy" authorityName="Kurile Islands, Chirpoy" box="[278,708,1700,1724]" class="Polyplacophora" family="Leptochitonidae" genus="Kaasichiton" kingdom="Animalia" order="Lepidopleurida" pageId="2" pageNumber="507" phylum="Mollusca" rank="species" species="lukini">
<emphasis id="F1BFEAD1B200FF8E055FF6EFFE2FF967" box="[278,463,1700,1724]" italics="true" pageId="2" pageNumber="507">Kaasichiton lukini</emphasis>
, Kurile Islands, Chirpoy
</taxonomicName>
Island, 20 m, holotype (ZIN 1919), BL 4.0 mm (AC, EG),
<taxonomicName id="04CB4D40B200FF8E016BF6EFFE4FF93B" authority=", Kurile Islands, Urup" authorityName="Kurile Islands, Urup" class="Polyplacophora" family="Leptochitonidae" genus="Kaasichiton" kingdom="Animalia" order="Lepidopleurida" pageId="2" pageNumber="507" phylum="Mollusca" rank="species" species="lukini">
<emphasis id="F1BFEAD1B200FF8E016BF6EFFF31F93B" italics="true" pageId="2" pageNumber="507">Kaasichiton lukini</emphasis>
, Kurile Islands, Urup
</taxonomicName>
Island, Van der Linde Cape, 20 m, paratype (ZIN 1920), BL 3.0 mm (D):
<emphasis id="F1BFEAD1B200FF8E00E0F683FB22F93B" bold="true" box="[1193,1218,1736,1760]" pageId="2" pageNumber="507">A.</emphasis>
Valve I, dorsal view;
<emphasis id="F1BFEAD1B200FF8E04DEF6A7FF4EF8DF" bold="true" box="[151,174,1772,1796]" pageId="2" pageNumber="507">B.</emphasis>
Valve V, dorsal view;
<emphasis id="F1BFEAD1B200FF8E05DEF6A7FE30F8DF" bold="true" box="[407,464,1772,1796]" pageId="2" pageNumber="507">C, D.</emphasis>
Valves VIII, dorsal views;
<emphasis id="F1BFEAD1B200FF8E06A0F6A7FCE1F8DF" bold="true" box="[745,769,1772,1796]" pageId="2" pageNumber="507">E.</emphasis>
Valve V, detail of tegmentum surface of central area;
<emphasis id="F1BFEAD1B200FF8E0165F6A7FADFF8DF" bold="true" box="[1324,1343,1772,1796]" pageId="2" pageNumber="507">F.</emphasis>
Valve V, rostral view;
<emphasis id="F1BFEAD1B200FF8E0554F75BFED7F8F3" bold="true" box="[285,311,1808,1832]" pageId="2" pageNumber="507">G.</emphasis>
Valve VIII, lateral view.
</paragraph>
</caption>
<subSubSection id="8BD16548B201FF88048EF0DCFE0FFEC6" lastPageId="4" lastPageNumber="509" pageId="3" pageNumber="508" type="discussion">
<paragraph id="C37436C3B201FF8F048EF0DCFE93FCEF" blockId="3.[151,1437,151,821]" pageId="3" pageNumber="508">
<emphasis id="F1BFEAD1B201FF8F048EF0DCFEDBFF6A" bold="true" box="[199,315,151,177]" pageId="3" pageNumber="508">Remarks.</emphasis>
The species-rich genus
<taxonomicName id="04CB4D40B201FF8F061EF0DCFD3DFF6A" authorityName="Gray" authorityYear="1847" box="[599,733,151,177]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="3" pageNumber="508" phylum="Mollusca" rank="genus">
<emphasis id="F1BFEAD1B201FF8F061EF0DCFD3DFF6A" box="[599,733,151,177]" italics="true" pageId="3" pageNumber="508">Leptochiton</emphasis>
</taxonomicName>
in its traditional composition contained species with a wide range of morphological characters. Recently several genera (
<taxonomicName id="04CB4D40B201FF8F0720F0F1FB77FF0E" authority="Sirenko, 1973" authorityName="Sirenko" authorityYear="1973" box="[873,1175,186,213]" class="Polyplacophora" family="Leptochitonidae" genus="Hanleyella" kingdom="Animalia" order="Lepidopleurida" pageId="3" pageNumber="508" phylum="Mollusca" rank="genus">
<emphasis id="F1BFEAD1B201FF8F0720F0F1FC04FF0F" box="[873,996,186,212]" italics="true" pageId="3" pageNumber="508">Hanleyella</emphasis>
<bibRefCitation id="A75A4B32B201FF8F07B8F0F1FB77FF0E" author="Sirenko, B. I." box="[1009,1175,186,213]" pageId="3" pageNumber="508" pagination="1569 - 1571" refId="ref8123" refString="Sirenko, B. I. (1973) A new genus of the family Lepidopleuridae (Neoloricata). Zoological journal, 52 (10), 1569 - 1571, figs 1 - 16. [in Russian]" type="journal article" year="1973">Sirenko, 1973</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="04CB4D40B201FF8F00E1F0F1FF2FFF23" authority="Sirenko, 1988" authorityName="Sirenko" authorityYear="1988" class="Polyplacophora" family="Abyssochitonidae" genus="Ferreiraella" kingdom="Animalia" order="Lepidopleurida" pageId="3" pageNumber="508" phylum="Mollusca" rank="genus">
<emphasis id="F1BFEAD1B201FF8F00E1F0F1FAD1FF0F" box="[1192,1329,186,212]" italics="true" pageId="3" pageNumber="508">Ferreiraella</emphasis>
Sirenko, 1988
</taxonomicName>
,
<taxonomicName id="04CB4D40B201FF8F0497F095FDC3FF23" authority="Sirenko, 1992" authorityName="Sirenko" authorityYear="1992" box="[222,547,222,248]" class="Polyplacophora" family="Nierstraszellidae" genus="Nierstraszella" kingdom="Animalia" order="Lepidopleurida" pageId="3" pageNumber="508" phylum="Mollusca" rank="genus">
<emphasis id="F1BFEAD1B201FF8F0497F095FE9BFF23" box="[222,379,222,248]" italics="true" pageId="3" pageNumber="508">Nierstraszella</emphasis>
<bibRefCitation id="A75A4B32B201FF8F05CBF095FDC3FF23" author="Sirenko, B. I." box="[386,547,222,248]" pageId="3" pageNumber="508" pagination="81 - 90" refId="ref8256" refString="Sirenko, B. I. (1992) Nierstraszellidae fam. nov. - a new family of chitons (Polyplacophora, Lepidopleurida) from the bathyal of Western Pacific. Ruthenica, 2, 81 - 90." type="journal article" year="1992">Sirenko, 1992</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="04CB4D40B201FF8F0617F095FB9FFF23" authority=", Sirenko, Saito &amp; Schwabe, 2022" authorityName=", Sirenko, Saito &amp; Schwabe" authorityYear="2022" box="[606,1151,222,249]" class="Polyplacophora" family="Leptochitonidae" genus="Belknapchiton" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidopleurida" pageId="3" pageNumber="508" phylum="Mollusca" rank="genus">
<emphasis id="F1BFEAD1B201FF8F0617F095FCE0FF23" box="[606,768,222,248]" italics="true" pageId="3" pageNumber="508">Belknapchiton</emphasis>
,
<bibRefCitation id="A75A4B32B201FF8F0747F095FB9FFF23" author="Sirenko, B. &amp; Saito, H. &amp; Schwabe, E." box="[782,1151,222,248]" pageId="3" pageNumber="508" pagination="101 - 124" refId="ref8575" refString="Sirenko, B. Saito, H. &amp; Schwabe, E. (2022) A redescription of Leptochiton belknapi Dall, 1878 (Mollusca: Polyplacophora: Leptochitonidae), the type species of the new genus Belknapchiton. Zootaxa, 5205 (2), 101 - 124. https: // doi. org / 10.11646 / zootaxa. 5205.2.1" type="journal article" year="2022">Sirenko, Saito &amp; Schwabe, 2022</bibRefCitation>
</taxonomicName>
) could be separated from
<taxonomicName id="04CB4D40B201FF8F04DEF149FEFDFEC7" authorityName="Gray" authorityYear="1847" box="[151,285,258,284]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="3" pageNumber="508" phylum="Mollusca" rank="genus">
<emphasis id="F1BFEAD1B201FF8F04DEF149FEFDFEC7" box="[151,285,258,284]" italics="true" pageId="3" pageNumber="508">Leptochiton</emphasis>
</taxonomicName>
. The genus
<taxonomicName id="04CB4D40B201FF8F05EEF149FD35FEC7" authority="Iredale, 1914" authorityName="Iredale" authorityYear="1914" box="[423,725,258,284]" class="Polyplacophora" family="Leptochitonidae" genus="Terenochiton" kingdom="Animalia" order="Lepidopleurida" pageId="3" pageNumber="508" phylum="Mollusca" rank="genus">
<emphasis id="F1BFEAD1B201FF8F05EEF149FDD9FEC7" box="[423,569,258,284]" italics="true" pageId="3" pageNumber="508">Terenochiton</emphasis>
Iredale, 1914
</taxonomicName>
could be maintained (
<bibRefCitation id="A75A4B32B201FF8F0799F149FB8BFEC7" author="Sirenko, B. I." box="[976,1131,258,284]" pageId="3" pageNumber="508" pagination="1569 - 1571" refId="ref8123" refString="Sirenko, B. I. (1973) A new genus of the family Lepidopleuridae (Neoloricata). Zoological journal, 52 (10), 1569 - 1571, figs 1 - 16. [in Russian]" type="journal article" year="1973">Sirenko 1973</bibRefCitation>
, 1988, 1993; Sirenko
<emphasis id="F1BFEAD1B201FF8F012DF148FA7DFEC7" box="[1380,1437,258,284]" italics="true" pageId="3" pageNumber="508">et al.</emphasis>
2022) and several groups of morphologically similar species were recognized (
<bibRefCitation id="A75A4B32B201FF8F07BCF16DFB70FE9B" author="Sirenko, B. I." box="[1013,1168,293,320]" pageId="3" pageNumber="508" pagination="24 - 31" refId="ref8335" refString="Sirenko, B. I. (2015) The enigmatic viviparous chiton Calloplax vivipara (Plate, 1899) (Mollusca: Polyplacophora) and a survey of the types of reproduction in chitons. Russian Journal of Marine Biology, 41 (1), 24 - 31. https: // doi. org / 10.1134 / S 1063074015010095" type="journal article" year="2015">Sirenko 2015</bibRefCitation>
). The genus
<taxonomicName id="04CB4D40B201FF8F015FF16DFA7CFE9B" authorityName="Gray" authorityYear="1847" box="[1302,1436,294,320]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="3" pageNumber="508" phylum="Mollusca" rank="genus">
<emphasis id="F1BFEAD1B201FF8F015FF16DFA7CFE9B" box="[1302,1436,294,320]" italics="true" pageId="3" pageNumber="508">Leptochiton</emphasis>
</taxonomicName>
in its traditional composition is polyphyletic according to molecular studies (
<bibRefCitation id="A75A4B32B201FF8F07B6F102FB3DFEB8" author="Sigwart, J. D. &amp; Schwabe, E. &amp; Saito, H. &amp; Samadi, S. &amp; Giribet, G." box="[1023,1245,329,355]" pageId="3" pageNumber="508" pagination="560 - 572" refId="ref8048" refString="Sigwart, J. D., Schwabe, E., Saito, H., Samadi, S. &amp; Giribet, G. (2011) Evolution in the deep sea: a combined analysis of the earliest diverging living chitons (Mollusca: Polyplacophora: Lepidopleurida). Invertebrate Systematics, 24 (6), 560 - 572. https: // doi. org / 10.1071 / IS 10028" type="journal article" year="2011">
Sigwart
<emphasis id="F1BFEAD1B201FF8F0029F101FB79FEB8" box="[1120,1177,329,355]" italics="true" pageId="3" pageNumber="508">et al.</emphasis>
2011
</bibRefCitation>
). The new genus can be compared with the remaining
<taxonomicName id="04CB4D40B201FF8F0674F126FCFBFE5C" box="[573,795,365,391]" pageId="3" pageNumber="508">
<emphasis id="F1BFEAD1B201FF8F0674F126FD23FE5C" box="[573,707,365,391]" italics="true" pageId="3" pageNumber="508">Leptochiton</emphasis>
species
</taxonomicName>
forming groups of species that are similar to each other. The new genus
<taxonomicName id="04CB4D40B201FF8F0500F1DAFE30FE70" authorityName="Sirenko" authorityYear="2024" box="[329,464,401,427]" class="Polyplacophora" family="Leptochitonidae" genus="Kaasichiton" kingdom="Animalia" order="Lepidopleurida" pageId="3" pageNumber="508" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="F1BFEAD1B201FF8F0500F1DAFE30FE70" box="[329,464,401,427]" italics="true" pageId="3" pageNumber="508">Kaasichiton</emphasis>
</taxonomicName>
<emphasis id="F1BFEAD1B201FF8F0591F1DAFDC7FE70" bold="true" box="[472,551,401,427]" pageId="3" pageNumber="508">
<taxonomicNameLabel id="EA8C57AAB201FF8F0591F1DAFDC7FE70" box="[472,551,401,427]" pageId="3" pageNumber="508" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
resembles the group of
<taxonomicName id="04CB4D40B201FF8F0771F1DAFBFBFE70" baseAuthorityName="Carpenter" baseAuthorityYear="1892" box="[824,1051,401,427]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="3" pageNumber="508" phylum="Mollusca" rank="species" species="rugatus">
<emphasis id="F1BFEAD1B201FF8F0771F1DAFBFBFE70" box="[824,1051,401,427]" italics="true" pageId="3" pageNumber="508">Leptochiton rugatus</emphasis>
</taxonomicName>
(Carpenter in
<bibRefCitation id="A75A4B32B201FF8F00F6F1DAFAB7FE70" author="Pilsbry, H. A." box="[1215,1367,401,427]" pageId="3" pageNumber="508" pagination="1 - 64" refId="ref7982" refString="Pilsbry, H. A. (1892) Monograph of the Polyplacophora. In: Tryon Jr., G. W. (Ed.), Manual of Conchology, structural and systematic. Series 1. 14 (53). Academy of Natural Sciences, Philadelphia, Pennsylvania, pp. 1 - 64, pls. 1 - 15." type="book chapter" year="1892">Pilsbry, 1892</bibRefCitation>
). This group of chitons consisting of six species inhabits the northern part of the Pacific Ocean (
<bibRefCitation id="A75A4B32B201FF8F00CBF1FFFA78FE14" author="Sirenko, B. I. &amp; Sigwart, J. D." box="[1154,1432,436,463]" pageId="3" pageNumber="508" refId="ref8511" refString="Sirenko, B. I. &amp; Sigwart, J. D. (2021) Leptochiton subrugatus sp. nov. (Mollusca: Polyplacophora) from low boreal waters of northern Pacific. Marine Biodiversity, 51 (64). [unknown pagination] https: // doi. org / 10.1007 / s 12526 - 021 - 01190 - z" type="journal volume" year="2021">Sirenko &amp; Sigwart 2021</bibRefCitation>
, 2022), including the areas inhabited by species of the new genus.
<taxonomicName id="04CB4D40B201FF8F07C9F193FBE7FE29" authorityName="Sirenko" authorityYear="2024" box="[896,1031,472,498]" class="Polyplacophora" family="Leptochitonidae" genus="Kaasichiton" kingdom="Animalia" order="Lepidopleurida" pageId="3" pageNumber="508" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="F1BFEAD1B201FF8F07C9F193FBE7FE29" box="[896,1031,472,498]" italics="true" pageId="3" pageNumber="508">Kaasichiton</emphasis>
</taxonomicName>
<emphasis id="F1BFEAD1B201FF8F0046F193FBBFFE29" bold="true" box="[1039,1119,472,498]" pageId="3" pageNumber="508">
<taxonomicNameLabel id="EA8C57AAB201FF8F0046F193FBBFFE29" box="[1039,1119,472,498]" pageId="3" pageNumber="508" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
differs from the
<taxonomicName id="04CB4D40B201FF8F0156F192FF38FDCC" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="3" pageNumber="508" phylum="Mollusca" rank="species" species="rugatusgroup">
<emphasis id="F1BFEAD1B201FF8F0156F192FF38FDCC" italics="true" pageId="3" pageNumber="508">L. rugatusgroup</emphasis>
</taxonomicName>
by having a head valve, which is always wider than the tail valve (
<emphasis id="F1BFEAD1B201FF8F07F0F1B6FC30FDCD" box="[953,976,509,534]" italics="true" pageId="3" pageNumber="508">vs</emphasis>
head valve is narrower than the tail valve in species of
<emphasis id="F1BFEAD1B201FF8F0579F26AFE4FFDE1" box="[304,431,545,570]" italics="true" pageId="3" pageNumber="508">L. rugatus-</emphasis>
group), a wide, flattened posterior mucro on the postmucronal area (
<emphasis id="F1BFEAD1B201FF8F0089F26AFB37FDE1" box="[1216,1239,545,570]" italics="true" pageId="3" pageNumber="508">vs</emphasis>
a narrow mucro, commonly on the anterior postmucronal area in species of the
<emphasis id="F1BFEAD1B201FF8F0713F20FFC37FD86" box="[858,983,580,605]" italics="true" pageId="3" pageNumber="508">L. rugatus-</emphasis>
group), a very narrow perinotum, which is 910 times narrower than the width of the fifth valve (
<emphasis id="F1BFEAD1B201FF8F0747F223FCC5FD5A" box="[782,805,616,641]" italics="true" pageId="3" pageNumber="508">vs</emphasis>
the perinotum is only 45 times narrower than the valve in species of the
<emphasis id="F1BFEAD1B201FF8F051DF2C7FE31FD7E" box="[340,465,652,677]" italics="true" pageId="3" pageNumber="508">L. rugatus-</emphasis>
group), a weak tegmentum sculpture (
<emphasis id="F1BFEAD1B201FF8F0735F2C7FC73FD7E" box="[892,915,652,677]" italics="true" pageId="3" pageNumber="508">vs</emphasis>
distinct sculpture in species of the
<taxonomicName id="04CB4D40B201FF8F0169F2C7FF3CFD12" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="3" pageNumber="508" phylum="Mollusca" rank="species" species="rugatusgroup">
<emphasis id="F1BFEAD1B201FF8F0169F2C7FF3CFD12" italics="true" pageId="3" pageNumber="508">L. rugatusgroup</emphasis>
</taxonomicName>
), and wing-shaped first lateral teeth (
<emphasis id="F1BFEAD1B201FF8F06C1F2E4FD7FFD13" box="[648,671,687,712]" italics="true" pageId="3" pageNumber="508">vs</emphasis>
narrow, oar-shaped first lateral teeth in species). Unlike species of the
<emphasis id="F1BFEAD1B201FF8F0488F298FEDDFD37" box="[193,317,723,748]" italics="true" pageId="3" pageNumber="508">L. rugatus-</emphasis>
group, all species of the new genus are brooding eggs and juveniles in their pallial groove. The new genus is well distinguished from the other species of the genus
<taxonomicName id="04CB4D40B201FF8F0723F2BDFC10FCCB" authorityName="Gray" authorityYear="1847" box="[874,1008,758,784]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="3" pageNumber="508" phylum="Mollusca" rank="genus">
<emphasis id="F1BFEAD1B201FF8F0723F2BDFC10FCCB" box="[874,1008,758,784]" italics="true" pageId="3" pageNumber="508">Leptochiton</emphasis>
</taxonomicName>
by morphological characters listed in the genus diagnosis
</paragraph>
<caption id="97B4664BB201FF8F04DEF7F0FD88F82C" ID-DOI="http://doi.org/10.5281/zenodo.13268890" ID-Zenodo-Dep="13268890" httpUri="https://zenodo.org/record/13268890/files/figure.png" pageId="3" pageNumber="508" startId="3.[151,250,1979,2003]" targetBox="[151,1435,844,1955]" targetPageId="3" targetType="figure">
<paragraph id="C37436C3B201FF8F04DEF7F0FD88F82C" blockId="3.[151,1436,1979,2039]" pageId="3" pageNumber="508">
<emphasis id="F1BFEAD1B201FF8F04DEF7F0FEF4F808" bold="true" box="[151,276,1979,2003]" pageId="3" pageNumber="508">FIGURE 2.</emphasis>
<taxonomicName id="04CB4D40B201FF8F0552F7F0FD54F809" authority=", Kurile Islands, Urup" authorityName="Kurile Islands, Urup" box="[283,692,1979,2003]" class="Polyplacophora" family="Leptochitonidae" genus="Kaasichiton" kingdom="Animalia" order="Lepidopleurida" pageId="3" pageNumber="508" phylum="Mollusca" rank="species" species="lukini">
<emphasis id="F1BFEAD1B201FF8F0552F7F0FE36F808" box="[283,470,1979,2003]" italics="true" pageId="3" pageNumber="508">Kaasichiton lukini</emphasis>
, Kurile Islands, Urup
</taxonomicName>
Island, Van der Linde Cape, 20 m, paratype (ZIN 1920), BL 3.0 mm:
<emphasis id="F1BFEAD1B201FF8F01CDF7F0FF50F82C" bold="true" pageId="3" pageNumber="508">A, C.</emphasis>
Dorsal spicules and needles;
<emphasis id="F1BFEAD1B201FF8F0595F794FDF2F82C" bold="true" box="[476,530,2015,2039]" pageId="3" pageNumber="508">B, D.</emphasis>
Radula.
</paragraph>
</caption>
<paragraph id="C37436C3B206FF880498F0DCFE0FFEC6" blockId="4.[151,1436,151,285]" pageId="4" pageNumber="509">
<taxonomicName id="04CB4D40B206FF880498F0DCFD7BFF6A" authority="Haddon, 1886" authorityName="Haddon" authorityYear="1886" box="[209,667,151,177]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="4" pageNumber="509" phylum="Mollusca" rank="species" species="kerguelensis">
<emphasis id="F1BFEAD1B206FF880498F0DCFE0CFF6A" box="[209,492,151,177]" italics="true" pageId="4" pageNumber="509">Leptochiton kerguelensis</emphasis>
<bibRefCitation id="A75A4B32B206FF8805BFF0DCFD7BFF6A" author="Haddon, A. C." box="[502,667,151,177]" pageId="4" pageNumber="509" pagination="1 - 50" refId="ref7814" refString="Haddon, A. C. (1886) Report on the Polyplacophora collected by H. M. S. &quot; Challenger &quot; during the years 1873 - 1876. Challenger Report, 15 (43), 1 - 50." type="journal article" year="1886">Haddon, 1886</bibRefCitation>
</taxonomicName>
, widespread in the seas of the Southern Ocean, is also a brooding species, and is morphological closest to the species of
<taxonomicName id="04CB4D40B206FF8806BFF0F0FC9DFF0E" authorityName="Sirenko" authorityYear="2024" box="[758,893,187,213]" class="Polyplacophora" family="Leptochitonidae" genus="Kaasichiton" kingdom="Animalia" order="Lepidopleurida" pageId="4" pageNumber="509" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="F1BFEAD1B206FF8806BFF0F0FC9DFF0E" box="[758,893,187,213]" italics="true" pageId="4" pageNumber="509">Kaasichiton</emphasis>
</taxonomicName>
<emphasis id="F1BFEAD1B206FF8807CDF0F0FC32FF0E" bold="true" box="[900,978,187,213]" pageId="4" pageNumber="509">
<taxonomicNameLabel id="EA8C57AAB206FF8807CDF0F0FC32FF0E" box="[900,978,187,213]" pageId="4" pageNumber="509" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
However, it differs from them by having an anterior mucro, a narrow, oar-shaped first lateral tooth of the radula and twice as many longitudinal ribs on the dorsal scales of the perinotum.
</paragraph>
</subSubSection>
</treatment>
</document>