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<document id="DFECBEE0DC80E2B8601331A2C1BC5F65" ID-CLB-Dataset="304926" ID-DOI="10.3897/aiep.54.135448" ID-GBIF-Dataset="22888b0d-a602-4918-9cc1-db34e6998697" ID-ZooBank="1F507CB1-0C06-4381-9CBF-CB0A9FFC8263" ID-publisher-id="135448" URI-arpha="470541C0-0A45-53EE-8F14-A86CE57F3B99" URI-zoobank="http://zoobank.org/1F507CB1-0C06-4381-9CBF-CB0A9FFC8263" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" article-type="research-article" checkinTime="1730398432430" checkinUser="pensoft" docAuthor="Dewa, Yuna &amp; Motomura, Hiroyuki" docDate="2024" docId="4B190EF571535E359FDD79EBC31536B7" docLanguage="en" docName="Acta Ichthyologica et Piscatoria 54: 261-268" docOrigin="Acta Ichthyologica et Piscatoria 54" docSource="https://aiep.pensoft.net/article/135448/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Enneapterygius rhothion Fricke 1997" docType="treatment" docVersion="4" dtd-version="3.0" lastPageNumber="268" masterDocId="470541C00A4553EE8F14A86CE57F3B99" masterDocTitle="First records of two triplefins, Enneapterygius rhothion and Enneapterygius olivaceus (Actinopterygii: Blenniiformes: Tripterygiidae), from Australia and Vanuatu" masterLastPageNumber="268" masterPageNumber="261" pageNumber="261" updateTime="1730408107366" updateUser="admin">
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<mods:title id="314671D4293306A1D2AD303E73EEA585">First records of two triplefins, Enneapterygius rhothion and Enneapterygius olivaceus (Actinopterygii: Blenniiformes: Tripterygiidae), from Australia and Vanuatu</mods:title>
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<mods:namePart id="985EE73F73F66CD5AE15E46E577C0D3C">Dewa, Yuna</mods:namePart>
<mods:nameIdentifier id="72D357EE0C30C79381BC216470E823D3" type="ORCID">0000-0002-8512-6782</mods:nameIdentifier>
<mods:affiliation id="6AC22E9837DB1A17167F850A53A489F2">United Graduate School of Agricultural Sciences, Kagoshima University, Kagoshima, Japan</mods:affiliation>
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<mods:namePart id="48AC34F2471B1E84D95E6BFB3C38EE34">Motomura, Hiroyuki</mods:namePart>
<mods:nameIdentifier id="B34E306AA0C8EB333BB3F6EEC700FE67" type="ORCID">0000-0002-7448-2482</mods:nameIdentifier>
<mods:affiliation id="0C21DEAAFF386DB158787230843C543E">Kagoshima University Museum, Kagoshima, Japan</mods:affiliation>
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<mods:date id="CCF4B15803077629302F3B3D2E659BF8">2024</mods:date>
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<treatment id="4B190EF571535E359FDD79EBC31536B7" ID-DOI="http://doi.org/10.5281/zenodo.14019627" ID-GBIF-Taxon="245302291" ID-Zenodo-Dep="14019627" ID-arpha="4B190EF5-7153-5E35-9FDD-79EBC31536B7" LSID="urn:lsid:plazi:treatment:4B190EF571535E359FDD79EBC31536B7" httpUri="http://treatment.plazi.org/id/4B190EF571535E359FDD79EBC31536B7">
<subSubSection id="FD54BD615C256E4F695C156BAD0454FF" type="nomenclature">
<paragraph id="863152495BFF3F32F303E2184B87FA6F">
<taxonomicName id="188E08A34F29B37B90BBDD0524365BB2" authority="Fricke, 1997" authorityName="Fricke" authorityYear="1997" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Blenniiformes" phylum="Chordata" rank="species" species="rhothion">
<emphasis id="F8ED228A4B86F7B7EC101C186E1155D8" italics="true">Enneapterygius rhothion</emphasis>
<bibRefCitation id="65BCFE7F530064D5EC205BD541C47001" author="Fricke R" journalOrPublisher="Koeltz Scientific Books, Königstein, Germany" refId="B4" refString="Fricke R (1997) Tripterygiid fishes of the western and central Pacific (Teleostei). Koeltz Scientific Books, Konigstein, Germany, 607 pp." title="Tripterygiid fishes of the western and central Pacific (Teleostei)." year="1997">Fricke, 1997</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="79C1C390AFB730001D55F4F25F63D7D0" type="description">
<paragraph id="5E0CAFE74FF8C3A1BD228F6922F45C81">
<figureCitation id="8DA340EA426852A5AAE6628F39BB979B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Photographs of preserved specimen of Enneapterygius rhothion (A: AMS I. 5088, male, 32.5 mm SL, Australia; B: SMNS 18322, holotype, male, 28.9 mm SL, New Caledonia, Photo by M. Meguro)." figureDoi="10.3897/aiep.54.135448.figure1" httpUri="https://binary.pensoft.net/fig/1165072">English name: New Caledonian blackhead surf triplefin Figs 1</figureCitation>
,
<figureCitation id="FFEDDBB08327BD6A67DEE5895A661C38" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photographs of A: dorsal and B: ventral views of head of Enneapterygius rhothion (AMS I. 5088, male, 32.5 mm SL, Australia), showing cephalic sensory system." figureDoi="10.3897/aiep.54.135448.figure2" httpUri="https://binary.pensoft.net/fig/1165073">2</figureCitation>
,
<figureCitation id="ADF243155DC737CD68F0C72EE4D82941" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distributional map of Enneapterygius rhothion (red circles) and E. olivaceus (blue triangles). Closed and open symbols indicate localities determined during the presently reported study and previous records, respectively. Black arrows indicate type localities of the two species." figureDoi="10.3897/aiep.54.135448.figure3" httpUri="https://binary.pensoft.net/fig/1165074">3</figureCitation>
;
<tableCitation id="56B1E86B5CEBEC3DEBD567E990644DDC" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements and counts of Enneapterygius rhothion. Means in parentheses." httpUri="http://table.plazi.org/id/75AA8939657332AF822205B274797CEA" tableUuid="75AA8939657332AF822205B274797CEA">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVGAC" type="material">
<paragraph id="80175656220C8CDD638743484BA10C54">
<heading id="64A30560974FC346F2B6CADFAE04BDCF" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="7D667749D5E08F99D41E36D846CADF99">
<materialsCitation id="3018D5640539C47640235D1C28D7ABB5" collectionCode="AMS" collectorName="C. Hedley" country="Australia" latitude="-17.230001" location="Green Island" longLatPrecision="21" longitude="145.96971" specimenCode="AMS I. 5088" specimenCount="1" specimenCount-male="1" stateProvince="Queensland">
<specimenCode id="214F3569AB881F2CB8E45D3BA9837B08">AMS I. 5088</specimenCode>
,
<specimenCount id="98788EA20A17033433B502EBDA760718" type="male">male</specimenCount>
,
<quantity id="B82AF4AD6175716622F35DF784DEBD96" metricMagnitude="-2" metricUnit="m" metricValue="3.25" unit="mm" value="32.5">32.5 mm</quantity>
SL,
<location id="54AC2FEDF6431419FB3DDF92EF413E45" LSID="urn:lsid:plazi:treatment:4B190EF571535E359FDD79EBC31536B7:54AC2FEDF6431419FB3DDF92EF413E45" country="Australia" latitude="-17.230001" longLatPrecision="21" longitude="145.96971" name="Green Island" stateProvince="Queensland">Green Island</location>
,
<collectingRegion id="3FF56CB3D6841F661C4992BFD68BF8A8" country="Australia" name="Queensland">Queensland</collectingRegion>
,
<collectingCountry id="1FECC8448BBB366B6EBD3099EA82F39A" name="Australia">Australia</collectingCountry>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E5GAC" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[145.969722,-17.230000]}">
<geoCoordinate id="79859C4A2170143F8DCEE4E6AE351AD3" degrees="17" direction="south" minutes="13" orientation="latitude" precision="15" seconds="48" value="-17.230001">17°13′48″S</geoCoordinate>
,
<geoCoordinate id="E5333EBE6E01C8ED0ACDEF735B16A494" degrees="145" direction="east" minutes="58" orientation="longitude" precision="15" seconds="11" value="145.96971">145°58′11″E</geoCoordinate>
</named-content>
, 1901,
<collectorName id="49CAE28D6E48518B0B169C124A9EEDEA">C. Hedley</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDHAC" type="description">
<paragraph id="EEC2F409F1C98A2F31ECD8776391B5D5">
<heading id="A60C26B2341945007884873560086D8D" reason="title">Description.</heading>
</paragraph>
<paragraph id="6B764D003F0FDEB47ADF026026EA7C46">
Counts and measurements given in Table
<tableCitation id="181729337AAE2F3C0EDFB509F74BD4E2" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements and counts of Enneapterygius rhothion. Means in parentheses." httpUri="http://table.plazi.org/id/75AA8939657332AF822205B274797CEA" tableUuid="75AA8939657332AF822205B274797CEA">1</tableCitation>
. Body moderately elongate, slightly compressed anteriorly, progressively more compressed posteriorly (Fig.
<figureCitation id="10F52D7BC0AA235268F786E0C25DDDD6" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Photographs of preserved specimen of Enneapterygius rhothion (A: AMS I. 5088, male, 32.5 mm SL, Australia; B: SMNS 18322, holotype, male, 28.9 mm SL, New Caledonia, Photo by M. Meguro)." figureDoi="10.3897/aiep.54.135448.figure1" httpUri="https://binary.pensoft.net/fig/1165072">1 A</figureCitation>
). Dorsal profile of snout straight, steep. Mouth slightly oblique; posterior margin of maxilla extending beyond anterior margin of pupil; anterior tip of upper jaw slightly above level of lower margin of orbit (lateral view). Medial supratemporal canal with two branches (Fig.
<figureCitation id="1AC53B3C869DF39C4D3BFBDCC83FDE38" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photographs of A: dorsal and B: ventral views of head of Enneapterygius rhothion (AMS I. 5088, male, 32.5 mm SL, Australia), showing cephalic sensory system." figureDoi="10.3897/aiep.54.135448.figure2" httpUri="https://binary.pensoft.net/fig/1165073">2 A</figureCitation>
). Anterior nostril a membranous tube, at mid-eye level, slightly closer to eye than to upper lip; nasal tentacle slender, unbranched, with pointed tip; posterior nostril opening circular, without membranous tube (Fig.
<figureCitation id="0BD85BBF1B59A24504BF793F5F1B683A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Photographs of A: dorsal and B: ventral views of head of Enneapterygius rhothion (AMS I. 5088, male, 32.5 mm SL, Australia), showing cephalic sensory system." figureDoi="10.3897/aiep.54.135448.figure2" httpUri="https://binary.pensoft.net/fig/1165073">2 A</figureCitation>
). Eye oriented dorsolaterally, with small, pointed tentacle, slightly longer than nasal tentacle, on posterodorsal margin. Interorbital space narrow, its width less than pupil diameter. Opercular margin slightly pointed, reaching below base of 3 rd spine of first dorsal fin.
</paragraph>
<caption id="75AA8939657332AF822205B274797CEA" ID-Table-UUID="75AA8939657332AF822205B274797CEA" httpUri="http://table.plazi.org/id/75AA8939657332AF822205B274797CEA" startId="T1" targetIsTable="true">
<paragraph id="C139DA416EEACCCAC2C1CD3DE4FD105E">
<label id="9784F43DDAA23A9324100798B236BB18">Table 1.</label>
</paragraph>
<paragraph id="6B0BC06B8A4814AB0EA21F18B5634F6C">
Measurements and counts of
<taxonomicName id="6CB9F2777FACF3F594D154C67DB1A862" authorityName="Fricke" authorityYear="1997" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="rhothion">
<emphasis id="6306E3512636DE23DBD77E5C99C86C0C" italics="true">Enneapterygius rhothion</emphasis>
</taxonomicName>
. Means in parentheses.
</paragraph>
</caption>
<paragraph id="B020410D545050561FDD97C8016EE61F">
<table id="TID0ERKAE" rules="all">
<tbody id="BA3CD2D10607E92C7E875773560DA7FC">
<tr id="AB784AE3BCC1F377704EA8D5F6583CCC">
<th id="C0896EADD3D5ECC53AD631CC194F68DF" colspan="1" rowspan="4">Character</th>
<th id="6B39625FA496E90D3D339BE0D91065FD" colspan="1" rowspan="1">Australia</th>
<th id="F181B4029AE8C845ACD6EDF23051CFB3" colspan="2" rowspan="1">New Caledonia and Vanuatu</th>
</tr>
<tr id="26EF8FFC09AC61B2EADCE2D6F1162E72">
<th id="B98A59F82C0D147CBCF8F0F465FCDCF1" colspan="1" rowspan="1">AMS I. 5088</th>
<th id="34FE9CABC9490226C0761719C4EE928E" colspan="1" rowspan="1">SMNS 18322</th>
<th id="85DD6628DD06774CED34C39AE16680F2" colspan="1" rowspan="1">
<emphasis id="1470C69294854BFFC9061828D367DC1D" italics="true">n</emphasis>
= 8
</th>
</tr>
<tr id="9A11AE9F6981CC4C0ECB9117AEFA0805">
<th id="6AA73922FF75F6C576D3B8FF55BFA3C5" colspan="1" rowspan="1">Non-type</th>
<th id="AE29DEB441DA02F9594C47DF2A073428" colspan="1" rowspan="1">Holotype</th>
<th id="A3BE71C87F8201504D0EBD587FF26308" colspan="1" rowspan="1">Paratypes</th>
</tr>
<tr id="988324AC3D73629F4E7D29488137DF9C">
<th id="58447EB81D1FF393EDFEDFB7C399BB0E" colspan="1" rowspan="1">Male</th>
<th id="592E8B0C622C8456CE50D539FC35285D" colspan="1" rowspan="1">Male</th>
<th id="4BC17DECFABC20E167C6BACE1B49FF48" colspan="1" rowspan="1">Males</th>
</tr>
<tr id="A7A29E3B7FC40DD8E5A701EEF68D06FF">
<td id="ACDDC2520913868027EE2C7EDA6B9D92" colspan="4" rowspan="1">
<emphasis id="F3AF8937C6F423AA1726F8F5CA3E5820" bold="true">Morphometric data; absolute value [mm</emphasis>
]
</td>
</tr>
<tr id="443B5BE59CE6948E6F402EDC3AB7773A">
<td id="2FBF5BEDA74C43BCA12D75DD52761C0D" colspan="1" rowspan="1">Standard length (SL)</td>
<td id="727E08CDA74BF0A82AC98E04A2D0F65A" colspan="1" rowspan="1">32.5</td>
<td id="B84A9458F7426351EE44F329235F3DBE" colspan="1" rowspan="1">28.9</td>
<td id="A7FE3CD452C4582C800F9AB4E36B4F8F" colspan="1" rowspan="1">23.4–27.8</td>
</tr>
<tr id="5CC9EABF0434848DC235A8C2B29CE45F">
<td id="FD65DFA6D4103BD84D3CF96AFD1808F2" colspan="4" rowspan="1">
<emphasis id="5F855CA26A881D0D08DA9EA725FBFD2F" bold="true">Morphometric data; relative values [% SL</emphasis>
]
</td>
</tr>
<tr id="015847BDF974038C17A7AA7B442AEE9C">
<td id="517DA051573D27E20CBDA36C3ABC498B" colspan="1" rowspan="1">Body depth</td>
<td id="15111B4749B71992F425E0E286B3992C" colspan="1" rowspan="1">20.5</td>
<td id="4921CEBB4E4F0AFAD3643F67B9DE37DE" colspan="1" rowspan="1">20.0</td>
<td id="D3B4E5F5C4E5061F57081D602BD5F29F" colspan="1" rowspan="1">18.5–20.4 (19.4)</td>
</tr>
<tr id="FDEC5CC4CEB1C322A02DE650007E0250">
<td id="0D0771F24FE6B84468957D61BA737411" colspan="1" rowspan="1">Body width</td>
<td id="3299B6244BFF6AC0CD10003E15A2593F" colspan="1" rowspan="1">18.4</td>
<td id="9BB0FCDB5B0D415798C788E76EEBEC21" colspan="1" rowspan="1">18.0</td>
<td id="F13133A0AA980F4F94EE4CB2FE04D067" colspan="1" rowspan="1">15.6–17.7 (16.8)</td>
</tr>
<tr id="82A850F77908803A364DB4D278870566">
<td id="F141360FF511EA2FF1D2682C9EFCE4AC" colspan="1" rowspan="1">Head length</td>
<td id="1EA29FD7BBFCECD93BC3DE350AE0F082" colspan="1" rowspan="1">29.3</td>
<td id="84B6C842B4A301E5672AD6C023A5BDD0" colspan="1" rowspan="1">26.7</td>
<td id="F0D0B6C3094DC1B931ECF6616F921223" colspan="1" rowspan="1">28.7–29.8 (29.2)</td>
</tr>
<tr id="C7D44893A64AB683042813011D4949AE">
<td id="9F631C73EB92F69C622E1365DAC1E811" colspan="1" rowspan="1">Snout length</td>
<td id="F15021005EBFF7DF9265809F2A1CC01A" colspan="1" rowspan="1">12.5</td>
<td id="9BA306C14FE82EE803BE650FFC72BC91" colspan="1" rowspan="1">9.2</td>
<td id="185EC9DC5BED5B0B2B9C4FA0B20EB732" colspan="1" rowspan="1">9.1–10.1 (9.6)</td>
</tr>
<tr id="7AAEBBDB2776F31B5F54F1AF37D327DA">
<td id="1D0A798D4F589BFCCB413541B733C8E5" colspan="1" rowspan="1">Orbital tentacle length</td>
<td id="0C4BD3A833A74DD5FC02BE1B21563BEB" colspan="1" rowspan="1">1.6</td>
<td id="D8E9AF3EBA58F7CE06B2EE46929789F1" colspan="1" rowspan="1">2.1</td>
<td id="A0DBB79E07F8ED5C8B7BF954F2C389D7" colspan="1" rowspan="1">—</td>
</tr>
<tr id="8FAFDC945C80C6D7294CEB06BEEC9492">
<td id="1211D6C6A981364E74533D2DFFD3694A" colspan="1" rowspan="1">Orbit diameter</td>
<td id="3FBB3C66E751B5B636CCB11E87A82AC8" colspan="1" rowspan="1">8.4</td>
<td id="58DDE019C37F212A25117A1ADF6C0BBD" colspan="1" rowspan="1">9.6</td>
<td id="DD9A286BAD882583678DEF27D1F6DBAA" colspan="1" rowspan="1">9.6–10.4 (10.0)</td>
</tr>
<tr id="556CC4A12DA9E1BEA58F6FA0EF68720F">
<td id="5CF1BEC837706C9F4F4B53DADA0B97DE" colspan="1" rowspan="1">Interorbital width</td>
<td id="242C6047586466FD23BB090D4A82717A" colspan="1" rowspan="1">2.3</td>
<td id="E57A4D4878002593787DB69E0993917E" colspan="1" rowspan="1">2.9</td>
<td id="19E979D91F32EAD1A5CAD97F0A0DE716" colspan="1" rowspan="1">2.4–2.9 (2.7)</td>
</tr>
<tr id="1F115618ACA12777F889DEFE64455CEF">
<td id="4AE31137D38038BE40EB2D7F60C6E8DE" colspan="1" rowspan="1">Upper-jaw length</td>
<td id="A551BB9E37068EC54654C73438BA5A3F" colspan="1" rowspan="1">11.6</td>
<td id="5167BABF95CA93688D196D322E70DDED" colspan="1" rowspan="1">11.4</td>
<td id="F28B5F3F3D6A6708158D98577485A588" colspan="1" rowspan="1">11.1–11.8 (11.5)</td>
</tr>
<tr id="A829FA5D8D215AA733C8B4A2E07751FF">
<td id="F83E0E4A3A402E0192039C6966CD61E8" colspan="1" rowspan="1">Postorbital length</td>
<td id="B567637C43B5BE4FB425F2BCDEBD9B06" colspan="1" rowspan="1">12.1</td>
<td id="B5C9159B4B78D124E553211654FAD23B" colspan="1" rowspan="1">12.0</td>
<td id="DD16D6EC0DA2BA14A6C1333705CE6DE8" colspan="1" rowspan="1">11.6–13.0 (12.5)</td>
</tr>
<tr id="C9C75F9E3BF21B6B1791B0723C13E839">
<td id="F1A7325B5F9F64243D218A1EF8E2DCD4" colspan="1" rowspan="1">Pre- 1 st-dorsal-fin length</td>
<td id="256B7EBCBA0A57C4B7D61F12654A7903" colspan="1" rowspan="1">25.4</td>
<td id="A4BB29D859DC42F994FF68EA0ECA34AF" colspan="1" rowspan="1">24.8</td>
<td id="7E26DB57893EF1F58ACC6C56365110A8" colspan="1" rowspan="1">25.2–27.3 (26.3)</td>
</tr>
<tr id="43DE5E3B87E4A7F94E180E0FFFE8D136">
<td id="C99B202401CE74FF8064ABD3C3673D82" colspan="1" rowspan="1">Pre- 2 nd-dorsal-fin length</td>
<td id="E6BEE83B7FDB24531406FAC436C1FDD1" colspan="1" rowspan="1">36.5</td>
<td id="9D3B102DCFA787C0B1DF2612A5FEB98F" colspan="1" rowspan="1">35.6</td>
<td id="85CD8357FB87887DD71C1F81BAFB565B" colspan="1" rowspan="1">35.5–38.6 (37.5)</td>
</tr>
<tr id="3C74D078AF2E05B9383CF6182A8C3373">
<td id="DBFF89875157D6FD7981CBDD75E46161" colspan="1" rowspan="1">Pre- 3 rd-dorsal-fin length</td>
<td id="46E053F83C711C54D66A72A3C4732E28" colspan="1" rowspan="1">69.2</td>
<td id="146F03DC55A38976BB3A1F677A6BE60A" colspan="1" rowspan="1">71.3</td>
<td id="7273F63FF250390555DE083FCC2C71CC" colspan="1" rowspan="1">69.7–74.6 (72.9)</td>
</tr>
<tr id="1B0D60F1882DEF8953B4BDB1B4C723EA">
<td id="00DC0E6548B29C733F9CDFB60CC7ADF5" colspan="1" rowspan="1">Pre-anal-fin length</td>
<td id="5A80AADD9CED105C24269ED5705ED8B6" colspan="1" rowspan="1">47.0</td>
<td id="BDC3D3E682619DEEF6C8570F882C9EBC" colspan="1" rowspan="1">48.2</td>
<td id="DB87A40EEAF323BD8DA9961B5D874082" colspan="1" rowspan="1">48.9–52.8 (50.7)</td>
</tr>
<tr id="650333EAEF72A3193EDE6D2B7A3DEBD0">
<td id="B28E60388A74D0906005154D1C42EE0F" colspan="1" rowspan="1">Anal-fin base length</td>
<td id="15C5368CE40480ADC9F22E7415639B6C" colspan="1" rowspan="1">41.1</td>
<td id="F23173076E2BDCF2E46FF0D81B8B96DD" colspan="1" rowspan="1">42.2</td>
<td id="C0E5F79E18A848002C08C76C175D7AF0" colspan="1" rowspan="1">37.6–42.7 (40.4)</td>
</tr>
<tr id="7CCABBCAF4DE3F9A0A1F43BF496EB291">
<td id="793B2F02FF5F90F08A15D1ADD6FFA693" colspan="1" rowspan="1">Pre-pectoral-fin length</td>
<td id="16219C8FF028C3417156A3CF27F84D75" colspan="1" rowspan="1">32.2</td>
<td id="BFDC4B52C65A059E5BE4BFFFB742AFFD" colspan="1" rowspan="1">31.3</td>
<td id="AAD48C1AB005673C969E6F7E1A808243" colspan="1" rowspan="1">29.3–33.1 (31.4)</td>
</tr>
<tr id="067621D98CF0B257F661E814DD068E91">
<td id="E37EFCD80379AE729EBD008C6D182195" colspan="1" rowspan="1">Pre-pelvic-fin length</td>
<td id="4D1876BDAC06A9AD3B64DD6DC7248EED" colspan="1" rowspan="1">21.3</td>
<td id="B5BBA573FDE9DB2290D0EAA6836DF0A0" colspan="1" rowspan="1">24.4</td>
<td id="185B3E252B4638873A477E6A9762FEBB" colspan="1" rowspan="1">20.1–24.6 (23.0)</td>
</tr>
<tr id="FF28E85A8DA96C64C75FFC82F5C2641C">
<td id="29EDA7020AB2750F43FD04AA02A9864C" colspan="1" rowspan="1">Caudal peduncle length</td>
<td id="FE98BF5131353B4998BB02C91F47A2D4" colspan="1" rowspan="1">10.5</td>
<td id="ED60C426BBC3121C6B87F30334C5CAC2" colspan="1" rowspan="1">10.9</td>
<td id="CCE70BF837D7D2857514922A4530E9BB" colspan="1" rowspan="1">10.9–12.3 (11.6)</td>
</tr>
<tr id="C5CAC58C28B25AB97429229B661A61EB">
<td id="081A906E5243804576F5DDF21E6664EE" colspan="1" rowspan="1">Caudal peduncle depth</td>
<td id="84E239F0EF7C1D88D7F00168E9587699" colspan="1" rowspan="1">9.1</td>
<td id="EFEC35FD1DE704580EAA7F342CD0475E" colspan="1" rowspan="1">8.0</td>
<td id="7AF3B4DB54AEA3C658AAEFECFAB63658" colspan="1" rowspan="1">7.6–8.9 (8.2)</td>
</tr>
<tr id="7E94733151F78A305253CF894598E6C7">
<td id="9E48F2316F81B508D5ACCED25A9E4791" colspan="1" rowspan="1">1 st spine length of 1 st dorsal fin</td>
<td id="A2C90D03157DFE18E0E7CF2834F18B43" colspan="1" rowspan="1">Damaged</td>
<td id="AE845E3D334CC99C81FD6A1485D6FB23" colspan="1" rowspan="1">9.0</td>
<td id="92F980D4509500203EE3FB5559FDE323" colspan="1" rowspan="1">9.8–10.5 (10.1)</td>
</tr>
<tr id="965AE15ACB61348DE58DBF80DBF3A32E">
<td id="BE6C0F6FAE3AD5B1886E645FA62BCA83" colspan="1" rowspan="1">2 nd spine length of 1 st dorsal fin</td>
<td id="7E1EEFD58FB8F71891BBDC17235E9F48" colspan="1" rowspan="1">10.4</td>
<td id="8309750F0D59EBBB95AC148A5690D1C6" colspan="1" rowspan="1">8.2</td>
<td id="25A0BA3110CFABA1530DA19AB8EB576E" colspan="1" rowspan="1">9.2–9.8 (9.4)</td>
</tr>
<tr id="3948DE7AD148170700E00903185ED8CE">
<td id="F5D9BA08707244873FB0F2A41D7823A3" colspan="1" rowspan="1">3 rd spine length of 1 st dorsal fin</td>
<td id="831342EA3B87411D229165F8EC1B2520" colspan="1" rowspan="1">8.9</td>
<td id="03695EEF4D808C88FDE44B7915566465" colspan="1" rowspan="1">7.0</td>
<td id="E3675D566622673F03CF269BFAFEC713" colspan="1" rowspan="1">7.8–9.0 (8.1)</td>
</tr>
<tr id="51F15DED5F7EB0E1CE01DF57A5B11AFA">
<td id="3B32810FE4DAC7982E46CA65E8326BF7" colspan="1" rowspan="1">1 st dorsal-fin base length</td>
<td id="29CD87431C060159E1CC7A7479BCFD2A" colspan="1" rowspan="1">5.6</td>
<td id="2A2A2BE37BA5D5214D73702F5CCA2315" colspan="1" rowspan="1">5.6</td>
<td id="8FC8A5420D9A6A6C1A42A337AFFBC312" colspan="1" rowspan="1">5.0–6.3 (5.8)</td>
</tr>
<tr id="A6FD2B2C2CEA966BCB783DC8729E011E">
<td id="2AC31E2C58919E2324227CF261989BDA" colspan="1" rowspan="1">1 st spine length of 2 nd dorsal fin</td>
<td id="71F34EA719AC2790AD7DCE37C0D1B545" colspan="1" rowspan="1">14.0</td>
<td id="9B1F109E89A8A2F9ECECD16C03A728AA" colspan="1" rowspan="1">14.2</td>
<td id="A745282E48F90CB5D7A07E4400416FA1" colspan="1" rowspan="1">14.1–15.5 (15.0)</td>
</tr>
<tr id="7181C1BC6B981464AEB1AD332477CA3C">
<td id="2628B17E4EDF616A31352E9F30D9C35B" colspan="1" rowspan="1">2 nd spine length of 2 nd dorsal fin</td>
<td id="F9C719CB377AC22F7E0FC1A03E41F2A9" colspan="1" rowspan="1">12.5</td>
<td id="C476D32AF5AEFC74E0CCCA08FA9B831E" colspan="1" rowspan="1">14.7</td>
<td id="9CABE29A32D37C1D980D7BB816D4AB1A" colspan="1" rowspan="1">14.6–16.6 (15.6)</td>
</tr>
<tr id="4C01B8038F44657137C609D88E29AED7">
<td id="E2B9DE0143551AA5EF9758C9B56BFEB2" colspan="1" rowspan="1">3 rd spine length of 2 nd dorsal fin</td>
<td id="7D7A3DE6F3350FB8A62B1DB22A370E59" colspan="1" rowspan="1">15.4</td>
<td id="6BB1FC46D3F3B012147FF9DE9B456CBA" colspan="1" rowspan="1">14.7</td>
<td id="F0B0A2DF0E6196447E72EFD0505BDD14" colspan="1" rowspan="1">14.5–16.5 (15.5)</td>
</tr>
<tr id="6B9D0F76D73D7863FBA3AC3A3718D4A2">
<td id="DDA74B05E1C7F0988D3578EC67C32E6E" colspan="1" rowspan="1">2 nd dorsal-fin base length</td>
<td id="5C553D4C510A45D7B1A974E5A90AE9DB" colspan="1" rowspan="1">30.5</td>
<td id="8768253F19B39D1EF07BA0199BD75AEA" colspan="1" rowspan="1">30.9</td>
<td id="1EF99A8EE29867DD405C291F17E380D7" colspan="1" rowspan="1">29.5–34.9 (31.9)</td>
</tr>
<tr id="658943FD6F19B9979DD60B073A9BBB98">
<td id="DC95EFFD0CA243B2E0F7B55A8953E891" colspan="1" rowspan="1">1 st ray length of 3 rd dorsal fin</td>
<td id="A4E47A40B8157F66A8AE23ACAFECCFB4" colspan="1" rowspan="1">Damaged</td>
<td id="F7FC4FE2CFFA3EA8602BD36156EE4D6B" colspan="1" rowspan="1">16.4</td>
<td id="BE2D7FBC194452F42153827275D64A95" colspan="1" rowspan="1">16.1–18.3 (17.0)</td>
</tr>
<tr id="5F7DB6C0B05AF2329543DCC90CB4279E">
<td id="69ED4F9EDED9A0E540080D9FE0A576B3" colspan="1" rowspan="1">2 nd ray length of 3 rd dorsal fin</td>
<td id="8AC090AA7DAA04A65773DDA4C828A952" colspan="1" rowspan="1">Damaged</td>
<td id="E0A268A7AAA0824BBFFCC3DDB830F7FE" colspan="1" rowspan="1">15.1</td>
<td id="EC18C03ADBAEEE24E782E3B2E0BE0951" colspan="1" rowspan="1">16.0–17.1 (16.6)</td>
</tr>
<tr id="E17C84D9CCEC53B27E6B142CC100782C">
<td id="830AB14850743B799FD306237155CC11" colspan="1" rowspan="1">3 rd ray length of 3 rd dorsal fin</td>
<td id="54D2C3A2FA4D9AE8228B8AA775A17A13" colspan="1" rowspan="1">Damaged</td>
<td id="4B2729C75C498D00D4CA7C948A176B56" colspan="1" rowspan="1">14.9</td>
<td id="53806297A5E1A2C860310C4841245E70" colspan="1" rowspan="1">9.5–16.1 (14.9)</td>
</tr>
<tr id="ABDC4783FB3923C0F5CCE665817CA249">
<td id="377CE6030EA407B81F8CD1086BB88678" colspan="1" rowspan="1">3 rd dorsal fin base length</td>
<td id="A4D7FB94E94889F8C9269B5CB66F5D20" colspan="1" rowspan="1">16.4</td>
<td id="3A20811C28699E63A6B259C9923376F2" colspan="1" rowspan="1">16.7</td>
<td id="884B31DE345BD96A5EE71F49D6A3BA97" colspan="1" rowspan="1">14.3–18.1 (16.4)</td>
</tr>
<tr id="CEB1019EE19F7731216015BA03E7666F">
<td id="0F7BD835F4A74E0E39D5696F9A86F86E" colspan="1" rowspan="1">Pectoral-fin length</td>
<td id="B88A3104D55C338360743212BDFB923F" colspan="1" rowspan="1">Damaged</td>
<td id="F8D9F6C8FEECFDD494A6C7887399B1D9" colspan="1" rowspan="1">27.0</td>
<td id="5F4187CE84DA09F1F1E151167DFEABAA" colspan="1" rowspan="1">29.8–34.8 (32.7)</td>
</tr>
<tr id="D4EE1D02120A63F10E852B22E4B2995A">
<td id="A4C9F53D8AE280297ECE36EDF61F13ED" colspan="1" rowspan="1">1 st ray length of pelvic fin</td>
<td id="4211AF3677C5F24C76D49A6DADCDCE15" colspan="1" rowspan="1">16.4</td>
<td id="8044CCA56027AE12CF3C1EA3885BB805" colspan="1" rowspan="1">16.0</td>
<td id="CD3FE57A3A70314D15F593E314B6B0DB" colspan="1" rowspan="1">15.6–19.3 (17.6)</td>
</tr>
<tr id="A6E4126FC0288FEDC33CA207BB367148">
<td id="8F063F8CD20C21C4804AE1940FA05847" colspan="1" rowspan="1">2 nd ray length of pelvic fin</td>
<td id="9C23EB73D39D76D63B90F5877CC0BD76" colspan="1" rowspan="1">Damaged</td>
<td id="4268530E432EEE23328C098E5B26A7DD" colspan="1" rowspan="1">19.3</td>
<td id="D54387E1D11AC7D5A050703054051D81" colspan="1" rowspan="1">22.6–25.1 (24.3)</td>
</tr>
<tr id="4909C8ACB27908CBEB561ABBC145BFA6">
<td id="D94C0DB29E5B1E1F10BF0EC576230892" colspan="4" rowspan="1">
<emphasis id="565189D04F4B2CB10E305937F7DD7721" bold="true">Meristic data (counts)</emphasis>
</td>
</tr>
<tr id="008D5FA0903C4AAD2FC225C0C5164630">
<td id="3700FC6C48C862BCC22B9D852ED11EEC" colspan="1" rowspan="1">Dorsal-fin rays</td>
<td id="BB5C47DF6B9D781D5971D1A60F383178" colspan="1" rowspan="1">III, XIII, 10</td>
<td id="A959A0E9368E26551297732D068728E4" colspan="1" rowspan="1">III, XIII, 9</td>
<td id="D9D253D1871F347DD3278DBB66A35AE4" colspan="1" rowspan="1">III, XIII or XIV, 9 or 10</td>
</tr>
<tr id="D6FEF649A17ECF3E6C837C9AF9C46BE5">
<td id="FBD3F6A16C9B6BC1A45F2DABE06768B3" colspan="1" rowspan="1">Anal-fin rays</td>
<td id="66B01B28F2CD2354948F56BF35D4B598" colspan="1" rowspan="1">I, 18</td>
<td id="6A8E664AC3FB7C6ABBE45D276170117B" colspan="1" rowspan="1">I, 18</td>
<td id="54ED23F1037D0EA72FA9142B1CCFB6A7" colspan="1" rowspan="1">I, 17 or 18</td>
</tr>
<tr id="D6C5C1AFFF20F3BE198A189F379B7FA1">
<td id="B9E78768CD6789F1F4C2097BA4AD3F61" colspan="1" rowspan="1">Pectoral-fin rays</td>
<td id="531B79406709345715FD9CA3CF3A47C4" colspan="1" rowspan="1">iii + damaged = 15</td>
<td id="2294DC4F0314C555C1CE40C2438BD87D" colspan="1" rowspan="1">ii + 6 + vii = 15</td>
<td id="5E5569A20BCAB3186EF96BCDDE6BAA96" colspan="1" rowspan="1">ii – iv + 4–6 + 6 or 7 = 14–16</td>
</tr>
<tr id="0A62A99E1BE44E47146CDB0918F8578D">
<td id="929547D1E70C82F38706514C3C259A9D" colspan="1" rowspan="1">Scale rows in longitudinal series</td>
<td id="F398557616C76068776A28798A26C6F3" colspan="1" rowspan="1">33</td>
<td id="3620E26CDDC0CB8F14C90534835F0873" colspan="1" rowspan="1">34</td>
<td id="627812CA3E5BFC17A5DA1648A001DB75" colspan="1" rowspan="1">34 or 35</td>
</tr>
<tr id="04D45036E0E1711A85FC32ED8FA5B121">
<td id="8C928E9228A4C90500BDB3BE9E40917E" colspan="1" rowspan="1">Pored lateral-line scales</td>
<td id="04DCDD5ED459A0EBD375B605837736D8" colspan="1" rowspan="1">16</td>
<td id="6A82F9F7C13C8F63B3F5477DFAB3DF19" colspan="1" rowspan="1">19</td>
<td id="F8CD8AD092FF644264191047EB4D9B47" colspan="1" rowspan="1">15–17</td>
</tr>
<tr id="F320B570977DC30288D66C29E38EB2A4">
<td id="0D8D80ADD75EB684E4414184FA269A04" colspan="1" rowspan="1">Notched lateral-line scales</td>
<td id="1EDC3948FFB84E88B1622ABD784C4350" colspan="1" rowspan="1">20</td>
<td id="3C39F3D4F438C15E7500C5A99E327EB4" colspan="1" rowspan="1">19</td>
<td id="B53D52A7776CCFB7D7527E70CD6EDC65" colspan="1" rowspan="1">19–21</td>
</tr>
<tr id="B0FADCBD78FDAC57F5F16A47F768761A">
<td id="F5237E24EEB0F5DF8467DEFD95555A37" colspan="1" rowspan="1">Scales above 1 st PLL</td>
<td id="567A5230E2F360CD0758EB9E115A23D0" colspan="1" rowspan="1">3</td>
<td id="E032F00764EF45A6BCC16073DF7F7C83" colspan="1" rowspan="1">4</td>
<td id="D43A0BAEB961EBE164064667BC64CE98" colspan="1" rowspan="1">4</td>
</tr>
<tr id="7FFFD093C1A2A1FFE0A2C1A44DA58E80">
<td id="84817E045B3DFBF768282D4C8FC2F9F0" colspan="1" rowspan="1">Scales below 2 nd dorsal fin</td>
<td id="96F1C6EB1E09727FE205009B11D45A7B" colspan="1" rowspan="1">3</td>
<td id="A6D6A28BBBB634AE0D13E0DB4BCF592D" colspan="1" rowspan="1">3 ½</td>
<td id="34B7CF9D3AD8CD5A4DBB2F0CF59763A2" colspan="1" rowspan="1">3 ½ or 4</td>
</tr>
<tr id="EBF8A004190FB3C9F7DC981E096C7896">
<td id="BD470077C13CA6497B2401B18A2D696D" colspan="1" rowspan="1">Scales below 1 st NLL</td>
<td id="6382C21035DA279D250B870BCD1FA524" colspan="1" rowspan="1">3 ½</td>
<td id="5F5D2E499039DF05056B59E6BAA5F9E6" colspan="1" rowspan="1">3 ½</td>
<td id="2BE829BBFE403E8A16AAB8E3E6BC51B8" colspan="1" rowspan="1">3 or 3 ½</td>
</tr>
<tr id="E3FD1A74BC7661787D1999061A609B13">
<td id="F1C7A4583BEF604E013395280A69081A" colspan="1" rowspan="1">Scales above last PLL</td>
<td id="FC836884D2EAB4314AFBE6FFA10A4EF6" colspan="1" rowspan="1">2 ½</td>
<td id="8A8C6E412C3DFE66203D883F3E922780" colspan="1" rowspan="1">3</td>
<td id="508FF6B92868CAF83F5BC4E94174D644" colspan="1" rowspan="1">2 ½ or 3</td>
</tr>
<tr id="6873E37DBBD2AEEEB7D4E7E50CDC7C82">
<td id="67A9D7FC786F1695762E7407AF1BE95B" colspan="1" rowspan="1">Mandibular pore formula</td>
<td id="01CBED7F8105C435C2B0F5EDDC00F917" colspan="1" rowspan="1">3 + 2 + 3</td>
<td id="9778D62F50C1EE21134B1D83ED6F1A37" colspan="1" rowspan="1">4 + 2 + 4</td>
<td id="F27FE8E5EC807D243992F0D310539DBA" colspan="1" rowspan="1">3 or 4 + 1 or 2 + 3 or 4</td>
</tr>
</tbody>
</table>
</paragraph>
<caption id="AF63FB552CD9A6FE72B09F520B50C9B4" ID-DOI="10.3897/aiep.54.135448.figure1" ID-arpha="00434F7C-24EC-58F0-9969-44798E369073" httpUri="https://binary.pensoft.net/fig/1165072" startId="F1">
<paragraph id="3CC4F1593375039C3E7949C343474AE4">
<label id="BB0002D85820C8053099E659795471B5">Figure 1.</label>
</paragraph>
<paragraph id="7C10961FEC5C2DA4D937AD5105346C71">
Photographs of preserved specimen of
<taxonomicName id="19D79F8E021830134ADEF8140F623C93" authorityName="Fricke" authorityYear="1997" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="rhothion">
<emphasis id="2EBCFA2ACA4F366C8B52068EAA6794AE" italics="true">Enneapterygius rhothion</emphasis>
</taxonomicName>
(
<emphasis id="22938FA5AE2EB584FAD9907B60E01B60" bold="true">A</emphasis>
: AMS I. 5088, male, 32.5 mm SL, Australia;
<emphasis id="03488693CD66AFAFBA2BF9701CFF69CC" bold="true">B</emphasis>
: SMNS 18322, holotype, male, 28.9 mm SL, New Caledonia, Photo by M. Meguro).
</paragraph>
</caption>
<caption id="F0466698719265489D2FBF1A0C681394" ID-DOI="10.3897/aiep.54.135448.figure2" ID-arpha="4CE11ABD-2A02-5138-B1FC-BE2938EEDDC7" httpUri="https://binary.pensoft.net/fig/1165073" startId="F2">
<paragraph id="4FC9AD6C09F0F2DF4F0832BE3C5677E7">
<label id="29BE866E916B63DC28F1495CC1DD25F5">Figure 2.</label>
</paragraph>
<paragraph id="9A4FD786AF251791145A2461E48C4873">
Photographs of
<emphasis id="8A08BAA0EAEB772E986A3D05A762D92F" bold="true">A</emphasis>
: dorsal and
<emphasis id="E793CFA42A2E87F6C9A198EF3A157C76" bold="true">B</emphasis>
: ventral views of head of
<taxonomicName id="DDD4DD7A8A7BB77CFF0A874DCC9BFEF5" authorityName="Fricke" authorityYear="1997" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="rhothion">
<emphasis id="DEFCA0DBFD109347AE99E5309AE4A211" italics="true">Enneapterygius rhothion</emphasis>
</taxonomicName>
(AMS I. 5088, male, 32.5 mm SL, Australia), showing cephalic sensory system.
</paragraph>
</caption>
<paragraph id="BD7A3A3C4A7B2383A581BB7EDD3178D0">Lateral line discontinuous, with anterior series of pored scales ending below base of 11 th spine of 2 nd dorsal fin; second scale of posterior series of notched scales below third scale from last pored scale, ending at caudal-fin base; body scales ctenoid; scales absent on head (including maxilla, interorbital space, preopercle, and opercle), pectoral-fin base, pre- and inter-pelvic-fin region, abdomen, pre-dorsal-fin region, and all fin membranes, except basal part of caudal fin.</paragraph>
<paragraph id="798AFDC9E10E5B22D9971198EFA228D2">First dorsal fin triangular, origin over and slightly forward of midpoint between pre-opercular and opercular margins; 1 st spine of first dorsal fin damaged, 2 nd spine longer than 3 rd spine. Origin of second dorsal fin just above 4 th pored lateral-line scale, 3 rd spine longest, spines thereafter becoming gradually shorter posteriorly, forming rounded margin. Third dorsal fin damaged, its origin just above 20 th scale row of longitudinal series. Anal-fin membranous margin deeply incised between rays; anal fin origin just below 7 th spine base of second dorsal fin, its posteriormost tip close to caudal-fin base. Pectoral fin damaged (posterior tip lost); upper and lowermost pectoral-fin base level with bases of 3 rd and 2 nd spines, respectively, of first dorsal fin. Pelvic fin origin vertically below base of 1 st spine of first dorsal fin. Caudal fin rounded, its length less than head length.</paragraph>
</subSubSection>
<subSubSection id="SECID0EVOAE" type="Nuptial male coloration">
<paragraph id="68CCF8F3D737D457183548C1132C9BD8">
<heading id="A476B83BB2F09773F6E400261A002C2C" reason="title">Nuptial male coloration</heading>
</paragraph>
<paragraph id="27EF9DF33DAF9D2CE19DE1E4C2736465">
<emphasis id="780248A75C4411879B77566237092E8C" bold="true">(in preservative).</emphasis>
Based on Fig.
<figureCitation id="8AC5D12A1FE88558A80190A95CE092B9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Photographs of preserved specimen of Enneapterygius rhothion (A: AMS I. 5088, male, 32.5 mm SL, Australia; B: SMNS 18322, holotype, male, 28.9 mm SL, New Caledonia, Photo by M. Meguro)." figureDoi="10.3897/aiep.54.135448.figure1" httpUri="https://binary.pensoft.net/fig/1165072">1 A</figureCitation>
. Body generally yellowish white; brownish pigmentation forming faint A- and X-shaped bars on lateral surface of body. Head, including snout, lips, cheek, and opercle, and pectoral-fin base brown with numerous melanophores. Pre-pelvic region brown (extending to just behind pelvic-fin base). Orbital tentacle pale brown. First and second dorsal fins dark brown. Third dorsal fin damaged, but brownish basally. Pectoral, pelvic, anal and caudal fins whitish.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBPAE" type="distribution">
<paragraph id="85B9375E30D3274E7816DF1BDE276BFB">
<heading id="9EA6FD1D6600889FD16C7C6896DD8F85" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="A3E8620F3ED83C5659CE97325EF13A37">
Currently known from
<collectingCountry id="4262B2784BE2FA0A34531297AA7398A8" name="New Caledonia">New Caledonia</collectingCountry>
(Chesterfield Islands, Grande Terre, Ile des Pins, and Loyalty Islands),
<collectingCountry id="2B2007A917BDFDE41A70ACCB170FB629" name="Vanuatu">Vanuatu</collectingCountry>
(Erromango Island), and
<collectingCountry id="CDB9BBCEEF062F8932FD39184028089D" name="Australia">Australia</collectingCountry>
(
<bibRefCitation id="CD7C729C015516FC970872570B820D69" author="Fricke" firstAuthor="Fricke" journalOrPublisher="Koeltz Scientific Books, Königstein, Germany" refId="B4" refString="Fricke R (1997) Tripterygiid fishes of the western and central Pacific (Teleostei). Koeltz Scientific Books, Königstein, Germany, 607 pp." title="Tripterygiid fishes of the western and central Pacific (Teleostei)." year="1997">Fricke 1997</bibRefCitation>
,
<bibRefCitation id="C8F45A42821BB794E30FB456E590B389" DOI="10.1023/A:1020076906430" author="Fricke" firstAuthor="Fricke" journalOrPublisher="Environmental Biology of Fishes" pagination="175-198" refId="B6" refString="Fricke R (2002) Tripterygiid fishes of New Caledonia, with zoogeographical remarks. Environmental Biology of Fishes 65: 175–198. https://doi.org/10.1023/A:1020076906430" title="Tripterygiid fishes of New Caledonia, with zoogeographical remarks." volume="65" year="2002">2002</bibRefCitation>
; this study) (Fig.
<figureCitation id="9DC124A19DA0E369F80F6F83484797A5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distributional map of Enneapterygius rhothion (red circles) and E. olivaceus (blue triangles). Closed and open symbols indicate localities determined during the presently reported study and previous records, respectively. Black arrows indicate type localities of the two species." figureDoi="10.3897/aiep.54.135448.figure3" httpUri="https://binary.pensoft.net/fig/1165074">3</figureCitation>
).
</paragraph>
<caption id="F0CA3228908DED096F781655B450A1D8" ID-DOI="10.3897/aiep.54.135448.figure3" ID-arpha="167181C2-D47E-54B4-AA4A-4A3DCF9E3B1E" httpUri="https://binary.pensoft.net/fig/1165074" startId="F3">
<paragraph id="242C947BF37F395E4E4BD19CC41F66AE">
<label id="61FA74006592FBEC1FDE1AD112D4E97A">Figure 3.</label>
</paragraph>
<paragraph id="D33BA5DAA7AD59DEE48DCA7FB414998D">
Distributional map of
<taxonomicName id="45681986D980182B024B7E90FF6FC4E5" authorityName="Fricke" authorityYear="1997" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="rhothion">
<emphasis id="2B992328DCE2EA8B56695A3AF7D918D1" italics="true">Enneapterygius rhothion</emphasis>
</taxonomicName>
(
<emphasis id="C6C4A865859D4704167867C4554862CA" italics="true">red circles</emphasis>
) and
<taxonomicName id="D0580D4ABBF555600A62628A421E9BFE" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="D33828099684BF831FFF076FAA1B0BD1" italics="true">E. olivaceus</emphasis>
</taxonomicName>
(
<emphasis id="5A2C261FE91BE95C392964E37ED7E797" italics="true">blue triangles</emphasis>
). Closed and open symbols indicate localities determined during the presently reported study and previous records, respectively.
<emphasis id="9B4661A7B007B77BBC4C8BB2660C958F" italics="true">Black arrows</emphasis>
indicate type localities of the two species.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E3QAE" type="remarks">
<paragraph id="8CA483767B949D54FAB070568BA042ED">
<heading id="B7FE1250A835B73C10F9EBA4A9D5EC94" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="0C8AF819CFA1642E601E164898E7230E">
The morphometric and meristic characters of the single specimen collected from
<collectingCountry id="AB4BFAA3342097CAD7274CAFA5369D1B" name="Australia">Australia</collectingCountry>
(AMS I. 5088, male,
<quantity id="DBD56DC78FA2CD3E12210D1BC86F37CA" metricMagnitude="-2" metricUnit="m" metricValue="3.25" unit="mm" value="32.5">32.5 mm</quantity>
SL) agreed closely with the type series of
<taxonomicName id="529F4FC1861E6AD9CDAD8A448BC5C19B" authorityName="Fricke" authorityYear="1997" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="rhothion">
<emphasis id="879AE793FA62F6A3A1466DB6B6A88BE8" italics="true">E. rhothion</emphasis>
</taxonomicName>
re-examined in this study (Table
<tableCitation id="BE592D8D3BC344D6790EFFB0AF0E65CE" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements and counts of Enneapterygius rhothion. Means in parentheses." httpUri="http://table.plazi.org/id/75AA8939657332AF822205B274797CEA" tableUuid="75AA8939657332AF822205B274797CEA">1</tableCitation>
), and much of the original description of the species provided by
<bibRefCitation id="A49591827EE7B4233A0DE60EBB793C1D" author="Fricke R" firstAuthor="Fricke" journalOrPublisher="Koeltz Scientific Books, Königstein, Germany" refId="B4" refString="Fricke R (1997) Tripterygiid fishes of the western and central Pacific (Teleostei). Koeltz Scientific Books, Königstein, Germany, 607 pp." title="Tripterygiid fishes of the western and central Pacific (Teleostei)." year="1997">Fricke (1997)</bibRefCitation>
. However, although
<bibRefCitation id="86763316E8BD85DDD689872D0475744B" author="Fricke R" firstAuthor="Fricke" journalOrPublisher="Koeltz Scientific Books, Königstein, Germany" refId="B4" refString="Fricke R (1997) Tripterygiid fishes of the western and central Pacific (Teleostei). Koeltz Scientific Books, Königstein, Germany, 607 pp." title="Tripterygiid fishes of the western and central Pacific (Teleostei)." year="1997">Fricke (1997)</bibRefCitation>
described the species as having a short orbital tentacle of length 0.6 % – 0.9 % of SL, both the presently reported specimen and
<typeStatus id="F5CD53CE1ADFD65C1090E0ECD9E06EB7" type="holotype">holotype</typeStatus>
of
<taxonomicName id="E797AE028D7E754B182A23E08FDF0A5E" authorityName="Fricke" authorityYear="1997" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="rhothion">
<emphasis id="F259EC1DD07BD84C1EF4358AAF49ACC1" italics="true">E. rhothion</emphasis>
</taxonomicName>
had relatively long orbital tentacles (1.6 % – 2.1 % of SL). In addition, the mandibular pore formula of the species should be revised, the examined specimens having 3–4 + 2 + 3–4 (total 8–10 pores) [vs. 4–5 + 2 + 4–5 (total 10–12) in the original description]. Individual mandibular pore counts were as follows: 4 + 2 + 4 (
<typeStatus id="57C084E973BFE1BC3EE689260CE6DA1E" type="holotype">holotype</typeStatus>
and
<specimenCount id="A9DDC03A386BC2F2225CEF5560E62DD9" count="3" type="generic" typeStatus="paratype">3 paratypes</specimenCount>
); 3 + 2 + 3 (
<specimenCount id="0C1FF675A24321DF16208F8DE6169588" count="4" type="generic" typeStatus="paratype">4 paratypes</specimenCount>
and non-type Australian specimen); 4 + 1 + 4 (single
<typeStatus id="8798FC9A2804CE83728CC56BE89D7668" type="paratype">paratype</typeStatus>
). The coloration of the presently reported specimen was consistent with that of
<taxonomicName id="B9A27BAAC0517888C9E484BEA62786EA" authorityName="Fricke" authorityYear="1997" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="rhothion">
<emphasis id="637524113E78E82F5BB54A0B4A3FA1DA" italics="true">E. rhothion</emphasis>
</taxonomicName>
shown in
<bibRefCitation id="5A95D7981DF6A82D8757468B5BFFE48D" author="Fricke R" firstAuthor="Fricke" journalOrPublisher="Koeltz Scientific Books, Königstein, Germany" refId="B4" refString="Fricke R (1997) Tripterygiid fishes of the western and central Pacific (Teleostei). Koeltz Scientific Books, Königstein, Germany, 607 pp." title="Tripterygiid fishes of the western and central Pacific (Teleostei)." year="1997">Fricke (1997)</bibRefCitation>
: head, including snout, lips, cheek, and opercle, and pectoral-fin base black; lateral surface of body with faint bars; first and second dorsal fins black; anal fin pale (without markings). Although the color pattern of the caudal fin (dorsally and ventrally with faint vertical dark streaks;
<bibRefCitation id="987953A17EEE6EC20C5D1A3D95C7560E" author="Fricke" firstAuthor="Fricke" journalOrPublisher="Koeltz Scientific Books, Königstein, Germany" refId="B4" refString="Fricke R (1997) Tripterygiid fishes of the western and central Pacific (Teleostei). Koeltz Scientific Books, Königstein, Germany, 607 pp." title="Tripterygiid fishes of the western and central Pacific (Teleostei)." year="1997">Fricke 1997</bibRefCitation>
) is one of the diagnostic characters for the species, such pigmentation was lost in the presently reported specimen after long-term preservation (Fig.
<figureCitation id="0A4BE75797582B57EC0763464A7CD440" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Photographs of preserved specimen of Enneapterygius rhothion (A: AMS I. 5088, male, 32.5 mm SL, Australia; B: SMNS 18322, holotype, male, 28.9 mm SL, New Caledonia, Photo by M. Meguro)." figureDoi="10.3897/aiep.54.135448.figure1" httpUri="https://binary.pensoft.net/fig/1165072">1 A</figureCitation>
).
</paragraph>
<paragraph id="426EDA8FF7771F72892CADB9F4681EB1">
<taxonomicName id="9D03DF2A3C836F5FBF517905F1415583" authorityName="Fricke" authorityYear="1997" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="rhothion">
<emphasis id="B885FD81FA2523208C516344E4C5A4F2" italics="true">Enneapterygius rhothion</emphasis>
</taxonomicName>
was originally described based on
<specimenCount id="1B98A3FA3AA5050EBA71A55626177E97" count="129" type="generic">129 specimens</specimenCount>
from
<collectingCountry id="8C885DDA520A7E4053C050278AFB1A6C" name="New Caledonia">New Caledonia</collectingCountry>
and
<collectingCountry id="55A8F7B302B8AE07BDBA599FAB000AB9" name="Vanuatu">Vanuatu</collectingCountry>
. Subsequently,
<bibRefCitation id="32FF989AFE1D641D179DACC8C191BF6A" DOI="10.1023/A:1020076906430" author="Fricke R" firstAuthor="Fricke" journalOrPublisher="Environmental Biology of Fishes" pagination="175-198" refId="B6" refString="Fricke R (2002) Tripterygiid fishes of New Caledonia, with zoogeographical remarks. Environmental Biology of Fishes 65: 175–198. https://doi.org/10.1023/A:1020076906430" title="Tripterygiid fishes of New Caledonia, with zoogeographical remarks." volume="65" year="2002">Fricke (2002)</bibRefCitation>
reported the species from Ile des Pins as an additional record of the species from
<collectingCountry id="359ABF6563C320189C0A42C2928794B0" name="New Caledonia">New Caledonia</collectingCountry>
. In addition,
<bibRefCitation id="E66919B982A188EB7331500A21E4AFCA" author="Randall JE" firstAuthor="Randall" journalOrPublisher="University of Hawai’i Press, Honolulu, HI, USA" refId="B12" refString="Randall JE (2005) Reef and shore fishes of the South Pacific. New Caledonia to Tahiti and the Pitcairn Islands. University of Hawai’i Press, Honolulu, HI, USA, 720 pp." title="Reef and shore fishes of the South Pacific. New Caledonia to Tahiti and the Pitcairn Islands." year="2005">Randall (2005)</bibRefCitation>
showed a fresh photograph of a female individual of
<taxonomicName id="776523F95414F4A0986872925B72A681" authorityName="Fricke" authorityYear="1997" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="rhothion">
<emphasis id="492B9843E1AEB28C005A92E36C7A55DF" italics="true">E. rhothion</emphasis>
</taxonomicName>
in his field guide. The species is one of the common triplefins in New Caledonian waters (
<bibRefCitation id="ED2878F5159578D30990B3E79825969E" author="Fricke" firstAuthor="Fricke" journalOrPublisher="Koeltz Scientific Books, Königstein, Germany" refId="B4" refString="Fricke R (1997) Tripterygiid fishes of the western and central Pacific (Teleostei). Koeltz Scientific Books, Königstein, Germany, 607 pp." title="Tripterygiid fishes of the western and central Pacific (Teleostei)." year="1997">Fricke 1997</bibRefCitation>
,
<bibRefCitation id="56E04AE5B44F9D864B7F3CF559CB0258" DOI="10.1023/A:1020076906430" author="Fricke" firstAuthor="Fricke" journalOrPublisher="Environmental Biology of Fishes" pagination="175-198" refId="B6" refString="Fricke R (2002) Tripterygiid fishes of New Caledonia, with zoogeographical remarks. Environmental Biology of Fishes 65: 175–198. https://doi.org/10.1023/A:1020076906430" title="Tripterygiid fishes of New Caledonia, with zoogeographical remarks." volume="65" year="2002">2002</bibRefCitation>
). In his taxonomic works on triplefins,
<bibRefCitation id="DB0DA5C59C1DB8F4C1B8C0F138014025" author="Fricke" firstAuthor="Fricke" journalOrPublisher="Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie)" pagination="1-9" refId="B5" refString="Fricke R (2001) Enneapterygius trisignatus, a new species from northern Grande Terre, with a key to New Caledonian tripterygiid fishes (Teleostei). Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie) 630: 1–9." title="Enneapterygius trisignatus, a new species from northern Grande Terre, with a key to New Caledonian tripterygiid fishes (Teleostei)." volume="630" year="2001">Fricke (2001</bibRefCitation>
,
<bibRefCitation id="EAA67FA1985C68D6E3AC98F3955504FA" DOI="10.1023/A:1020076906430" author="Fricke" firstAuthor="Fricke" journalOrPublisher="Environmental Biology of Fishes" pagination="175-198" refId="B6" refString="Fricke R (2002) Tripterygiid fishes of New Caledonia, with zoogeographical remarks. Environmental Biology of Fishes 65: 175–198. https://doi.org/10.1023/A:1020076906430" title="Tripterygiid fishes of New Caledonia, with zoogeographical remarks." volume="65" year="2002">2002</bibRefCitation>
) provided identification keys to the tripterygiid species in New Caledonian waters. Following those keys, the presently reported Australian specimen (AMS I. 5088, male,
<quantity id="BF9137BF759391586216A7CD78A44332" metricMagnitude="-2" metricUnit="m" metricValue="3.25" unit="mm" value="32.5">32.5 mm</quantity>
SL) was identified as
<taxonomicName id="681F22869CD039FAC23AEA7A9E32B0DC" authority="(Waite, 1904)" baseAuthorityName="Waite" baseAuthorityYear="1904" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="rufopileus">
<emphasis id="B0272B521C8A26D13F6149186DC22FB9" italics="true">Enneapterygius rufopileus</emphasis>
(Waite, 1904)
</taxonomicName>
. However, the keys described the number of symphyseal mandibular pores and the coloration of the second dorsal fin of
<taxonomicName id="76DE0804823AE2581123AA44BB865EC6" authorityName="Fricke" authorityYear="1997" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="rhothion">
<emphasis id="3316CE8EB4CAA3F33742DB149FFA47AD" italics="true">E. rhothion</emphasis>
</taxonomicName>
erroneously [symphyseal mandibular pore 1, and second dorsal fin pale (rarely spotted) in males, respectively] (see
<bibRefCitation id="67A4B61E4DCBFD2642BDCA3C6F3E1AE6" author="Fricke" firstAuthor="Fricke" journalOrPublisher="Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie)" pagination="1-9" refId="B5" refString="Fricke R (2001) Enneapterygius trisignatus, a new species from northern Grande Terre, with a key to New Caledonian tripterygiid fishes (Teleostei). Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie) 630: 1–9." title="Enneapterygius trisignatus, a new species from northern Grande Terre, with a key to New Caledonian tripterygiid fishes (Teleostei)." volume="630" year="2001">Fricke 2001</bibRefCitation>
,
<bibRefCitation id="39A8A86AF20C0F9D210D73EB01E084C1" DOI="10.1023/A:1020076906430" author="Fricke" firstAuthor="Fricke" journalOrPublisher="Environmental Biology of Fishes" pagination="175-198" refId="B6" refString="Fricke R (2002) Tripterygiid fishes of New Caledonia, with zoogeographical remarks. Environmental Biology of Fishes 65: 175–198. https://doi.org/10.1023/A:1020076906430" title="Tripterygiid fishes of New Caledonia, with zoogeographical remarks." volume="65" year="2002">2002</bibRefCitation>
). Although
<taxonomicName id="FE30DE03BB5474BC7EB10741FB751957" authorityName="Fricke" authorityYear="1997" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="rhothion">
<emphasis id="0DA757C017EFB6175332C0A15A810D1B" italics="true">E. rhothion</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="01AB6F24B4104DE4FB81928A48F3D550" baseAuthorityName="Waite" baseAuthorityYear="1904" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="rufopileus">
<emphasis id="7E0D62E46C4D29E3879E34524894FE20" italics="true">E. rufopileus</emphasis>
</taxonomicName>
in scale counts and overall appearance, the former differs in having the following characters: symphyseal mandibular pore single (vs. double in
<taxonomicName id="8B19E26E226E51966E70EC2E613243F6" baseAuthorityName="Waite" baseAuthorityYear="1904" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="rufopileus">
<emphasis id="54998A8A890334685D13FB3E618A8CE5" italics="true">E. rufopileus</emphasis>
</taxonomicName>
), head (including maxillary) and pectoral-fin base entirely black (posterior two-thirds of jaws white, pectoral-fin base with black blotch), body with 7 bands (5 bands); first and second dorsal fin black in nuptial males (translucent white); third dorsal fin with black pigmentation in both sexes (not pigmented in either sex); dorsal and ventral parts of caudal fin with vertical streaks in both sexes (caudal fin pale in both sexes) (
<bibRefCitation id="61E745C7E71B770E664FB0EC45AC2085" author="Fricke" firstAuthor="Fricke" journalOrPublisher="Koeltz Scientific Books, Königstein, Germany" refId="B4" refString="Fricke R (1997) Tripterygiid fishes of the western and central Pacific (Teleostei). Koeltz Scientific Books, Königstein, Germany, 607 pp." title="Tripterygiid fishes of the western and central Pacific (Teleostei)." year="1997">Fricke 1997</bibRefCitation>
; this study).
</paragraph>
<paragraph id="DB09C1A587445FE6269F670D7D99C605">
<taxonomicName id="749BEBB8ACEED174BFE16AE0F269FA80" authorityName="Fricke" authorityYear="1997" family="Tripterygiidae" genus="Enneapterygius" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="rhothion">
<emphasis id="7C2DCB2F06D271DBB0607301A7DF907F" italics="true">Enneapterygius rhothion</emphasis>
</taxonomicName>
has previously been recorded only from
<collectingCountry id="988BA2BCBC599E40C3D48BDE5551B9C5" name="New Caledonia">New Caledonia</collectingCountry>
(Chesterfield Islands, Grande Terre, Ile des Pins, and Loyalty Islands) and
<collectingCountry id="7CB33D4005538C05A7AC5F1664571F8A" name="Vanuatu">Vanuatu</collectingCountry>
(Erromango Island), and was considered endemic to those areas (
<bibRefCitation id="B072155B0E7ACC54FBA5D641DEDCCD3C" author="Fricke" firstAuthor="Fricke" journalOrPublisher="Koeltz Scientific Books, Königstein, Germany" refId="B4" refString="Fricke R (1997) Tripterygiid fishes of the western and central Pacific (Teleostei). Koeltz Scientific Books, Königstein, Germany, 607 pp." title="Tripterygiid fishes of the western and central Pacific (Teleostei)." year="1997">Fricke 1997</bibRefCitation>
,
<bibRefCitation id="EE76F34902DA17F2B08B3FD2ADF9655B" DOI="10.1023/A:1020076906430" author="Fricke" firstAuthor="Fricke" journalOrPublisher="Environmental Biology of Fishes" pagination="175-198" refId="B6" refString="Fricke R (2002) Tripterygiid fishes of New Caledonia, with zoogeographical remarks. Environmental Biology of Fishes 65: 175–198. https://doi.org/10.1023/A:1020076906430" title="Tripterygiid fishes of New Caledonia, with zoogeographical remarks." volume="65" year="2002">2002</bibRefCitation>
). The presently reported specimen represents the first record of the species from
<collectingCountry id="D0B23EB141B6FFBA64F3C43EF5A669C2" name="Australia">Australia</collectingCountry>
, and the new northernmost record.
</paragraph>
</subSubSection>
</treatment>
</document>
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