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<document id="30E43C94B6CA7B2CD24FADDF073F08A7" ID-DOI="10.5281/zenodo.5161214" ID-ISSN="1942-1354" ID-Zenodo-Dep="5161214" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628116654208" checkinUser="felipe" docAuthor="Mawdsley, Jonathan R." docDate="2011" docId="4B0D3211FFBBFF90FF49FE9839512E5E" docLanguage="en" docName="InsectaMundi.0191.1-13.pdf" docOrigin="Insecta Mundi 2011 (191)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Chaetodera andriana" docType="treatment" docVersion="3" lastPageNumber="7" masterDocId="B7344A69FFB3FF98FF89FFFA3B7B2A65" masterDocTitle="Taxonomy, identification, and phylogeny of the African and Madagascan species of the tiger beetle genus Chaetodera Jeannel 1946 (Coleoptera: Cicindelidae)" masterLastPageNumber="13" masterPageNumber="1" pageNumber="7" updateTime="1699035013485" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="6D9E95787A7699B925C762804B38F5FE">Taxonomy, identification, and phylogeny of the African and Madagascan species of the tiger beetle genus Chaetodera Jeannel 1946 (Coleoptera: Cicindelidae)</mods:title>
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<mods:namePart id="9DFA681988C1D8ABB85F523A96D2287E">Mawdsley, Jonathan R.</mods:namePart>
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<mods:date id="ECA1BDBB685541EF9CC7BF2E96866B0E">2011</mods:date>
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<mods:number id="CC1769E8DA62B3E6D9F9B2DCD3050C0C">2011-09-02</mods:number>
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<mods:number id="F176737C40438ED5BCCBC4785DA667C4">2011</mods:number>
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<mods:number id="B39EA1527B9DFE0BC23965C762213D1D">191</mods:number>
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<mods:identifier id="FBA4A4AEAAFEBBC06A50B675F5904EB9" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="E3AF1E8C080BA7F849794BD3113F18AA" type="Zenodo-Dep">5161214</mods:identifier>
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<treatment id="4B0D3211FFBBFF90FF49FE9839512E5E" LSID="urn:lsid:plazi:treatment:4B0D3211FFBBFF90FF49FE9839512E5E" httpUri="http://treatment.plazi.org/id/4B0D3211FFBBFF90FF49FE9839512E5E" lastPageNumber="7" pageId="8" pageNumber="7">
<subSubSection id="8BBED08CFFBBFF90FF49FE9839CD2B1E" box="[192,694,354,379]" pageId="8" pageNumber="7" type="nomenclature">
<paragraph id="C31B8307FFBBFF90FF49FE9839CD2B1E" blockId="8.[192,694,354,411]" box="[192,694,354,379]" pageId="8" pageNumber="7">
<heading id="9853346BFFBBFF90FF49FE9839CD2B1E" bold="true" box="[192,694,354,379]" fontSize="10" level="2" pageId="8" pageNumber="7" reason="3">
<taxonomicName id="04A4F884FFBBFF90FF49FE9839CD2B1E" ID-CoL="TBP9" authority="(Alluaud 1900)" baseAuthorityName="Alluaud" baseAuthorityYear="1900" box="[192,694,354,379]" class="Insecta" family="Carabidae" genus="Chaetodera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="andriana">
<emphasis id="F1D05F15FFBBFF90FF49FE9839CD2B1E" bold="true" box="[192,694,354,379]" pageId="8" pageNumber="7">
<emphasis id="F1D05F15FFBBFF90FF49FE983A912B1E" bold="true" box="[192,490,354,379]" italics="true" pageId="8" pageNumber="7">Chaetodera andriana</emphasis>
(
<bibRefCitation id="A735FEF6FFBBFF90FE70FE9839D62B1E" author="Alluaud, C." box="[505,685,354,379]" pageId="8" pageNumber="7" pagination="509" refId="ref9544" refString="Alluaud, C. 1900. Liste des Insectes Coleopteres de la Region Malgache. p. vii + 509. In: A. Grandidier (ed.). Histoire Physique, Naturelle et Politique de Madagascar. Volume XXI. Histoire Naturelle des Coleopteres. I (l), p. vii + 509." type="book chapter" year="1900">Alluaud 1900</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8BBED08CFFBBFF90FF49FE793A052BFE" box="[192,382,387,411]" pageId="8" pageNumber="7" type="description">
<paragraph id="C31B8307FFBBFF90FF49FE793A052BFE" blockId="8.[192,694,354,411]" box="[192,382,387,411]" pageId="8" pageNumber="7">
<figureCitation id="5B9F9F82FFBBFF90FF49FE793A2F2BFE" box="[192,340,387,411]" captionStart="Figures 1-15" captionStartId="6.[192,286,1663,1685]" captionTargetBox="[334,1296,192,1635]" captionTargetId="figure-20@6.[334,1296,192,1635]" captionTargetPageId="6" captionText="Figures 1-15. Dorsal habitus images of Afrotropical Chaetodera species. 1) C. antatsima, syntype male, Madagascar Sud, Bassin du Mandraré. 2) C. antatsima, syntype female of C. antatsima var. fotsy, Mananpatra, Madagascar. 3) C. perrieri, male, Analaleva, Madagascar. 4) C. perrieri, syntype male, Madagascar. 5) C. andriana, syntype female, Madagascar (Sud), Bassin du Mandraré. 6) C. andriana, female, Ambositra, Madagascar. 7) C. maheva, syntype male, without locality label. 8) C. maheva, female, Majunga, Madagascar. 9) C. maheva, female, Maevatanana, Madagascar. 10) C. blanchardi, syntype male, Ouarsanguelis, Somalia. 11) C. regalis, male, Senegal. 12) C. regalis, female, Tana-Galla, Kenya. 13) C. regalis, paratype male of C. regalis bremeri Mandl, El Geneina, Darfur, Sudan. 14-15) Adult males of C. regalis photographed on sandbars of the Sabie River, Kruger National Park, South Africa." figureDoi="http://doi.org/10.5281/zenodo.5161216" httpUri="https://zenodo.org/record/5161216/files/figure.png" pageId="8" pageNumber="7">Figures 5, 6</figureCitation>
,
<figureCitation id="5B9F9F82FFBBFF90FEE9FE793A052BFE" box="[352,382,387,411]" captionStart="Figures 16-17" captionStartId="7.[192,289,891,913]" captionTargetBox="[244,1359,193,856]" captionTargetId="figure-234@7.[192,1440,192,876]" captionTargetPageId="7" captionText="Figures 16-17. Distribution of Chaetodera species in Africa and Madagascar. 16) Distribution of C. antatsima (dark circles), C. perrieri (dark triangles), and C. blanchardi (dark squares). 17) Distribution of C. andriana (open circles), C. maheva (open stars), and C. regalis (dark stars)." figureDoi="http://doi.org/10.5281/zenodo.5161218" httpUri="https://zenodo.org/record/5161218/files/figure.png" pageId="8" pageNumber="7">17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8BBED08CFFBBFF90FF49FE3939812BBE" box="[192,762,451,475]" pageId="8" pageNumber="7" type="reference_group">
<paragraph id="C31B8307FFBBFF90FF49FE3939812BBE" blockId="8.[192,762,451,475]" box="[192,762,451,475]" pageId="8" pageNumber="7">
<treatmentCitationGroup id="E3B4A429FFBBFF90FF49FE3939812BBE" box="[192,762,451,475]" pageId="8" pageNumber="7">
<treatmentCitation id="4205A516FFBBFF90FF49FE3939F12BBE" author="Alluaud, C." box="[192,650,451,475]" page="18" pageId="8" pageNumber="7" year="1900">
<taxonomicName id="04A4F884FFBBFF90FF49FE3939F12BBE" ID-CoL="V8L6" authority="Alluaud (1900: 18" authorityName="Alluaud" authorityYear="1900" box="[192,650,451,475]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="andriana">
<emphasis id="F1D05F15FFBBFF90FF49FE393AD62BBE" box="[192,429,451,475]" italics="true" pageId="8" pageNumber="7">Cicindela andriana</emphasis>
<bibRefCitation id="A735FEF6FFBBFF90FE3BFE3939F12BBE" author="Alluaud, C." box="[434,650,451,475]" pageId="8" pageNumber="7" pagination="509" refId="ref9544" refString="Alluaud, C. 1900. Liste des Insectes Coleopteres de la Region Malgache. p. vii + 509. In: A. Grandidier (ed.). Histoire Physique, Naturelle et Politique de Madagascar. Volume XXI. Histoire Naturelle des Coleopteres. I (l), p. vii + 509." type="book chapter" year="1900">Alluaud (1900: 18</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figure 2)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="8BBED08CFFBBFF90FF49FDF93A3F285E" pageId="8" pageNumber="7" type="materials_examined">
<paragraph id="C31B8307FFBBFF90FF49FDF93A3F285E" blockId="8.[192,1440,515,571]" pageId="8" pageNumber="7">
<emphasis id="F1D05F15FFBBFF90FF49FDF93AFB287E" bold="true" box="[192,384,515,539]" pageId="8" pageNumber="7">Type material.</emphasis>
<typeStatus id="1C1F3DA5FFBBFF90FE0CFDF93A9C287E" box="[389,487,515,539]" pageId="8" pageNumber="7" type="syntype">Syntype</typeStatus>
female, labeled “
<collectingCountry id="BBB3C397FFBBFF90FD22FDF93847287E" box="[683,828,515,539]" name="Madagascar" pageId="8" pageNumber="7">Madagascar</collectingCountry>
(Sud)/Bassin du Mandraré/
<bibRefCitation id="A735FEF6FFBBFF90FBF7FDF93E67287E" author="Alluaud, C." box="[1150,1308,515,539]" pageId="8" pageNumber="7" pagination="509" refId="ref9544" refString="Alluaud, C. 1900. Liste des Insectes Coleopteres de la Region Malgache. p. vii + 509. In: A. Grandidier (ed.). Histoire Physique, Naturelle et Politique de Madagascar. Volume XXI. Histoire Naturelle des Coleopteres. I (l), p. vii + 509." type="book chapter" year="1900">Alluaud 1900</bibRefCitation>
48” (
<collectionCode id="A5B51BC2FFBBFF90FADDFDF93EE0287E" box="[1364,1435,515,539]" pageId="8" pageNumber="7">DEIC</collectionCode>
, examined).
</paragraph>
</subSubSection>
<subSubSection id="8BBED08CFFBBFF90FF49FD99398B293E" pageId="8" pageNumber="7" type="diagnosis">
<paragraph id="C31B8307FFBBFF90FF49FD99398B293E" blockId="8.[192,1440,611,859]" pageId="8" pageNumber="7">
<emphasis id="F1D05F15FFBBFF90FF49FD993A2A281E" bold="true" box="[192,337,611,635]" pageId="8" pageNumber="7">Diagnosis.</emphasis>
Length 14.0-17.0 mm. Dorsal coloration as shown in
<figureCitation id="5B9F9F82FFBBFF90FC7BFD993FF2281E" box="[1010,1161,611,635]" captionStart="Figures 1-15" captionStartId="6.[192,286,1663,1685]" captionTargetBox="[334,1296,192,1635]" captionTargetId="figure-20@6.[334,1296,192,1635]" captionTargetPageId="6" captionText="Figures 1-15. Dorsal habitus images of Afrotropical Chaetodera species. 1) C. antatsima, syntype male, Madagascar Sud, Bassin du Mandraré. 2) C. antatsima, syntype female of C. antatsima var. fotsy, Mananpatra, Madagascar. 3) C. perrieri, male, Analaleva, Madagascar. 4) C. perrieri, syntype male, Madagascar. 5) C. andriana, syntype female, Madagascar (Sud), Bassin du Mandraré. 6) C. andriana, female, Ambositra, Madagascar. 7) C. maheva, syntype male, without locality label. 8) C. maheva, female, Majunga, Madagascar. 9) C. maheva, female, Maevatanana, Madagascar. 10) C. blanchardi, syntype male, Ouarsanguelis, Somalia. 11) C. regalis, male, Senegal. 12) C. regalis, female, Tana-Galla, Kenya. 13) C. regalis, paratype male of C. regalis bremeri Mandl, El Geneina, Darfur, Sudan. 14-15) Adult males of C. regalis photographed on sandbars of the Sabie River, Kruger National Park, South Africa." figureDoi="http://doi.org/10.5281/zenodo.5161216" httpUri="https://zenodo.org/record/5161216/files/figure.png" pageId="8" pageNumber="7">Figures 5, 6</figureCitation>
. This species is easily distinguished from the Madagascan
<taxonomicName id="04A4F884FFBBFF90FD0BFD7E387F28FE" baseAuthorityName="Kunckel d'Herculais" baseAuthorityYear="1887" box="[642,772,643,667]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="maheva">
<emphasis id="F1D05F15FFBBFF90FD0BFD7E387F28FE" box="[642,772,643,667]" italics="true" pageId="8" pageNumber="7">C. maheva</emphasis>
</taxonomicName>
by its broader yellow markings that run more directly across the width of the elytra (more transverse) than those of
<taxonomicName id="04A4F884FFBBFF90FC11FD5E3F6C28DE" baseAuthorityName="Kunckel d'Herculais" baseAuthorityYear="1887" box="[920,1047,675,699]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="maheva">
<emphasis id="F1D05F15FFBBFF90FC11FD5E3F6C28DE" box="[920,1047,675,699]" italics="true" pageId="8" pageNumber="7">C. maheva</emphasis>
</taxonomicName>
, particularly on the back (apical) half of the elytra. From the continental African
<taxonomicName id="04A4F884FFBBFF90FC8BFD3E380E28BE" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[770,885,707,731]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="regalis">
<emphasis id="F1D05F15FFBBFF90FC8BFD3E380E28BE" box="[770,885,707,731]" italics="true" pageId="8" pageNumber="7">C. regalis</emphasis>
</taxonomicName>
, it differs in having the dark elytral markings black when viewed directly from above (these markings are typically metallic blue in
<taxonomicName id="04A4F884FFBBFF90FB58FD1E3E3E289E" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[1233,1349,739,763]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="regalis">
<emphasis id="F1D05F15FFBBFF90FB58FD1E3E3E289E" box="[1233,1349,739,763]" italics="true" pageId="8" pageNumber="7">C. regalis</emphasis>
</taxonomicName>
, but on rare occasions can be black) and in having a much longer yellow marking along the basal portion of the elytral suture. The black elytral markings of
<taxonomicName id="04A4F884FFBBFF90FD62FCDE38FB295E" authorityName="Alluaud" authorityYear="1900" box="[747,896,803,827]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="andriana">
<emphasis id="F1D05F15FFBBFF90FD62FCDE38FB295E" box="[747,896,803,827]" italics="true" pageId="8" pageNumber="7">C. andriana</emphasis>
</taxonomicName>
occasionally will have a metallic blue sheen, especially when viewed obliquely (at an angle).
</paragraph>
</subSubSection>
<subSubSection id="8BBED08CFFBBFF90FF49FC7939512E5E" pageId="8" pageNumber="7" type="materials_examined">
<paragraph id="C31B8307FFBBFF90FF49FC7939512E5E" blockId="8.[192,1440,899,1083]" pageId="8" pageNumber="7">
<emphasis id="F1D05F15FFBBFF90FF49FC793AB429FE" bold="true" box="[192,463,899,923]" pageId="8" pageNumber="7">Material examined.</emphasis>
<materialsCitation id="73CC895AFFBBFF90FE5EFC793FA729FE" box="[471,1244,899,923]" collectionCode="DEIC, NMNH" country="Madagascar" location="Ambositra" pageId="8" pageNumber="7" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<collectingCountry id="BBB3C397FFBBFF90FE5EFC7939DF29FE" box="[471,676,899,923]" name="Madagascar" pageId="8" pageNumber="7">MADAGASCAR</collectingCountry>
:
<location id="C67BD5DCFFBBFF90FD26FC79384A29FE" LSID="urn:lsid:plazi:treatment:4B0D3211FFBBFF90FF49FE9839512E5E:C67BD5DCFFBBFF90FD26FC79384A29FE" box="[687,817,899,923]" country="Madagascar" name="Ambositra" pageId="8" pageNumber="7">Ambositra</location>
(
<specimenCount id="D5A2488EFFBBFF90FCCDFC7938E229FE" box="[836,921,899,923]" pageId="8" pageNumber="7" type="male">1 male</specimenCount>
,
<collectionCode id="A5B51BC2FFBBFF90FC2CFC79389429FE" box="[933,1007,899,923]" pageId="8" pageNumber="7">DEIC</collectionCode>
;
<specimenCount id="D5A2488EFFBBFF90FC72FC793F1329FE" box="[1019,1128,899,923]" pageId="8" pageNumber="7" type="female">1 female</specimenCount>
,
<collectionCode id="A5B51BC2FFBBFF90FBFCFC793FAF29FE" box="[1141,1236,899,923]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="7" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="73CC895AFFBBFF90FB60FC793A1529DE" collectionCode="AMNH" country="Madagascar" location="Anakazoabo" pageId="8" pageNumber="7" specimenCount="1" specimenCount-male="1">
<location id="C67BD5DCFFBBFF90FB60FC793EFB29FE" LSID="urn:lsid:plazi:treatment:4B0D3211FFBBFF90FF49FE9839512E5E:C67BD5DCFFBBFF90FB60FC793EFB29FE" box="[1257,1408,899,923]" country="Madagascar" name="Anakazoabo" pageId="8" pageNumber="7">Anakazoabo</location>
(
<specimenCount id="D5A2488EFFBBFF90FA1AFC793B8429DE" pageId="8" pageNumber="7" type="male">1 male</specimenCount>
,
<collectionCode id="A5B51BC2FFBBFF90FE83FC593A1C29DE" box="[266,359,931,955]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="8" pageNumber="7">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="73CC895AFFBBFF90FEF1FC5939A529DE" box="[376,734,931,955]" collectionCode="AMNH" country="Madagascar" location="Analalava" pageId="8" pageNumber="7" specimenCount="1" specimenCount-female="1">
<location id="C67BD5DCFFBBFF90FEF1FC593A8D29DE" LSID="urn:lsid:plazi:treatment:4B0D3211FFBBFF90FF49FE9839512E5E:C67BD5DCFFBBFF90FEF1FC593A8D29DE" box="[376,502,931,955]" country="Madagascar" name="Analalava" pageId="8" pageNumber="7">Analalava</location>
(
<specimenCount id="D5A2488EFFBBFF90FD81FC59390B29DE" box="[520,624,931,955]" pageId="8" pageNumber="7" type="female">1 female</specimenCount>
,
<collectionCode id="A5B51BC2FFBBFF90FDF3FC5939AC29DE" box="[634,727,931,955]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="8" pageNumber="7">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="73CC895AFFBBFF90FD61FC593F2C29DE" box="[744,1111,931,955]" collectionCode="CMNH" country="Madagascar" location="Ankazoabo" pageId="8" pageNumber="7" specimenCount="1" specimenCount-female="1">
<location id="C67BD5DCFFBBFF90FD61FC59380B29DE" LSID="urn:lsid:plazi:treatment:4B0D3211FFBBFF90FF49FE9839512E5E:C67BD5DCFFBBFF90FD61FC59380B29DE" box="[744,880,931,955]" country="Madagascar" name="Ankazoabo" pageId="8" pageNumber="7">Ankazoabo</location>
(
<specimenCount id="D5A2488EFFBBFF90FC08FC59389229DE" box="[897,1001,931,955]" pageId="8" pageNumber="7" type="female">1 female</specimenCount>
,
<collectionCode id="A5B51BC2FFBBFF90FC7AFC593F2B29DE" box="[1011,1104,931,955]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="8" pageNumber="7" type="Museum">CMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="73CC895AFFBBFF90FBEBFC593A6729BE" collectingDate="1987-01" collectionCode="TMSA" country="Madagascar" location="Antsalova" pageId="8" pageNumber="7" specimenCount="1" specimenCount-male="1">
<location id="C67BD5DCFFBBFF90FBEBFC593F9A29DE" LSID="urn:lsid:plazi:treatment:4B0D3211FFBBFF90FF49FE9839512E5E:C67BD5DCFFBBFF90FBEBFC593F9A29DE" box="[1122,1249,931,955]" country="Madagascar" name="Antsalova" pageId="8" pageNumber="7">Antsalova</location>
,
<date id="B71AA5C7FFBBFF90FB62FC593E4229DE" box="[1259,1337,931,955]" pageId="8" pageNumber="7" value="1987-01">
<collectingDate id="A75E5C2FFFBBFF90FB62FC593E4229DE" box="[1259,1337,931,955]" pageId="8" pageNumber="7" value="1987-01">I.1987</collectingDate>
</date>
(
<specimenCount id="D5A2488EFFBBFF90FAC3FC593EE029DE" box="[1354,1435,931,955]" pageId="8" pageNumber="7" type="male">1 male</specimenCount>
,
<collectionCode id="A5B51BC2FFBBFF90FF49FC393A6F29BE" box="[192,276,963,987]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="8" pageNumber="7">TMSA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="73CC895AFFBBFF90FEAFFC39390E29BE" box="[294,629,963,987]" collectionCode="CMNH" country="Madagascar" location="Befotaka" pageId="8" pageNumber="7" specimenCount="2" specimenCount-male="2">
<location id="C67BD5DCFFBBFF90FEAFFC393AED29BE" LSID="urn:lsid:plazi:treatment:4B0D3211FFBBFF90FF49FE9839512E5E:C67BD5DCFFBBFF90FEAFFC393AED29BE" box="[294,406,963,987]" country="Madagascar" name="Befotaka" pageId="8" pageNumber="7">Befotaka</location>
(
<specimenCount id="D5A2488EFFBBFF90FE2DFC39397D29BE" box="[420,518,963,987]" pageId="8" pageNumber="7" type="male">2 males</specimenCount>
,
<collectionCode id="A5B51BC2FFBBFF90FD99FC39391629BE" box="[528,621,963,987]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="8" pageNumber="7" type="Museum">CMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="73CC895AFFBBFF90FD09FC393F6329BE" box="[640,1048,963,987]" collectingDate="1933-01" collectionCode="DEIC" country="Madagascar" location="Mandrare" pageId="8" pageNumber="7" specimenCount="1" specimenCount-male="1">
<location id="C67BD5DCFFBBFF90FD09FC39398429BE" LSID="urn:lsid:plazi:treatment:4B0D3211FFBBFF90FF49FE9839512E5E:C67BD5DCFFBBFF90FD09FC39398429BE" box="[640,767,963,987]" country="Madagascar" name="Mandrare" pageId="8" pageNumber="7">Mandrare</location>
,
<date id="B71AA5C7FFBBFF90FC83FC39382329BE" box="[778,856,963,987]" pageId="8" pageNumber="7" value="1933-01">
<collectingDate id="A75E5C2FFFBBFF90FC83FC39382329BE" box="[778,856,963,987]" pageId="8" pageNumber="7" value="1933-01">I.1933</collectingDate>
</date>
(
<specimenCount id="D5A2488EFFBBFF90FCE0FC3938C729BE" box="[873,956,963,987]" pageId="8" pageNumber="7" type="male">1 male</specimenCount>
,
<collectionCode id="A5B51BC2FFBBFF90FC4FFC393F6B29BE" box="[966,1040,963,987]" pageId="8" pageNumber="7">DEIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="73CC895AFFBBFF90FBAAFC393A6B299E" collectingDate="1931-03" collectionCode="DEIC" country="Madagascar" location="Morondava" pageId="8" pageNumber="7" specimenCount="1" specimenCount-female="1">
<location id="C67BD5DCFFBBFF90FBAAFC393FC829BE" LSID="urn:lsid:plazi:treatment:4B0D3211FFBBFF90FF49FE9839512E5E:C67BD5DCFFBBFF90FBAAFC393FC829BE" box="[1059,1203,963,987]" country="Madagascar" name="Morondava" pageId="8" pageNumber="7">Morondava</location>
,
<date id="B71AA5C7FFBBFF90FB34FC393E5929BE" box="[1213,1314,963,987]" pageId="8" pageNumber="7" value="1931-03">
<collectingDate id="A75E5C2FFFBBFF90FB34FC393E5929BE" box="[1213,1314,963,987]" pageId="8" pageNumber="7" value="1931-03">III.1931</collectingDate>
</date>
(
<specimenCount id="D5A2488EFFBBFF90FABAFC393EE029BE" box="[1331,1435,963,987]" pageId="8" pageNumber="7" type="female">1 female</specimenCount>
,
<collectionCode id="A5B51BC2FFBBFF90FF49FC193A73299E" box="[192,264,995,1019]" pageId="8" pageNumber="7">DEIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="73CC895AFFBBFF90FE93FC1938AA299E" box="[282,977,995,1020]" collectionCode="DEIC" country="Madagascar" location="Plateau de l'Andray - Reg. d'Ambovombe" pageId="8" pageNumber="7" specimenCount="1" specimenCount-female="1">
<location id="C67BD5DCFFBBFF90FE93FC193878299E" LSID="urn:lsid:plazi:treatment:4B0D3211FFBBFF90FF49FE9839512E5E:C67BD5DCFFBBFF90FE93FC193878299E" box="[282,771,995,1020]" country="Madagascar" name="Plateau de l'Andray - Reg. d'Ambovombe" pageId="8" pageNumber="7">Plateau de lAndray Reg. dAmbovombe</location>
(
<specimenCount id="D5A2488EFFBBFF90FC9BFC193803299E" box="[786,888,995,1019]" pageId="8" pageNumber="7" type="female">1 female</specimenCount>
,
<collectionCode id="A5B51BC2FFBBFF90FC08FC1938B2299E" box="[897,969,995,1019]" pageId="8" pageNumber="7">DEIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="73CC895AFFBBFF90FC52FC1938BE2E7E" collectingDate="1925-05" collectionCode="CMNH, DEIC, V" country="Madagascar" location="Tananarive" pageId="8" pageNumber="7" specimenCount="5" specimenCount-female="3" specimenCount-male="2">
<location id="C67BD5DCFFBBFF90FC52FC193F1C299E" LSID="urn:lsid:plazi:treatment:4B0D3211FFBBFF90FF49FE9839512E5E:C67BD5DCFFBBFF90FC52FC193F1C299E" box="[987,1127,995,1019]" country="Madagascar" name="Tananarive" pageId="8" pageNumber="7">Tananarive</location>
(
<specimenCount id="D5A2488EFFBBFF90FBFEFC193FA6299E" box="[1143,1245,995,1019]" pageId="8" pageNumber="7" type="female">1 female</specimenCount>
,
<collectionCode id="A5B51BC2FFBBFF90FB6FFC193E3B299E" box="[1254,1344,995,1019]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="8" pageNumber="7" type="Museum">CMNH</collectionCode>
;
<specimenCount id="D5A2488EFFBBFF90FAC3FC193EE0299E" box="[1354,1435,995,1019]" pageId="8" pageNumber="7" type="male">1 male</specimenCount>
,
<specimenCount id="D5A2488EFFBBFF90FF49FBF93A522E7E" box="[192,297,1027,1051]" pageId="8" pageNumber="7" type="female">1 female</specimenCount>
,
<collectionCode id="A5B51BC2FFBBFF90FEBBFBF93A002E7E" box="[306,379,1027,1051]" pageId="8" pageNumber="7">DEIC</collectionCode>
), 1924 (
<specimenCount id="D5A2488EFFBBFF90FE51FBF939442E7E" box="[472,575,1027,1051]" pageId="8" pageNumber="7" type="female">1 female</specimenCount>
,
<collectionCode id="A5B51BC2FFBBFF90FDC1FBF939EA2E7E" box="[584,657,1027,1051]" pageId="8" pageNumber="7">DEIC</collectionCode>
),
<date id="B71AA5C7FFBBFF90FD2BFBF9398C2E7E" box="[674,759,1027,1051]" pageId="8" pageNumber="7" value="1925-05">
<collectingDate id="A75E5C2FFFBBFF90FD2BFBF9398C2E7E" box="[674,759,1027,1051]" pageId="8" pageNumber="7" value="1925-05">
<collectionCode id="A5B51BC2FFBBFF90FD2BFBF939C32E7E" box="[674,696,1027,1051]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="7" type="Museum">V</collectionCode>
.1925
</collectingDate>
</date>
(
<specimenCount id="D5A2488EFFBBFF90FC81FBF9382C2E7E" box="[776,855,1027,1051]" pageId="8" pageNumber="7" type="male">1 male</specimenCount>
,
<collectionCode id="A5B51BC2FFBBFF90FCEBFBF938C62E7E" box="[866,957,1027,1051]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="8" pageNumber="7" type="Museum">CMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="73CC895AFFBBFF90FC46FBF9395D2E5E" collectionCode="DEIC" country="Madagascar" location="Tulear" pageId="8" pageNumber="7" specimenCount="5" specimenCount-female="1" specimenCount-male="4">
<location id="C67BD5DCFFBBFF90FC46FBF93F582E7E" LSID="urn:lsid:plazi:treatment:4B0D3211FFBBFF90FF49FE9839512E5E:C67BD5DCFFBBFF90FC46FBF93F582E7E" box="[975,1059,1027,1051]" country="Madagascar" name="Tulear" pageId="8" pageNumber="7">Tuléar</location>
, 1930 (
<specimenCount id="D5A2488EFFBBFF90FBFFFBF93FA32E7E" box="[1142,1240,1027,1051]" pageId="8" pageNumber="7" type="male">3 males</specimenCount>
,
<specimenCount id="D5A2488EFFBBFF90FB54FBF93E3A2E7E" box="[1245,1345,1027,1051]" pageId="8" pageNumber="7" type="female">1 female</specimenCount>
,
<collectionCode id="A5B51BC2FFBBFF90FAC3FBF93EEF2E7E" box="[1354,1428,1027,1051]" pageId="8" pageNumber="7">DEIC</collectionCode>
). “Madagascar” (
<specimenCount id="D5A2488EFFBBFF90FEF3FBD93AB02E5E" box="[378,459,1059,1083]" pageId="8" pageNumber="7" type="male">1 male</specimenCount>
,
<collectionCode id="A5B51BC2FFBBFF90FE5DFBD939652E5E" box="[468,542,1059,1083]" pageId="8" pageNumber="7">DEIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>