189 lines
19 KiB
XML
189 lines
19 KiB
XML
<document id="5BCDA0462DD232D2F32E4AD976EC4B08" ID="10.11646/zootaxa.4178.3.7" ID-DOI="10.11646/zootaxa.4178.3.7" ID-GBIF-Dataset="cc051f0e-8ad3-4da6-bfff-6034096af9d1" ID-ISSN="1175-5326" ID-Zenodo-Dep="272510" ID-ZooBank="5D757FFE-7367-47A4-9523-9A0CEA3B2763" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1477477146551" checkinUser="plazi" docAuthor="Marusik, Yuri M. & Fomichev, Alexander A." docDate="2016" docId="4B072842FFFB2B631DAB3BBAFC06FA17" docLanguage="en" docName="zootaxa.4178.3.7.pdf" docOrigin="Zootaxa 4178 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Synaphosus makhambetensis Ponomarev 2008" docType="treatment" docVersion="9" lastPageNumber="437" masterDocId="B73E503AFFF22B6A1D3C387CFFF9FFE2" masterDocTitle="A survey of East Palaearctic Gnaphosidae (Araneae). 5. On Synaphosus from Central Asia" masterLastPageNumber="442" masterPageNumber="428" pageNumber="437" updateTime="1698390873933" updateUser="plazi">
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<mods:titleInfo id="B51F7EB34D1B14106FD4A8A333E8D4C3">
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<mods:title id="9B1A2822AEE35FFAA2392464534CB17C">A survey of East Palaearctic Gnaphosidae (Araneae). 5. On Synaphosus from Central Asia</mods:title>
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<mods:namePart id="FEABC4A2CA8A00446833185248F4694E">Marusik, Yuri M.</mods:namePart>
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<mods:name id="FD88E006A15094D0D19C6D7D28C09D2D" type="personal">
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<mods:roleTerm id="108094EE6D186669FF2F5807FB80B73A">Author</mods:roleTerm>
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<mods:namePart id="735986B586FD8BBC50693386B2930EB5">Fomichev, Alexander A.</mods:namePart>
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<mods:typeOfResource id="B667FFC877AA5901FA8014BAB65AB7AE">text</mods:typeOfResource>
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<mods:title id="EE5740DEAA42B82BDFC9103A18BD1230">Zootaxa</mods:title>
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<mods:date id="6003848415DD1474522B15E6FE3A5C38">2016</mods:date>
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<mods:number id="29552BBB1647B265B6F20E3BCFA33619">4178</mods:number>
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<mods:identifier id="234874BAE6C51C87C6216999032170CB" type="DOI">10.11646/zootaxa.4178.3.7</mods:identifier>
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<treatment id="4B072842FFFB2B631DAB3BBAFC06FA17" ID-DOI="http://doi.org/10.5281/zenodo.5671449" ID-GBIF-Taxon="124558200" ID-Zenodo-Dep="5671449" LSID="urn:lsid:plazi:treatment:4B072842FFFB2B631DAB3BBAFC06FA17" httpUri="http://treatment.plazi.org/id/4B072842FFFB2B631DAB3BBAFC06FA17" lastPageNumber="437" pageId="9" pageNumber="437">
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<subSubSection id="8BB4CADFFFFB2B631DAB3BBAFEC9FBE0" pageId="9" pageNumber="437" type="nomenclature">
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<paragraph id="C3119954FFFB2B631DAB3BBAFD2EFC02" blockId="9.[151,727,966,1026]" box="[151,727,966,992]" pageId="9" pageNumber="437">
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<heading id="98592E38FFFB2B631DAB3BBAFD2EFC02" bold="true" box="[151,727,966,992]" fontSize="11" level="1" pageId="9" pageNumber="437" reason="1">
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<taxonomicName id="04AEE2D7FFFB2B631DAB3BBAFD2EFC02" ID-CoL="53TRW" authority="Ponomarev, 2008" authorityName="Ponomarev" authorityYear="2008" box="[151,727,966,992]" class="Arachnida" family="Gnaphosidae" genus="Synaphosus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="437" phylum="Arthropoda" rank="species" species="makhambetensis">
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<emphasis id="F1DA4546FFFB2B631DAB3BBAFD2EFC02" bold="true" box="[151,727,966,992]" pageId="9" pageNumber="437">
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<emphasis id="F1DA4546FFFB2B631DAB3BBAFE00FC02" bold="true" box="[151,505,966,992]" italics="true" pageId="9" pageNumber="437">Synaphosus makhambetensis</emphasis>
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Ponomarev, 2008
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</emphasis>
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||
</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="C3119954FFFB2B631DAB3B95FEC9FBE0" blockId="9.[151,727,966,1026]" box="[151,304,1001,1026]" pageId="9" pageNumber="437">
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<figureCitation id="5B9585D1FFFB2B631DAB3B95FF27FBE0" box="[151,222,1001,1026]" captionStart="FIGURES 1 ‒ 6" captionStartId="3.[151,264,1789,1812]" captionTargetBox="[151,1436,473,1768]" captionTargetId="figure@3.[151,1436,470,1768]" captionTargetPageId="3" captionText="FIGURES 1 ‒ 6. Dorsal view of Synaphosus ovtsharenkoi sp. n. (1), S. mongolicus sp. n. (2), S. shmakovi sp. n. (3), S. palearcticus (4, from Tajikistan), S. saidovi sp. n. (5) and S. makhambetensis (6). 1 a, 4 a male; 1 b, 2 ‒ 3, 4 b, 5 ‒ 6 female. Scale = 0.5 mm." httpUri="https://zenodo.org/record/272511/files/figure.png" pageId="9" pageNumber="437">Figs 6</figureCitation>
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,
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<figureCitation id="5B9585D1FFFB2B631DD63B96FEF0FBE0" box="[234,265,1002,1026]" captionStart="FIGURES 31 ‒ 35" captionStartId="8.[151,264,1201,1224]" captionTargetBox="[151,1435,193,1179]" captionTargetId="figure@8.[151,1436,193,1180]" captionTargetPageId="8" captionText="FIGURES 31 ‒ 35. Epigynes of Synaphosus turanicus (31 ‒ 33), S. makhambetensis (34) and S. mongolicus sp. n. (35). 31, 34 intact, ventral; 32 ‒ 33, 35 b after maceration, dorsal; 35 a after maceration, ventral. Scale = 0.2 mm. Abbreviations: Ar anterior pair of receptacles, Cg club-like glands, Sm " sea mine " - like structure" httpUri="https://zenodo.org/record/272515/files/figure.png" pageId="9" pageNumber="437">34</figureCitation>
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,
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<figureCitation id="5B9585D1FFFB2B631C283B95FEC9FBE0" box="[276,304,1001,1026]" captionStart="FIGURE 51" captionStartId="14.[151,250,957,979]" captionTargetBox="[168,1436,308,916]" captionTargetId="figure@14.[168,1436,308,916]" captionTargetPageId="14" captionText="FIGURE 51. Collecting localities of specimens examined in this study of seven Synaphosus species." httpUri="https://zenodo.org/record/272518/files/figure.png" pageId="9" pageNumber="437">51</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="8BB4CADFFFFB2B631DAB3C4DFCE9FBA5" box="[151,784,1073,1095]" pageId="9" pageNumber="437" type="reference_group">
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||
<paragraph id="C3119954FFFB2B631DAB3C4DFCE9FBA5" blockId="9.[151,784,1073,1095]" box="[151,784,1073,1095]" pageId="9" pageNumber="437">
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<taxonomicName id="04AEE2D7FFFB2B631DAB3C4DFD5EFBA5" ID-CoL="53TRW" authority="Ponomarev, 2008a: 58" authorityName="Ponomarev" authorityPageNumber="58" authorityYear="2008" box="[151,679,1073,1095]" class="Arachnida" family="Gnaphosidae" genus="Synaphosus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="437" phylum="Arthropoda" rank="species" species="makhambetensis">
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<emphasis id="F1DA4546FFFB2B631DAB3C4DFE45FBA5" box="[151,444,1073,1095]" italics="true" pageId="9" pageNumber="437">Synaphosus makhambetensis</emphasis>
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Ponomarev, 2008a: 58
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</taxonomicName>
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, f. 30 (
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<specimenCount id="D5A852DDFFFB2B631FCD3C4DFCFBFBA5" box="[753,770,1073,1095]" pageId="9" pageNumber="437" type="female">♀</specimenCount>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="8BB4CADFFFFB2B631DAB3C08FD97FB52" pageId="9" pageNumber="437" type="materials_examined">
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||
<paragraph id="C3119954FFFB2B631DAB3C08FD97FB52" blockId="9.[151,1436,1140,1525]" pageId="9" pageNumber="437">
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<materialsCitation id="73C69309FFFB2B631DAB3C08FD97FB52" ID-GBIF-Occurrence="1324201451" collectingDate="1978-07" collectionCode="ZMMU" collectorName="F. A. Saraev" country="Kazakhstan" location="Ural River" pageId="9" pageNumber="437" specimenCount="1" specimenCount-female="1" stateProvince="Atyrau Province" typeStatus="holotype">
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<emphasis id="F1DA4546FFFB2B631DAB3C08FE78FB6F" bold="true" box="[151,385,1140,1165]" pageId="9" pageNumber="437">Material examined.</emphasis>
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<typeStatus id="1C1527F6FFFB2B631CB43C08FE09FB6F" box="[392,496,1140,1165]" pageId="9" pageNumber="437" type="holotype">Holotype</typeStatus>
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<specimenCount id="D5A852DDFFFB2B631CCB3C09FDF5FB6E" box="[503,524,1141,1164]" pageId="9" pageNumber="437" type="female">♀</specimenCount>
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||
(
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<collectionCode id="A5BF0191FFFB2B631F273C08FD81FB6E" box="[539,632,1140,1164]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="9" pageNumber="437">ZMMU</collectionCode>
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),
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<collectingCountry id="BBB9D9C4FFFB2B631FB03C08FCB5FB6E" box="[652,844,1140,1164]" name="Kazakhstan" pageId="9" pageNumber="437">KAZAKHSTAN</collectingCountry>
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,
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<collectingRegion id="016A57B6FFFB2B631E6B3C08FBEAFB6F" box="[855,1043,1140,1165]" country="Kazakhstan" name="Atyrau Oblysy" pageId="9" pageNumber="437">
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<emphasis id="F1DA4546FFFB2B631E6B3C08FC5EFB6F" bold="true" box="[855,935,1140,1165]" italics="true" pageId="9" pageNumber="437">Atyrau</emphasis>
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Province
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</collectingRegion>
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, environs of Makhambet Vil.,
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<location id="C671CF8FFFFB2B6318563C08FF2CFB52" LSID="urn:lsid:plazi:treatment:4B072842FFFB2B631DAB3BBAFC06FA17:C671CF8FFFFB2B6318563C08FF2CFB52" country="Kazakhstan" name="Ural River" pageId="9" pageNumber="437" stateProvince="Atyrau Province">Ural River</location>
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right bank,
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<collectingDate id="A754467CFFFB2B631C603CEBFE33FB52" box="[348,458,1175,1200]" pageId="9" pageNumber="437" value="1978-07">
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<date id="B710BF94FFFB2B631C603CEBFE33FB52" box="[348,458,1175,1200]" pageId="9" pageNumber="437" value="1978-07">July 1978</date>
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</collectingDate>
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(
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<collectorName id="6E5BFC82FFFB2B631CE63CE4FD9BFB52" box="[474,610,1176,1200]" pageId="9" pageNumber="437">F.A. Saraev</collectorName>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="8BB4CADFFFFB2B631DFB3CC0FE5EFAA2" pageId="9" pageNumber="437" type="diagnosis">
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<paragraph id="C3119954FFFB2B631DFB3CC0FE5EFAA2" blockId="9.[151,1436,1140,1525]" pageId="9" pageNumber="437">
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<emphasis id="F1DA4546FFFB2B631DFB3CC0FEC3FB37" bold="true" box="[199,314,1212,1237]" pageId="9" pageNumber="437">Diagnosis</emphasis>
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. This species is similar to
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<taxonomicName id="04AEE2D7FFFB2B631F5C3CC0FD1CFB36" box="[608,741,1212,1236]" class="Arachnida" family="Gnaphosidae" genus="Synaphosus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="437" phylum="Arthropoda" rank="species" species="shmakovi" status="sp. nov.">
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<emphasis id="F1DA4546FFFB2B631F5C3CC0FD1CFB36" box="[608,741,1212,1236]" italics="true" pageId="9" pageNumber="437">S. shmakovi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="EAE9F83DFFFB2B631FD73CC0FCD3FB37" box="[747,810,1212,1237]" pageId="9" pageNumber="437" rank="species">
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<emphasis id="F1DA4546FFFB2B631FD73CC0FCD3FB37" bold="true" box="[747,810,1212,1237]" pageId="9" pageNumber="437">sp. n.</emphasis>
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</taxonomicNameLabel>
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and
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<taxonomicName id="04AEE2D7FFFB2B631E623CC0FC02FB36" box="[862,1019,1212,1236]" class="Arachnida" family="Gnaphosidae" genus="Synaphosus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="437" phylum="Arthropoda" rank="species" species="mongolicus" status="sp. nov.">
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<emphasis id="F1DA4546FFFB2B631E623CC0FC02FB36" box="[862,1019,1212,1236]" italics="true" pageId="9" pageNumber="437">S. mongolicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="EAE9F83DFFFB2B63193D3CC0FBB9FB37" box="[1025,1088,1212,1237]" pageId="9" pageNumber="437" rank="species">
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<emphasis id="F1DA4546FFFB2B63193D3CC0FBB9FB37" bold="true" box="[1025,1088,1212,1237]" pageId="9" pageNumber="437">sp. n.</emphasis>
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</taxonomicNameLabel>
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; it differs from
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<taxonomicName id="04AEE2D7FFFB2B6319D33CC0FA8DFB36" box="[1263,1396,1212,1236]" class="Arachnida" family="Gnaphosidae" genus="Synaphosus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="437" phylum="Arthropoda" rank="species" species="shmakovi" status="sp. nov.">
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<emphasis id="F1DA4546FFFB2B6319D33CC0FA8DFB36" box="[1263,1396,1212,1236]" italics="true" pageId="9" pageNumber="437">S. shmakovi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="EAE9F83DFFFB2B6318463CC0FF57FB1A" pageId="9" pageNumber="437" rank="species">
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<emphasis id="F1DA4546FFFB2B6318463CC0FF57FB1A" bold="true" pageId="9" pageNumber="437">sp. n.</emphasis>
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</taxonomicNameLabel>
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by the pocket thinner than the receptacle (pocket wider than receptacle in
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<taxonomicName id="04AEE2D7FFFB2B6319103C9CFB4EFB1A" box="[1068,1207,1248,1272]" class="Arachnida" family="Gnaphosidae" genus="Synaphosus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="437" phylum="Arthropoda" rank="species" species="shmakovi" status="sp. nov.">
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<emphasis id="F1DA4546FFFB2B6319103C9CFB4EFB1A" box="[1068,1207,1248,1272]" italics="true" pageId="9" pageNumber="437">S. shmakovi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="EAE9F83DFFFB2B6319FE3CA3FAFFFB1A" box="[1218,1286,1247,1272]" pageId="9" pageNumber="437" rank="species">
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<emphasis id="F1DA4546FFFB2B6319FE3CA3FAFFFB1A" bold="true" box="[1218,1286,1247,1272]" pageId="9" pageNumber="437">sp. n.</emphasis>
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</taxonomicNameLabel>
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) and can be separated from
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<taxonomicName id="04AEE2D7FFFB2B631C7B3D78FE1CFAFE" box="[327,485,1284,1308]" class="Arachnida" family="Gnaphosidae" genus="Synaphosus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="437" phylum="Arthropoda" rank="species" species="mongolicus" status="sp. nov.">
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<emphasis id="F1DA4546FFFB2B631C7B3D78FE1CFAFE" box="[327,485,1284,1308]" italics="true" pageId="9" pageNumber="437">S. mongolicus</emphasis>
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</taxonomicName>
|
||
<taxonomicNameLabel id="EAE9F83DFFFB2B631CD13D78FDD7FAFF" box="[493,558,1284,1309]" pageId="9" pageNumber="437" rank="species">
|
||
<emphasis id="F1DA4546FFFB2B631CD13D78FDD7FAFF" bold="true" box="[493,558,1284,1309]" pageId="9" pageNumber="437">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
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by the anterior loops not extending beyond pocket (extending beyond pocket in
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<taxonomicName id="04AEE2D7FFFB2B631D883D54FEA8FAA2" box="[180,337,1320,1344]" class="Arachnida" family="Gnaphosidae" genus="Synaphosus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="437" phylum="Arthropoda" rank="species" species="mongolicus" status="sp. nov.">
|
||
<emphasis id="F1DA4546FFFB2B631D883D54FEA8FAA2" box="[180,337,1320,1344]" italics="true" pageId="9" pageNumber="437">S. mongolicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EAE9F83DFFFB2B631C643D5BFE6EFAA2" box="[344,407,1319,1344]" pageId="9" pageNumber="437" rank="species">
|
||
<emphasis id="F1DA4546FFFB2B631C643D5BFE6EFAA2" bold="true" box="[344,407,1319,1344]" pageId="9" pageNumber="437">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8BB4CADFFFFB2B631DFB3D30FD2AFA86" box="[199,723,1356,1381]" pageId="9" pageNumber="437" type="description">
|
||
<paragraph id="C3119954FFFB2B631DFB3D30FD2AFA86" blockId="9.[151,1436,1140,1525]" box="[199,723,1356,1381]" pageId="9" pageNumber="437">
|
||
<emphasis id="F1DA4546FFFB2B631DFB3D30FEA8FA87" bold="true" box="[199,337,1356,1381]" pageId="9" pageNumber="437">Description</emphasis>
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||
. Described by Ponomarev (2008).
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</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="8BB4CADFFFFB2B631DFB3D13FACBFA32" pageId="9" pageNumber="437" type="discussion">
|
||
<paragraph id="C3119954FFFB2B631DFB3D13FACBFA32" blockId="9.[151,1436,1140,1525]" pageId="9" pageNumber="437">
|
||
<emphasis id="F1DA4546FFFB2B631DFB3D13FEB1FA6A" bold="true" box="[199,328,1391,1416]" pageId="9" pageNumber="437">Comments</emphasis>
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. The species is known by a
|
||
<typeStatus id="1C1527F6FFFB2B631F9D3D13FCFAFA6A" box="[673,771,1391,1416]" pageId="9" pageNumber="437" type="holotype">holotype</typeStatus>
|
||
female and was never revised since the description. We provide figures to demonstrate differences with the two sibling new species
|
||
<taxonomicName id="04AEE2D7FFFB2B631ECA3DE8FB87FA4E" box="[1014,1150,1428,1452]" class="Arachnida" family="Gnaphosidae" genus="Synaphosus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="437" phylum="Arthropoda" rank="species" species="shmakovi" status="sp. nov.">
|
||
<emphasis id="F1DA4546FFFB2B631ECA3DE8FB87FA4E" box="[1014,1150,1428,1452]" italics="true" pageId="9" pageNumber="437">S. shmakovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EAE9F83DFFFB2B6319B93DE8FB3FFA4F" box="[1157,1222,1428,1453]" pageId="9" pageNumber="437" rank="species">
|
||
<emphasis id="F1DA4546FFFB2B6319B93DE8FB3FFA4F" bold="true" box="[1157,1222,1428,1453]" pageId="9" pageNumber="437">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="04AEE2D7FFFB2B6319C23DE8FA65FA4E" box="[1278,1436,1428,1452]" class="Arachnida" family="Gnaphosidae" genus="Synaphosus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="437" phylum="Arthropoda" rank="species" species="mongolicus" status="sp. nov.">
|
||
<emphasis id="F1DA4546FFFB2B6319C23DE8FA65FA4E" box="[1278,1436,1428,1452]" italics="true" pageId="9" pageNumber="437">S. mongolicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EAE9F83DFFFB2B631DAB3DCBFF2FFA32" box="[151,214,1463,1488]" pageId="9" pageNumber="437" rank="species">
|
||
<emphasis id="F1DA4546FFFB2B631DAB3DCBFF2FFA32" bold="true" box="[151,214,1463,1488]" pageId="9" pageNumber="437">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
It is worth mentioning that original figure of the epigyne is very schematic and provides no details.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8BB4CADFFFFB2B631DFB3DA0FC06FA17" box="[199,1023,1500,1525]" pageId="9" pageNumber="437" type="distribution">
|
||
<paragraph id="C3119954FFFB2B631DFB3DA0FC06FA17" blockId="9.[151,1436,1140,1525]" box="[199,1023,1500,1525]" pageId="9" pageNumber="437">
|
||
<emphasis id="F1DA4546FFFB2B631DFB3DA0FEA0FA17" bold="true" box="[199,345,1500,1525]" pageId="9" pageNumber="437">Distribution</emphasis>
|
||
. This species is known from the
|
||
<typeStatus id="1C1527F6FFFB2B631FF73DA1FD02FA17" box="[715,763,1501,1525]" pageId="9" pageNumber="437">type</typeStatus>
|
||
locality only (
|
||
<figureCitation id="5B9585D1FFFB2B631EA33DA0FC0BFA17" box="[927,1010,1500,1525]" captionStart="FIGURE 51" captionStartId="14.[151,250,957,979]" captionTargetBox="[168,1436,308,916]" captionTargetId="figure@14.[168,1436,308,916]" captionTargetPageId="14" captionText="FIGURE 51. Collecting localities of specimens examined in this study of seven Synaphosus species." httpUri="https://zenodo.org/record/272518/files/figure.png" pageId="9" pageNumber="437">Fig. 51</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |