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<document id="151CE8E93105D78CA8A2F624BF99437D" ID-DOI="10.5852/ejt.2023.915.2379" ID-ISSN="2118-9773" ID-Zenodo-Dep="10410126" ID-ZooBank="78346F83-1177-4EAE-AFFD-D4DDCC1BFB83" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1703073859257" checkinUser="plazi" docAuthor="Balkenohl, Michael" docDate="2023" docId="4A6487ECCF17FFEDFD9AFDE7FC742805" docLanguage="en" docName="EJT.2023.915.1-57.pdf" docOrigin="European Journal of Taxonomy 915 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2379/10451" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Sulciclivina mikirensis Balkenohl 2023, sp. nov." docType="treatment" docUuid="32AC25F6-7416-475D-A8D7-8942944D531B" docUuidSource="ZooBank" docVersion="2" lastPageNumber="19" masterDocId="B65DFF94CF07FFFFFFC5FFC1FF8F2A5D" masterDocTitle="Revision of the genus Sulciclivina Balkenohl, 2022 from Asia (Coleoptera, Carabidae, Clivinini)" masterLastPageNumber="57" masterPageNumber="1" pageNumber="17" updateTime="1703074388710" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="189EA4F17E65D650E75B128A591C4FFD">Revision of the genus Sulciclivina Balkenohl, 2022 from Asia (Coleoptera, Carabidae, Clivinini)</mods:title>
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<mods:namePart id="F9BB1CD9A121CDEC601DCA2756773CC3">Balkenohl, Michael</mods:namePart>
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<mods:title id="39EF21EEC853A7E385AE3DE1FDD1C3A7">European Journal of Taxonomy</mods:title>
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<mods:date id="B50AF52C5FE84729BCA31BCA4A397F3D">2023</mods:date>
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<treatment id="4A6487ECCF17FFEDFD9AFDE7FC742805" ID-DOI="http://doi.org/10.5281/zenodo.10410134" ID-Zenodo-Dep="10410134" LSID="urn:lsid:zoobank.org:act:32AC25F6-7416-475D-A8D7-8942944D531B" httpUri="http://treatment.plazi.org/id/4A6487ECCF17FFEDFD9AFDE7FC742805" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
<subSubSection id="8AD76571CF17FFEFFD9AFDE7FB4F283B" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="C27236FACF17FFEFFD9AFDE7FC5A281C" blockId="16.[371,1216,550,650]" box="[607,981,550,577]" pageId="16" pageNumber="17">
<heading id="993A8196CF17FFEFFD9AFDE7FC5A281C" box="[607,981,550,577]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="17" reason="2">
<taxonomicName id="05CD4D79CF17FFEFFD9AFDE7FCFB281C" authority="Balkenohl, 2023" authorityName="Balkenohl" authorityYear="2023" box="[607,884,550,577]" class="Insecta" family="Carabidae" genus="Sulciclivina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mikirensis" status="sp. nov.">
<emphasis id="F0B9EAE8CF17FFEFFD9AFDE7FCFB281C" bold="true" box="[607,884,550,577]" italics="true" pageId="16" pageNumber="17">Sulciclivina mikirensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EB8A5793CF17FFEFFCBFFDE6FC5A281C" box="[890,981,551,577]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
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</paragraph>
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<subSubSection id="8AD76571CF17FFEFFD0CFDAEFCE528D7" box="[713,874,623,650]" pageId="16" pageNumber="17" type="description">
<paragraph id="C27236FACF17FFEFFD0CFDAEFCE528D7" blockId="16.[371,1216,550,650]" box="[713,874,623,650]" pageId="16" pageNumber="17">
<figureCitation id="5AF62A7FCF17FFEFFD0CFDAEFC9C28D4" box="[713,787,623,649]" captionStart="Figs 411" captionStartId="11.[189,241,1857,1883]" captionTargetBox="[204,1380,703,1763]" captionTargetId="figure-113@11.[189,1398,670,1822]" captionTargetPageId="11" captionText="Figs 411. Sulciclivina Balkenohl, 2022, habitus, dorsal view. 45. S. attenuata attenuata (Herbst, 1806). 4. ♀, lectotype (MFNB). 5. ♂ from Swat (CMBB). 6. S. attenuata semireticulata subsp. nov., ♂, holotype (MNHN). 7. S. bhamoensis (Bates, 1892), ♀, paralectotype (RBINS). 8. S. mikirensis sp. nov., ♀, holotype (NHMB). 9. S. basiangusta sp. nov., ♂, holotype (NHMB). 10. S. coxisetosa sp. nov., ♀, holotype (NHMB). 11. S. splendida sp. nov., ♂, holotype (CADW)." figureDoi="http://doi.org/10.5281/zenodo.10410132" httpUri="https://zenodo.org/record/10410132/files/figure.png" pageId="16" pageNumber="17">Figs 8</figureCitation>
, 39, 53
</paragraph>
</subSubSection>
<subSubSection id="8AD76571CF17FFEFFF78FD70FB482E62" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="C27236FACF17FFEFFF78FD70FEB92896" blockId="16.[189,310,689,715]" box="[189,310,689,715]" pageId="16" pageNumber="17">
<heading id="993A8196CF17FFEFFF78FD70FEB92896" bold="true" box="[189,310,689,715]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="F0B9EAE8CF17FFEFFF78FD70FEB92896" bold="true" box="[189,310,689,715]" pageId="16" pageNumber="17">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF17FFEFFF78FD20FB482E62" blockId="16.[187,1399,737,1088]" pageId="16" pageNumber="17">
A larger-sized species with a peltate-like pronotum, with interval eight of the elytra reticulated. The proepisternum is swollen laterally but appears more edged than in the other species. Mainly distinguished from the similar species
<taxonomicName id="05CD4D79CF17FFEFFE1AFCE8FD07291E" box="[479,648,809,835]" class="Insecta" family="Carabidae" genus="Eoclivina" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bhamoensis">
<emphasis id="F0B9EAE8CF17FFEFFE1AFCE8FD07291E" box="[479,648,809,835]" italics="true" pageId="16" pageNumber="17">S. bhamoensis</emphasis>
</taxonomicName>
by the reticulation on the elytron, which is restricted to interval eight (females). In
<taxonomicName id="05CD4D79CF17FFEFFE65FC8CFDBA293A" authority="Balkenohl, 2023" authorityName="Balkenohl" authorityYear="2023" box="[416,565,845,871]" class="Insecta" family="Carabidae" genus="Sulciclivina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mikirensis" status="sp. nov.">
<emphasis id="F0B9EAE8CF17FFEFFE65FC8CFDBA293A" box="[416,565,845,871]" italics="true" pageId="16" pageNumber="17">S. mikirensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EB8A5793CF17FFEFFDFBFC8CFD13293A" box="[574,668,845,871]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
, the lobe of the mentum is flattened with a small carina at the middle. The median tooth is wide, distinctly less projecting anteriorly than the lobe, flattened apically and rounded off. In
<taxonomicName id="05CD4D79CF17FFEFFEAAFC54FD9B29F2" box="[367,532,917,943]" class="Insecta" family="Carabidae" genus="Eoclivina" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bhamoensis">
<emphasis id="F0B9EAE8CF17FFEFFEAAFC54FD9B29F2" box="[367,532,917,943]" italics="true" pageId="16" pageNumber="17">S. bhamoensis</emphasis>
</taxonomicName>
however, the lobe of the mentum shows askew carinae, the median tooth is wide and nearly as far protruding anteriorly as the lobe. Distinguished from
<taxonomicName id="05CD4D79CF17FFEFFBECFC78FAA7298E" baseAuthorityName="Herbst" baseAuthorityYear="1806" box="[1065,1320,953,979]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="attenuata" subSpecies="attenuata">
<emphasis id="F0B9EAE8CF17FFEFFBECFC78FAA7298E" box="[1065,1320,953,979]" italics="true" pageId="16" pageNumber="17">S. attenuata attenuata</emphasis>
</taxonomicName>
by the pronotum with a distinctly straight lateral margin, the basolateral impression with an elongated group of numerous punctures, and with the paramedially situated longitudinal cloud of medium-sized punctures reaching from middle of pronotum up to base. In addition, the gonocoxites are different.
</paragraph>
</subSubSection>
<subSubSection id="8AD76571CF17FFEFFF78FBA9FB1E2EEC" box="[189,1169,1128,1202]" pageId="16" pageNumber="17" type="etymology">
<paragraph id="C27236FACF17FFEFFF78FBA9FECB2EDF" blockId="16.[189,324,1128,1154]" box="[189,324,1128,1154]" pageId="16" pageNumber="17">
<heading id="993A8196CF17FFEFFF78FBA9FECB2EDF" bold="true" box="[189,324,1128,1154]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="F0B9EAE8CF17FFEFFF78FBA9FECB2EDF" bold="true" box="[189,324,1128,1154]" pageId="16" pageNumber="17">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF17FFEFFF78FB56FB1E2EEC" blockId="16.[189,1169,1175,1202]" box="[189,1169,1175,1202]" pageId="16" pageNumber="17">
The name refers to the Mikir Hills in
<collectingRegion id="0009F818CF17FFEFFDB6FB56FD492EEC" box="[627,710,1175,1201]" country="India" name="Assam" pageId="16" pageNumber="17">Assam</collectingRegion>
, the collection locality of the
<typeStatus id="1D768858CF17FFEFFBE7FB56FB022EEC" box="[1058,1165,1175,1201]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="8AD76571CF17FFEFFF78FB18FEFA2FA8" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="C27236FACF17FFEFFF78FB18FEE32EAE" blockId="16.[189,364,1241,1267]" box="[189,364,1241,1267]" pageId="16" pageNumber="17">
<heading id="993A8196CF17FFEFFF78FB18FEE32EAE" bold="true" box="[189,364,1241,1267]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="F0B9EAE8CF17FFEFFF78FB18FEE32EAE" bold="true" box="[189,364,1241,1267]" pageId="16" pageNumber="17">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF17FFEFFF2FFAC8FED32F7E" blockId="16.[189,1398,1289,1426]" box="[234,348,1289,1315]" pageId="16" pageNumber="17">
<emphasis id="F0B9EAE8CF17FFEFFF2FFAC8FED32F7E" bold="true" box="[234,348,1289,1315]" pageId="16" pageNumber="17">
<typeStatus id="1D768858CF17FFEFFF2FFAC8FED32F7E" box="[234,348,1289,1315]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="C27236FACF17FFEFFF78FAECFBF42FCD" blockId="16.[189,1398,1289,1426]" pageId="16" pageNumber="17">
<materialsCitation id="72A53CA7CF17FFEFFF78FAECFBF42FCD" collectingDate="1961-05" collectionCode="NHMB" collectorName="G. Scherer &amp; Museum Frey Tutzing" country="India" location="ASSAM" pageId="16" pageNumber="17" specimenCount="1" specimenCount-female="1" stateProvince="Assam" typeStatus="holotype">
<collectingCountry id="BADA766ACF17FFEFFF78FAECFE802F1A" box="[189,271,1325,1351]" name="India" pageId="16" pageNumber="17">INDIA</collectingCountry>
<specimenCount id="D4CBFD73CF17FFEFFEE7FAECFEB72F1A" box="[290,312,1325,1351]" count="1" pageId="16" pageNumber="17" type="female"></specimenCount>
; with white, black printed label and data: “
<collectingRegion id="0009F818CF17FFEFFCDFFAECFC0C2F1A" box="[794,899,1325,1351]" country="India" name="Assam" pageId="16" pageNumber="17">ASSAM</collectingRegion>
, Kaziranga nördl. Mikir-Hills Brahmaputra,
<date id="B673103ACF17FFEFFF78FA90FEAF2F36" box="[189,288,1361,1387]" pageId="16" pageNumber="17" value="1961-05">
<collectingDate id="A637E9D2CF17FFEFFF78FA90FEAF2F36" box="[189,288,1361,1387]" pageId="16" pageNumber="17" value="1961-05">V. 1961</collectingDate>
</date>
, leg.
<collectorName id="6F38532CCF17FFEFFE9CFA90FE582F36" box="[345,471,1361,1387]" pageId="16" pageNumber="17">G.Scherer</collectorName>
” / white, black printed “
<collectorName id="6F38532CCF17FFEFFCCEFA90FBAC2F36" box="[779,1059,1361,1387]" pageId="16" pageNumber="17">Museum Frey Tutzing</collectorName>
” / white, black printed and handwritten in black ink “
<taxonomicName id="05CD4D79CF17FFEFFE28FAB4FD722FD2" authority="Hbst." authorityName="Hbst." box="[493,765,1397,1424]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="attenuata">Clivina attenuata Hbst.</taxonomicName>
det. ING.JEDLIČKA”;
<collectionCode id="A4DCAE3FCF17FFEFFBDEFAB7FBF82FCD" box="[1051,1143,1398,1424]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="16" pageNumber="17">NHMB</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="C27236FACF17FFEFFF78FA76FEFA2FA8" blockId="16.[187,1399,1463,1526]" pageId="16" pageNumber="17">
<emphasis id="F0B9EAE8CF17FFEFFF78FA76FEA52F8C" bold="true" box="[189,298,1463,1489]" pageId="16" pageNumber="17">Remark.</emphasis>
Three terminal antennomeres at the left side and two apical tarsomeres of the right intermediate leg are missing.
</paragraph>
</subSubSection>
<subSubSection id="8AD76571CF17FFEDFF78F9DCFEB62BBA" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17" type="description">
<paragraph id="C27236FACF17FFEFFF78F9DCFEC12C6A" blockId="16.[189,334,1565,1591]" box="[189,334,1565,1591]" pageId="16" pageNumber="17">
<heading id="993A8196CF17FFEFFF78F9DCFEC12C6A" bold="true" box="[189,334,1565,1591]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="F0B9EAE8CF17FFEFFF78F9DCFEC12C6A" bold="true" box="[189,334,1565,1591]" pageId="16" pageNumber="17">Description</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF17FFEFFF78F98CFF772CD6" blockId="16.[189,1399,1613,1675]" pageId="16" pageNumber="17">
MEASUREMENTS.
<typeStatus id="1D768858CF17FFEFFE46F98CFE7A2C3A" box="[387,501,1613,1639]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
: body length
<quantity id="05359B1FCF17FFEFFD56F98CFD702C35" box="[659,767,1613,1640]" metricMagnitude="-3" metricUnit="m" metricValue="7.69" pageId="16" pageNumber="17" unit="mm" value="7.69">7.69 mm</quantity>
, width
<quantity id="05359B1FCF17FFEFFC90F98CFC3D2C35" box="[853,946,1613,1640]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="16" pageNumber="17" unit="mm" value="2.2">2.2 mm</quantity>
; L/W of pronotum 0.98; L/W of elytra 1.94.
</paragraph>
<paragraph id="C27236FACF17FFEFFF78F975FB432CAF" blockId="16.[189,1398,1715,1778]" pageId="16" pageNumber="17">COLOUR. Glossy. Piceous. Labrum, intermediate and hind leg dark-fuscous, antenna and tarsomeres of front leg fuscous. Supraantennal plate translucent-fuscous up to the top of the convexity.</paragraph>
<paragraph id="C27236FACF17FFEEFF78F8DBFBC12891" blockId="16.[189,1399,1818,2025]" lastBlockId="17.[188,1400,265,716]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
HEAD. A third narrower than pronotum. Clypeus with central part moderately bilobed anteriorly; lateral lobe almost fused with wing, wing indistinctly traceable; supraantennal plate laterally with less convex margin, distinctly less projecting laterally than eye; clypeus and wing slightly, and supraantennal plate distinctly reflexed margined. Supraantennal plate overlapping eye anteriorly by around ten percent (dorsal view), with supraorbital keel extended up to the anterior third of the eye, separated from wing by sharp notch. Clypeus transverse, moderately convex, separated from frons by distinct step and deepened sulcus, step and sulcus in form like an inverted slightly rounded flat V. Frons moderately convex, with two small and distinct askew impressions at middle, with supraorbital carina up to posterior eye-level. Clypeus smooth, frons with minute and medium-sized punctures at gena-level, punctures larger laterally; frons separated from supraantennal plate and supraorbital carina by deep moderately wide furrow, furrow slightly widening at the posterior end; with two supraorbital setigerous punctures at middle and posterior end of the eye. Neck constriction missing. Eye semi-hemispherical projecting in dorsal view. Main part of eye situated ventrally where it is also globose. Due to the overlapping supraantennal plate and the gena, the globosity of the eye appears in lateral view somewhat narrowed. Gena distinct, slightly convex, covering less than a quarter of posterior eye in ventral view. Antenna relatively short, reaching middle of pronotum, antennomeres five to ten of moderate length (L/W around 1.2). Labrum straight anteriorly, with isodiametric reticulation, six setose.
<collectingRegion id="0009F818CF16FFEEFD61FDAAFC8828D8" box="[676,775,619,645]" country="Myanmar" name="Chin" pageId="17" pageNumber="18">Mentum</collectingRegion>
with transverse reticulation at middle, lobe flattened, shape of lobe semi-ovoid, with isodiametric reticulation, with small carina at middle, median tooth wide, distinctly less projecting anteriorly as lobe, flattened apically and rounded off.
</paragraph>
<paragraph id="C27236FACF16FFEEFF78FD32FB972ECF" blockId="17.[187,1400,755,1171]" pageId="17" pageNumber="18">PRONOTUM. Disk slightly convex in lateral and distinctly convex in frontal view. Indistinctly wider than long. Reflexed lateral margin smooth, attenuating in anterior two-thirds with lateral margin extraordinarily straight, widest behind middle; anterior angle slightly projecting, posterior angle missing; lateral channel moderately narrow, irregularly reticulated, completely and regularly rounded off at level of posterior setigerous puncture and continuing up to base. Median line sharp, crenulated, complete.Anterior transverse line indistinctly punctured, joining median line, finely joining anterior margin. Surface glossy, anterior margin reticulated laterally, basolaterally with impression and elongated group of distinct punctures, paramedially with distinct long-oval group of dense punctures of moderate size reaching from middle of pronotum up to base, with smaller group of punctures in continuation of the paramedian group posteriorly to the anterior transverse line, with few transverse wrinkles and with microscopic punctures on whole disk. Base distinctly marked, slightly wider than lateral channel. Proepisternum distinctly visible in dorsal view, appearing more edged as in other species where it is more rounded.</paragraph>
<paragraph id="C27236FACF16FFEEFF78FB7BFB1C2F91" blockId="17.[188,1399,1210,1484]" pageId="17" pageNumber="18">ELYTRON. Shape elongate. Disk flattened in anterior third in lateral view, distinctly and regularly convex in frontal view. Less than twice as long as wide, with maximum width slightly behind middle. Reflexed lateral margin smooth. Scutellar striole missing; setigerous tubercle at base of first stria, with small but distinct tubercle at base of third interval. Humeral tooth situated at base of fourth stria. Striae moderately deep, punctuation of striae distinctly impressed, one to three free at base, four reaching humerus, one and two, three and four, and five and six joining apically. Intervals moderately convex, more convex laterally, interval eight carinate apically. Setigerous punctures missing on intervals. Surface of intervals glossy, interval eight with isodiametric reticulation, tip of apex with interval seven smooth.</paragraph>
<paragraph id="C27236FACF16FFEEFF78FA35FC372C50" blockId="17.[189,952,1523,1550]" box="[189,952,1523,1550]" pageId="17" pageNumber="18">
HIND WINGS. Alae fully developed (on the basis of the
<typeStatus id="1D768858CF16FFEEFC85FA32FC232C50" box="[832,940,1523,1549]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="C27236FACF16FFEEFF78F9F4FAAE2CCB" blockId="17.[189,1400,1589,1686]" pageId="17" pageNumber="18">LOWER SURFACE. Proepisternum covered laterally with distinct isodiametric reticulation, with dense transverse wrinkles and medium-sized punctures. Sternite of abdomen covered nearly completely with moderately large punctures, with caudal margin smooth; with isodiametric reticulation laterally.</paragraph>
<paragraph id="C27236FACF16FFEEFF78F97FFCB52CA6" blockId="17.[189,1399,1725,1787]" pageId="17" pageNumber="18">LEGS. Tibia covered with isodiametric reticulation. Protibia with three spines of moderate length, not sulcate dorsally, movable spur apically gently arcuate.</paragraph>
<paragraph id="C27236FACF16FFEEFF78F8E2FD852D61" blockId="17.[189,522,1826,1853]" box="[189,522,1826,1853]" pageId="17" pageNumber="18">MALE GENITALIA. Unknown.</paragraph>
<paragraph id="C27236FACF16FFEEFF78F8A5FB382DB5" blockId="17.[187,1399,1892,2025]" pageId="17" pageNumber="18">FEMALE GONOCOXITES AND EPIPLEURITE (Fig. 39). Gonocoxites foliform, distinctly widened and somewhat jolted, flattened dorsally and ventrally, with four long and one small nematiform setae positioned at the median side, seta generating punctures not tuberculate, with one apical nematiform seta arising from rounded apex. Epipleurite elongate-rectangular, with distinctly widened angulated rod.</paragraph>
<paragraph id="C27236FACF15FFEDFF78FEC9FEBC2B7F" blockId="18.[189,307,264,290]" box="[189,307,264,290]" pageId="18" pageNumber="19">
<heading id="993A8196CF15FFEDFF78FEC9FEBC2B7F" bold="true" box="[189,307,264,290]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="F0B9EAE8CF15FFEDFF78FEC9FEBC2B7F" bold="true" box="[189,307,264,290]" pageId="18" pageNumber="19">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF15FFEDFF78FEF9FE2D2B2B" blockId="18.[189,1398,312,374]" pageId="18" pageNumber="19">The number of moderately-sized punctures on the disk of the pronotum varies on both sides. Intraspecific variation unknown.</paragraph>
<paragraph id="C27236FACF15FFEDFF78FE5FFE3F2BE5" blockId="18.[189,432,414,440]" box="[189,432,414,440]" pageId="18" pageNumber="19">
<heading id="993A8196CF15FFEDFF78FE5FFE3F2BE5" bold="true" box="[189,432,414,440]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="F0B9EAE8CF15FFEDFF78FE5FFE3F2BE5" bold="true" box="[189,432,414,440]" pageId="18" pageNumber="19">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF15FFEDFF78FE0CFEB62BBA" blockId="18.[189,313,461,487]" box="[189,313,461,487]" pageId="18" pageNumber="19">Unknown.</paragraph>
</subSubSection>
<subSubSection id="8AD76571CF15FFEDFF78FDCEFC742805" box="[189,1019,527,601]" pageId="18" pageNumber="19" type="distribution">
<paragraph id="C27236FACF15FFEDFF78FDCEFED92874" blockId="18.[189,342,527,553]" box="[189,342,527,553]" pageId="18" pageNumber="19">
<heading id="993A8196CF15FFEDFF78FDCEFED92874" bold="true" box="[189,342,527,553]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="F0B9EAE8CF15FFEDFF78FDCEFED92874" bold="true" box="[189,342,527,553]" pageId="18" pageNumber="19">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C27236FACF15FFEDFF78FDFEFC742805" blockId="18.[189,1019,574,601]" box="[189,1019,574,601]" pageId="18" pageNumber="19">
Known from the
<typeStatus id="1D768858CF15FFEDFE43FDFEFE362804" box="[390,441,575,601]" pageId="18" pageNumber="19">type</typeStatus>
locality north of the Mikir-Hills in
<collectingRegion id="0009F818CF15FFEDFC9DFDFEFC242804" box="[856,939,575,601]" country="India" name="Assam" pageId="18" pageNumber="19">Assam</collectingRegion>
,
<collectingCountry id="BADA766ACF15FFEDFC72FDFFFC782805" box="[951,1015,574,600]" name="India" pageId="18" pageNumber="19">India</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>