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<mods:title id="2F9AFEF5C1B95A41970E050F9235AEAC">A review of the families and genera of the superfamily PLATYSCELOIDEA Bowman &amp; Gruner, 1973 (Crustacea: Amphipoda: Hyperiidea), together with keys to the families, genera and species</mods:title>
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<paragraph id="C272A4021828FFE1FF5EF9A0FDFEF984" blockId="98.[151,631,1628,1654]" box="[151,631,1628,1654]" pageId="98" pageNumber="99">
<heading id="993A136E1828FFE1FF5EF9A0FDFEF984" bold="true" box="[151,631,1628,1654]" fontSize="11" level="1" pageId="98" pageNumber="99" reason="1">
<emphasis id="F0B978101828FFE1FF5EF9A0FDFEF984" bold="true" box="[151,631,1628,1654]" pageId="98" pageNumber="99">
Family
<taxonomicName id="05CDDF811828FFE1FF3FF9A0FDFEF984" ID-CoL="7NKJB" authority="Bate, 1862" authorityName="Bate" authorityYear="1862" box="[246,631,1628,1654]" class="Malacostraca" family="Platyscelidae" kingdom="Animalia" order="Amphipoda" pageId="98" pageNumber="99" phylum="Arthropoda" rank="family">
PLATYSCELIDAE
<bibRefCitation id="A65CD9F31828FFE1FE3DF9A0FDFEF984" author="Bate" box="[500,631,1628,1654]" pageId="98" pageNumber="99" refString="Bate, C. S. (1862) Catalogue of the specimens of Amphipodous Crustacea in the collection of the British Museum. British Museum, Natural History, London, 399 pp., plates 1 - 58." type="book chapter" year="1862">Bate, 1862</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8AD7F7891828FFE0FF5EF95EFF5FFD02" lastPageId="99" lastPageNumber="100" pageId="98" pageNumber="99" type="diagnosis">
<paragraph id="C272A4021828FFE1FF5EF95EFA14F828" blockId="98.[151,1437,1698,2011]" pageId="98" pageNumber="99">
<emphasis id="F0B978101828FFE1FF5EF95EFE98F949" bold="true" box="[151,273,1698,1723]" pageId="98" pageNumber="99">Diagnosis.</emphasis>
Body length up to
<quantity id="053509E71828FFE1FE31F95EFDC5F948" box="[504,588,1698,1723]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="98" pageNumber="99" unit="mm" value="25.0">25 mm</quantity>
, rolled into ball, the enlarged basis of P5 &amp; 6 completely covering other pereopoda, with telson fitting neatly into telsonic groove of basis of P6, slightly flattened dorso-ventrally. Head relatively short, with beak between first antennae, fitting neatly between distal margins of basis of P6 when rolled up. Eyes in single (
<taxonomicName id="05CDDF811828FFE1FEB7F8F2FE77F8D5" box="[382,510,1806,1831]" class="Malacostraca" family="Platyscelidae" genus="Tetrathyrus" kingdom="Animalia" order="Amphipoda" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis id="F0B978101828FFE1FEB7F8F2FE77F8D5" box="[382,510,1806,1831]" italics="true" pageId="98" pageNumber="99">Tetrathyrus</emphasis>
</taxonomicName>
), or double groups, occupying most of head surface. Coxae 16 separate from pereonites; coxa 7 almost totally fused with pereonite (except
<taxonomicName id="05CDDF811828FFE1FCA2F8CEFC63F8B9" box="[875,1002,1842,1867]" class="Malacostraca" family="Platyscelidae" genus="Platyscelus" kingdom="Animalia" order="Amphipoda" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis id="F0B978101828FFE1FCA2F8CEFC63F8B9" box="[875,1002,1842,1867]" italics="true" pageId="98" pageNumber="99">Platyscelus</emphasis>
</taxonomicName>
), with slight posterior notch between coxa and pereonite. Antennae 1 of females with 23 articulate peduncle, first flagellar article (callynophore) with slight bulge proximally (
<taxonomicName id="05CDDF811828FFE1FE71F886FDB1F861" box="[440,568,1914,1939]" class="Malacostraca" family="Platyscelidae" genus="Tetrathyrus" kingdom="Animalia" order="Amphipoda" pageId="98" pageNumber="99" phylum="Arthropoda" rank="genus">
<emphasis id="F0B978101828FFE1FE71F886FDB1F861" box="[440,568,1914,1939]" italics="true" pageId="98" pageNumber="99">Tetrathyrus</emphasis>
</taxonomicName>
), or distally, with two much narrower, smaller articles inserted terminally, Antennae 1 of males with 12 articulate peduncle and enlarged, curved first flagellar article (callynophore), with two-field brush of aesthetascs medially, and with three smaller, slender articles inserted on antero-distal corner.
</paragraph>
<paragraph id="C272A4021829FFE0FF5EFF64FC31FD5A" blockId="99.[151,1437,151,2013]" pageId="99" pageNumber="100">
Antennae 2 of females absent (
<taxonomicName id="05CDDF811829FFE0FE3BFF6BFDFBFF42" box="[498,626,151,176]" class="Malacostraca" family="Platyscelidae" genus="Tetrathyrus" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F0B978101829FFE0FE3BFF6BFDFBFF42" box="[498,626,151,176]" italics="true" pageId="99" pageNumber="100">Tetrathyrus</emphasis>
</taxonomicName>
), or of five slender articles, the terminal one often very small. Antennae 2 of males with five slender articles, folded back on one another underneath head and pereon, basal article the shortest and slightly more robust than following articles, articles 2 &amp; 3 sub-equal in length; articles 4 &amp; 5 are also sub-equal in length but distinctly shorter than previous pair, article 5 sometimes slightly shorter than article 4. Mouthparts in form of short, broad cylinder. Mandibles with 3-articulate palp in males, absent in females; incisor relatively broad, with slightly enlarged tooth laterally. Maxillae 1 reduced in size, consisting of small, elongate lobes, with oblique distal margin armed with few robust setae, or two, or four, bifid teeth. Maxillae 2 reduced to small, rounded or pointed lobes, slightly curved with rounded bulge medially. Gnathopoda simple, sub-chelate, or chelate. Pereopods 5 &amp; 6 with basis transformed into broad operculum, always longer and larger in P6; distal articles inserted sub-terminally on basis of P5, more proximally for P6. Pereopod 6; basis with, or without (
<taxonomicName id="05CDDF811829FFE0FF69FE03FEA9FDEA" box="[160,288,511,536]" class="Malacostraca" family="Platyscelidae" genus="Tetrathyrus" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F0B978101829FFE0FF69FE03FEA9FDEA" box="[160,288,511,536]" italics="true" pageId="99" pageNumber="100">Tetrathyrus</emphasis>
</taxonomicName>
), fissure, postero-distal corner with ridge-groove locking mechanism with opposing P6, posterior margin with telsonic groove. Pereopod 7 reduced in size, usually consisting of basis and 12 tiny articles, rarely with full complement of articles (abnormal). Uropod 3; endopod fused with peduncle, rarely articulated, or endopod of U2 also fused (
<taxonomicName id="05CDDF811829FFE0FE11FD90FDD1FD77" box="[472,600,620,645]" class="Malacostraca" family="Platyscelidae" genus="Tetrathyrus" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F0B978101829FFE0FE11FD90FDD1FD77" box="[472,600,620,645]" italics="true" pageId="99" pageNumber="100">Tetrathyrus</emphasis>
</taxonomicName>
). Telson triangular, relatively broad, fused with double urosomite. Gills with, or without, folds on pereonites 26. Oostegites on pereonites 25.
</paragraph>
<paragraph id="C272A4021829FFE0FF0EFD48FF5FFD02" blockId="99.[151,1437,151,2013]" pageId="99" pageNumber="100">
<emphasis id="F0B978101829FFE0FF0EFD48FEAFFD3F" bold="true" box="[199,294,692,717]" pageId="99" pageNumber="100">Genera.</emphasis>
<taxonomicName id="05CDDF811829FFE0FEFBFD48FDB6FD3F" authority="Bate, 1862" authorityName="Bate" authorityYear="1862" box="[306,575,692,717]" class="Malacostraca" family="Platyscelidae" genus="Platyscelus" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F0B978101829FFE0FEFBFD48FE38FD3F" box="[306,433,692,717]" italics="true" pageId="99" pageNumber="100">Platyscelus</emphasis>
<bibRefCitation id="A65CD9F31829FFE0FE74FD49FDB6FD3F" author="Bate" box="[445,575,692,717]" pageId="99" pageNumber="100" refString="Bate, C. S. (1862) Catalogue of the specimens of Amphipodous Crustacea in the collection of the British Museum. British Museum, Natural History, London, 399 pp., plates 1 - 58." type="book chapter" year="1862">Bate, 1862</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="05CDDF811829FFE0FD86FD48FCE1FD3E" authority="Claus, 1879" authorityName="Claus" authorityYear="1879" box="[591,872,692,717]" class="Malacostraca" family="Platyscelidae" genus="Hemityphis" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F0B978101829FFE0FD86FD48FD47FD3F" box="[591,718,692,717]" italics="true" pageId="99" pageNumber="100">Hemityphis</emphasis>
<bibRefCitation id="A65CD9F31829FFE0FD13FD48FCE1FD3E" author="Claus" box="[730,872,692,717]" pageId="99" pageNumber="100" refString="Claus, C. (1879) Die Gattungen und Arten der Platysceliden in Systematischer Ubersicht. Arbeiten aus dem Zoologischen Institut der Universitat zu Wien und der Zoologischen Station Triest, 2, 1 - 52 (147 - 198)." type="journal article" year="1879">Claus, 1879</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="05CDDF811829FFE0FCB1FD48FB00FD3E" authority="Claus, 1879" authorityName="Claus" authorityYear="1879" box="[888,1161,692,717]" class="Malacostraca" family="Platyscelidae" genus="Paratyphis" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F0B978101829FFE0FCB1FD48FC7AFD3F" box="[888,1011,692,717]" italics="true" pageId="99" pageNumber="100">Paratyphis</emphasis>
<bibRefCitation id="A65CD9F31829FFE0FC36FD48FB00FD3E" author="Claus" box="[1023,1161,692,717]" pageId="99" pageNumber="100" refString="Claus, C. (1879) Die Gattungen und Arten der Platysceliden in Systematischer Ubersicht. Arbeiten aus dem Zoologischen Institut der Universitat zu Wien und der Zoologischen Station Triest, 2, 1 - 52 (147 - 198)." type="journal article" year="1879">Claus, 1879</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="05CDDF811829FFE0FB00FD48FF5BFD02" authority="Claus, 1879" authorityName="Claus" authorityYear="1879" class="Malacostraca" family="Platyscelidae" genus="Tetrathyrus" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F0B978101829FFE0FB00FD48FAC0FD3F" box="[1225,1353,692,717]" italics="true" pageId="99" pageNumber="100">Tetrathyrus</emphasis>
<bibRefCitation id="A65CD9F31829FFE0FA9CFD48FF5BFD02" author="Claus" pageId="99" pageNumber="100" refString="Claus, C. (1879) Die Gattungen und Arten der Platysceliden in Systematischer Ubersicht. Arbeiten aus dem Zoologischen Institut der Universitat zu Wien und der Zoologischen Station Triest, 2, 1 - 52 (147 - 198)." type="journal article" year="1879">Claus, 1879</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="8AD7F7891829FFE7FF0EFD00FE9FFD77" lastPageId="100" lastPageNumber="101" pageId="99" pageNumber="100" type="discussion">
<paragraph id="C272A4021829FFE0FF0EFD00FCD0FBE2" blockId="99.[151,1437,151,2013]" pageId="99" pageNumber="100">
<emphasis id="F0B978101829FFE0FF0EFD00FEB2FCE7" bold="true" box="[199,315,764,789]" pageId="99" pageNumber="100">Remarks.</emphasis>
In the past this family has also been known as Typhidae
<bibRefCitation id="A65CD9F31829FFE0FC04FD01FBD9FCE7" author="Dana" box="[973,1104,764,789]" pageId="99" pageNumber="100" refString="Dana, J. D. (1853) Crustacea, Part II. United States Exploring Expedition, 14, 689 - 1618. [Plates 1 - 96 published in 1855]" type="journal article" year="1853">Dana, 1853</bibRefCitation>
(
<bibRefCitation id="A65CD9F31829FFE0FBA9FD00FB6CFCE6" author="Claus" box="[1120,1253,764,789]" pageId="99" pageNumber="100" refString="Claus, C. (1879) Die Gattungen und Arten der Platysceliden in Systematischer Ubersicht. Arbeiten aus dem Zoologischen Institut der Universitat zu Wien und der Zoologischen Station Triest, 2, 1 - 52 (147 - 198)." type="journal article" year="1879">Claus 1879</bibRefCitation>
,
<bibRefCitation id="A65CD9F31829FFE0FB3BFD00FA11FCE6" author="Stebbing" box="[1266,1432,764,789]" pageId="99" pageNumber="100" refString="Stebbing, T. R. R. (1888) Report on the Amphipoda collected by H. M. S. ' Challenger' during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. ' Challenger' during the years 1873 - 76. Zoology, 29, i - xxiv &amp; 1 - 1737, plates 1 - 210." type="journal article" year="1888">Stebbing 1888</bibRefCitation>
,
<bibRefCitation id="A65CD9F31829FFE0FF5EFCE3FECFFCCA" author="Chevreux" box="[151,326,799,824]" pageId="99" pageNumber="100" refString="Chevreux, E. (1900) Amphipodes provenant des campagnes de I'Hirondelle (1885 - 1888). Resultats des Campagnes scientifiques accomplies sur son Yacht, par Albert 1 er, Prince Souverain de Monaco, 16 (i - iv), 1 - 195, plates 1 - 18." type="journal article" year="1900">Chevreux 1900</bibRefCitation>
,
<bibRefCitation id="A65CD9F31829FFE0FE98FCE3FDD3FCCA" author="Chevreux" box="[337,602,799,824]" pageId="99" pageNumber="100" refString="Chevreux, E. &amp; Fage, L. (1925) Amphipodes. Faune de France, 9, 488 pp., 438 text Figs" type="book" year="1925">Chevreux &amp; Fage 1925</bibRefCitation>
) and Eutyphidae
<bibRefCitation id="A65CD9F31829FFE0FCE9FCE3FC5CFCCA" author="Bovallius" box="[800,981,799,824]" pageId="99" pageNumber="100" refString="Bovallius, C. (1887) Systematical list of the Amphipoda Hyperiidea. Bihang till Kungliga Vetenskaps-Akademiens Handlingar, 11 (16), 1 - 50." type="journal article" year="1887">Bovallius, 1887</bibRefCitation>
. The name Typhidae is derived from the genus
<taxonomicName id="05CDDF811829FFE0FF29FCB8FE3EFCAF" authority="Risso, 1816" authorityName="Risso" authorityYear="1816" box="[224,439,836,861]" class="Gastropoda" family="Muricidae" genus="Typhis" kingdom="Animalia" order="Neogastropoda" pageId="99" pageNumber="100" phylum="Mollusca" rank="genus">
<emphasis id="F0B978101829FFE0FF29FCB8FEA0FCAF" box="[224,297,836,861]" italics="true" pageId="99" pageNumber="100">Typhis</emphasis>
<bibRefCitation id="A65CD9F31829FFE0FEF8FCB8FE3EFCAF" author="Risso" box="[305,439,836,861]" pageId="99" pageNumber="100" refString="Risso, A. (1816) Histoire naturelle des Crustaces des environs de Nice. Librairie Grecque-Latine-Allemande: Paris, 175 pp., 3 plates." type="book" year="1816">Risso, 1816</bibRefCitation>
</taxonomicName>
which, although being an older name, is preoccupied by a genus of molluscs (
<bibRefCitation id="A65CD9F31829FFE0FAFCFCB8FF5AFC72" author="Montfort" pageId="99" pageNumber="100" refString="Montfort, P. Denys de (1810) Conchyliologie Systematique, et classification methodique des Coquilles; offrant leurs figures, leur arrangement generique, leurs descriptions caracteristiques, leurs noms; ainsi que leur synonymie en plusieurs langues. Tome Second. Paris: F. Schoell, 1808 - 10, 676 pp." type="book chapter" year="1810">Montfort 1810</bibRefCitation>
). Consequently
<bibRefCitation id="A65CD9F31829FFE0FE41FC94FD84FC72" author="Bate" box="[392,525,871,896]" pageId="99" pageNumber="100" refString="Bate, C. S. (1862) Catalogue of the specimens of Amphipodous Crustacea in the collection of the British Museum. British Museum, Natural History, London, 399 pp., plates 1 - 58." type="book chapter" year="1862">Bate (1862)</bibRefCitation>
proposed the name
<taxonomicName id="05CDDF811829FFE0FD39FC9BFC0EFC72" box="[752,903,871,896]" class="Malacostraca" family="Platyscelidae" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="family">Platyscelidae</taxonomicName>
, derived from the next oldest, available generic name,
<taxonomicName id="05CDDF811829FFE0FF2BFC70FE6CFC57" authority="Bate, 1861" authorityName="Bate" authorityYear="1861" box="[226,485,908,933]" class="Malacostraca" family="Platyscelidae" genus="Platyscelus" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F0B978101829FFE0FF2BFC70FEE8FC57" box="[226,353,908,933]" italics="true" pageId="99" pageNumber="100">Platyscelus</emphasis>
<bibRefCitation id="A65CD9F31829FFE0FEA0FC71FE6CFC57" author="Bate" box="[361,485,908,933]" pageId="99" pageNumber="100" refString="Bate, C. S. (1861) On the morphology of some Amphipoda of the Division Hyperina. Annals and Magazine of Natural History, series 3, 8 (43), 1 - 16, plates 1 - 2." type="journal article" year="1861">Bate, 1861</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="A65CD9F31829FFE0FE38FC70FD0AFC56" author="Claus" box="[497,643,908,933]" pageId="99" pageNumber="100" refString="Claus, C. (1879) Die Gattungen und Arten der Platysceliden in Systematischer Ubersicht. Arbeiten aus dem Zoologischen Institut der Universitat zu Wien und der Zoologischen Station Triest, 2, 1 - 52 (147 - 198)." type="journal article" year="1879">Claus (1879)</bibRefCitation>
incorrectly proposed a new genus
<taxonomicName id="05CDDF811829FFE0FBC2FC70FBE4FC57" box="[1035,1133,908,933]" class="Malacostraca" family="Platyscelidae" genus="Eutyphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F0B978101829FFE0FBC2FC70FBE4FC57" box="[1035,1133,908,933]" italics="true" pageId="99" pageNumber="100">Eutyphis</emphasis>
</taxonomicName>
to encompass a number of older generic names including
<taxonomicName id="05CDDF811829FFE0FE3BFC53FDFBFC3A" box="[498,626,943,968]" class="Malacostraca" family="Platyscelidae" genus="Platyscelus" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F0B978101829FFE0FE3BFC53FDFBFC3A" box="[498,626,943,968]" italics="true" pageId="99" pageNumber="100">Platyscelus</emphasis>
</taxonomicName>
.
<bibRefCitation id="A65CD9F31829FFE0FD49FC53FCB7FC3A" author="Bovallius" box="[640,830,943,968]" pageId="99" pageNumber="100" refString="Bovallius, C. (1887) Systematical list of the Amphipoda Hyperiidea. Bihang till Kungliga Vetenskaps-Akademiens Handlingar, 11 (16), 1 - 50." type="journal article" year="1887">Bovallius (1887)</bibRefCitation>
following Claus, proposed the family Eutyphidae but incorrectly attributed it to
<bibRefCitation id="A65CD9F31829FFE0FE04FC29FDE8FC1F" author="Dana" box="[461,609,980,1005]" pageId="99" pageNumber="100" refString="Dana, J. D. (1853) Crustacea, Part II. United States Exploring Expedition, 14, 689 - 1618. [Plates 1 - 96 published in 1855]" type="journal article" year="1853">Dana (1853)</bibRefCitation>
. Thus,
<taxonomicName id="05CDDF811829FFE0FD73FC28FCB0FC1F" box="[698,825,980,1005]" class="Malacostraca" family="Platyscelidae" genus="Platyscelus" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F0B978101829FFE0FD73FC28FCB0FC1F" box="[698,825,980,1005]" italics="true" pageId="99" pageNumber="100">Platyscelus</emphasis>
</taxonomicName>
is the oldest available generic name and the family should be known as
<taxonomicName id="05CDDF811829FFE0FEB5FC0BFD9AFBE2" box="[380,531,1015,1040]" class="Malacostraca" family="Platyscelidae" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="family">Platyscelidae</taxonomicName>
, as proposed by
<bibRefCitation id="A65CD9F31829FFE0FD05FC04FCDDFBE2" author="Bate" box="[716,852,1015,1040]" pageId="99" pageNumber="100" refString="Bate, C. S. (1862) Catalogue of the specimens of Amphipodous Crustacea in the collection of the British Museum. British Museum, Natural History, London, 399 pp., plates 1 - 58." type="book chapter" year="1862">Bate (1862)</bibRefCitation>
.
</paragraph>
<paragraph id="C272A4021829FFE0FF0EFBE0FEAAFA6E" blockId="99.[151,1437,151,2013]" pageId="99" pageNumber="100">
This family is very similar to
<taxonomicName id="05CDDF811829FFE0FDD9FBE0FD16FBC7" box="[528,671,1052,1077]" class="Malacostraca" family="Parascelidae" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="family">Parascelidae</taxonomicName>
, but is distinguished by the shape of the mouthparts, which are in the form of a broad, rounded cylinder (sharp, pointed cone in
<taxonomicName id="05CDDF811829FFE0FCE0FBC3FC33FBAA" box="[809,954,1087,1112]" class="Malacostraca" family="Parascelidae" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="family">Parascelidae</taxonomicName>
), and pereopod 7 which is usually reduced to the basis and a few vestigial, terminal articles. Also, the basis of pereopod 5 is more elongate in
<taxonomicName id="05CDDF811829FFE0FACBFB98FA10FB8F" box="[1282,1433,1124,1149]" class="Malacostraca" family="Platyscelidae" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="family">Platyscelidae</taxonomicName>
, and in
<taxonomicName id="05CDDF811829FFE0FF21FB7BFEFDFB52" box="[232,372,1159,1184]" class="Malacostraca" family="Parascelidae" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="family">Parascelidae</taxonomicName>
the basis of pereopod 6 is more prominently narrowed for the distal half. The presence of a fissure on the basis of pereopod 6 can also be a useful character to distinguish the genera of the two families. In the family
<taxonomicName id="05CDDF811829FFE0FF2FFB33FEF3FB1A" box="[230,378,1231,1256]" class="Malacostraca" family="Platyscelidae" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="family">Platyscelidae</taxonomicName>
a fissure is only absent in
<taxonomicName id="05CDDF811829FFE0FD69FB33FCA9FB1A" box="[672,800,1231,1256]" class="Malacostraca" family="Platyscelidae" genus="Tetrathyrus" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F0B978101829FFE0FD69FB33FCA9FB1A" box="[672,800,1231,1256]" italics="true" pageId="99" pageNumber="100">Tetrathyrus</emphasis>
</taxonomicName>
, which is readily distinguished by the morphology of the gnathopoda, and the shape of the basis of pereopod 6. In all other genera of
<taxonomicName id="05CDDF811829FFE0FC28FB08FBF1FAFF" box="[993,1144,1268,1293]" class="Malacostraca" family="Platyscelidae" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="family">Platyscelidae</taxonomicName>
, a fissure is present on the basis of pereopod 6, and is located just below the notch for the first urosomite (beginning of telsonic groove). In the family
<taxonomicName id="05CDDF811829FFE0FF21FAC0FEFDFAA7" box="[232,372,1340,1365]" class="Malacostraca" family="Parascelidae" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="family">Parascelidae</taxonomicName>
a fissure is absent in
<taxonomicName id="05CDDF811829FFE0FDA4FAC0FD63FAA7" box="[621,746,1340,1365]" class="Malacostraca" family="Parascelidae" genus="Parascelus" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F0B978101829FFE0FDA4FAC0FD63FAA7" box="[621,746,1340,1365]" italics="true" pageId="99" pageNumber="100">Parascelus</emphasis>
</taxonomicName>
and
<taxonomicName id="05CDDF811829FFE0FCEBFAC0FC0FFAA7" box="[802,902,1340,1365]" class="Insecta" family="Attelabidae" genus="Euscelus" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F0B978101829FFE0FCEBFAC0FC0FFAA7" box="[802,902,1340,1365]" italics="true" pageId="99" pageNumber="100">Euscelus</emphasis>
</taxonomicName>
, but is present in
<taxonomicName id="05CDDF811829FFE0FB9DFAC0FB34FAA7" box="[1108,1213,1340,1365]" class="Malacostraca" family="Parascelidae" genus="Thyropus" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F0B978101829FFE0FB9DFAC0FB34FAA7" box="[1108,1213,1340,1365]" italics="true" pageId="99" pageNumber="100">Thyropus</emphasis>
</taxonomicName>
, where it is located well above the telsonic groove, and in
<taxonomicName id="05CDDF811829FFE0FD8AFAA3FD58FA8A" box="[579,721,1375,1400]" class="Malacostraca" family="Parascelidae" genus="Schizoscelus" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="F0B978101829FFE0FD8AFAA3FD58FA8A" box="[579,721,1375,1400]" italics="true" pageId="99" pageNumber="100">Schizoscelus</emphasis>
</taxonomicName>
where it is very long and extends above and below the notch for urosomite 1.
</paragraph>
<paragraph id="C272A4021829FFE7FF0EFA5BFAB2FE5F" blockId="99.[151,1437,151,2013]" lastBlockId="100.[151,1436,151,645]" lastPageId="100" lastPageNumber="101" pageId="99" pageNumber="100">
Members of this family are usually found rolled into a ball; the enlarged basis of pereopods 5 and 6 completely covering the gnathopoda and other pereopoda, with the telson fitting neatly into a telsonic groove on the basis of pereopod 6 (
<figureCitation id="5AF6B8871829FFE0FEE3FA13FE09F9FA" box="[298,384,1519,1544]" captionStart="FIGURE 49" captionStartId="101.[151,250,1937,1959]" captionTargetBox="[235,1360,208,1908]" captionTargetId="figure@101.[206,1380,193,1916]" captionTargetPageId="101" captionText="FIGURE 49. Platyscelus ovoides (Risso, 1816), female (17.5 mm) and male (15.2 mm), Tasman Sea, SAMA C 4224. A C, habitus, ventral view when rolled up, and anterior view of head of female. Scale bars = 2.0 mm (A C), 1.0 mm (G 1 &amp; 2), 0.2 mm (remainder)." httpUri="https://zenodo.org/record/166468/files/figure.png" pageId="99" pageNumber="100">Fig. 49</figureCitation>
B). Although the ability to roll into a ball has evolved in other families of the superfamily Platysceloidea, none have perfected the habit to the extent found in the families
<taxonomicName id="05CDDF811829FFE0FBF3F9E8FB47F9DF" box="[1082,1230,1556,1581]" class="Malacostraca" family="Platyscelidae" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="family">Platyscelidae</taxonomicName>
and
<taxonomicName id="05CDDF811829FFE0FAC0F9E8FA10F9DF" box="[1289,1433,1556,1581]" class="Malacostraca" family="Parascelidae" kingdom="Animalia" order="Amphipoda" pageId="99" pageNumber="100" phylum="Arthropoda" rank="family">Parascelidae</taxonomicName>
. The morphological characters that allow individuals to achieve this have not been described in any detail previously. These characters vary slightly, in detail, at the generic and specific level but are basically the same for all members of the two families as follows. The basis of pereopod 5 is articulated proximally with the basis of pereopod 6 by means of a small knob, located medially, near the proximal anterior corner of the basis of pereopod 6, which articulates in a groove on pereopod 5, produced by an elongate process on the medial surface of the coxae and a small, adjacent knob on the basis (
<figureCitation id="5AF6B8871829FFE0FDADF910FD30F8F7" box="[612,697,1772,1797]" captionStart="FIGURE 52" captionStartId="106.[151,250,1971,1993]" captionTargetBox="[251,1309,197,1909]" captionTargetId="figure@106.[231,1333,193,1916]" captionTargetPageId="106" captionText="FIGURE 52. Hemityphis tenuimanus Claus, 1879, male (6.8 mm), Tasman Sea, SAMA C 5363. Scale bars = 0.1 mm (Mxp), 0.5 mm (remainder)." httpUri="https://zenodo.org/record/166471/files/figure.png" pageId="99" pageNumber="100">Fig. 52</figureCitation>
). The basis of pereopod 5 has a bevelled anterior margin that fits under the coxae (14), and distally in a groove under the head, next to the second antennae. The posterior margin of the basis of pereopod 5 overlaps with the anterior margin of the basis of pereopod 6, which is bevelled to ensure a neat overlap. The posterior margin of the basis of pereopod 6 has a distinct notch (or shoulder) for urosomite 1, followed by a telsonic groove, which neatly accommodates the double urosomite and the telson. Distal to the telsonic groove is a locking mechanism for opposing pereopoda, consisting of a ridge/groove arrangement,
<collectingMethod id="1B8CDC151829FFE0FA87F85CFEA0F82F" pageId="99" pageNumber="100">similar to the funnel</collectingMethod>
locking mechanism found in squid, with the left usually overlapping the right, suggesting they are predominantly left handed (in 160 specimens of
<taxonomicName id="05CDDF81182EFFE7FCCDFF6BFC60FF42" box="[772,1001,151,176]" class="Malacostraca" family="Platyscelidae" genus="Platyscelus" kingdom="Animalia" order="Amphipoda" pageId="100" pageNumber="101" phylum="Arthropoda" rank="species" species="ovoides">
<emphasis id="F0B97810182EFFE7FCCDFF6BFC60FF42" box="[772,1001,151,176]" italics="true" pageId="100" pageNumber="101">Platyscelus ovoides</emphasis>
</taxonomicName>
examined only three had the right overlapping the left). In addition, the distal margin of the basis of pereopod 6, distal and anterior to the locking mechanism, is grooved so that the margins of opposing pereopods can overlap thus assisting in holding them together. The head has a central beak, which fits neatly with the antero-distal corner of the basis of pereopod 6 (see
<figureCitation id="5AF6B887182EFFE7FF07FEDBFEADFEB2" box="[206,292,295,320]" captionStart="FIGURE 49" captionStartId="101.[151,250,1937,1959]" captionTargetBox="[235,1360,208,1908]" captionTargetId="figure@101.[206,1380,193,1916]" captionTargetPageId="101" captionText="FIGURE 49. Platyscelus ovoides (Risso, 1816), female (17.5 mm) and male (15.2 mm), Tasman Sea, SAMA C 4224. A C, habitus, ventral view when rolled up, and anterior view of head of female. Scale bars = 2.0 mm (A C), 1.0 mm (G 1 &amp; 2), 0.2 mm (remainder)." httpUri="https://zenodo.org/record/166468/files/figure.png" pageId="100" pageNumber="101">Fig. 49</figureCitation>
B). Finally the epimeral plates have lateral ridges, which fit neatly against the proximal part of the posterior margin of the basis of pereopod 6, when the animal is curled. Thus, these animals are able to achieve an almost smooth, tightly packed, globular shape. The reasons for adopting this habit are not known. A spherical shape would sink more rapidly, and this may assist in the avoidance of predators, or in diurnal migrations.
</paragraph>
<paragraph id="C272A402182EFFE7FF0EFE4BFDB5FE07" blockId="100.[151,1436,151,645]" pageId="100" pageNumber="101">
The family
<taxonomicName id="05CDDF81182EFFE7FE84FE4BFE68FE22" box="[333,481,439,464]" class="Malacostraca" family="Platyscelidae" kingdom="Animalia" order="Amphipoda" pageId="100" pageNumber="101" phylum="Arthropoda" rank="family">Platyscelidae</taxonomicName>
has not been reviewed since the works of
<bibRefCitation id="A65CD9F3182EFFE7FC04FE4BFBD3FE22" author="Claus" box="[973,1114,439,464]" pageId="100" pageNumber="101" refString="Claus, C. (1879) Die Gattungen und Arten der Platysceliden in Systematischer Ubersicht. Arbeiten aus dem Zoologischen Institut der Universitat zu Wien und der Zoologischen Station Triest, 2, 1 - 52 (147 - 198)." type="journal article" year="1879">Claus (1879</bibRefCitation>
,
<bibRefCitation id="A65CD9F3182EFFE7FBAEFE44FB29FE22" author="Claus" box="[1127,1184,440,464]" pageId="100" pageNumber="101" refString="Claus, C. (1887) Die Platysceliden. Alfred Holder, Vienna, 77 pp., plates 1 - 25." type="book chapter" year="1887">1887</bibRefCitation>
) and all genera are in need of thorough taxonomic revision.
</paragraph>
<paragraph id="C272A402182EFFE7FF0EFE03FE9FFD77" blockId="100.[151,1436,151,645]" pageId="100" pageNumber="101">
Prior to this review
<taxonomicName id="05CDDF81182EFFE7FE6CFE03FDBDFDEA" box="[421,564,511,536]" class="Malacostraca" family="Platyscelidae" genus="Amphithyrus" kingdom="Animalia" order="Amphipoda" pageId="100" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F0B97810182EFFE7FE6CFE03FDBDFDEA" box="[421,564,511,536]" italics="true" pageId="100" pageNumber="101">Amphithyrus</emphasis>
</taxonomicName>
was included in this family, but this genus differs significantly from the other genera of
<taxonomicName id="05CDDF81182EFFE7FEC5FDD8FE2AFDCF" box="[268,419,548,573]" class="Malacostraca" family="Platyscelidae" kingdom="Animalia" order="Amphipoda" pageId="100" pageNumber="101" phylum="Arthropoda" rank="family">Platyscelidae</taxonomicName>
, and is accommodated in the new family, Amphithyridae, together with
<taxonomicName id="05CDDF81182EFFE7FB20FDD8FAE3FDCF" box="[1257,1386,548,573]" class="Malacostraca" family="Pronoidae" genus="Paralycaea" kingdom="Animalia" order="Amphipoda" pageId="100" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F0B97810182EFFE7FB20FDD8FAE3FDCF" box="[1257,1386,548,573]" italics="true" pageId="100" pageNumber="101">Paralycaea</emphasis>
</taxonomicName>
and
<taxonomicName id="05CDDF81182EFFE7FF5EFDBBFECFFD92" box="[151,326,583,608]" class="Malacostraca" family="Pronoidae" genus="Amphithyropsis" kingdom="Animalia" order="Amphipoda" pageId="100" pageNumber="39" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="F0B97810182EFFE7FF5EFDBBFECFFD92" box="[151,326,583,608]" italics="true" pageId="100" pageNumber="101">Amphithyropsis</emphasis>
</taxonomicName>
, with which it shares a number of significant characters. Thus, only four genera are recognised in this family.
</paragraph>
</subSubSection>
</treatment>
</document>