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<document id="37943DAE07E52195A1EA3E90AB4E953B" ID-DOI="10.11646/zootaxa.4040.1.5" ID-GBIF-Dataset="c486d3d3-9944-4850-af11-c1fbc61bdaac" ID-ISSN="1175-5326" ID-Zenodo-Dep="289985" ID-ZooBank="703E3F2B-69F7-4781-ACDE-51C6291544BF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461333711373" checkinUser="plazi" docAuthor="Henao, Efrain R., Carneiro, Eduardo, Mielke, Olaf H. H. &amp; Casagrade, Mirna M." docDate="2015" docId="4A5E87B9FFB79A12FF19B01F6A3EB502" docLanguage="en" docName="zt04040p073.pdf" docOrigin="Zootaxa 4040 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Wahydra subhebetis Steinhauser 1991" docType="treatment" docVersion="6" lastPageNumber="66" masterDocId="B667FFC1FFB19A15FF8EB75B680AB72F" masterDocTitle="Taxonomic notes on Wahydra Steinhauser (Hesperiidae, Hesperiinae, Anthoptini) with description of four new species" masterLastPageNumber="73" masterPageNumber="59" pageNumber="65" updateTime="1698618892071" updateUser="plazi">
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<mods:title id="954E18C6B12F7B22AA2400020252C918">Taxonomic notes on Wahydra Steinhauser (Hesperiidae, Hesperiinae, Anthoptini) with description of four new species</mods:title>
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<mods:namePart id="CC7DE8AAE7EA54433BC44B8E2E3B34F9">Henao, Efrain R.</mods:namePart>
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<mods:namePart id="B066FD8E8BD85CA570C0223B8465D6DE">Carneiro, Eduardo</mods:namePart>
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<mods:namePart id="15FB2ADB09D32E370FCF20E52290B9B4">Mielke, Olaf H. H.</mods:namePart>
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<mods:namePart id="21D330625DE10F78D952B96CBB20D91B">Casagrade, Mirna M.</mods:namePart>
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<mods:date id="0558E0B41F56FBAA016691638C9538D4">2015</mods:date>
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<treatment id="4A5E87B9FFB79A12FF19B01F6A3EB502" ID-DOI="http://doi.org/10.5281/zenodo.6110611" ID-GBIF-Taxon="127694068" ID-Zenodo-Dep="6110611" LSID="urn:lsid:plazi:treatment:4A5E87B9FFB79A12FF19B01F6A3EB502" httpUri="http://treatment.plazi.org/id/4A5E87B9FFB79A12FF19B01F6A3EB502" lastPageId="7" lastPageNumber="66" pageId="6" pageNumber="65">
<subSubSection id="8AED6524FFB79A13FF19B01F6966B050" pageId="6" pageNumber="65" type="nomenclature">
<paragraph id="C24836AFFFB79A13FF19B01F6A61B071" blockId="6.[151,619,1860,1919]" box="[151,619,1860,1886]" pageId="6" pageNumber="65">
<heading id="990081C3FFB79A13FF19B01F6A61B071" bold="true" box="[151,619,1860,1886]" fontSize="11" level="1" pageId="6" pageNumber="65" reason="1">
<taxonomicName id="05F74D2CFFB79A13FF19B01F6A61B071" authority="Steinhauser, 1991" authorityName="Steinhauser" authorityYear="1991" box="[151,619,1860,1886]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="subhebetis">
<emphasis id="F083EABDFFB79A13FF19B01F6A61B071" bold="true" box="[151,619,1860,1886]" pageId="6" pageNumber="65">
<emphasis id="F083EABDFFB79A13FF19B01F6980B071" bold="true" box="[151,394,1860,1886]" italics="true" pageId="6" pageNumber="65">Wahydra subhebetis</emphasis>
<bibRefCitation id="A6664B5EFFB79A13FE1FB01F6A61B071" author="Steinhauser" box="[401,619,1860,1886]" pageId="6" pageNumber="65" refString="Steinhauser, S. R. (1991) Taxonomic notes and descriptions of new taxa in the Neotropical Hesperiidae. Part 2, Heteropterinae and Hesperiinae, Vinius group. Bulletin of the Allyn Museum, 132, 1 - 79." type="journal article" year="1991">Steinhauser, 1991</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="C24836AFFFB79A13FF19B03D6966B050" blockId="6.[151,619,1860,1919]" box="[151,364,1894,1919]" pageId="6" pageNumber="65">
(
<figureCitation id="5ACC2A2AFFB79A13FF11B03D691BB050" box="[159,273,1894,1919]" captionStart="FIGURES 1 26" captionStartId="4.[151,264,1812,1834]" captionTargetBox="[188,1391,195,1811]" captionTargetId="figure@4.[188,1398,195,1907]" captionTargetPageId="4" captionText="FIGURES 1 26. Dorsal and ventral views of Wahydra Steinhauser, 1991 species. Figs 1 2: Wahydra kenava (Butler, 1870); Figs 3 4: Wahydra bella (Hayward, 1939); Figs 5 6: Wahydra nieblensis Steinhauser, 1991; Figs 7 8: Wahydra ekka (Evans, 1955) comb. nov.; Figs 9 10: Wahydra subhebetis Steinhauser, 1991; Figs 11 12: Wahydra tassa (Evans, 1955); Figs 13 16: Wahydra vola (Evans, 1955) (male left, female right); Figs 17 18: Wahydra trinitas Henao, Carneiro, Mielke &amp; Casagrande sp. nov. (Holotype).; Figs 19 20: Wahydra curtis Mielke, Henao, Carneiro &amp; Casagrande sp. nov. (Paratypus); Figs 21 22: Wahydra shueyi Carneiro, Henao, Mielke &amp; Casagrande sp. nov. (Paratypus).; Figs 23 24: Wahydra mapiriensis Casagrande, Henao, Carneiro &amp; Mielke sp. nov. (Holotype).; Figs 25 26: Wahydra sp. (unidentified female)." httpUri="https://zenodo.org/record/289986/files/figure.png" pageId="6" pageNumber="65">Figs 9, 10</figureCitation>
,
<figureCitation id="5ACC2A2AFFB79A13FE93B03C6931B050" box="[285,315,1895,1919]" captionStart="FIGURES 27 40" captionStartId="5.[151,264,1801,1823]" captionTargetBox="[197,1344,199,1760]" captionTargetId="figure@5.[191,1359,193,1770]" captionTargetPageId="5" captionText="FIGURES 27 40. Right dorsal forewing of the male of Wahydra Steinhauser, 1991 species, showing general aspect of brands (27 39). Fig. 27: Wahydra kenava (Butler, 1870); Fig. 28: Wahydra bella (Hayward, 1939); Fig. 29: Wahydra nieblensis Steinhauser, 1991; Fig. 30: Wahydra ekka (Evans, 1955) comb. nov.; Fig. 31: Wahydra subhebetis Steinhauser, 1991; Fig. 32: Wahydra tassa (Evans, 1955); Fig. 33: Wahydra vola (Evans, 1955); Fig. 34: Wahydra trinitas Carneiro, Mielke &amp; Casagrande sp. nov. (Holotype); Fig. 35: Wahydra curtis Mielke, Henao, Carneiro &amp; Casagrande sp. nov. (Paratype); Fig. 36: Wahydra shueyi Carneiro, Henao, Mielke &amp; Casagrande sp. nov. (Holotype).; Fig. 37: Wahydra mapiriensis Casagrande, Henao, Carneiro &amp; Mielke sp. nov. (Holotype); Fig. 38: Wahydra dores (Bell, 1959); Fig. 39: Wahydra thisbe (Hayward, 1942); Fig. 40: Wahydra sp. (unidentified female) dorsal discal spots." httpUri="https://zenodo.org/record/289987/files/figure.png" pageId="6" pageNumber="65">31</figureCitation>
,
<figureCitation id="5ACC2A2AFFB79A13FEC9B03D696EB050" box="[327,356,1894,1919]" captionStart="FIGURES 41 46" captionStartId="6.[151,264,1696,1718]" captionTargetBox="[163,1390,204,1633]" captionTargetId="figure@6.[151,1436,193,1675]" captionTargetPageId="6" captionText="FIGURES 41 46. Male genitalia of Wahydra Steinhauser, 1991 species. Fig. 41: Wahydra kenava (Butler, 1870); Fig. 42: Wahydra bella (Hayward, 1939); Fig. 43: Wahydra nieblensis Steinhauser, 1991; Fig. 44: Wahydra ekka (Evans, 1955) comb. nov.; Fig. 45: Wahydra subhebetis Steinhauser, 1991; Fig. 46: Wahydra tassa (Evans, 1955)." httpUri="https://zenodo.org/record/289988/files/figure.png" pageId="6" pageNumber="65">45</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="8AED6524FFB79A12FF19B0F46A49B625" lastPageId="7" lastPageNumber="66" pageId="6" pageNumber="65" type="reference_group">
<paragraph id="C24836AFFFB79A12FF19B0F46A49B625" blockId="6.[151,1436,1967,2019]" lastBlockId="7.[199,1436,152,266]" lastPageId="7" lastPageNumber="66" pageId="6" pageNumber="65">
<taxonomicName id="05F74D2CFFB79A13FF19B0F46A2BB0EA" authority="Steinhauser, 1991" authorityName="Steinhauser" authorityYear="1991" box="[151,545,1967,1989]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="subhebetis">
<emphasis id="F083EABDFFB79A13FF19B0F46969B0EA" box="[151,355,1967,1989]" italics="true" pageId="6" pageNumber="65">Wahydra subhebetis</emphasis>
<bibRefCitation id="A6664B5EFFB79A13FEE4B0F46A2BB0EA" author="Steinhauser" box="[362,545,1967,1989]" pageId="6" pageNumber="65" refString="Steinhauser, S. R. (1991) Taxonomic notes and descriptions of new taxa in the Neotropical Hesperiidae. Part 2, Heteropterinae and Hesperiinae, Vinius group. Bulletin of the Allyn Museum, 132, 1 - 79." type="journal article" year="1991">Steinhauser, 1991</bibRefCitation>
</taxonomicName>
.
<emphasis id="F083EABDFFB79A13FDA2B0F46B00B0EA" bold="true" box="[556,778,1967,1989]" pageId="6" pageNumber="65">Bull. Allyn Mus. 132</emphasis>
: 33, figs 3841 (male, female d, v), 88 (male gen.), 109 (female gen.);
<typeStatus id="1D4C880DFFB79A13FE86B096696BB0CC" box="[264,353,1997,2019]" pageId="6" pageNumber="65" type="holotype">holotype</typeStatus>
male,
<date id="B649106FFFB79A13FE2BB0966A02B0CC" box="[421,520,1997,2019]" pageId="6" pageNumber="65" value="1974-05-03">3-V-1974</date>
, La Linea (Quindio Pass), Tolima-Quindio,
<collectingCountry id="BAE0763FFFB79A13FC46B0966C24B0CC" box="[968,1070,1997,2019]" name="Colombia" pageId="6" pageNumber="65">Colombia</collectingCountry>
,
<quantity id="050F9B4AFFB79A13FBB7B0966C82B0CC" box="[1081,1160,1997,2019]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="6" pageNumber="65" unit="m" value="3100.0">3100 m</quantity>
, S. R. &amp; L. M. Steinhauser leg.; AME.—
<bibRefCitation id="A6664B5EFFB69A12FEDCB7C369E8B781" author="Bridges" box="[338,482,152,174]" pageId="7" pageNumber="66" refString="Bridges, C. A. (1994) Catalogue of the family-group, genus-group and species-group names of the Hesperioidea (Lepidoptera) of the world. Urbana, 598 pp." type="book" year="1994">Bridges, 1994</bibRefCitation>
.
<emphasis id="F083EABDFFB69A12FE65B7C36CD3B781" bold="true" box="[491,1241,152,174]" pageId="7" pageNumber="66">Cat. Fam.-Group, Gen.-Group, Sp. Group Nam. Hesp. (Lep.) World 8</emphasis>
, p. 215;
<emphasis id="F083EABDFFB69A12FABFB7C36D34B781" bold="true" box="[1329,1342,152,174]" pageId="7" pageNumber="66">9</emphasis>
, p. 74.—
<bibRefCitation id="A6664B5EFFB69A12FF49B7ED6958B7E3" author="Mielke" box="[199,338,182,204]" pageId="7" pageNumber="66" refString="Mielke, O. H. H. (2004) Hesperiidae. In: Lamas, G. (Ed.), Checklist: Part 4 A. Hesperioidea - Papilionoidea. In: Heppner, J. B. (Ed.), Atlas of Neotropical Lepidoptera. Vol. 5 A. Association for Tropical Lepidoptera, Scientific Publishers, Gainesville, pp. 25 - 86." type="book chapter" year="2004">Mielke, 2004</bibRefCitation>
. Hesperioidea, p. 84,
<emphasis id="F083EABDFFB69A12FDBDB7EC6A42B7E3" bold="true" box="[563,584,183,204]" italics="true" pageId="7" pageNumber="66">in</emphasis>
Lamas (ed.).
<emphasis id="F083EABDFFB69A12FD56B7ED6CD6B7E3" bold="true" box="[728,1244,182,204]" pageId="7" pageNumber="66">Checklist: Part 4A, Hesperioidea-Papilionoidea</emphasis>
,
<emphasis id="F083EABDFFB69A12FB65B7EC6D0AB7E3" bold="true" box="[1259,1280,183,204]" italics="true" pageId="7" pageNumber="66">in</emphasis>
Heppner (ed.).
<emphasis id="F083EABDFFB69A12FF49B78E69CAB7C4" bold="true" box="[199,448,213,235]" pageId="7" pageNumber="66">
<taxonomicName id="05F74D2CFFB69A12FF49B78E6991B7C4" authority="Neotrop. Lep." authorityName="Neotrop. Lep." box="[199,411,213,235]" class="Insecta" genus="Atlas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="genus">Atlas Neotrop. Lep.</taxonomicName>
5A
</emphasis>
.—
<bibRefCitation id="A6664B5EFFB69A12FE50B78E6A63B7C4" author="Mielke" box="[478,617,213,235]" pageId="7" pageNumber="66" refString="Mielke, O. H. H. (2005) Catalogue of the American Hesperioidea: Hesperiidae (Lepidoptera) 5: Hesperiinae 2. In: Megaleas - Zenis. Sociedade Brasileira de Zoologia, Curitiba, pp. 775 - 1055." type="book chapter" year="2005">Mielke, 2005</bibRefCitation>
.
<emphasis id="F083EABDFFB69A12FDFDB78E6B85B7C4" bold="true" box="[627,911,213,235]" pageId="7" pageNumber="66">Cat. Amer. Hesperioidea 5</emphasis>
, p. 1363.—González M. &amp; Andrade-C., 2008.
<emphasis id="F083EABDFFB69A12FAE3B78E69D8B625" bold="true" pageId="7" pageNumber="66">Rev. Acad. Colomb. Cienc. 32</emphasis>
(124): 430.
</paragraph>
</subSubSection>
<subSubSection id="8AED6524FFB69A12FF19B66D6CEFB6EE" pageId="7" pageNumber="66" type="diagnosis">
<paragraph id="C24836AFFFB69A12FF19B66D6CEFB6EE" blockId="7.[151,1436,310,557]" pageId="7" pageNumber="66">
<emphasis id="F083EABDFFB69A12FF19B66D6954B660" bold="true" box="[151,350,310,335]" pageId="7" pageNumber="66">Diagnosis. Male:</emphasis>
Forewing length
<quantity id="050F9B4AFFB69A12FDABB66D6AA4B660" box="[549,686,310,335]" metricMagnitude="-2" metricUnit="m" metricValue="1.175" metricValueMax="1.2" metricValueMin="1.15" pageId="7" pageNumber="66" unit="mm" value="11.75" valueMax="12.0" valueMin="11.5">11.512mm</quantity>
. Nudum 13. DFW brand thin, subapical spot yellowish, reduced in R4R5, but developed in R5-M1. VFW costal area, apical spot and marginal band homogeneous yellowish. VHW with fainted reddish discal spots and streak in CuA2-2A. Median apophyses of tegumen developed, thin, simple; lateral processes pointed. Harpe large, sharply angled. Ventro-posterior end of aedeagus truncated.
</paragraph>
</subSubSection>
<subSubSection id="8AED6524FFB69A12FF49B6976A3EB502" pageId="7" pageNumber="66" type="materials_examined">
<paragraph id="C24836AFFFB69A12FF49B6976C86B527" blockId="7.[151,1436,310,557]" pageId="7" pageNumber="66">
<emphasis id="F083EABDFFB69A12FF49B6976A0DB6CA" bold="true" box="[199,519,460,485]" pageId="7" pageNumber="66">
Studied material:
<collectingCountry id="BAE0763FFFB69A12FE12B6976A08B6CA" box="[412,514,460,485]" name="Ecuador" pageId="7" pageNumber="66">Ecuador</collectingCountry>
,
</emphasis>
Tungurahua, Rio Blanco,
<date id="B649106FFFB69A12FCA3B6966B89B6CA" box="[813,899,461,485]" pageId="7" pageNumber="66" value="1977-05">V-1977</date>
, Lafebre leg,
<specimenCount id="D4F1FD26FFB69A12FB95B6966C62B6CA" box="[1051,1128,460,485]" pageId="7" pageNumber="66" type="male">1 male</specimenCount>
,
<collectionCode id="A4E6AE6AFFB69A12FBFDB6966CB6B6CB" box="[1139,1212,461,484]" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="7" pageNumber="66">DZUP</collectionCode>
29.902;
<emphasis id="F083EABDFFB69A12FA90B6976D96B6CA" bold="true" box="[1310,1436,460,485]" pageId="7" pageNumber="66">
<collectingCountry id="BAE0763FFFB69A12FA90B6976D9DB6CA" box="[1310,1431,460,485]" name="Colombia" pageId="7" pageNumber="66">Colombia</collectingCountry>
:
</emphasis>
Tolima-Quindio, La Linea,
<quantity id="050F9B4AFFB69A12FE45B6AB6A16B527" box="[459,540,496,520]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="7" pageNumber="66" unit="m" value="3100.0">3100m</quantity>
,
<date id="B649106FFFB69A12FDA9B6AB6A9FB527" box="[551,661,496,520]" pageId="7" pageNumber="66" value="1974-05-03">3-V-1974</date>
. S. R. &amp; L. M. Steinhuaser,
<specimenCount id="D4F1FD26FFB69A12FC41B6AB6C22B526" box="[975,1064,496,521]" pageId="7" pageNumber="66" type="male">3 males</specimenCount>
,
<collectionCode id="A4E6AE6AFFB69A12FBBAB6AB6C82B527" box="[1076,1160,496,520]" pageId="7" pageNumber="66">MGCL</collectionCode>
.
</paragraph>
<paragraph id="C24836AFFFB69A12FF49B54F6A3EB502" blockId="7.[151,1436,310,557]" box="[199,564,532,557]" pageId="7" pageNumber="66">
<emphasis id="F083EABDFFB69A12FF49B54F6900B502" bold="true" box="[199,266,532,557]" pageId="7" pageNumber="66">
<typeStatus id="1D4C880DFFB69A12FF49B54F690CB502" box="[199,262,532,557]" pageId="7" pageNumber="66">Type</typeStatus>
:
</emphasis>
<typeStatus id="1D4C880DFFB69A12FE9CB54F6970B502" box="[274,378,532,557]" pageId="7" pageNumber="66" type="holotype">Holotype</typeStatus>
male in
<collectionCode id="A4E6AE6AFFB69A12FE55B54E6A25B502" box="[475,559,533,557]" pageId="7" pageNumber="66">MGCL</collectionCode>
.
</paragraph>
</subSubSection>
</treatment>
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