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<mods:title id="FA7AFE2BC77BC4AC774346B170D7D3D4">Taxonomic notes on Wahydra Steinhauser (Hesperiidae, Hesperiinae, Anthoptini) with description of four new species</mods:title>
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<mods:namePart id="050320658D8506CA82525455B1A030A3">Henao, Efrain R.</mods:namePart>
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<mods:namePart id="A145677D16D6E587C89EE86763273CDD">Carneiro, Eduardo</mods:namePart>
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<mods:namePart id="7413D9FAC28F4AB147381DC29055A4C0">Mielke, Olaf H. H.</mods:namePart>
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<mods:namePart id="09385197D8731B0B2D73F347D57255E0">Casagrade, Mirna M.</mods:namePart>
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<treatment id="4A5E87B9FFB69A1DFF19B1A26A25B5B5" ID-DOI="http://doi.org/10.5281/zenodo.6110616" ID-GBIF-Taxon="127694069" ID-Zenodo-Dep="6110616" LSID="urn:lsid:plazi:treatment:4A5E87B9FFB69A1DFF19B1A26A25B5B5" httpUri="http://treatment.plazi.org/id/4A5E87B9FFB69A1DFF19B1A26A25B5B5" lastPageId="8" lastPageNumber="67" pageId="7" pageNumber="66">
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<paragraph id="C24836AFFFB69A12FF19B1A269E7B03B" blockId="7.[151,503,1785,1845]" box="[151,493,1785,1812]" pageId="7" pageNumber="66">
<heading id="990081C3FFB69A12FF19B1A269E7B03B" bold="true" box="[151,493,1785,1812]" fontSize="11" level="1" pageId="7" pageNumber="66" reason="1">
<taxonomicName id="05F74D2CFFB69A12FF19B1A269E7B03B" authority="Evans, 1955" authorityName="Evans" authorityYear="1955" box="[151,493,1785,1812]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="vola">
<emphasis id="F083EABDFFB69A12FF19B1A269E7B03B" bold="true" box="[151,493,1785,1812]" pageId="7" pageNumber="66">
<emphasis id="F083EABDFFB69A12FF19B1A26934B03C" bold="true" box="[151,318,1785,1811]" italics="true" pageId="7" pageNumber="66">Wahydra vola</emphasis>
(
<bibRefCitation id="A6664B5EFFB69A12FEC3B1A169EFB03B" author="Evans" box="[333,485,1786,1812]" pageId="7" pageNumber="66" refString="Evans, W. H. (1955) A Catalogue of the American Hesperiidae: indicating the classification and nomenclature adopted in the British Museum (Natural History). British Museum (Natural History), London. IV, 469 pp." type="book" year="1955">Evans, 1955</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="C24836AFFFB69A12FF19B04769FDB01A" blockId="7.[151,503,1785,1845]" box="[151,503,1820,1845]" pageId="7" pageNumber="66">
(
<figureCitation id="5ACC2A2AFFB69A12FF11B047692AB01A" box="[159,288,1820,1845]" captionStart="FIGURES 1 26" captionStartId="4.[151,264,1812,1834]" captionTargetBox="[188,1391,195,1811]" captionTargetId="figure@4.[188,1398,195,1907]" captionTargetPageId="4" captionText="FIGURES 1 26. Dorsal and ventral views of Wahydra Steinhauser, 1991 species. Figs 1 2: Wahydra kenava (Butler, 1870); Figs 3 4: Wahydra bella (Hayward, 1939); Figs 5 6: Wahydra nieblensis Steinhauser, 1991; Figs 7 8: Wahydra ekka (Evans, 1955) comb. nov.; Figs 9 10: Wahydra subhebetis Steinhauser, 1991; Figs 11 12: Wahydra tassa (Evans, 1955); Figs 13 16: Wahydra vola (Evans, 1955) (male left, female right); Figs 17 18: Wahydra trinitas Henao, Carneiro, Mielke &amp; Casagrande sp. nov. (Holotype).; Figs 19 20: Wahydra curtis Mielke, Henao, Carneiro &amp; Casagrande sp. nov. (Paratypus); Figs 21 22: Wahydra shueyi Carneiro, Henao, Mielke &amp; Casagrande sp. nov. (Paratypus).; Figs 23 24: Wahydra mapiriensis Casagrande, Henao, Carneiro &amp; Mielke sp. nov. (Holotype).; Figs 25 26: Wahydra sp. (unidentified female)." httpUri="https://zenodo.org/record/289986/files/figure.png" pageId="7" pageNumber="66">Figs 1316</figureCitation>
,
<figureCitation id="5ACC2A2AFFB69A12FEA5B0466943B01A" box="[299,329,1821,1845]" captionStart="FIGURES 27 40" captionStartId="5.[151,264,1801,1823]" captionTargetBox="[197,1344,199,1760]" captionTargetId="figure@5.[191,1359,193,1770]" captionTargetPageId="5" captionText="FIGURES 27 40. Right dorsal forewing of the male of Wahydra Steinhauser, 1991 species, showing general aspect of brands (27 39). Fig. 27: Wahydra kenava (Butler, 1870); Fig. 28: Wahydra bella (Hayward, 1939); Fig. 29: Wahydra nieblensis Steinhauser, 1991; Fig. 30: Wahydra ekka (Evans, 1955) comb. nov.; Fig. 31: Wahydra subhebetis Steinhauser, 1991; Fig. 32: Wahydra tassa (Evans, 1955); Fig. 33: Wahydra vola (Evans, 1955); Fig. 34: Wahydra trinitas Carneiro, Mielke &amp; Casagrande sp. nov. (Holotype); Fig. 35: Wahydra curtis Mielke, Henao, Carneiro &amp; Casagrande sp. nov. (Paratype); Fig. 36: Wahydra shueyi Carneiro, Henao, Mielke &amp; Casagrande sp. nov. (Holotype).; Fig. 37: Wahydra mapiriensis Casagrande, Henao, Carneiro &amp; Mielke sp. nov. (Holotype); Fig. 38: Wahydra dores (Bell, 1959); Fig. 39: Wahydra thisbe (Hayward, 1942); Fig. 40: Wahydra sp. (unidentified female) dorsal discal spots." httpUri="https://zenodo.org/record/289987/files/figure.png" pageId="7" pageNumber="66">33</figureCitation>
,
<figureCitation id="5ACC2A2AFFB69A12FEDBB0466991B01A" box="[341,411,1820,1845]" captionStart="FIGURES 47 53" captionStartId="9.[151,264,1636,1658]" captionTargetBox="[152,1427,193,1636]" captionTargetId="figure@9.[151,1436,171,1639]" captionTargetPageId="9" captionText="FIGURES 47 53. Male genitalia of Wahydra Steinhauser, 1991 species. Fig. 47: Wahydra vola (Evans, 1955); Fig. 48: Wahydra trinitas Henao, Carneiro, Mielke &amp; Casagrande sp. nov. (Paratype); Fig. 49: Wahydra curtis Mielke, Henao, Carneiro &amp; Casagrande sp. nov. (Paratype); Fig. 50: Wahydra shueyi Carneiro, Henao, Mielke &amp; Casagrande sp. nov. (Paratype); Fig. 51: Wahydra mapiriensis Casagrande, Henao, Carneiro &amp; Mielke sp. nov. (Holotype). Figs 52 to 53: Female genitalia of Wahydra Steinhauser, 1991 species. Fig. 52: Wahydra vola (Evans, 1955); Fig. 53: Wahydra sp. (unidentified female)." httpUri="https://zenodo.org/record/289989/files/figure.png" pageId="7" pageNumber="66">
47,
<specimenCount id="D4F1FD26FFB69A12FEF1B0476991B01A" box="[383,411,1820,1845]" pageId="7" pageNumber="66" type="female">52</specimenCount>
</figureCitation>
female)
</paragraph>
</subSubSection>
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<paragraph id="C24836AFFFB69A12FF19B03F6AB3B098" blockId="7.[151,1437,1892,2037]" pageId="7" pageNumber="66">
<taxonomicName id="05F74D2CFFB69A12FF19B03F69FCB055" ID-CoL="5LTRK" authority="Evans, 1955" authorityName="Evans" authorityYear="1955" box="[151,502,1892,1914]" class="Insecta" family="Hesperiidae" genus="Zalomes" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="subSpecies" species="kenava" subSpecies="vola">
<emphasis id="F083EABDFFB69A12FF19B03F6964B055" box="[151,366,1892,1914]" italics="true" pageId="7" pageNumber="66">Zalomes kenava vola</emphasis>
<bibRefCitation id="A6664B5EFFB69A12FEFBB03E69FCB055" author="Evans" box="[373,502,1892,1914]" pageId="7" pageNumber="66" refString="Evans, W. H. (1955) A Catalogue of the American Hesperiidae: indicating the classification and nomenclature adopted in the British Museum (Natural History). British Museum (Natural History), London. IV, 469 pp." type="book" year="1955">Evans, 1955</bibRefCitation>
</taxonomicName>
.
<emphasis id="F083EABDFFB69A12FD8EB03F6AC4B055" bold="true" box="[512,718,1892,1914]" pageId="7" pageNumber="66">Cat. Amer. Hesp. 4</emphasis>
, p. 69; [holo]
<typeStatus id="1D4C880DFFB69A12FCD6B03F6B8CB055" box="[856,902,1892,1914]" pageId="7" pageNumber="66">type</typeStatus>
male,
<date id="B649106FFFB69A12FC45B03F6C28B055" box="[971,1058,1892,1914]" pageId="7" pageNumber="66" value="1904-04">IV-1904</date>
, dry season, Carabaya, S. E.
<collectingCountry id="BAE0763FFFB69A12FAC4B03E6D71B055" box="[1354,1403,1893,1914]" name="Peru" pageId="7" pageNumber="66">Peru</collectingCountry>
, G. Ockenden. leg.; BM(NH).—
<bibRefCitation id="A6664B5EFFB69A12FE66B0D86A70B0B6" author="Bridges" box="[488,634,1923,1945]" pageId="7" pageNumber="66" refString="Bridges, C. A. (1983) Lepidoptera: Hesperiidae. Notes on species-group names. Urbana, 13 pp." type="book" year="1983">Bridges, 1983</bibRefCitation>
.
<emphasis id="F083EABDFFB69A12FD0AB0D86B00B0B6" bold="true" box="[644,778,1923,1945]" pageId="7" pageNumber="66">Lep. Hesp. 1</emphasis>
, p. 116;
<emphasis id="F083EABDFFB69A12FCEDB0D86B65B0B6" bold="true" box="[867,879,1923,1945]" pageId="7" pageNumber="66">2</emphasis>
, p. 40.—
<bibRefCitation id="A6664B5EFFB69A12FC40B0D86C6AB0B6" author="Bridges" box="[974,1120,1923,1945]" pageId="7" pageNumber="66" refString="Bridges, C. A. (1988) Catalogue of Hesperiidae (Lepidoptera: Rhopalocera). Urbana, 10 pp." type="book" year="1988">Bridges, 1988</bibRefCitation>
.
<emphasis id="F083EABDFFB69A12FBE7B0D86CE4B0B6" bold="true" box="[1129,1262,1923,1945]" pageId="7" pageNumber="66">Cat. Hesp. 1</emphasis>
, p. 199;
<emphasis id="F083EABDFFB69A12FAC9B0D86D5EB0B6" bold="true" box="[1351,1364,1923,1945]" pageId="7" pageNumber="66">2</emphasis>
, p. 66. (no genus)
<taxonomicName id="05F74D2CFFB69A12FE87B0FA693FB098" box="[265,309,1953,1975]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="vola">
<emphasis id="F083EABDFFB69A12FE87B0FA693FB098" box="[265,309,1953,1975]" italics="true" pageId="7" pageNumber="66">vola</emphasis>
</taxonomicName>
;
<bibRefCitation id="A6664B5EFFB69A12FECCB0FA69C6B098" author="Beattie" box="[322,460,1953,1975]" pageId="7" pageNumber="66" refString="Beattie, J. R. (1976) The Rhopalocera directory. Insecta directory. Berkeley, JB Indexes, 365 pp." type="book" year="1976">Beattie, 1976</bibRefCitation>
.
<emphasis id="F083EABDFFB69A12FE5BB0FA6A6AB098" bold="true" box="[469,608,1953,1975]" pageId="7" pageNumber="66">Rhop. Direct</emphasis>
., p. 286.
</paragraph>
<paragraph id="C24836AFFFB69A1DFF19B09B6CF5B7C4" blockId="7.[151,1437,1892,2037]" lastBlockId="8.[199,1436,152,235]" lastPageId="8" lastPageNumber="67" pageId="7" pageNumber="66">
<taxonomicName id="05F74D2CFFB69A12FF19B09B692CB0F9" box="[151,294,1984,2006]" class="Insecta" family="Hesperiidae" genus="Wahydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="vola">
<emphasis id="F083EABDFFB69A12FF19B09B692CB0F9" box="[151,294,1984,2006]" italics="true" pageId="7" pageNumber="66">Wahydra vola</emphasis>
</taxonomicName>
;
<bibRefCitation id="A6664B5EFFB69A12FEB8B09B69FAB0F9" author="Steinhauser" box="[310,496,1984,2006]" pageId="7" pageNumber="66" refString="Steinhauser, S. R. (1991) Taxonomic notes and descriptions of new taxa in the Neotropical Hesperiidae. Part 2, Heteropterinae and Hesperiinae, Vinius group. Bulletin of the Allyn Museum, 132, 1 - 79." type="journal article" year="1991">Steinhauser, 1991</bibRefCitation>
.
<emphasis id="F083EABDFFB69A12FE72B09B6AEBB0F9" bold="true" box="[508,737,1984,2006]" pageId="7" pageNumber="66">Bull. Allyn Mus. 132</emphasis>
: 32.—
<bibRefCitation id="A6664B5EFFB69A12FCA7B09B6BB4B0F9" author="Bridges" box="[809,958,1984,2006]" pageId="7" pageNumber="66" refString="Bridges, C. A. (1994) Catalogue of the family-group, genus-group and species-group names of the Hesperioidea (Lepidoptera) of the world. Urbana, 598 pp." type="book" year="1994">Bridges, 1994</bibRefCitation>
.
<emphasis id="F083EABDFFB69A12FC44B09B69FCB0DA" bold="true" pageId="7" pageNumber="66">Cat. Fam.-Group, Gen.-Group, Sp. Group Nam. Hesp. (Lep.) World 8</emphasis>
, p. 239;
<emphasis id="F083EABDFFB69A12FDD9B0846A6EB0DA" bold="true" box="[599,612,2015,2037]" pageId="7" pageNumber="66">9</emphasis>
, p. 74.—Lamas; Grados &amp; Valencia, 1999.
<emphasis id="F083EABDFFB69A12FBA4B0846D1FB0DA" bold="true" box="[1066,1301,2015,2037]" pageId="7" pageNumber="66">Rev. peruana Ent. 41</emphasis>
: 3.—
<bibRefCitation id="A6664B5EFFB69A1DFADEB08468F7B781" author="Lamas" lastPageId="8" lastPageNumber="67" pageId="7" pageNumber="66" refString="Lamas, G. (2003) Las Mariposas de Machu Picchu. Guia ilustrada de las mariposas del Santuario Historico Machu Picchu, Cuzco, Peru. Lima, Profonanpe, 221 pp." type="book" year="2003">Lamas, 2003</bibRefCitation>
.
<emphasis id="F083EABDFFB99A1DFE84B7C369F7B781" bold="true" box="[266,509,152,174]" pageId="8" pageNumber="67">Marip. Machu Picchu</emphasis>
, p. 30, 178, 189, 214, fig. 57 (d).—
<bibRefCitation id="A6664B5EFFB99A1DFC0EB7C36C1BB781" author="Mielke" box="[896,1041,152,174]" pageId="8" pageNumber="67" refString="Mielke, O. H. H. (2004) Hesperiidae. In: Lamas, G. (Ed.), Checklist: Part 4 A. Hesperioidea - Papilionoidea. In: Heppner, J. B. (Ed.), Atlas of Neotropical Lepidoptera. Vol. 5 A. Association for Tropical Lepidoptera, Scientific Publishers, Gainesville, pp. 25 - 86." type="book chapter" year="2004">Mielke, 2004</bibRefCitation>
. Hesperioidea, p. 84,
<emphasis id="F083EABDFFB99A1DFB74B7C36D05B782" bold="true" box="[1274,1295,152,173]" italics="true" pageId="8" pageNumber="67">in</emphasis>
Lamas (ed.).
<emphasis id="F083EABDFFB99A1DFF49B7ED6ADFB7E3" bold="true" box="[199,725,182,204]" pageId="8" pageNumber="67">Checklist: Part 4A, Hesperioidea-Papilionoidea</emphasis>
,
<emphasis id="F083EABDFFB99A1DFD68B7EC6AF1B7E3" bold="true" box="[742,763,183,204]" italics="true" pageId="8" pageNumber="67">in</emphasis>
Heppner (ed.).
<emphasis id="F083EABDFFB99A1DFC24B7ED6CBBB7E3" bold="true" box="[938,1201,182,204]" pageId="8" pageNumber="67">
<taxonomicName id="05F74D2CFFB99A1DFC24B7ED6C8DB7E3" authority="Neotrop. Lep." authorityName="Neotrop. Lep." box="[938,1159,182,204]" class="Insecta" genus="Atlas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="67" phylum="Arthropoda" rank="genus">Atlas Neotrop. Lep.</taxonomicName>
5A
</emphasis>
.—
<bibRefCitation id="A6664B5EFFB99A1DFB5EB7ED6D6AB7E3" author="Mielke" box="[1232,1376,182,204]" pageId="8" pageNumber="67" refString="Mielke, O. H. H. (2005) Catalogue of the American Hesperioidea: Hesperiidae (Lepidoptera) 5: Hesperiinae 2. In: Megaleas - Zenis. Sociedade Brasileira de Zoologia, Curitiba, pp. 775 - 1055." type="book chapter" year="2005">Mielke, 2005</bibRefCitation>
.
<emphasis id="F083EABDFFB99A1DFAE0B7ED69BAB7C4" bold="true" pageId="8" pageNumber="67">Cat. Amer. Hesperioidea 5</emphasis>
, p. 1364.—Gareca
<emphasis id="F083EABDFFB99A1DFDF8B78E6AA9B7C4" bold="true" box="[630,675,212,235]" italics="true" pageId="8" pageNumber="67">et al</emphasis>
., 2006,
<emphasis id="F083EABDFFB99A1DFD7BB78E6B00B7C5" bold="true" box="[757,778,213,234]" italics="true" pageId="8" pageNumber="67">in</emphasis>
Gareca &amp; Reichle.
<emphasis id="F083EABDFFB99A1DFC5CB78E6CB2B7C4" bold="true" box="[978,1208,213,235]" pageId="8" pageNumber="67">
Marip. diurn.
<collectingCountry id="BAE0763FFFB99A1DFBE2B78E6CB2B7C4" box="[1132,1208,213,235]" name="Bolivia" pageId="8" pageNumber="67">Bolivia</collectingCountry>
</emphasis>
, p. 25.
</paragraph>
</subSubSection>
<subSubSection id="8AED6524FFB99A1DFF19B64C6CECB6C8" pageId="8" pageNumber="67" type="diagnosis">
<paragraph id="C24836AFFFB99A1DFF19B64C6CECB6C8" blockId="8.[151,1436,279,667]" pageId="8" pageNumber="67">
<emphasis id="F083EABDFFB99A1DFF19B64C696BB61F" bold="true" box="[151,353,279,304]" pageId="8" pageNumber="67">Diagnosis. Male:</emphasis>
Forewing length
<quantity id="050F9B4AFFB99A1DFDBCB64C6AB1B61F" box="[562,699,279,304]" metricMagnitude="-2" metricUnit="m" metricValue="1.275" metricValueMax="1.4" metricValueMin="1.15" pageId="8" pageNumber="67" unit="mm" value="12.75" valueMax="14.0" valueMin="11.5">11.514mm</quantity>
. Nudum 13. DFW brand thin; subapical spot orange, in R5-M1. VFW costal area, apical spot and marginal band homogeneous yellowish. VHW with faint yellowish discal bands and reddish discal spots. Median apophyses of tegumen developed, large, proximally constricted, simple; lateral projections pointed. Harpe thin, smoothly angled. Ventro-posterior end of aedeagus triangular. Lamella antevaginalis projected below ostium bursae, and with symmetrical triangular posterior projections besides the ostium bursae. Lobes of posterior margin of lamella postvaginalis large, medially shallow divided.
</paragraph>
</subSubSection>
<subSubSection id="8AED6524FFB99A1DFF49B6A96A25B5B5" pageId="8" pageNumber="67" type="materials_examined">
<paragraph id="C24836AFFFB99A1DFF49B6A96ABEB558" blockId="8.[151,1436,279,667]" pageId="8" pageNumber="67">
<emphasis id="F083EABDFFB99A1DFF49B6A969D0B524" bold="true" box="[199,474,498,523]" pageId="8" pageNumber="67">
Studied material:
<collectingCountry id="BAE0763FFFB99A1DFE2FB6A969D0B524" box="[417,474,498,523]" name="Peru" pageId="8" pageNumber="67">Peru</collectingCountry>
</emphasis>
, Cuzco, Machu Picchu, Camino Inca,
<quantity id="050F9B4AFFB99A1DFC18B6A86BEEB524" box="[918,996,499,523]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="8" pageNumber="67" unit="m" value="2400.0">2400m</quantity>
<date id="B649106FFFB99A1DFC62B6A96C85B524" box="[1004,1167,498,523]" pageId="8" pageNumber="67" value="2001-10-22" valueMax="2001-10-26" valueMin="2001-10-22">22-26-X-2001</date>
, Mielke leg,
<specimenCount id="D4F1FD26FFB99A1DFAA8B6A86D8BB524" box="[1318,1409,498,523]" pageId="8" pageNumber="67" type="male">9 males</specimenCount>
,
<specimenCount id="D4F1FD26FFB99A1DFA00B6A868FBB500" pageId="8" pageNumber="67" type="female">2 females</specimenCount>
, OM 55.848, OM 55.960, OM 55.967, OM 56.016, OM 56.093, OM 56.107, OM 56.128, OM 56.135, OM 56.142, OM 56.247, OM 56.339; Urubamba,
<date id="B649106FFFB99A1DFD2BB5606B35B57C" box="[677,831,571,595]" pageId="8" pageNumber="67" value="1973-08-28">28-VIII-1973</date>
,
<specimenCount id="D4F1FD26FFB99A1DFCC2B5606BB9B57C" box="[844,947,570,595]" pageId="8" pageNumber="67" type="female">1 female</specimenCount>
,
<collectionCode id="A4E6AE6AFFB99A1DFC4FB5606C00B57D" box="[961,1034,571,594]" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="8" pageNumber="67">DZUP</collectionCode>
29.912; Puno, Carabaya, Limbani, dry S.
<quantity id="050F9B4AFFB99A1DFF6FB505693EB558" box="[225,308,606,631]" metricMagnitude="3" metricUnit="m" metricValue="2.8956" pageId="8" pageNumber="67" unit="ft" value="9500.0">9500 ft</quantity>
, 04-04-?, G. Ockenden in
<collectionCode id="A4E6AE6AFFB99A1DFDD6B53B6ABAB558" box="[600,688,608,631]" pageId="8" pageNumber="67">BMNH</collectionCode>
.
</paragraph>
<paragraph id="C24836AFFFB99A1DFF49B5D96A25B5B5" blockId="8.[151,1436,279,667]" box="[199,559,642,667]" pageId="8" pageNumber="67">
<emphasis id="F083EABDFFB99A1DFF49B5D96900B5B4" bold="true" box="[199,266,642,667]" pageId="8" pageNumber="67">
<typeStatus id="1D4C880DFFB99A1DFF49B5D9690CB5B4" box="[199,262,642,667]" pageId="8" pageNumber="67">Type</typeStatus>
:
</emphasis>
<typeStatus id="1D4C880DFFB99A1DFE9CB5D96970B5B4" box="[274,378,642,667]" pageId="8" pageNumber="67" type="holotype">Holotype</typeStatus>
male in
<collectionCode id="A4E6AE6AFFB99A1DFE55B5D86A25B5B5" box="[475,559,643,666]" pageId="8" pageNumber="67">BMNH</collectionCode>
</paragraph>
</subSubSection>
</treatment>
</document>