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<document id="CFBB0FD9DAF11E931EF2BF194C9586DF" ID-CLB-Dataset="22090" ID-DOI="10.5852/ejt.2017.331" ID-GBIF-Dataset="acf565ee-29ba-45e7-9ef3-d1e85dec9ae7" ID-ISSN="2118-9773" ID-Zenodo-Dep="3829776" ID-ZooBank="3FD5D922-B91F-42BB-9D7B-7DFB98841732" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1589572797790" checkinUser="carolina" docAuthor="Ikeda, Hiroshi &amp; Yoshitomi, Hiroyuki" docDate="2017" docId="4A5D7055450DFFF3FDE3BAEBFE4FFB63" docLanguage="en" docName="ejt-2017-331.pdf.imf" docOrigin="European Journal of Taxonomy 331" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Intybia tsushimensis" docType="treatment" docVersion="7" lastPageNumber="18" masterDocId="B664082D4503FFE2FFE6BC65FF8CFFBE" masterDocTitle="Revision of the genus Intybia (Coleoptera, Malachiidae) from Japan" masterLastPageNumber="31" masterPageNumber="1" pageNumber="15" updateTime="1698839968837" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="EEABDCB0DF362DF71DE95BC7F0E6504E">Revision of the genus Intybia (Coleoptera, Malachiidae) from Japan</mods:title>
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<mods:namePart id="1E24593A31DB85A7CB7F53327C1EEF87">Ikeda, Hiroshi</mods:namePart>
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<mods:namePart id="0545743612F7FFB28486CE3DB8971342">Yoshitomi, Hiroyuki</mods:namePart>
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<mods:affiliation id="55DD4F8AFCEA4EEFA3FB24F0E99333F1">Email: hymushi @ agr. ehime-u. ac. jp &amp; urn: lsid: zoobank. org: author: 744 FA 6 FC- 2 D 84 - 41 B 7 - 8 A 6 C-C 1611 CC 68 EE 1</mods:affiliation>
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<treatment id="4A5D7055450DFFF3FDE3BAEBFE4FFB63" ID-DOI="http://doi.org/10.5281/zenodo.3848092" ID-GBIF-Taxon="164161045" ID-Zenodo-Dep="3848092" LSID="urn:lsid:plazi:treatment:4A5D7055450DFFF3FDE3BAEBFE4FFB63" httpUri="http://treatment.plazi.org/id/4A5D7055450DFFF3FDE3BAEBFE4FFB63" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15">
<subSubSection id="8AEE92C8450DFFECFDE3BAEBFBCEF972" pageId="14" pageNumber="15" type="nomenclature">
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<taxonomicName id="05F4BAC0450DFFECFDE3BAEBFBA2F917" authority="(Sato &amp; Ohbayashi, 1968)" baseAuthorityName="Sato &amp; Ohbayashi" baseAuthorityYear="1968" box="[517,1070,1678,1705]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="tsushimensis">
<emphasis id="F0801D51450DFFECFDE3BAEBFD7FF916" box="[517,755,1678,1704]" italics="true" pageId="14" pageNumber="15">Intybia tsushimensis</emphasis>
(Satô &amp; Ohbayashi, 1968)
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[Japanese name: Tsushima-hiro-obi-jôkai-modoki]
</paragraph>
</subSubSection>
<subSubSection id="8AEE92C8450DFFECFDD4BAB0FB8EF951" box="[562,1026,1749,1776]" pageId="14" pageNumber="15" type="description">
<paragraph id="C24BC143450DFFECFDD4BAB0FB8EF951" blockId="14.[497,1090,1678,1776]" box="[562,1026,1749,1776]" pageId="14" pageNumber="15">
<figureCitation id="5ACFDDC6450DFFECFDD4BAB0FD1CF951" box="[562,656,1749,1775]" captionStart="Fig" captionStartId="4.[189,232,1893,1919]" captionTargetBox="[198,1389,740,1831]" captionTargetId="figure@4.[189,1398,723,1853]" captionTargetPageId="4" captionText="Fig. 1. Dorsal habitus Intybia spp., ♂. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov., holotype. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955). H. I. kawasakii (Nakane, 1956), holotype. Scales = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3829778" httpUri="https://zenodo.org/record/3829778/files/figure.png" pageId="14" pageNumber="15">Figs 1B</figureCitation>
,
<figureCitation id="5ACFDDC6450DFFECFD7BBAB0FD4EF951" box="[669,706,1749,1775]" captionStart="Fig" captionStartId="5.[190,233,1894,1920]" captionTargetBox="[195,1390,781,1842]" captionTargetId="figure@5.[190,1399,771,1854]" captionTargetPageId="5" captionText="Fig. 2. Dosal habitus of Intybia spp., ♀. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov., paratype. D. I. pelegrini pelegrini (Pic, 1910). E. I.niponica (Lewis, 1895). F. I.takaraensis (Nakane, 1955).G. I. kishiii (Nakane, 1955). H. I. kawasakii (Nakane, 1956), paratype. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3829782" httpUri="https://zenodo.org/record/3829782/files/figure.png" pageId="14" pageNumber="15">2B</figureCitation>
,
<figureCitation id="5ACFDDC6450DFFECFD28BAB0FD7EF951" box="[718,754,1749,1775]" captionStart="Fig" captionStartId="7.[190,233,1895,1921]" captionTargetBox="[265,1321,271,1836]" captionTargetId="figure@7.[246,1331,265,1859]" captionTargetPageId="7" captionText="Fig. 4. Right antennae of Intybia spp., ♂. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955). H. I. kawasakii (Nakane, 1956). Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3829786" httpUri="https://zenodo.org/record/3829786/files/figure.png" pageId="14" pageNumber="15">4B</figureCitation>
,
<figureCitation id="5ACFDDC6450DFFECFD19BAB0FCAFF94E" box="[767,803,1749,1776]" captionStart="Fig" captionStartId="8.[190,233,1894,1920]" captionTargetBox="[285,1298,276,1852]" captionTargetId="figure@8.[273,1302,265,1859]" captionTargetPageId="8" captionText="Fig. 5. Right antennae of Intybia spp., ♀. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955). H. I. kawasakii (Nakane, 1956). Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3829788" httpUri="https://zenodo.org/record/3829788/files/figure.png" pageId="14" pageNumber="15">5B</figureCitation>
,
<figureCitation id="5ACFDDC6450DFFECFCD6BAB0FCD9F951" box="[816,853,1749,1775]" captionStart="Fig" captionStartId="9.[190,233,1895,1921]" captionTargetBox="[263,1324,265,1858]" captionTargetId="figure@9.[263,1324,265,1859]" captionTargetPageId="9" captionText="Fig. 6. SEM photographs of right male antennomere III of Intybia spp. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C, H. I. donan sp. nov., showing projection (H). D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955)." figureDoi="http://doi.org/10.5281/zenodo.3829790" httpUri="https://zenodo.org/record/3829790/files/figure.png" pageId="14" pageNumber="15">6B</figureCitation>
,
<figureCitation id="5ACFDDC6450DFFECFC87BAB0FC09F951" box="[865,901,1749,1775]" captionStart="Fig" captionStartId="12.[190,233,1893,1919]" captionTargetBox="[191,1394,543,1850]" captionTargetId="figure@12.[189,1398,538,1851]" captionTargetPageId="12" captionText="Fig. 9. Spinous area and spinous plate of Intybia spp. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica(Lewis,1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955).Abbreviations: sa = spinous area; sp = spinous plate. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829796" httpUri="https://zenodo.org/record/3829796/files/figure.png" pageId="14" pageNumber="15">9B</figureCitation>
,
<figureCitation id="5ACFDDC6450DFFECFC74BAB0FC48F951" box="[914,964,1749,1775]" captionStart="Fig" captionStartId="13.[190,233,1893,1919]" captionTargetBox="[216,1374,635,1827]" captionTargetId="figure@13.[188,1397,614,1853]" captionTargetPageId="13" captionText="Fig. 10. Endophallic sclerites of Intybia spp. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955). Abbreviations: gp = gonoporal piece; l = ligula; sgp = semi-gonoporal piece. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829798" httpUri="https://zenodo.org/record/3829798/files/figure.png" pageId="14" pageNumber="15">10B</figureCitation>
,
<figureCitation id="5ACFDDC6450DFFECFC37BAB0FB8EF951" box="[977,1026,1749,1775]" captionStart="Fig" captionStartId="15.[190,233,1895,1921]" captionTargetBox="[196,1385,1174,1842]" captionTargetId="figure@15.[192,1400,1167,1854]" captionTargetPageId="15" captionText="Fig. 11.Map showing thedistribution of Intybia spp.A.The histriogroup:●:Intybia histrio(Kiesenwetter, 1874), ▲: I. tsushimensis (Satô &amp; Ohbayashi, 1968), ■: I. donan sp. nov.). B. The pelegrini group: ○: I. pelegrini (Pic, 1910), △: I. niponica (Lewis, 1895), □: I. takaraensis (Nakane, 1955), ×: I. kishiii (Nakane, 1955), +: I. kawasakii (Nakane, 1956)." figureDoi="http://doi.org/10.5281/zenodo.3829800" httpUri="https://zenodo.org/record/3829800/files/figure.png" pageId="14" pageNumber="15">11A</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="C24BC143450DFFECFF5BBB72FD6CF88F" blockId="14.[189,1258,1814,1841]" box="[189,736,1814,1841]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="E2E4E66D450DFFECFF5BBB72FD6CF88F" box="[189,736,1814,1841]" pageId="14" pageNumber="15">
<treatmentCitation id="4355E752450DFFECFF5BBB72FD6CF88F" box="[189,736,1814,1841]" pageId="14" pageNumber="15">
<taxonomicName id="05F4BAC0450DFFECFF5BBB72FD6CF88F" authority="Sato &amp; Ohbayashi, 1968: 5" authorityName="Sato &amp; Ohbayashi" authorityPageNumber="5" authorityYear="1968" box="[189,736,1814,1841]" class="Insecta" family="Melyridae" genus="Laius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="tsushimensis">
<emphasis id="F0801D51450DFFECFF5BBB72FE16F88E" box="[189,410,1814,1840]" italics="true" pageId="14" pageNumber="15">Laius tsushimensis</emphasis>
<bibRefCitation id="A665BCB2450DFFECFE47BB73FD6CF88F" box="[417,736,1814,1841]" pageId="14" pageNumber="15">Satô &amp; Ohbayashi, 1968: 5</bibRefCitation>
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</paragraph>
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<subSubSection id="8AEE92C8450DFFECFD01BB72FB66F88F" box="[743,1258,1814,1841]" pageId="14" pageNumber="15" type="type_taxon">
<paragraph id="C24BC143450DFFECFD01BB72FB66F88F" blockId="14.[189,1258,1814,1841]" box="[743,1258,1814,1841]" pageId="14" pageNumber="15">[type locality: Mt. Ariake-yama, Tsushima].</paragraph>
</subSubSection>
<subSubSection id="8AEE92C8450DFFECFF5BBB3CFEDCF806" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="C24BC143450DFFECFF5BBB3CFB79F8CC" blockId="14.[189,1398,1879,1976]" box="[189,1269,1879,1906]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="E2E4E66D450DFFECFF5BBB3CFB79F8CC" box="[189,1269,1879,1906]" pageId="14" pageNumber="15">
<taxonomicName id="05F4BAC0450DFFECFF5BBB3CFE16F8CC" authorityName="Sato &amp; Ohbayashi" authorityYear="1968" box="[189,410,1880,1906]" class="Insecta" family="Melyridae" genus="Laius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="tsushimensis">
<emphasis id="F0801D51450DFFECFF5BBB3CFE16F8CC" box="[189,410,1880,1906]" italics="true" pageId="14" pageNumber="15">Laius tsushimensis</emphasis>
</taxonomicName>
–
<treatmentCitation id="4355E752450DFFECFE59BB3DFD1AF8CC" author="Nakane T." box="[447,662,1880,1906]" page="164" pageId="14" pageNumber="15" year="1983">
<bibRefCitation id="A665BCB2450DFFECFE59BB3DFD1AF8CC" author="Nakane T." box="[447,662,1880,1906]" pageId="14" pageNumber="15" pagination="161 - 166" refId="ref17940" refString="Nakane T. 1983. Notes on Japanese Melyridae (Coleoptera). Kitakyushu no Kontyu 30 (3): 161 - 166." type="journal article" year="1983">Nakane 1983: 164</bibRefCitation>
</treatmentCitation>
.—
<treatmentCitation id="4355E752450DFFECFD24BB3DFCFAF8CC" author="Sato M." box="[706,886,1879,1906]" page="164" pageId="14" pageNumber="15" year="1985">
<bibRefCitation id="A665BCB2450DFFECFD24BB3DFCFAF8CC" author="Sato M." box="[706,886,1879,1906]" pageId="14" pageNumber="15" pagination="164 - 165" refId="ref18206" refString="Sato M. 1985. Melyridae. In: Ueno S. Kurosawa Y. &amp; Sato M. (eds), The Coleoptera of Japan in Color, 3: 164 - 165, pl. 26. Hoikusha, Osaka. [In Japanese.]" type="journal article" year="1985">Satô 1985: 164</bibRefCitation>
</treatmentCitation>
, pl. 26, fig. 6.—
<treatmentCitation id="4355E752450DFFECFBDABB3DFB7DF8CC" author="Sato M." box="[1084,1265,1880,1906]" page="366" pageId="14" pageNumber="15" year="1989">
<bibRefCitation id="A665BCB2450DFFECFBDABB3DFB7DF8CC" author="Sato M." box="[1084,1265,1880,1906]" pageId="14" pageNumber="15" pagination="366" refId="ref18255" refString="Sato M. 1989. Melyridae. In: Hirahima Y. (et al.) A Check List of Japanese Insects 1: 366. [In Japanese.]" type="journal article" year="1989">Satô 1989: 366</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="C24BC143450DFFECFF5BBB1EFEDCF806" blockId="14.[189,1398,1879,1976]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="E2E4E66D450DFFECFF5BBB1EFEDCF806" pageId="14" pageNumber="15">
<taxonomicName id="05F4BAC0450DFFECFF5BBB1EFE21F82B" baseAuthorityName="Sato &amp; Ohbayashi" baseAuthorityYear="1968" box="[189,429,1915,1941]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="tsushimensis">
<emphasis id="F0801D51450DFFECFF5BBB1EFE21F82B" box="[189,429,1915,1941]" italics="true" pageId="14" pageNumber="15">Intybia tsushimensis</emphasis>
</taxonomicName>
–
<treatmentCitation id="4355E752450DFFECFE28BB1EFD22F82B" author="Wittmer W." box="[462,686,1915,1941]" page="191" pageId="14" pageNumber="15" year="1997">
<bibRefCitation id="A665BCB2450DFFECFE28BB1EFD22F82B" author="Wittmer W." box="[462,686,1915,1941]" pageId="14" pageNumber="15" pagination="181 - 211" refId="ref18419" refString="Wittmer W. 1997. Zur Kenntnis der Gattungen Intybia Pascoe und Stenolaius Wittmer (Coleoptera, Malachiidae). Japanese Journal of Systematic Entomology 3 (2): 181 - 211." type="journal article" year="1997">Wittmer 1997: 191</bibRefCitation>
</treatmentCitation>
.—
<treatmentCitation id="4355E752450DFFECFD3BBB1EFC20F82B" author="Mayor A. J." box="[733,940,1915,1941]" page="417" pageId="14" pageNumber="15" year="2007">
<bibRefCitation id="A665BCB2450DFFECFD3BBB1EFC20F82B" author="Mayor A. J." box="[733,940,1915,1941]" pageId="14" pageNumber="15" pagination="415 - 454" refId="ref17633" refString="Mayor A. J. 2007. Malachiidae. In: Lobl I. &amp; Smetana A. (ed.) Catalogue of Palaeartic Coleoptera, Vol. 4, pp. 415 - 454. Apollo Books, Vester Skerninge. https: // doi. org / 10.1163 / 9789004260894" type="journal article" year="2007">Mayor 2007: 417</bibRefCitation>
</treatmentCitation>
[catalogued].—
<treatmentCitation id="4355E752450DFFECFB92BB1EFEC0F806" author="Yoshitomi H. &amp; Hayashi N." page="19" pageId="14" pageNumber="15" year="2011">
<bibRefCitation id="A665BCB2450DFFECFB92BB1EFEC0F806" author="Yoshitomi H. &amp; Hayashi N." pageId="14" pageNumber="15" pagination="18 - 24" refId="ref18569" refString="Yoshitomi H. &amp; Hayashi N. 2011. List of the family Melyridae and its related families in Japan. Sayabane, New Series, Tokyo (2): 18 - 24. [In Japanese.]" type="journal article" year="2011">Yoshitomi &amp; Hayashi 2011: 19</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="8AEE92C8450CFFEDFF5BBD6EFD29FE03" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="C24BC143450CFFEDFF5BBD6EFEBAFE9B" blockId="15.[189,310,267,293]" box="[189,310,267,293]" pageId="15" pageNumber="16">
<heading id="9903762F450CFFEDFF5BBD6EFEBAFE9B" bold="true" box="[189,310,267,293]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="F0801D51450CFFEDFF5BBD6EFEBAFE9B" bold="true" box="[189,310,267,293]" pageId="15" pageNumber="16">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="C24BC143450CFFEDFF5BBD5CFD29FE03" blockId="15.[189,1398,313,445]" pageId="15" pageNumber="16">Elytra with a pale yellow fascia, just before the middle, which is narrowed at suture; legs fully black; antennomere III cylinder-like shaped, shallowly concave in dorso-mesal portion, and provided with a tongue-like projection near behind; gonoporal piece pluri-curved, slender; semi-gonoporal piece slightly curved; ligula short; spinous plate absent.</paragraph>
</subSubSection>
<subSubSection id="8AEE92C8450CFFEDFF5BBD81FA81FBDC" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="C24BC143450CFFEDFF5BBD81FE26FE40" blockId="15.[189,426,484,510]" box="[189,426,484,510]" pageId="15" pageNumber="16">
<heading id="9903762F450CFFEDFF5BBD81FE26FE40" bold="true" box="[189,426,484,510]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="F0801D51450CFFEDFF5BBD81FE26FE40" bold="true" box="[189,426,484,510]" pageId="15" pageNumber="16">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="729CCB1E450CFFEDFF0CBE76FBB3FDEE" ID-GBIF-Occurrence="2620311301" collectionCode="EUMJ" collectorName="Is. Tsushima &amp; July &amp; M. Sato" country="Japan" location="Mt. Ariake" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<paragraph id="C24BC143450CFFEDFF0CBE76FED0FD93" blockId="15.[189,1087,531,593]" box="[234,348,531,557]" pageId="15" pageNumber="16">
<emphasis id="F0801D51450CFFEDFF0CBE76FED0FD93" bold="true" box="[234,348,531,557]" pageId="15" pageNumber="16">
<typeStatus id="1D4F7FE1450CFFEDFF0CBE76FED0FD93" box="[234,348,531,557]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="C24BC143450CFFEDFF5BBE53FBB3FDEE" blockId="15.[189,1087,531,593]" box="[189,1087,566,593]" pageId="15" pageNumber="16">
<collectingCountry id="BAE381D3450CFFEDFF5BBE53FE96FDEE" box="[189,282,566,592]" name="Japan" pageId="15" pageNumber="16">JAPAN</collectingCountry>
:
<specimenCount id="D4F20ACA450CFFEDFEC3BE52FEB1FDEF" box="[293,317,567,593]" pageId="15" pageNumber="16" type="male">♂</specimenCount>
, [
<collectorName id="6F01A495450CFFEDFEB4BE52FE66FDEE" box="[338,490,566,593]" pageId="15" pageNumber="16">Is. Tsushima</collectorName>
,
<location id="C72B9798450CFFEDFE13BE52FDF4FDEE" LSID="urn:lsid:plazi:treatment:4A5D7055450DFFF3FDE3BAEBFE4FFB63:C72B9798450CFFEDFE13BE52FDF4FDEE" box="[501,632,566,593]" country="Japan" name="Mt. Ariake" pageId="15" pageNumber="16">Mt. Ariake</location>
,
<collectorName id="6F01A495450CFFEDFD65BE53FD39FDEE" box="[643,693,566,592]" pageId="15" pageNumber="16">July</collectorName>
, 16. 1960,
<collectorName id="6F01A495450CFFEDFCD0BE52FC1FFDEE" box="[822,915,566,593]" pageId="15" pageNumber="16">M. Satô</collectorName>
leg.] (
<collectionCode id="A4E55986450CFFEDFC39BE53FBBEFDEE" box="[991,1074,566,592]" collectionName="Japan, Matsuyama, Ehime University" country="Japan" name="Ehime University" pageId="15" pageNumber="16">EUMJ</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="C24BC143450CFFEDFF0CBE1DFEEBFD2C" blockId="15.[189,1399,632,728]" box="[234,359,632,658]" pageId="15" pageNumber="16">
<emphasis id="F0801D51450CFFEDFF0CBE1DFEEBFD2C" bold="true" box="[234,359,632,658]" pageId="15" pageNumber="16">
<typeStatus id="1D4F7FE1450CFFEDFF0CBE1DFEEBFD2C" box="[234,359,632,658]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="C24BC143450CFFEDFF5BBEFEFEA9FD66" blockId="15.[189,1399,632,728]" pageId="15" pageNumber="16">
<collectingCountry id="BAE381D3450CFFEDFF5BBEFEFE96FD0B" box="[189,282,667,693]" name="Japan" pageId="15" pageNumber="16">JAPAN</collectingCountry>
:
<specimenCount id="D4F20ACA450CFFEDFEC3BEFEFEDEFD0B" box="[293,338,667,693]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
,
<specimenCount id="D4F20ACA450CFFEDFEB8BEFEFE06FD0B" box="[350,394,667,693]" pageId="15" pageNumber="16" type="female">1 ♀</specimenCount>
, same data as for the
<typeStatus id="1D4F7FE1450CFFEDFD60BEFEFD7CFD0B" box="[646,752,667,693]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
;
<specimenCount id="D4F20ACA450CFFEDFD1ABEFEFCABFD0B" box="[764,807,667,693]" pageId="15" pageNumber="16" type="female">1 ♀</specimenCount>
, Izuhara, Tsushima,
<date id="B64AE783450CFFEDFBFFBEFEFB36FD0B" box="[1049,1210,667,693]" pageId="15" pageNumber="16" value="1968-08-21">21 Aug. 1968</date>
, S. Nomoto leg. (
<collectionCode id="A4E55986450CFFEDFF23BEDBFE94FD66" box="[197,280,702,728]" country="Japan" name="Ehime University" pageId="15" pageNumber="16">EUMJ</collectionCode>
).
</paragraph>
<paragraph id="C24BC143450CFFEDFF5BBE9AFE45FCA7" blockId="15.[189,457,767,793]" box="[189,457,767,793]" pageId="15" pageNumber="16">
<heading id="9903762F450CFFEDFF5BBE9AFE45FCA7" bold="true" box="[189,457,767,793]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="F0801D51450CFFEDFF5BBE9AFE45FCA7" bold="true" box="[189,457,767,793]" pageId="15" pageNumber="16">Additional specimens</emphasis>
</heading>
</paragraph>
<paragraph id="C24BC143450CFFEDFF5BBF4BFA81FBDC" blockId="15.[189,1399,814,1122]" pageId="15" pageNumber="16">
<collectingCountry id="BAE381D3450CFFEDFF5BBF4BFE95FCF6" box="[189,281,814,840]" name="Japan" pageId="15" pageNumber="16">JAPAN</collectingCountry>
, Tsushima Is.:
<specimenCount id="D4F20ACA450CFFEDFE23BF4BFE7EFCF7" box="[453,498,814,841]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
, Ariake-san,
<date id="B64AE783450CFFEDFD6DBF4BFC90FCF7" box="[651,796,814,841]" pageId="15" pageNumber="16" value="1956-08-06">6 Aug. 1956</date>
, Y. Miyake leg. (
<collectionCode id="A4E55986450CFFEDFC06BF4BFBBFFCF6" box="[992,1075,814,840]" country="Japan" name="Ehime University" pageId="15" pageNumber="16">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA450CFFEDFBA0BF4BFBFFFCF7" box="[1094,1139,814,841]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
, Sasuna,
<date id="B64AE783450CFFEDFB06BF4BFAFDFCF6" box="[1248,1393,814,840]" pageId="15" pageNumber="16" value="1977-07-31">31 Jul. 1977</date>
, K. Matsuda leg. (
<collectionCode id="A4E55986450CFFEDFE6FBF34FE50FCD5" box="[393,476,849,875]" country="Japan" name="Ehime University" pageId="15" pageNumber="16">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA450CFFEDFE17BF34FD91FCD2" box="[497,541,849,876]" pageId="15" pageNumber="16" type="female">1 ♀</specimenCount>
, same data,
<date id="B64AE783450CFFEDFD56BF34FCCFFCD5" box="[688,835,849,875]" pageId="15" pageNumber="16" value="1977-08-02">2 Aug. 1977</date>
, K. Matsuda leg. (
<collectionCode id="A4E55986450CFFEDFBFABF34FBE3FCD5" box="[1052,1135,849,875]" country="Japan" name="Ehime University" pageId="15" pageNumber="16">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA450CFFEDFB62BF34FB3CFCD2" box="[1156,1200,849,876]" pageId="15" pageNumber="16" type="female">1 ♀</specimenCount>
, Mitake,
<date id="B64AE783450CFFEDFAC7BF34FF76FC30" pageId="15" pageNumber="16" value="1980-06-29">29 Jun. 1980</date>
, N. Ohbayashi leg. (
<collectionCode id="A4E55986450CFFEDFE0EBF11FDB7FC30" box="[488,571,884,910]" country="Japan" name="Ehime University" pageId="15" pageNumber="16">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA450CFFEDFDA9BF11FDF0FC31" box="[591,636,884,911]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
, Sasuna,
<date id="B64AE783450CFFEDFD0CBF11FCF7FC31" box="[746,891,884,911]" pageId="15" pageNumber="16" value="1985-07-14">14 Jul. 1985</date>
, K. Ando leg. (
<collectionCode id="A4E55986450CFFEDFBCDBF11FBF2FC30" box="[1067,1150,884,910]" country="Japan" name="Ehime University" pageId="15" pageNumber="16">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA450CFFEDFB74BF11FB4CFC31" box="[1170,1216,884,911]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
, Mokkoku-san, Izuhara-chô,
<date id="B64AE783450CFFEDFEBCBFFDFE7EFC0C" box="[346,498,919,946]" pageId="15" pageNumber="16" value="2000-07-13">13 Jul. 2000</date>
, Y. Sugiura leg. (
<collectionCode id="A4E55986450CFFEDFD25BFFDFC9AFC0C" box="[707,790,920,946]" country="Japan" name="Ehime University" pageId="15" pageNumber="16">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA450CFFEDFCCABFFDFCD1FC0C" box="[812,861,920,946]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
, Tsushima,
<date id="B64AE783450CFFEDFC0ABFFDFB0CFC0F" box="[1004,1152,919,946]" pageId="15" pageNumber="16" value="1969-07-11">11 Jul. 1969</date>
(
<collectionCode id="A4E55986450CFFEDFB77BFFDFB68FC0C" box="[1169,1252,920,946]" country="Japan" name="Ehime University" pageId="15" pageNumber="16">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA450CFFEDFB1DBFFDFAA7FC0C" box="[1275,1323,920,946]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
, same data,
<date id="B64AE783450CFFEDFF19BFDEFE1BFC6B" box="[255,407,955,981]" pageId="15" pageNumber="16" value="1969-07-14">14 Jul. 1969</date>
, S. Miyamato leg. (
<collectionCode id="A4E55986450CFFEDFD6EBFDEFD57FC6B" box="[648,731,955,981]" country="Japan" name="Ehime University" pageId="15" pageNumber="16">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA450CFFEDFD14BFDEFCADFC6B" box="[754,801,955,981]" pageId="15" pageNumber="16" type="female">1 ♀</specimenCount>
, Izuhara-Sasutôge,
<date id="B64AE783450CFFEDFBE9BFDEFB3AFC6B" box="[1039,1206,955,981]" pageId="15" pageNumber="16" value="1941-08-24">24 Aug. 1941</date>
, T. Shirôzu leg. (
<collectionCode id="A4E55986450CFFEDFF23BFBBFE94FC46" box="[197,280,990,1016]" pageId="15" pageNumber="16">SEHU</collectionCode>
);
<specimenCount id="D4F20ACA450CFFEDFECABFBBFEE2FC47" box="[300,366,990,1017]" pageId="15" pageNumber="16" type="male">2 ♂♂</specimenCount>
, Shitasaki, Kamitsuhisma-machi,
<date id="B64AE783450CFFEDFCE6BFBBFC15FC46" box="[768,921,990,1017]" pageId="15" pageNumber="16" value="2002-06-21">21 Jun. 2002</date>
, T. Kurihara leg. (
<collectionCode id="A4E55986450CFFEDFB8ABFBBFB33FC46" box="[1132,1215,990,1016]" country="Japan" name="Ehime University" pageId="15" pageNumber="16">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA450CFFEDFB35BFBBFA8DFC47" box="[1235,1281,990,1017]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
, Waniura, Kamitsushima-machi,
<date id="B64AE783450CFFEDFE2CB864FDE6FBA5" box="[458,618,1025,1052]" pageId="15" pageNumber="16" value="2002-06-21">21 Jun. 2002</date>
, T. Kurihara leg. (
<collectionCode id="A4E55986450CFFEDFCADB864FC12FBA5" box="[843,926,1025,1051]" country="Japan" name="Ehime University" pageId="15" pageNumber="16">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA450CFFEDFC53B864FC6AFBA2" box="[949,998,1025,1052]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
, Kechi, Mitsushima-chô,
<date id="B64AE783450CFFEDFAF9B864FF77FB80" pageId="15" pageNumber="16" value="2002-06-22">22 Jun. 2002</date>
, T. Kurihara leg. (
<collectionCode id="A4E55986450CFFEDFE33B841FDA4FB80" box="[469,552,1060,1086]" country="Japan" name="Ehime University" pageId="15" pageNumber="16">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA450CFFEDFDD8B841FDE1FB81" box="[574,621,1060,1087]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
, Sumo, Mitsushima-chô,
<date id="B64AE783450CFFEDFC79B841FBB1FB80" box="[927,1085,1060,1087]" pageId="15" pageNumber="16" value="2002-06-15">15 Jun. 2002</date>
, T. Kurihara leg. (
<collectionCode id="A4E55986450CFFEDFAF0B841FAE5FB80" box="[1302,1385,1060,1086]" country="Japan" name="Ehime University" pageId="15" pageNumber="16">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA450CFFEDFF5BB82DFF65FBDC" box="[189,233,1096,1122]" pageId="15" pageNumber="16" type="female">1 ♀</specimenCount>
, Ayumodoshi-shizenkoen, Izuhara-machi, Tsushima-shi,
<date id="B64AE783450CFFEDFC6FB82DFB90FBDC" box="[905,1052,1096,1122]" pageId="15" pageNumber="16" value="2013-08-08">8 Aug. 2013</date>
, N. Tsuji leg. (
<collectionCode id="A4E55986450CFFEDFB2EB82DFA8CFBDC" box="[1224,1280,1096,1122]" pageId="15" pageNumber="16">NTJ</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="968B91CB450CFFEDFF58BB02FD7BF855" ID-DOI="http://doi.org/10.5281/zenodo.3829800" ID-Zenodo-Dep="3829800" httpUri="https://zenodo.org/record/3829800/files/figure.png" pageId="15" pageNumber="16" startId="15.[190,233,1895,1921]" targetBox="[196,1385,1174,1842]" targetPageId="15">
<paragraph id="C24BC143450CFFEDFF58BB02FD7BF855" blockId="15.[190,1398,1894,2027]" pageId="15" pageNumber="16">
<emphasis id="F0801D51450CFFEDFF58BB02FE9AF83F" bold="true" box="[190,278,1895,1921]" pageId="15" pageNumber="16">Fig. 11.</emphasis>
Map showing the distribution of
<taxonomicName id="05F4BAC0450CFFEDFD6FBB02FD50F83F" authorityName="Pascoe" authorityYear="1866" box="[649,732,1895,1921]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="F0801D51450CFFEDFD6FBB02FD50F83F" box="[649,732,1895,1921]" italics="true" pageId="15" pageNumber="16">Intybia</emphasis>
</taxonomicName>
spp.
<emphasis id="F0801D51450CFFEDFCF5BB02FCA4F83F" bold="true" box="[787,808,1895,1921]" pageId="15" pageNumber="16">A</emphasis>
.The
<taxonomicName id="05F4BAC0450CFFEDFC83BB02FC3EF83F" baseAuthorityName="Kiesenwetter" baseAuthorityYear="1874" box="[869,946,1895,1921]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="histrio">
<emphasis id="F0801D51450CFFEDFC83BB02FC3EF83F" box="[869,946,1895,1921]" italics="true" pageId="15" pageNumber="16">histrio</emphasis>
</taxonomicName>
group:●:
<taxonomicName id="05F4BAC0450CFFEDFBC5BB02FE8AF81A" authority="(Kiesenwetter, 1874)" baseAuthorityName="Kiesenwetter" baseAuthorityYear="1874" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="histrio">
<emphasis id="F0801D51450CFFEDFBC5BB02FB4BF83F" box="[1059,1223,1895,1921]" italics="true" pageId="15" pageNumber="16">Intybia histrio</emphasis>
(
<bibRefCitation id="A665BCB2450CFFEDFB34BB02FF70F81A" author="Kiesenwetter E. A. H. von" pageId="15" pageNumber="16" pagination="241 - 288" refId="ref17457" refString="Kiesenwetter E. A. H. von. 1874. Die Malacodermen Japans nach dem Ergebnisse der Sammulungen des Herrn G. Lewis wahread der Jahre 1869 - 1871. Berliner Entomologische Zeitschrift 18: 241 - 288. https: // doi. org / 10.1002 / mmnd. 18740180302" type="journal article" year="1874">Kiesenwetter, 1874</bibRefCitation>
)
</taxonomicName>
, ▲:
<taxonomicName id="05F4BAC0450CFFEDFEA2BBEEFCB1F81B" authority="(Sato &amp; Ohbayashi, 1968)" baseAuthorityName="Sato &amp; Ohbayashi" baseAuthorityYear="1968" box="[324,829,1930,1957]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="tsushimensis">
<emphasis id="F0801D51450CFFEDFEA2BBEEFE7FF81A" box="[324,499,1930,1956]" italics="true" pageId="15" pageNumber="16">I. tsushimensis</emphasis>
(Satô &amp; Ohbayashi, 1968)
</taxonomicName>
, ■:
<taxonomicName id="05F4BAC0450CFFEDFC96BBEEFC59F81A" authority="Ikeda &amp; Yoshitomi, 2017" authorityName="Ikeda &amp; Yoshitomi" authorityYear="2017" box="[880,981,1930,1956]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="donan" status="sp. nov.">
<emphasis id="F0801D51450CFFEDFC96BBEEFC59F81A" box="[880,981,1930,1956]" italics="true" pageId="15" pageNumber="16">I. donan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EBB3A02A450CFFEDFC39BBEEFBCDF81B" box="[991,1089,1931,1957]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
).
<emphasis id="F0801D51450CFFEDFBBEBBEFFBE0F81A" bold="true" box="[1112,1132,1930,1956]" pageId="15" pageNumber="16">B</emphasis>
. The
<taxonomicName id="05F4BAC0450CFFEDFB52BBEFFA92F81A" baseAuthorityName="Pic" baseAuthorityYear="1910" box="[1204,1310,1930,1956]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="pelegrini">
<emphasis id="F0801D51450CFFEDFB52BBEFFA92F81A" box="[1204,1310,1930,1956]" italics="true" pageId="15" pageNumber="16">pelegrini</emphasis>
</taxonomicName>
group: ○:
<taxonomicName id="05F4BAC0450CFFEDFF06BBCBFE7CF876" authority="(Pic, 1910)" baseAuthorityName="Pic" baseAuthorityYear="1910" box="[224,496,1965,1992]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="pelegrini">
<emphasis id="F0801D51450CFFEDFF06BBCBFEEFF879" box="[224,355,1965,1991]" italics="true" pageId="15" pageNumber="16">I. pelegrini</emphasis>
(
<bibRefCitation id="A665BCB2450CFFEDFE95BBC8FE64F876" author="Pic M." box="[371,488,1965,1992]" pageId="15" pageNumber="16" pagination="60 - 63" refId="ref17992" refString="Pic M. 1910. Coleopteres exotiques nouveaux ou peu connus. L' Echange, Revue Linneenne 26: 60 - 63." type="journal article" year="1910">Pic, 1910</bibRefCitation>
)
</taxonomicName>
, △:
<taxonomicName id="05F4BAC0450CFFEDFDCFBBCBFCD4F876" authority="(Lewis, 1895)" baseAuthorityName="Lewis" baseAuthorityYear="1895" box="[553,856,1965,1992]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="niponica">
<emphasis id="F0801D51450CFFEDFDCFBBCBFD25F879" box="[553,681,1966,1991]" italics="true" pageId="15" pageNumber="16">I. niponica</emphasis>
(
<bibRefCitation id="A665BCB2450CFFEDFD5FBBC8FCC3F876" author="Lewis G." box="[697,847,1965,1992]" pageId="15" pageNumber="16" pagination="98 - 122" refId="ref17589" refString="Lewis G. 1895. On the Daschillidae and malacoderm Coleoptera of Japan. The Annals and Magazine of Natural History (6) 16: 98 - 122, pl. 6. https: // doi. org / 10.1080 / 00222939508680233" type="journal article" year="1895">Lewis, 1895</bibRefCitation>
)
</taxonomicName>
, □:
<taxonomicName id="05F4BAC0450CFFEDFC60BBCBFB64F876" authority="(Nakane, 1955)" baseAuthorityName="Nakane" baseAuthorityYear="1955" box="[902,1256,1965,1992]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="takaraensis">
<emphasis id="F0801D51450CFFEDFC60BBCBFBAAF879" box="[902,1062,1965,1991]" italics="true" pageId="15" pageNumber="16">I. takaraensis</emphasis>
(Nakane, 1955)
</taxonomicName>
, ×:
<taxonomicName id="05F4BAC0450CFFEDFAF3BBCBFEF4F855" authority="(Nakane, 1955)" baseAuthorityName="Nakane" baseAuthorityYear="1955" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="kishiii">
<emphasis id="F0801D51450CFFEDFAF3BBCBFAF9F879" box="[1301,1397,1965,1991]" italics="true" pageId="15" pageNumber="16">I. kishiii</emphasis>
(Nakane, 1955)
</taxonomicName>
, +:
<taxonomicName id="05F4BAC0450CFFEDFE45BBB7FD7FF855" authority="(Nakane, 1956)" baseAuthorityName="Nakane" baseAuthorityYear="1956" box="[419,755,2000,2027]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="kawasakii">
<emphasis id="F0801D51450CFFEDFE45BBB7FDBDF855" box="[419,561,2001,2027]" italics="true" pageId="15" pageNumber="16">I. kawasakii</emphasis>
(
<bibRefCitation id="A665BCB2450CFFEDFDA7BBB4FD66F855" author="Nakane T." box="[577,746,2000,2027]" pageId="15" pageNumber="16" pagination="159 - 174" refId="ref17827" refString="Nakane T. 1956. New or little-known Coleoptera from Japan and its abject regions, XIII. The scientific Reports of the Saikyo University (Natural Science and Living Science) (A) 2: 29 - 44, 159 - 174." type="journal article" year="1956">Nakane, 1956</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="8AEE92C84513FFF3FF5BBD6EFAD0FCA6" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
<paragraph id="C24BC1434513FFF2FF5BBD6EFEE7FE9B" blockId="16.[189,363,267,293]" box="[189,363,267,293]" pageId="16" pageNumber="17">
<heading id="9903762F4513FFF2FF5BBD6EFEE7FE9B" bold="true" box="[189,363,267,293]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="F0801D514513FFF2FF5BBD6EFEE7FE9B" bold="true" box="[189,363,267,293]" pageId="16" pageNumber="17">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="C24BC1434513FFF2FF0CBD5FFEA6FEEA" blockId="16.[189,1399,314,445]" box="[234,298,314,340]" pageId="16" pageNumber="17">
<emphasis id="F0801D514513FFF2FF0CBD5FFEA6FEEA" bold="true" box="[234,298,314,340]" pageId="16" pageNumber="17">Male</emphasis>
</paragraph>
<paragraph id="C24BC1434513FFF2FF5BBD39FE16FE03" blockId="16.[189,1399,314,445]" pageId="16" pageNumber="17">
Body (
<figureCitation id="5ACFDDC64513FFF2FEF6BD39FEE1FEC9" box="[272,365,348,375]" captionStart="Fig" captionStartId="4.[189,232,1893,1919]" captionTargetBox="[198,1389,740,1831]" captionTargetId="figure@4.[189,1398,723,1853]" captionTargetPageId="4" captionText="Fig. 1. Dorsal habitus Intybia spp., ♂. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov., holotype. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955). H. I. kawasakii (Nakane, 1956), holotype. Scales = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3829778" httpUri="https://zenodo.org/record/3829778/files/figure.png" pageId="16" pageNumber="17">Fig. 1B</figureCitation>
) oblong oval, shining, densely covered with yellow pubescence. Coloration of head, pronotum and elytra black; antennae yellow, with apical 6 or 7 segments black; elytral fascia pale yellow; legs black.
</paragraph>
<paragraph id="C24BC1434513FFF3FF5BBDB0FCA3FEF6" blockId="16.[189,1399,469,777]" lastBlockId="17.[189,1398,267,328]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
Head sparsely and minutely punctate; eyes normal, moderately prominent; HL/HW 0.59–0.89 (0.71). Antennae (
<figureCitation id="5ACFDDC64513FFF2FEDBBD9DFE1AFDAD" box="[317,406,504,531]" captionStart="Fig" captionStartId="7.[190,233,1895,1921]" captionTargetBox="[265,1321,271,1836]" captionTargetId="figure@7.[246,1331,265,1859]" captionTargetPageId="7" captionText="Fig. 4. Right antennae of Intybia spp., ♂. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955). H. I. kawasakii (Nakane, 1956). Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3829786" httpUri="https://zenodo.org/record/3829786/files/figure.png" pageId="16" pageNumber="17">Fig. 4B</figureCitation>
) long, nearly reaching base of elytra; antennomere III (
<figureCitation id="5ACFDDC64513FFF2FBFFBD9DFBFEFDAD" box="[1049,1138,504,531]" captionStart="Fig" captionStartId="9.[190,233,1895,1921]" captionTargetBox="[263,1324,265,1858]" captionTargetId="figure@9.[263,1324,265,1859]" captionTargetPageId="9" captionText="Fig. 6. SEM photographs of right male antennomere III of Intybia spp. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C, H. I. donan sp. nov., showing projection (H). D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955)." figureDoi="http://doi.org/10.5281/zenodo.3829790" httpUri="https://zenodo.org/record/3829790/files/figure.png" pageId="16" pageNumber="17">Fig. 6B</figureCitation>
) cylinder-like shaped, shallowly concave in dorso-mesal portion, with a tongue-like projection posteriorly; L/W = 1.47; approximate ratio of each antennomere (n = 1) as 13.0:1.0: 14.7:2.7: 3.3: 3.7: 4.0: 4.0: 4.0:3.7:5.0. Pronotum slightly broader than long, sparsely and minutely punctate; PW/PL 1.06–1.17 (1.14). Elytra oblong-ovate, slightly broadened posteriorly, shallowly and somewhat densely punctate; elytral fascia situated just before the middle, narrowed at suture; EL/EW 1.27–1.71 (1.49); EL/PL 2.35–2.94 (2.80); EW/PW 1.53–1.83 (1.69); TL/EW 2.17–2.72 (2.41). Aedeagus about 1.00 mm in AL. Gonoporal piece (
<figureCitation id="5ACFDDC64513FFF2FF20BE8AFEA3FCB7" box="[198,303,751,777]" captionStart="Fig" captionStartId="13.[190,233,1893,1919]" captionTargetBox="[216,1374,635,1827]" captionTargetId="figure@13.[188,1397,614,1853]" captionTargetPageId="13" captionText="Fig. 10. Endophallic sclerites of Intybia spp. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955). Abbreviations: gp = gonoporal piece; l = ligula; sgp = semi-gonoporal piece. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829798" httpUri="https://zenodo.org/record/3829798/files/figure.png" pageId="16" pageNumber="17">Fig. 10B</figureCitation>
) winding, slender, bearing minute spines in middle part,
<quantity id="050C6CA64513FFF2FC2BBE8AFBBAFCB7" box="[973,1078,751,777]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="16" pageNumber="17" unit="mm" value="0.32">0.32 mm</quantity>
in GL; GL/AL 0.32. Semi- gonoporal piece slightly curved,
<quantity id="050C6CA64512FFF3FDD8BD6EFD2AFE98" box="[574,678,267,294]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="18" unit="mm" value="0.2">0.20 mm</quantity>
in SGL. Ligula short,
<quantity id="050C6CA64512FFF3FC48BD6EFB9AFE98" box="[942,1046,267,294]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="18" unit="mm" value="0.05">0.05 mm</quantity>
in LL. Spinous area (
<figureCitation id="5ACFDDC64512FFF3FAF2BD6EFAE1FE9B" box="[1300,1389,267,293]" captionStart="Fig" captionStartId="12.[190,233,1893,1919]" captionTargetBox="[191,1394,543,1850]" captionTargetId="figure@12.[189,1398,538,1851]" captionTargetPageId="12" captionText="Fig. 9. Spinous area and spinous plate of Intybia spp. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica(Lewis,1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955).Abbreviations: sa = spinous area; sp = spinous plate. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829796" httpUri="https://zenodo.org/record/3829796/files/figure.png" pageId="17" pageNumber="18">Fig. 9B</figureCitation>
) bearing short triangular spines, lacking spinous plate.
</paragraph>
<caption id="968B91CB4513FFF2FF58BB03FD2EF854" ID-DOI="http://doi.org/10.5281/zenodo.3829802" ID-Zenodo-Dep="3829802" httpUri="https://zenodo.org/record/3829802/files/figure.png" pageId="16" pageNumber="17" startId="16.[190,233,1894,1920]" targetBox="[194,1397,892,1845]" targetPageId="16">
<paragraph id="C24BC1434513FFF2FF58BB03FD2EF854" blockId="16.[190,1397,1894,2026]" pageId="16" pageNumber="17">
<emphasis id="F0801D514513FFF2FF58BB03FEADF83E" bold="true" box="[190,289,1894,1920]" pageId="16" pageNumber="17">Fig. 12.</emphasis>
Habitat and habits of
<taxonomicName id="05F4BAC04513FFF2FDA1BB03FD16F83E" authorityName="Pascoe" authorityYear="1866" box="[583,666,1894,1920]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="F0801D514513FFF2FDA1BB03FD16F83E" box="[583,666,1894,1920]" italics="true" pageId="16" pageNumber="17">Intybia</emphasis>
</taxonomicName>
spp.
<emphasis id="F0801D514513FFF2FD00BB03FD77F83E" bold="true" box="[742,763,1894,1920]" pageId="16" pageNumber="17">A</emphasis>
.
<taxonomicName id="05F4BAC04513FFF2FCF7BB03FC33F83E" baseAuthorityName="Kiesenwetter" baseAuthorityYear="1874" box="[785,959,1894,1920]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="histrio">
<emphasis id="F0801D514513FFF2FCF7BB03FC33F83E" box="[785,959,1894,1920]" italics="true" pageId="16" pageNumber="17">Intybia histrio</emphasis>
</taxonomicName>
, ♂ (at Awa-shima, Kagawa Pref.).
<emphasis id="F0801D514513FFF2FF58BBEFFF5EF81A" bold="true" box="[190,210,1930,1956]" pageId="16" pageNumber="17">B</emphasis>
.
<taxonomicName id="05F4BAC04513FFF2FF02BBEFFEEBF81D" baseAuthorityName="Lewis" baseAuthorityYear="1895" box="[228,359,1930,1955]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="niponica">
<emphasis id="F0801D514513FFF2FF02BBEFFEEBF81D" box="[228,359,1930,1955]" italics="true" pageId="16" pageNumber="17">I. niponica</emphasis>
</taxonomicName>
, ♀ (at Saijo-shi, Ehime Pref.), photographs by HY.
<emphasis id="F0801D514513FFF2FC03BBECFBBFF81A" bold="true" box="[997,1075,1929,1956]" pageId="16" pageNumber="17">C – D</emphasis>
. Type locality of
<taxonomicName id="05F4BAC04513FFF2FAF6BBEFFAF9F81D" authority="Ikeda &amp; Yoshitomi, 2017" authorityName="Ikeda &amp; Yoshitomi" authorityYear="2017" box="[1296,1397,1929,1955]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="donan" status="sp. nov.">
<emphasis id="F0801D514513FFF2FAF6BBEFFAF9F81D" box="[1296,1397,1929,1955]" italics="true" pageId="16" pageNumber="17">I. donan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EBB3A02A4513FFF2FF58BBC8FE95F879" box="[190,281,1965,1991]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
(Mantabaru-shinrinkôen, Yonaguni-chô, Yaeyama-gun, Okinawa Pref.), showing the flower of
<taxonomicName id="05F4BAC04513FFF2FF58BBB5FE0CF854" box="[190,384,2000,2026]" class="Magnoliopsida" family="Primulaceae" genus="Ardisia" higherTaxonomySource="CoL" kingdom="Plantae" order="Ericales" pageId="16" pageNumber="17" phylum="Magnoliopsida" rank="species" species="sieboldii">
<emphasis id="F0801D514513FFF2FF58BBB5FE0CF854" box="[190,384,2000,2026]" italics="true" pageId="16" pageNumber="17">Ardisia sieboldii</emphasis>
</taxonomicName>
(D), photographs by HI.
</paragraph>
</caption>
<paragraph id="C24BC1434512FFF3FF5BBD0CFE94FE3D" blockId="17.[189,280,361,387]" box="[189,280,361,387]" pageId="17" pageNumber="18">
<heading id="9903762F4512FFF3FF5BBD0CFE94FE3D" bold="true" box="[189,280,361,387]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="F0801D514512FFF3FF5BBD0CFE94FE3D" bold="true" box="[189,280,361,387]" pageId="17" pageNumber="18">Female</emphasis>
</heading>
</paragraph>
<paragraph id="C24BC1434512FFF3FF5BBDF2FD7FFD80" blockId="17.[189,1399,407,574]" pageId="17" pageNumber="18">
Similar to male in general appearance (
<figureCitation id="5ACFDDC64512FFF3FD27BDF2FCAAFE0F" box="[705,806,407,433]" captionStart="Fig" captionStartId="5.[190,233,1894,1920]" captionTargetBox="[195,1390,781,1842]" captionTargetId="figure@5.[190,1399,771,1854]" captionTargetPageId="5" captionText="Fig. 2. Dosal habitus of Intybia spp., ♀. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov., paratype. D. I. pelegrini pelegrini (Pic, 1910). E. I.niponica (Lewis, 1895). F. I.takaraensis (Nakane, 1955).G. I. kishiii (Nakane, 1955). H. I. kawasakii (Nakane, 1956), paratype. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3829782" httpUri="https://zenodo.org/record/3829782/files/figure.png" pageId="17" pageNumber="18">Fig. 2B</figureCitation>
); HL/HW 0.57–0.79 (0.67); PW/PL 1.03–1.21 (1.14); EL/EW 1.36–1.71 (1.52); EL/PL 2.76–3.19 (3.01); EW/PW 1.51–1.87 (1.74); TL/ EW 2.11–2.68 (2.40). Antennae long (
<figureCitation id="5ACFDDC64512FFF3FD57BDB8FC9BFE46" box="[689,791,477,504]" captionStart="Fig" captionStartId="8.[190,233,1894,1920]" captionTargetBox="[285,1298,276,1852]" captionTargetId="figure@8.[273,1302,265,1859]" captionTargetPageId="8" captionText="Fig. 5. Right antennae of Intybia spp., ♀. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955). H. I. kawasakii (Nakane, 1956). Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3829788" httpUri="https://zenodo.org/record/3829788/files/figure.png" pageId="17" pageNumber="18">Fig. 5B</figureCitation>
); antennomere III distinctly expanded laterally, shallowly concave in dorsal portion, L/W = 2.30; approximate ratio of each antennomere (n = 1) as 9.7: 1.0: 7.7:3.7: 3.7: 3.7: 4.0: 4.0: 4.0:3.3: 4.0.
</paragraph>
<paragraph id="C24BC1434512FFF3FF5BBE3BFEF9FDC6" blockId="17.[189,373,606,632]" box="[189,373,606,632]" pageId="17" pageNumber="18">
<heading id="9903762F4512FFF3FF5BBE3BFEF9FDC6" bold="true" box="[189,373,606,632]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="F0801D514512FFF3FF5BBE3BFEF9FDC6" bold="true" box="[189,373,606,632]" pageId="17" pageNumber="18">Measurements</emphasis>
</heading>
</paragraph>
<paragraph id="C24BC1434512FFF3FF5BBEE9FAD0FD74" blockId="17.[189,1398,652,792]" pageId="17" pageNumber="18">Male (n = 12): TL 3.07–3.64 (3.39) mm; HL 0.47–0.63 (0.55) mm; HW 0.57–0.87 (0.78) mm; PW 0.75–0.88 (0.84) mm; PL 0.67–0.81 (0.75) mm; EL 1.90–2.32 (2.09) mm; EW 1.19–1.54 (1.41) mm.</paragraph>
<paragraph id="C24BC1434512FFF3FF5BBEBFFAD0FCA6" blockId="17.[189,1398,652,792]" pageId="17" pageNumber="18">Female (n = 6): TL 3.38–4.05 (3.58) mm; HL 0.50–0.70 (0.56) mm; HW 0.77–0.89 (0.83) mm; PW 0.79–0.95 (0.86) mm; PL 0.70–0.80 (0.75) mm; EL 2.10–2.55 (2.25) mm; EW 1.28–1.74 (1.49) mm.</paragraph>
</subSubSection>
<subSubSection id="8AEE92C84512FFF3FF5BBF52FE23FC3E" pageId="17" pageNumber="18" type="distribution">
<paragraph id="C24BC1434512FFF3FF5BBF52FE55FCEF" blockId="17.[189,473,823,849]" box="[189,473,823,849]" pageId="17" pageNumber="18">
<emphasis id="F0801D514512FFF3FF5BBF52FEDAFCEF" bold="true" box="[189,342,823,849]" pageId="17" pageNumber="18">Distribution</emphasis>
(
<figureCitation id="5ACFDDC64512FFF3FE80BF52FE5CFCEF" box="[358,464,823,849]" captionStart="Fig" captionStartId="15.[190,233,1895,1921]" captionTargetBox="[196,1385,1174,1842]" captionTargetId="figure@15.[192,1400,1167,1854]" captionTargetPageId="15" captionText="Fig. 11.Map showing thedistribution of Intybia spp.A.The histriogroup:●:Intybia histrio(Kiesenwetter, 1874), ▲: I. tsushimensis (Satô &amp; Ohbayashi, 1968), ■: I. donan sp. nov.). B. The pelegrini group: ○: I. pelegrini (Pic, 1910), △: I. niponica (Lewis, 1895), □: I. takaraensis (Nakane, 1955), ×: I. kishiii (Nakane, 1955), +: I. kawasakii (Nakane, 1956)." figureDoi="http://doi.org/10.5281/zenodo.3829800" httpUri="https://zenodo.org/record/3829800/files/figure.png" pageId="17" pageNumber="18">Fig. 11A</figureCitation>
)
</paragraph>
<paragraph id="C24BC1434512FFF3FF5BBF03FE23FC3E" blockId="17.[189,431,870,896]" box="[189,431,870,896]" pageId="17" pageNumber="18">
<collectingCountry id="BAE381D34512FFF3FF5BBF03FE8CFC3E" box="[189,256,870,896]" name="Japan" pageId="17" pageNumber="18">Japan</collectingCountry>
(Tsushima Is.)
</paragraph>
</subSubSection>
<subSubSection id="8AEE92C84512FFF3FF5BBFC5FEE2FBEC" pageId="17" pageNumber="18" type="discussion">
<paragraph id="C24BC1434512FFF3FF5BBFC5FEA3FC04" blockId="17.[189,303,928,954]" box="[189,303,928,954]" pageId="17" pageNumber="18">
<heading id="9903762F4512FFF3FF5BBFC5FEA3FC04" bold="true" box="[189,303,928,954]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="F0801D514512FFF3FF5BBFC5FEA3FC04" bold="true" box="[189,303,928,954]" pageId="17" pageNumber="18">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C24BC1434512FFF3FF5BBFABFEE2FBEC" blockId="17.[189,1399,974,1106]" pageId="17" pageNumber="18">
The species is similar to
<taxonomicName id="05F4BAC04512FFF3FE04BFABFC04FC56" authority="(Kiesenwetter, 1874)" baseAuthorityName="Kiesenwetter" baseAuthorityYear="1874" box="[482,904,974,1000]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="histrio">
<emphasis id="F0801D514512FFF3FE04BFABFDB9FC56" box="[482,565,974,1000]" italics="true" pageId="17" pageNumber="18">Intybia</emphasis>
<emphasis id="F0801D514512FFF3FDDBBFABFD06FC56" box="[573,650,974,1000]" italics="true" pageId="17" pageNumber="18">histrio</emphasis>
(
<bibRefCitation id="A665BCB24512FFF3FD7CBFABFC0CFC56" author="Kiesenwetter E. A. H. von" box="[666,896,974,1000]" pageId="17" pageNumber="18" pagination="241 - 288" refId="ref17457" refString="Kiesenwetter E. A. H. von. 1874. Die Malacodermen Japans nach dem Ergebnisse der Sammulungen des Herrn G. Lewis wahread der Jahre 1869 - 1871. Berliner Entomologische Zeitschrift 18: 241 - 288. https: // doi. org / 10.1002 / mmnd. 18740180302" type="journal article" year="1874">Kiesenwetter, 1874</bibRefCitation>
)
</taxonomicName>
in the shape of the male antennomere III, elytral coloration and endophallic structures, and differs from it in the coloration of the legs and antennae, and in the shapes of the semi-gonoporal piece and ligula. This species seems to be endemic to Tsushima Is.
</paragraph>
</subSubSection>
<subSubSection id="8AEE92C84512FFF3FF5BB814FE4FFB63" pageId="17" pageNumber="18" type="biology_ecology">
<paragraph id="C24BC1434512FFF3FF5BB814FE0DFB32" blockId="17.[189,385,1137,1164]" box="[189,385,1137,1164]" pageId="17" pageNumber="18">
<heading id="9903762F4512FFF3FF5BB814FE0DFB32" bold="true" box="[189,385,1137,1164]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="F0801D514512FFF3FF5BB814FE0DFB32" bold="true" box="[189,385,1137,1164]" pageId="17" pageNumber="18">Biological notes</emphasis>
</heading>
</paragraph>
<paragraph id="C24BC1434512FFF3FF5BB8C5FE4FFB63" blockId="17.[189,1398,1184,1245]" pageId="17" pageNumber="18">This species inhabits forest edges and grasslands, and can be collected from shrubs and bushes by sweeping and beating.</paragraph>
</subSubSection>
</treatment>
</document>
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