479 lines
78 KiB
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479 lines
78 KiB
XML
<document id="07CE66C24A3399DDCDBC86457A8274C3" ID-CLB-Dataset="20422" ID-DOI="10.5852/ejt.2022.798.1669" ID-GBIF-Dataset="31fec661-5dac-4626-856d-4ba4680efee7" ID-ISSN="2118-9773" ID-Zenodo-Dep="6323002" ID-ZooBank="50692D26-A41C-4F85-B207-A6747FD07470" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1646225219873" checkinUser="felipe" docAuthor="Vagalinski, Boyan, Borissov, Simeon, Bobeva, Aneliya, Canciani, Giacomo & Antić, Dragan Ž." docDate="2022" docId="4A307579CF480D0BFE50F96DFCFEFDBB" docLanguage="en" docName="ejt-798_30-69.pdf" docOrigin="European Journal of Taxonomy 798" docSource="http://dx.doi.org/10.5852/ejt.2022.798.1669" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Stygiiulus fimbriatus Vagalinski & Borissov & Bobeva & Canciani & Antić 2022, comb. et stat. nov." docType="treatment" docVersion="11" lastPageNumber="41" masterDocId="B6090D01CF4D0D00FF95FFEAFFB8FF83" masterDocTitle="The mostly cavernicolous millipede genus Stygiiulus Verhoeff, 1929, stat. nov.: taxonomy, distribution and phylogenetic relationships (Diplopoda, Julida, Julidae)" masterLastPageNumber="69" masterPageNumber="30" pageNumber="35" updateTime="1699300459248" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A852243001DD5DFA1D2DAF3A3AE1D6A1">The mostly cavernicolous millipede genus Stygiiulus Verhoeff, 1929, stat. nov.: taxonomy, distribution and phylogenetic relationships (Diplopoda, Julida, Julidae)</mods:title>
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<mods:namePart id="5A06CFACB7203D7521D970488B5C5BEA">Vagalinski, Boyan</mods:namePart>
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<mods:affiliation id="CE2954409AA0EF331DD0518BFA92B8F4">Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 2 Gagarin Street, 1113, Sofia, Bulgaria.</mods:affiliation>
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<mods:nameIdentifier id="7E3FC1A5531825B784C7EA2B69E0A906" type="email">boyanv84@gmail.com</mods:nameIdentifier>
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<mods:namePart id="8679A59FD0C56D60D7C583B68DC393D7">Borissov, Simeon</mods:namePart>
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<mods:affiliation id="97618BA9A46E34C32110353479965D2D">Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 2 Gagarin Street, 1113, Sofia, Bulgaria.</mods:affiliation>
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<mods:nameIdentifier id="7F3D3CCF503BA690A5B2377E774E9EBB" type="email">borissovsb@gmail.com</mods:nameIdentifier>
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<mods:namePart id="4694DAF4EBFD8DE2F77AD38C376852C8">Bobeva, Aneliya</mods:namePart>
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<mods:affiliation id="FBE04CFE8BF1D7CDDADFEFC89C3F5E2B">Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 2 Gagarin Street, 1113, Sofia, Bulgaria.</mods:affiliation>
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<mods:nameIdentifier id="E33662FCFEC251EE97D6E3D629179908" type="email">aneliabobeva@gmail.com</mods:nameIdentifier>
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<mods:namePart id="7A258A87A5BC3CFCEBFF17391BDE4E70">Canciani, Giacomo</mods:namePart>
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<mods:affiliation id="0130AB099CC7C2D10E6878712127A8EB">Jamarski Klub Kraški Krti / G. S. Talpe del Carso, loc. San Michele del Carso / Vrh, I- 34070 Savogna d’Isonzo (GO), Italy. & Associazione Speleologica Forum Julii Speleo, Via Carraria 99, I- 33043 Cividale del Friuli (UD), Italy.</mods:affiliation>
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<mods:nameIdentifier id="7346AC167B589A36985DF24B93E2CDF0" type="email">giacomo.canciani90@gmail.com</mods:nameIdentifier>
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<mods:namePart id="771C1AA0C07DE06F29998D842C2BAAAA">Antić, Dragan Ž.</mods:namePart>
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<mods:affiliation id="A802F1FF5C281D79EDF4895400ED6120">University of Belgrade - Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. & Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia.</mods:affiliation>
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<mods:nameIdentifier id="4825CC7F74EC330D16B68766C4BF54F3" type="email">dragan.antic@bio.bg.ac.rs</mods:nameIdentifier>
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<mods:title id="3638A0227E9A3AE2B18D3DA592A0CACC">European Journal of Taxonomy</mods:title>
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<mods:date id="91D1F2A21AD96718651BFB42364B79EF">2022</mods:date>
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<treatment id="4A307579CF480D0BFE50F96DFCFEFDBB" ID-DOI="http://doi.org/10.5281/zenodo.6323850" ID-GBIF-Taxon="193854117" ID-Zenodo-Dep="6323850" LSID="urn:lsid:plazi:treatment:4A307579CF480D0BFE50F96DFCFEFDBB" httpUri="http://treatment.plazi.org/id/4A307579CF480D0BFE50F96DFCFEFDBB" lastPageId="11" lastPageNumber="41" pageId="5" pageNumber="35">
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<heading id="996E7303CF480D05FE50F96DFBD6F921" box="[453,1134,1671,1698]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="35" reason="2">
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<taxonomicName id="0599BFECCF480D05FE50F96DFC34F921" ID-CoL="8YKQM" ID-ENA="2917786" authority="(Strasser, 1971)" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Strasser" baseAuthorityYear="1971" box="[453,908,1671,1698]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="fimbriatus" status="comb. et stat. nov.">
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<emphasis id="F0ED187DCF480D05FE50F96DFD7FF921" bold="true" box="[453,711,1671,1698]" italics="true" pageId="5" pageNumber="35">Stygiiulus fimbriatus</emphasis>
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(Strasser, 1971)
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<taxonomicNameLabel id="EBDEA506CF480D05FC01F962FBD6F921" box="[916,1134,1672,1698]" pageId="5" pageNumber="35">comb. et stat. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="8A8397E4CF480D05FD5AF941FCDDF946" box="[719,869,1707,1733]" pageId="5" pageNumber="35" type="description">
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<paragraph id="C226C46FCF480D05FD5AF941FCDDF946" blockId="5.[453,1134,1671,1733]" box="[719,869,1707,1733]" pageId="5" pageNumber="35">
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<figureCitation id="5AA2D8EACF480D05FD5AF941FC81F946" box="[719,825,1707,1733]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="6.[189,232,1857,1883]" captionStartId-1="8.[189,232,1787,1813]" captionStartId-2="10.[189,232,1822,1848]" captionTargetBox-0="[189,1398,340,1798]" captionTargetBox-1="[209,1381,264,1720]" captionTargetBox-2="[243,1336,266,1776]" captionTargetId-0="figure-11@6.[189,1398,340,1798]" captionTargetId-1="figure-12@8.[209,1381,264,1720]" captionTargetId-2="figure-12@10.[234,1353,262,1782]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionTargetPageId-2="10" captionText-0="Fig. 1. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., ♂♂ (A–H) and ♀ (I) from Foran di Landri Cave (IZB). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Labrum, frontodorsal view. D. Labrum, dorsal view. E. Pleurotergum 7, lateral view. F. Mid-body rings, lateral view. G. Telson, lateral view. H. Posterior node of hypopharynx, fronto-ventral view. I. Lobe of body ring 3, lateral view. Scale bars: A = 1 mm; B–G, I = 0.5 mm; H = 0.1 mm." captionText-1="Fig. 2. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., ♂ from from Foran di Landri Cave (IBER). A. Gnathochilarium, ventral view. B. Gnathal lobe of left mandible, ventral view. C. Left promere, caudal view. D. Left mesomere, oral view. E. Distal part of left opisthomere, mesal view. Abbreviations: ab = anterior branch of solenomere; el = external lobe; et = external tooth; il = internal lobe; it = internal tooth; mp = mollar plate; pb = posterior branch of solenomere; ph = posterior hump; pl = pectinate lamellae; pm = promentum; sf = setiform filaments; v = velum. Scale bars: A = 0.2 mm; B = 0.1 mm; C–D = 0.05 mm; E = 0.02 mm." captionText-2="Fig. 3. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., holotype ♂ (A, B) (MHNG), ♂ (C) (IZB) and ♀ (D) (IBER) from Foran di Landri Cave. A. Leg-pair 1, caudal view. B. Penis, oral/caudal view. C. Left gonopods, mesal view. D. Left vulva, caudal, somewhat mesal view. Abbreviations: ab = anterior branch of solenomere; bs = basal spine; fc = flagellum channel; lt = lateral tube; m = mesomere; mt = median tube; op = operculum; p = promere; pb = posterior branch of solenomere; ph = posterior hump; v = velum. Scale bars = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6323004" figureDoi-1="http://doi.org/10.5281/zenodo.6323008" figureDoi-2="http://doi.org/10.5281/zenodo.6323014" httpUri-0="https://zenodo.org/record/6323004/files/figure.png" httpUri-1="https://zenodo.org/record/6323008/files/figure.png" httpUri-2="https://zenodo.org/record/6323014/files/figure.png" pageId="5" pageNumber="35">Figs 1–3</figureCitation>
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,
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<figureCitation id="5AA2D8EACF480D05FCD2F941FCDDF946" box="[839,869,1707,1733]" captionStart="Fig" captionStartId="33.[189,232,1998,2024]" captionTargetBox="[189,1398,1099,1960]" captionTargetId="figure-332@33.[189,1398,1099,1960]" captionTargetPageId="33" captionText="Fig. 13. General distribution of the species of the genus Stygiiulus stat. nov." figureDoi="http://doi.org/10.5281/zenodo.6323034" httpUri="https://zenodo.org/record/6323034/files/figure.png" pageId="5" pageNumber="35">13</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="8A8397E4CF480D05FF28F906FB7CF8CB" pageId="5" pageNumber="35" type="reference_group">
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<paragraph id="C226C46FCF480D05FF28F906FC25F884" blockId="5.[189,925,1772,1799]" box="[189,925,1772,1799]" pageId="5" pageNumber="35">
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<treatmentCitationGroup id="E289E341CF480D05FF28F906FC25F884" box="[189,925,1772,1799]" pageId="5" pageNumber="35">
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<treatmentCitation id="4338E27ECF480D05FF28F906FC84F884" author="Strasser K." box="[189,828,1772,1799]" page="14" pageId="5" pageNumber="35" year="1971">
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<taxonomicName id="0599BFECCF480D05FF28F906FC84F884" ID-CoL="5F9XV" authority="Strasser, 1971 a: 13 - 14" authorityName="Strasser" authorityPageNumber="13 - 14" authorityYear="1971" box="[189,828,1772,1799]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="ausugi" subSpecies="fimbriatus">
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<emphasis id="F0ED187DCF480D05FF28F906FD99F885" box="[189,545,1772,1798]" italics="true" pageId="5" pageNumber="35">Typhloiulus ausugi fimbriatus</emphasis>
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<bibRefCitation id="A608B99ECF480D05FDBCF907FC84F884" author="Strasser K." box="[553,828,1773,1799]" pageId="5" pageNumber="35" pagination="1 - 21" refId="ref22340" refString="Strasser K. 1971 a. Uber italienische, besonders kavernikole Diplopoden. Memorie del Museo civico di Storia naturale di Verona 19: 1 - 21." type="journal article" year="1971">Strasser, 1971a: 13–14</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, fig. 14.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="C226C46FCF480D05FF28F8C4FB7CF8CB" blockId="5.[189,1220,1838,1865]" box="[189,1220,1838,1865]" pageId="5" pageNumber="35">
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<treatmentCitationGroup id="E289E341CF480D05FF28F8C4FB7CF8CB" box="[189,1220,1838,1865]" pageId="5" pageNumber="35">
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<taxonomicName id="0599BFECCF480D05FF28F8C4FD99F8CB" ID-CoL="5F9XV" authorityName="Strasser" authorityYear="1971" box="[189,545,1838,1864]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="ausugi" subSpecies="fimbriatus">
|
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<emphasis id="F0ED187DCF480D05FF28F8C4FD99F8CB" box="[189,545,1838,1864]" italics="true" pageId="5" pageNumber="35">Typhloiulus ausugi fimbriatus</emphasis>
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</taxonomicName>
|
||
–
|
||
<treatmentCitation id="4338E27ECF480D05FDD5F8C4FCB2F8CB" author="Minelli A." box="[576,778,1838,1865]" page="10" pageId="5" pageNumber="35" year="1985">
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<bibRefCitation id="A608B99ECF480D05FDD5F8C4FCB2F8CB" author="Minelli A." box="[576,778,1838,1865]" pageId="5" pageNumber="35" pagination="1 - 50" refId="ref21769" refString="Minelli A. 1985. Catalogo dei Diplopodi e dei Chilopodi cavernicoli italiani. Memorie del Museo civico di Storia naturale di Verona, Serie 2 (Sezione scienze della Vita) 4: 1 - 50." type="journal article" year="1985">Minelli 1985: 10</bibRefCitation>
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</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="4338E27ECF480D05FCABF8C4FB79F8CB" author="Vagalinski B. & Stoev P. & Enghoff H." box="[830,1217,1838,1865]" page="336" pageId="5" pageNumber="35" year="2015">
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<bibRefCitation id="A608B99ECF480D05FCABF8C4FB79F8CB" author="Vagalinski B. & Stoev P. & Enghoff H." box="[830,1217,1838,1865]" pageId="5" pageNumber="35" pagination="334 - 362" refId="ref22493" refString="Vagalinski B., Stoev P. & Enghoff H. 2015. A review of the millipede genus Typhloiulus Latzel, 1884 (Diplopoda: Julida: Julidae), with a description of three new species from Bulgaria and Greece. Zootaxa 3999 (3): 334 - 362. https: // doi. org / 10.11646 / zootaxa. 3999.3.2" type="journal article" year="2015">
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||
Vagalinski
|
||
<emphasis id="F0ED187DCF480D05FC51F8C5FBB8F8CB" box="[964,1024,1838,1864]" italics="true" pageId="5" pageNumber="35">et al.</emphasis>
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2015: 336–337
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</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="8A8397E4CF480D07FF28F885FCEBFE50" lastPageId="7" lastPageNumber="37" pageId="5" pageNumber="35" type="diagnosis">
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<paragraph id="C226C46FCF480D05FF28F885FE8CF80A" blockId="5.[189,308,1903,1929]" box="[189,308,1903,1929]" pageId="5" pageNumber="35">
|
||
<heading id="996E7303CF480D05FF28F885FE8CF80A" bold="true" box="[189,308,1903,1929]" fontSize="11" level="3" pageId="5" pageNumber="35" reason="3">
|
||
<emphasis id="F0ED187DCF480D05FF28F885FE8CF80A" bold="true" box="[189,308,1903,1929]" pageId="5" pageNumber="35">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF480D07FF28F874FCEBFE50" blockId="5.[189,1398,1950,2012]" lastBlockId="7.[188,1399,265,467]" lastPageId="7" lastPageNumber="37" pageId="5" pageNumber="35">
|
||
A species of
|
||
<taxonomicName id="0599BFECCF480D05FECAF874FE69F83B" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" box="[351,465,1950,1976]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="35" phylum="Arthropoda" rank="genus" status="stat. nov.">
|
||
<emphasis id="F0ED187DCF480D05FECAF874FE69F83B" box="[351,465,1950,1976]" italics="true" pageId="5" pageNumber="35">Stygiiulus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBDEA506CF480D05FE48F874FDF2F83B" box="[477,586,1950,1976]" pageId="5" pageNumber="35">stat. nov.</taxonomicNameLabel>
|
||
with modified mouthparts. Differs from its most similar congener,
|
||
<taxonomicName id="0599BFECCF480D05FF28F82BFEF4F858" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Strasser" baseAuthorityYear="1971" box="[189,332,1985,2011]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="gentianae" status="comb. et stat. nov.">
|
||
<emphasis id="F0ED187DCF480D05FF28F82BFEF4F858" box="[189,332,1985,2011]" italics="true" pageId="5" pageNumber="35">S. gentianae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBDEA506CF480D05FEC7F82BFD9EF85F" box="[338,550,1985,2012]" pageId="5" pageNumber="35">comb. et stat. nov.</taxonomicNameLabel>
|
||
, by gonopod details, viz. both promere and mesomere being apically bent frontad rather than turned towards one another, velum being marginally serrated all along rather than having a smooth anterior margin; by the more pronounced posterior part of pleurotergal flange
|
||
<quantity id="0561698ACF4A0D07FB6FFEC7FA9DFEC5" box="[1274,1317,300,327]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="7" pageNumber="37" unit="in" value="7.0">7 in</quantity>
|
||
males; by the vulval operculum being concave rather than convex, exceeding bursa by ca ¼ rather than
|
||
<superScript id="35EC6927CF4A0D07FA92FEA5FAA8FEDE" attach="right" box="[1287,1296,335,349]" fontSize="6" pageId="7" pageNumber="37">2</superScript>
|
||
⁄
|
||
<subScript id="5E1DC62ACF4A0D07FA83FEB3FAA7FEE4" attach="left" box="[1302,1311,345,359]" fontSize="6" pageId="7" pageNumber="37">5</subScript>
|
||
of total height of vulva; by the presence of a small, roundish anterior lobe on ventral margin of body ring
|
||
<quantity id="0561698ACF4A0D07FAD2FE99FACEFE0F" box="[1351,1398,370,397]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="7" pageNumber="37" unit="in" value="3.0">3 in</quantity>
|
||
females, vs ventral margin of body ring
|
||
<quantity id="0561698ACF4A0D07FD01FE7CFD79FE33" box="[660,705,406,432]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="7" pageNumber="37" unit="in" value="3.0">3 in</quantity>
|
||
females being almost entirely subtriangular in
|
||
<taxonomicName id="0599BFECCF4A0D07FB72FE7CFACFFE33" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Strasser" baseAuthorityYear="1971" box="[1255,1399,406,432]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="37" phylum="Arthropoda" rank="species" species="gentianae" status="comb. et stat. nov.">
|
||
<emphasis id="F0ED187DCF4A0D07FB72FE7CFACFFE33" box="[1255,1399,406,432]" italics="true" pageId="7" pageNumber="37">S. gentianae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBDEA506CF4A0D07FF2BFE53FE2BFE50" box="[190,403,441,467]" pageId="7" pageNumber="37">comb. et stat. nov.</taxonomicNameLabel>
|
||
, and by the somewhat longer epiproct.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="96E694E7CF4B0D06FF28F8ABFC75F86A" ID-DOI="http://doi.org/10.5281/zenodo.6323004" ID-Zenodo-Dep="6323004" httpUri="https://zenodo.org/record/6323004/files/figure.png" pageId="6" pageNumber="36" startId="6.[189,232,1857,1883]" targetBox="[189,1398,340,1798]" targetPageId="6">
|
||
<paragraph id="C226C46FCF4B0D06FF28F8ABFC75F86A" blockId="6.[189,1399,1857,2025]" pageId="6" pageNumber="36">
|
||
<emphasis id="F0ED187DCF4B0D06FF28F8ABFEB5F8D8" bold="true" box="[189,269,1857,1883]" pageId="6" pageNumber="36">Fig. 1.</emphasis>
|
||
<taxonomicName id="0599BFECCF4B0D06FE82F8ABFD68F8DF" authority="(Strasser, 1971)" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Strasser" baseAuthorityYear="1971" box="[279,720,1857,1884]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="36" phylum="Arthropoda" rank="species" species="fimbriatus" status="comb. et stat. nov.">
|
||
<emphasis id="F0ED187DCF4B0D06FE82F8ABFDB3F8D8" box="[279,523,1857,1883]" italics="true" pageId="6" pageNumber="36">Stygiiulus fimbriatus</emphasis>
|
||
(Strasser, 1971)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBDEA506CF4B0D06FD4FF8ABFC01F8DF" box="[730,953,1857,1884]" pageId="6" pageNumber="36">comb. et stat. nov.</taxonomicNameLabel>
|
||
, ♂♂ (A–H) and ♀ (I) from Foran di Landri Cave (IZB).
|
||
<emphasis id="F0ED187DCF4B0D06FE3EF88EFE07F8FD" bold="true" box="[427,447,1892,1918]" pageId="6" pageNumber="36">A</emphasis>
|
||
. Habitus, lateral view.
|
||
<emphasis id="F0ED187DCF4B0D06FD44F88FFD5DF8FC" bold="true" box="[721,741,1893,1919]" pageId="6" pageNumber="36">B</emphasis>
|
||
. Anterior part of body, lateral view.
|
||
<emphasis id="F0ED187DCF4B0D06FB00F88EFB11F8FD" bold="true" box="[1173,1193,1892,1918]" pageId="6" pageNumber="36">C</emphasis>
|
||
. Labrum, frontodorsal view.
|
||
<emphasis id="F0ED187DCF4B0D06FEC0F862FED1F821" bold="true" box="[341,361,1928,1954]" pageId="6" pageNumber="36">D</emphasis>
|
||
. Labrum, dorsal view.
|
||
<emphasis id="F0ED187DCF4B0D06FDE8F862FD28F821" bold="true" box="[637,656,1928,1954]" pageId="6" pageNumber="36">E</emphasis>
|
||
. Pleurotergum 7, lateral view.
|
||
<emphasis id="F0ED187DCF4B0D06FC6BF862FBB7F821" bold="true" box="[1022,1039,1928,1954]" pageId="6" pageNumber="36">F</emphasis>
|
||
. Mid-body rings, lateral view.
|
||
<emphasis id="F0ED187DCF4B0D06FF2BF841FF6CF846" bold="true" box="[190,212,1963,1989]" pageId="6" pageNumber="36">G</emphasis>
|
||
. Telson, lateral view.
|
||
<emphasis id="F0ED187DCF4B0D06FE43F841FE54F846" bold="true" box="[470,492,1963,1989]" pageId="6" pageNumber="36">H</emphasis>
|
||
. Posterior node of hypopharynx, fronto-ventral view.
|
||
<emphasis id="F0ED187DCF4B0D06FBFCF841FBCCF846" bold="true" box="[1129,1140,1963,1989]" pageId="6" pageNumber="36">I</emphasis>
|
||
. Lobe of body ring 3, lateral view. Scale bars: A = 1 mm; B–G, I = 0.5 mm; H = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="8A8397E4CF4A0D07FF28FE10FCE5FCDB" pageId="7" pageNumber="37" type="materials_examined">
|
||
<paragraph id="C226C46FCF4A0D07FF28FE10FE10FD97" blockId="7.[189,424,506,532]" box="[189,424,506,532]" pageId="7" pageNumber="37">
|
||
<heading id="996E7303CF4A0D07FF28FE10FE10FD97" bold="true" box="[189,424,506,532]" fontSize="11" level="3" pageId="7" pageNumber="37" reason="3">
|
||
<emphasis id="F0ED187DCF4A0D07FF28FE10FE10FD97" bold="true" box="[189,424,506,532]" pageId="7" pageNumber="37">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF4A0D07FF7FFDC3FACFFD2F" blockId="7.[189,1399,553,685]" pageId="7" pageNumber="37">
|
||
<materialsCitation id="72F1CE32CF4A0D07FF7FFDC3FACFFD2F" ID-GBIF-Occurrence="3504034326" collectorName="San Pietro & Natisone & Grotta & di San Giovanni d'Antro" country="Italy" location="San Pietro al Natisone" pageId="7" pageNumber="37" specimenCode="MHNG-ARTO-26720" specimenCount="1" specimenCount-male="1" stateProvince="Friuli-Venezia Giulia" typeStatus="holotype">
|
||
<emphasis id="F0ED187DCF4A0D07FF7FFDC3FEE3FDC0" bold="true" box="[234,347,553,579]" pageId="7" pageNumber="37">
|
||
<typeStatus id="1D227ACDCF4A0D07FF7FFDC3FEE3FDC0" box="[234,347,553,579]" pageId="7" pageNumber="37" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="BA8E84FFCF4A0D07FF28FDA6FEB7FDE5" box="[189,271,588,614]" name="Italy" pageId="7" pageNumber="37">ITALY</collectingCountry>
|
||
•
|
||
<specimenCount id="D49F0FE6CF4A0D07FEA0FDA7FEF6FDE4" box="[309,334,589,615]" pageId="7" pageNumber="37" type="male">♂</specimenCount>
|
||
; slide preparation plus ethanol sample;
|
||
<collectingRegion id="005D0A8DCF4A0D07FCD1FDA6FBFCFDE5" box="[836,1092,588,614]" country="Italy" name="Friuli-Venezia Giulia" pageId="7" pageNumber="37">Friuli-Venezia Giulia</collectingRegion>
|
||
,
|
||
<location id="C74692B4CF4A0D07FBC2FDA6FACAFDE5" LSID="urn:lsid:plazi:treatment:4A307579CF480D0BFE50F96DFCFEFDBB:C74692B4CF4A0D07FBC2FDA6FACAFDE5" box="[1111,1394,588,614]" country="Italy" name="San Pietro al Natisone" pageId="7" pageNumber="37" stateProvince="Friuli-Venezia Giulia">San Pietro al Natisone</location>
|
||
,
|
||
<location id="C74692B4CF4A0D07FF28FD85FD33FD0A" LSID="urn:lsid:plazi:treatment:4A307579CF480D0BFE50F96DFCFEFDBB:C74692B4CF4A0D07FF28FD85FD33FD0A" box="[189,651,623,650]" country="Italy" name="Grotta [cave] di San Giovanni d'Antro" pageId="7" pageNumber="37" stateProvince="Friuli-Venezia Giulia">Grotta [cave] di San Giovanni d’Antro</location>
|
||
; slide no. 1003: gonopods, antennae, leg-pairs 1, 2, 3, 7, penis, gnathochilarium, flanges of pleurotergum 7; ethanol sample: broken in six pieces;
|
||
<specimenCode id="923F6C14CF4A0D07FBF8FD78FACBFD2F" box="[1133,1395,658,684]" pageId="7" pageNumber="37">MHNG-ARTO-26720</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF4A0D07FF7FFD3EFE1CFD6D" blockId="7.[189,1398,724,856]" box="[234,420,724,750]" pageId="7" pageNumber="37">
|
||
<emphasis id="F0ED187DCF4A0D07FF7FFD3EFE1CFD6D" bold="true" box="[234,420,724,750]" pageId="7" pageNumber="37">Other material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF4A0D07FF28FD1DFCE5FCDB" blockId="7.[189,1398,724,856]" pageId="7" pageNumber="37">
|
||
<materialsCitation id="72F1CE32CF4A0D07FF28FD1DFCA0FCB6" ID-GBIF-Occurrence="3504034322" collectingDate="2019-07-10" collectionCode="IBER" collectorName="G. Canciani & M. Colautti" country="Italy" location="Foran di Landri Cave" municipality="Prestento di Torreano" pageId="7" pageNumber="37" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Friuli-Venezia Giulia">
|
||
<collectingCountry id="BA8E84FFCF4A0D07FF28FD1DFEB7FC92" box="[189,271,759,785]" name="Italy" pageId="7" pageNumber="37">ITALY</collectingCountry>
|
||
•
|
||
<specimenCount id="D49F0FE6CF4A0D07FEB9FD1DFECAFC91" box="[300,370,759,786]" pageId="7" pageNumber="37" type="male">3 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="D49F0FE6CF4A0D07FE14FD1DFE08FC91" box="[385,432,759,786]" pageId="7" pageNumber="37" type="female">1 ♀</specimenCount>
|
||
;
|
||
<collectingRegion id="005D0A8DCF4A0D07FE54FD1DFD03FC92" box="[449,699,759,785]" country="Italy" name="Friuli-Venezia Giulia" pageId="7" pageNumber="37">Friuli-Venezia Giulia</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="22425E15CF4A0D07FD5FFD12FC72FC91" box="[714,970,759,786]" pageId="7" pageNumber="37">Prestento di Torreano</collectingMunicipality>
|
||
,
|
||
<location id="C74692B4CF4A0D07FC4DFD12FB61FC92" LSID="urn:lsid:plazi:treatment:4A307579CF480D0BFE50F96DFCFEFDBB:C74692B4CF4A0D07FC4DFD12FB61FC92" box="[984,1241,759,786]" country="Italy" municipality="Prestento di Torreano" name="Foran di Landri Cave" pageId="7" pageNumber="37" stateProvince="Friuli-Venezia Giulia">Foran di Landri Cave</location>
|
||
(11 / 46FR);
|
||
<date id="B627E2AFCF4A0D07FF28FCF0FEF4FCB7" box="[189,332,794,820]" pageId="7" pageNumber="37" value="2019-07-10">
|
||
<collectingDate id="A6631B47CF4A0D07FF28FCF0FEF4FCB7" box="[189,332,794,820]" pageId="7" pageNumber="37" value="2019-07-10">10 Jul. 2019</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="6F6CA1B9CF4A0D07FEC3FCF0FE67FCB7" box="[342,479,794,820]" pageId="7" pageNumber="37">G. Canciani</collectorName>
|
||
and
|
||
<collectorName id="6F6CA1B9CF4A0D07FD81FCF1FD21FCB7" box="[532,665,794,821]" pageId="7" pageNumber="37">M. Colautti</collectorName>
|
||
leg.;
|
||
<collectionCode id="A4885CAACF4A0D07FD43FCF1FCA0FCB6" box="[726,792,795,821]" pageId="7" pageNumber="37">IBER</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="72F1CE32CF4A0D07FCBBFCF0FF54FCDB" ID-GBIF-Occurrence="3504034335" collectingDate="2019-07-10" collectionCode="IZB" collectorName="G. Canciani & M. Colautti" country="Italy" location="Foran di Landri Cave" municipality="Prestento di Torreano" pageId="7" pageNumber="37" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Friuli-Venezia Giulia">
|
||
<specimenCount id="D49F0FE6CF4A0D07FCBBFCF0FCC8FCB6" box="[814,880,794,821]" pageId="7" pageNumber="37" type="male">4 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="D49F0FE6CF4A0D07FCEFFCF0FC03FCB6" box="[890,955,794,821]" pageId="7" pageNumber="37" type="female">2 ♀♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<collectionCode id="A4885CAACF4A0D07FF28FCD4FF54FCDB" box="[189,236,830,856]" pageId="7" pageNumber="37">IZB</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="72F1CE32CF4A0D07FE91FCD4FCE1FCDB" ID-GBIF-Occurrence="3504034310" box="[260,857,829,856]" collectingDate="2019-07-10" collectionCode="NHMD" collectorName="G. Canciani & M. Colautti" country="Italy" location="Foran di Landri Cave" municipality="Prestento di Torreano" pageId="7" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Friuli-Venezia Giulia">
|
||
<specimenCount id="D49F0FE6CF4A0D07FE91FCD4FE8BFCDB" box="[260,307,830,856]" pageId="7" pageNumber="37" type="male">1 ♂</specimenCount>
|
||
; same collection data as for preceding;
|
||
<collectionCode id="A4885CAACF4A0D07FD69FCD4FCE1FCDB" box="[764,857,830,856]" pageId="7" pageNumber="37">NHMD</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8A8397E4CF4A0D07FF28FC95FC33FB8D" pageId="7" pageNumber="37" type="discussion">
|
||
<paragraph id="C226C46FCF4A0D07FF28FC95FE80FC1A" blockId="7.[189,312,895,921]" box="[189,312,895,921]" pageId="7" pageNumber="37">
|
||
<heading id="996E7303CF4A0D07FF28FC95FE80FC1A" bold="true" box="[189,312,895,921]" fontSize="11" level="3" pageId="7" pageNumber="37" reason="3">
|
||
<emphasis id="F0ED187DCF4A0D07FF28FC95FE80FC1A" bold="true" box="[189,312,895,921]" pageId="7" pageNumber="37">Comment</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF4A0D07FF28FC47FC33FB8D" blockId="7.[189,1399,941,1038]" pageId="7" pageNumber="37">
|
||
Since the original description was based upon
|
||
<specimenCount id="D49F0FE6CF4A0D07FD71FC44FCE8FC44" box="[740,848,941,968]" pageId="7" pageNumber="37" type="male">one male</specimenCount>
|
||
which was not designated by
|
||
<bibRefCitation id="A608B99ECF4A0D07FB2CFC44FACFFC4B" author="Strasser K." box="[1209,1399,942,968]" pageId="7" pageNumber="37" pagination="1 - 21" refId="ref22340" refString="Strasser K. 1971 a. Uber italienische, besonders kavernikole Diplopoden. Memorie del Museo civico di Storia naturale di Verona 19: 1 - 21." type="journal article" year="1971">Strasser (1971a)</bibRefCitation>
|
||
as
|
||
<typeStatus id="1D227ACDCF4A0D07FF74FC3BFEF2FC68" box="[225,330,977,1003]" pageId="7" pageNumber="37" type="holotype">holotype</typeStatus>
|
||
, we here consider that male as a
|
||
<typeStatus id="1D227ACDCF4A0D07FD7AFC3BFCEDFC68" box="[751,853,977,1003]" pageId="7" pageNumber="37" type="holotype">holotype</typeStatus>
|
||
fixed by monotypy, in order to stabilize the nomenclature of the species under Article 73.1.2 of the ICZN.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8A8397E4CF4A0D09FF28FBDFFD30F931" lastPageId="9" lastPageNumber="39" pageId="7" pageNumber="37" type="description">
|
||
<paragraph id="C226C46FCF4A0D07FF28FBDFFED2FBCC" blockId="7.[189,362,1077,1103]" box="[189,362,1077,1103]" pageId="7" pageNumber="37">
|
||
<heading id="996E7303CF4A0D07FF28FBDFFED2FBCC" bold="true" box="[189,362,1077,1103]" fontSize="11" level="3" pageId="7" pageNumber="37" reason="3">
|
||
<emphasis id="F0ED187DCF4A0D07FF28FBDFFED2FBCC" bold="true" box="[189,362,1077,1103]" pageId="7" pageNumber="37">Redescription</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF4A0D07FF28FB8EFB28FB21" blockId="7.[189,1399,1124,1186]" pageId="7" pageNumber="37">
|
||
SIZE AND NUMBER OF BODY RINGS. Non-type
|
||
<specimenCount id="D49F0FE6CF4A0D07FD57FB8EFD56FBFD" box="[706,750,1124,1150]" pageId="7" pageNumber="37" type="male">♂♂</specimenCount>
|
||
with BRF 26–31 +0–1 +T, L =
|
||
<quantity id="0561698ACF4A0D07FBE0FB8EFB41FBFD" box="[1141,1273,1124,1150]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="2.0" metricValueMin="1.4" pageId="7" pageNumber="37" unit="mm" value="17.0" valueMax="20.0" valueMin="14.0">14–20 mm</quantity>
|
||
, H =
|
||
<quantity id="0561698ACF4A0D07FAD1FB8EFE90FB21" metricMagnitude="-3" metricUnit="m" metricValue="1.175" metricValueMax="1.25" metricValueMin="1.1" pageId="7" pageNumber="37" unit="mm" value="1.175" valueMax="1.25" valueMin="1.1">1.1– 1.25 mm</quantity>
|
||
; non-type
|
||
<specimenCount id="D49F0FE6CF4A0D07FE37FB62FE76FB21" box="[418,462,1160,1186]" pageId="7" pageNumber="37" type="female">♀♀</specimenCount>
|
||
with BRF 28–32 +0–1 +T, L =
|
||
<quantity id="0561698ACF4A0D07FCA8FB6DFC05FB21" box="[829,957,1159,1186]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.1" metricValueMin="1.6" pageId="7" pageNumber="37" unit="mm" value="18.5" valueMax="21.0" valueMin="16.0">16–21 mm</quantity>
|
||
, H =
|
||
<quantity id="0561698ACF4A0D07FC68FB6DFB34FB21" box="[1021,1164,1159,1186]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="7" pageNumber="37" unit="mm" value="1.45" valueMax="1.6" valueMin="1.3">1.3–1.6 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF4A0D07FF28FB23FC70FA85" blockId="7.[188,1399,1224,1286]" pageId="7" pageNumber="37">
|
||
COLOURATION (
|
||
<figureCitation id="5AA2D8EACF4A0D07FEF6FB22FD9CFB60" box="[355,548,1224,1251]" captionStart="Fig" captionStartId="6.[189,232,1857,1883]" captionTargetBox="[189,1398,340,1798]" captionTargetId="figure-11@6.[189,1398,340,1798]" captionTargetPageId="6" captionText="Fig. 1. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., ♂♂ (A–H) and ♀ (I) from Foran di Landri Cave (IZB). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Labrum, frontodorsal view. D. Labrum, dorsal view. E. Pleurotergum 7, lateral view. F. Mid-body rings, lateral view. G. Telson, lateral view. H. Posterior node of hypopharynx, fronto-ventral view. I. Lobe of body ring 3, lateral view. Scale bars: A = 1 mm; B–G, I = 0.5 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6323004" httpUri="https://zenodo.org/record/6323004/files/figure.png" pageId="7" pageNumber="37">Fig. 1A–B, E–G</figureCitation>
|
||
). Generally brown-beige; metazonae with a narrow transverse dark brown band before their last third, the band gradually narrowing ventrally.
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF4A0D07FF28FAC4FEE5FA74" blockId="7.[189,1401,1325,1527]" pageId="7" pageNumber="37">
|
||
EXTERNAL STRUCTURES. Head with 2 vertigial, 2+2 supralabral and 16–18 labral setae. Antennae (
|
||
<figureCitation id="5AA2D8EACF4A0D07FA80FAC7FAD5FAC4" box="[1301,1389,1325,1351]" captionStart="Fig" captionStartId="6.[189,232,1857,1883]" captionTargetBox="[189,1398,340,1798]" captionTargetId="figure-11@6.[189,1398,340,1798]" captionTargetPageId="6" captionText="Fig. 1. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., ♂♂ (A–H) and ♀ (I) from Foran di Landri Cave (IZB). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Labrum, frontodorsal view. D. Labrum, dorsal view. E. Pleurotergum 7, lateral view. F. Mid-body rings, lateral view. G. Telson, lateral view. H. Posterior node of hypopharynx, fronto-ventral view. I. Lobe of body ring 3, lateral view. Scale bars: A = 1 mm; B–G, I = 0.5 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6323004" httpUri="https://zenodo.org/record/6323004/files/figure.png" pageId="7" pageNumber="37">Fig. 1B</figureCitation>
|
||
) 2.3 times as long as head and 2.3–2.4 times as long as H in males, and 2 times and 1.85–1.9 times, respectively, in females; antennomere 5 3.12–3.15 times as long as broad; antennomeres 2–5 subequal in length, 1.3–1.6 times as long as 6. Antennomeres V and VI each with a terminal corolla of large sensilla basiconica bacilliformia; antennomere VII with a terminal corolla of small sensilla basiconica bacilliformia.
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF4A0D07FF28F9F5FB7FF8D1" blockId="7.[188,1400,1566,1874]" pageId="7" pageNumber="37">
|
||
MOUTHPARTS (
|
||
<figureCitation id="5AA2D8EACF4A0D07FECBF9F5FDBBF9BA" box="[350,515,1567,1593]" captionStart="Fig" captionStartId="6.[189,232,1857,1883]" captionTargetBox="[189,1398,340,1798]" captionTargetId="figure-11@6.[189,1398,340,1798]" captionTargetPageId="6" captionText="Fig. 1. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., ♂♂ (A–H) and ♀ (I) from Foran di Landri Cave (IZB). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Labrum, frontodorsal view. D. Labrum, dorsal view. E. Pleurotergum 7, lateral view. F. Mid-body rings, lateral view. G. Telson, lateral view. H. Posterior node of hypopharynx, fronto-ventral view. I. Lobe of body ring 3, lateral view. Scale bars: A = 1 mm; B–G, I = 0.5 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6323004" httpUri="https://zenodo.org/record/6323004/files/figure.png" pageId="7" pageNumber="37">Figs 1C–D, H</figureCitation>
|
||
,
|
||
<figureCitation id="5AA2D8EACF4A0D07FD86F9F5FDE0F9BA" box="[531,600,1567,1593]" captionStart="Fig" captionStartId="8.[189,232,1787,1813]" captionTargetBox="[209,1381,264,1720]" captionTargetId="figure-12@8.[209,1381,264,1720]" captionTargetPageId="8" captionText="Fig. 2. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., ♂ from from Foran di Landri Cave (IBER). A. Gnathochilarium, ventral view. B. Gnathal lobe of left mandible, ventral view. C. Left promere, caudal view. D. Left mesomere, oral view. E. Distal part of left opisthomere, mesal view. Abbreviations: ab = anterior branch of solenomere; el = external lobe; et = external tooth; il = internal lobe; it = internal tooth; mp = mollar plate; pb = posterior branch of solenomere; ph = posterior hump; pl = pectinate lamellae; pm = promentum; sf = setiform filaments; v = velum. Scale bars: A = 0.2 mm; B = 0.1 mm; C–D = 0.05 mm; E = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.6323008" httpUri="https://zenodo.org/record/6323008/files/figure.png" pageId="7" pageNumber="37">2A–B</figureCitation>
|
||
). With moderate hydrophilous modifications (sensu
|
||
<bibRefCitation id="A608B99ECF4A0D07FB56F9F5FADCF9BA" author="Enghoff H." box="[1219,1380,1566,1593]" pageId="7" pageNumber="37" pagination="67 - 77" refId="ref20760" refString="Enghoff H. 1985. Modified mouthparts in hydrophilous cave millipedes (Diplopoda). In: Ellis W. N., Jeekel C. A. W. & Pieters F. F. J. M. (eds) Proceedings of the 6 th International Congress of Myriapodology. Amsterdam, 12 - 17 April 1984. Bijdragen tot de Dierkunde 55 (1): 67 - 77." type="journal article" year="1985">Enghoff 1985</bibRefCitation>
|
||
): Labrum either edentate (
|
||
<figureCitation id="5AA2D8EACF4A0D07FE4AF9A8FD83F9DF" box="[479,571,1602,1628]" captionStart="Fig" captionStartId="6.[189,232,1857,1883]" captionTargetBox="[189,1398,340,1798]" captionTargetId="figure-11@6.[189,1398,340,1798]" captionTargetPageId="6" captionText="Fig. 1. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., ♂♂ (A–H) and ♀ (I) from Foran di Landri Cave (IZB). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Labrum, frontodorsal view. D. Labrum, dorsal view. E. Pleurotergum 7, lateral view. F. Mid-body rings, lateral view. G. Telson, lateral view. H. Posterior node of hypopharynx, fronto-ventral view. I. Lobe of body ring 3, lateral view. Scale bars: A = 1 mm; B–G, I = 0.5 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6323004" httpUri="https://zenodo.org/record/6323004/files/figure.png" pageId="7" pageNumber="37">Fig. 1D</figureCitation>
|
||
) or with three well-developed labral teeth (
|
||
<figureCitation id="5AA2D8EACF4A0D07FBABF9A8FB22F9DF" box="[1086,1178,1602,1628]" captionStart="Fig" captionStartId="6.[189,232,1857,1883]" captionTargetBox="[189,1398,340,1798]" captionTargetId="figure-11@6.[189,1398,340,1798]" captionTargetPageId="6" captionText="Fig. 1. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., ♂♂ (A–H) and ♀ (I) from Foran di Landri Cave (IZB). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Labrum, frontodorsal view. D. Labrum, dorsal view. E. Pleurotergum 7, lateral view. F. Mid-body rings, lateral view. G. Telson, lateral view. H. Posterior node of hypopharynx, fronto-ventral view. I. Lobe of body ring 3, lateral view. Scale bars: A = 1 mm; B–G, I = 0.5 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6323004" httpUri="https://zenodo.org/record/6323004/files/figure.png" pageId="7" pageNumber="37">Fig. 1C</figureCitation>
|
||
). Gnathochilarium (
|
||
<figureCitation id="5AA2D8EACF4A0D07FF52F98FFE99F9FC" box="[199,289,1637,1663]" captionStart="Fig" captionStartId="8.[189,232,1787,1813]" captionTargetBox="[209,1381,264,1720]" captionTargetId="figure-12@8.[209,1381,264,1720]" captionTargetPageId="8" captionText="Fig. 2. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., ♂ from from Foran di Landri Cave (IBER). A. Gnathochilarium, ventral view. B. Gnathal lobe of left mandible, ventral view. C. Left promere, caudal view. D. Left mesomere, oral view. E. Distal part of left opisthomere, mesal view. Abbreviations: ab = anterior branch of solenomere; el = external lobe; et = external tooth; il = internal lobe; it = internal tooth; mp = mollar plate; pb = posterior branch of solenomere; ph = posterior hump; pl = pectinate lamellae; pm = promentum; sf = setiform filaments; v = velum. Scale bars: A = 0.2 mm; B = 0.1 mm; C–D = 0.05 mm; E = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.6323008" httpUri="https://zenodo.org/record/6323008/files/figure.png" pageId="7" pageNumber="37">Fig. 2A</figureCitation>
|
||
) rather short and distally markedly broad, with the 3 long distal setae on each stipes usual for the family, stipites medially each with 4–10 short setae arranged longitudinally, stipital palps conspicuously large; promentum large (pm), completely separating lingual lamellae, the latter each with 2–3 proximal and 1 distal seta. Gnathal lobes of mandibles (
|
||
<figureCitation id="5AA2D8EACF4A0D07FD77F925FC85F96A" box="[738,829,1743,1769]" captionStart="Fig" captionStartId="8.[189,232,1787,1813]" captionTargetBox="[209,1381,264,1720]" captionTargetId="figure-12@8.[209,1381,264,1720]" captionTargetPageId="8" captionText="Fig. 2. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., ♂ from from Foran di Landri Cave (IBER). A. Gnathochilarium, ventral view. B. Gnathal lobe of left mandible, ventral view. C. Left promere, caudal view. D. Left mesomere, oral view. E. Distal part of left opisthomere, mesal view. Abbreviations: ab = anterior branch of solenomere; el = external lobe; et = external tooth; il = internal lobe; it = internal tooth; mp = mollar plate; pb = posterior branch of solenomere; ph = posterior hump; pl = pectinate lamellae; pm = promentum; sf = setiform filaments; v = velum. Scale bars: A = 0.2 mm; B = 0.1 mm; C–D = 0.05 mm; E = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.6323008" httpUri="https://zenodo.org/record/6323008/files/figure.png" pageId="7" pageNumber="37">Fig. 2B</figureCitation>
|
||
) with the external (et) and the internal tooth (it) slightly to moderately reduced; molar plate (mp) relatively small (but larger compared to that in
|
||
<taxonomicName id="0599BFECCF4A0D07FA99F918FACDF88F" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Manfredi" baseAuthorityYear="1953" box="[1292,1397,1778,1804]" class="Diplopoda" family="Julidae" genus="Typhloiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="37" phylum="Arthropoda" rank="species" species="ausugi" status="comb. nov.">
|
||
<emphasis id="F0ED187DCF4A0D07FA99F918FACDF88F" box="[1292,1397,1778,1804]" italics="true" pageId="7" pageNumber="37">S. ausugi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBDEA506CF4A0D07FF28F8FFFE87F8B3" box="[189,319,1813,1840]" pageId="7" pageNumber="37" rank="species">comb. nov.</taxonomicNameLabel>
|
||
); the four pectinate lamellae (pl) consisting of slender and very densely set teeth. Hypopharynx distally densely ciliate, posterior node reduced, with roundish posterior margin (
|
||
<figureCitation id="5AA2D8EACF4A0D07FBC8F8D2FB01F8D1" box="[1117,1209,1848,1874]" captionStart="Fig" captionStartId="6.[189,232,1857,1883]" captionTargetBox="[189,1398,340,1798]" captionTargetId="figure-11@6.[189,1398,340,1798]" captionTargetPageId="6" captionText="Fig. 1. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., ♂♂ (A–H) and ♀ (I) from Foran di Landri Cave (IZB). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Labrum, frontodorsal view. D. Labrum, dorsal view. E. Pleurotergum 7, lateral view. F. Mid-body rings, lateral view. G. Telson, lateral view. H. Posterior node of hypopharynx, fronto-ventral view. I. Lobe of body ring 3, lateral view. Scale bars: A = 1 mm; B–G, I = 0.5 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6323004" httpUri="https://zenodo.org/record/6323004/files/figure.png" pageId="7" pageNumber="37">Fig. 1H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF4A0D09FF28F890FE99FE0F" blockId="7.[189,1399,1914,2011]" lastBlockId="9.[188,1399,265,396]" lastPageId="9" lastPageNumber="39" pageId="7" pageNumber="37">
|
||
COLLUM. Entirely smooth, its frontolateral margin bent outwards. Body rings considerably vaulted. Prozonae smooth. Metazonae with well-developed striation only ventrally, dorsally and laterally with faint and short striae only in anterior parts; setae ca 20% of H (
|
||
<figureCitation id="5AA2D8EACF4A0D07FC3CF82AFBB9F859" box="[937,1025,1984,2010]" captionStart="Fig" captionStartId="6.[189,232,1857,1883]" captionTargetBox="[189,1398,340,1798]" captionTargetId="figure-11@6.[189,1398,340,1798]" captionTargetPageId="6" captionText="Fig. 1. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., ♂♂ (A–H) and ♀ (I) from Foran di Landri Cave (IZB). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Labrum, frontodorsal view. D. Labrum, dorsal view. E. Pleurotergum 7, lateral view. F. Mid-body rings, lateral view. G. Telson, lateral view. H. Posterior node of hypopharynx, fronto-ventral view. I. Lobe of body ring 3, lateral view. Scale bars: A = 1 mm; B–G, I = 0.5 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6323004" httpUri="https://zenodo.org/record/6323004/files/figure.png" pageId="7" pageNumber="37">Fig. 1F</figureCitation>
|
||
), arranged in moderately dense whorl. Ozopores very small, placed behind pro-metazonal suture at ca
|
||
<superScript id="35EC6927CF440D09FC62FEE3FBB8FE94" attach="right" box="[1015,1024,265,279]" fontSize="6" pageId="9" pageNumber="39">2</superScript>
|
||
⁄
|
||
<subScript id="5E1DC62ACF440D09FB93FEF9FBB7FEA2" attach="left" box="[1030,1039,275,289]" fontSize="6" pageId="9" pageNumber="39">5</subScript>
|
||
of metazonal length measured from front to back. Tarsus of mid-body legs 2.2–2.4 times as long as tibia and 2.8–3.7 times as long as apical claw; accessory claw absent. Mid-body legs 1.5–1.6 times as long as H in males, 1.4 times in females.
|
||
</paragraph>
|
||
<caption id="96E694E7CF450D08FF28F911FD70F86A" ID-DOI="http://doi.org/10.5281/zenodo.6323008" ID-Zenodo-Dep="6323008" httpUri="https://zenodo.org/record/6323008/files/figure.png" pageId="8" pageNumber="38" startId="8.[189,232,1787,1813]" targetBox="[209,1381,264,1720]" targetPageId="8">
|
||
<paragraph id="C226C46FCF450D08FF28F911FD70F86A" blockId="8.[189,1399,1787,2025]" pageId="8" pageNumber="38">
|
||
<emphasis id="F0ED187DCF450D08FF28F911FEB5F896" bold="true" box="[189,269,1787,1813]" pageId="8" pageNumber="38">Fig. 2.</emphasis>
|
||
<taxonomicName id="0599BFECCF450D08FE82F911FD6AF896" authority="(Strasser, 1971)" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Strasser" baseAuthorityYear="1971" box="[279,722,1787,1813]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="8" pageNumber="38" phylum="Arthropoda" rank="species" species="fimbriatus" status="comb. et stat. nov.">
|
||
<emphasis id="F0ED187DCF450D08FE82F911FDB4F896" box="[279,524,1787,1813]" italics="true" pageId="8" pageNumber="38">Stygiiulus fimbriatus</emphasis>
|
||
(Strasser, 1971)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBDEA506CF450D08FD49F911FC05F895" box="[732,957,1787,1814]" pageId="8" pageNumber="38">comb. et stat. nov.</taxonomicNameLabel>
|
||
, ♂ from from Foran di Landri Cave (IBER).
|
||
<emphasis id="F0ED187DCF450D08FEB2F8F4FE84F8BB" bold="true" box="[295,316,1822,1848]" pageId="8" pageNumber="38">A</emphasis>
|
||
. Gnathochilarium, ventral view.
|
||
<emphasis id="F0ED187DCF450D08FD45F8F4FD5CF8BB" bold="true" box="[720,740,1822,1848]" pageId="8" pageNumber="38">B</emphasis>
|
||
. Gnathal lobe of left mandible, ventral view.
|
||
<emphasis id="F0ED187DCF450D08FA88F8F4FA8AF8BB" bold="true" box="[1309,1330,1822,1848]" pageId="8" pageNumber="38">C</emphasis>
|
||
. Left promere, caudal view.
|
||
<emphasis id="F0ED187DCF450D08FE47F8A8FE5FF8DF" bold="true" box="[466,487,1858,1884]" pageId="8" pageNumber="38">D</emphasis>
|
||
. Left mesomere, oral view.
|
||
<emphasis id="F0ED187DCF450D08FCD4F8A8FCEDF8DF" bold="true" box="[833,853,1858,1884]" pageId="8" pageNumber="38">E</emphasis>
|
||
. Distal part of left opisthomere, mesal view. Abbreviations: ab = anterior branch of solenomere; el = external lobe; et = external tooth; il = internal lobe; it = internal tooth; mp = mollar plate; pb = posterior branch of solenomere; ph = posterior hump; pl = pectinate lamellae; pm = promentum; sf = setiform filaments; v = velum. Scale bars: A = 0.2 mm; B = 0.1 mm; C–D = 0.05 mm; E = 0.02 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C226C46FCF440D09FF28FE5FFBB0FDBB" blockId="9.[189,1399,436,568]" pageId="9" pageNumber="39">
|
||
TELSON (
|
||
<figureCitation id="5AA2D8EACF440D09FEB0FE5EFE3CFE4D" box="[293,388,436,463]" captionStart="Fig" captionStartId="6.[189,232,1857,1883]" captionTargetBox="[189,1398,340,1798]" captionTargetId="figure-11@6.[189,1398,340,1798]" captionTargetPageId="6" captionText="Fig. 1. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., ♂♂ (A–H) and ♀ (I) from Foran di Landri Cave (IZB). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Labrum, frontodorsal view. D. Labrum, dorsal view. E. Pleurotergum 7, lateral view. F. Mid-body rings, lateral view. G. Telson, lateral view. H. Posterior node of hypopharynx, fronto-ventral view. I. Lobe of body ring 3, lateral view. Scale bars: A = 1 mm; B–G, I = 0.5 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6323004" httpUri="https://zenodo.org/record/6323004/files/figure.png" pageId="9" pageNumber="39">Fig. 1G</figureCitation>
|
||
). Epiproct short and stout, straight all along, ending with a short hyaline tip turned more or less dorsad, barely protruding behind caudal contour of paraprocts. Hypoproct broadly rounded, tightly fitting under paraprocts, with rather evenly setose ventral face. Paraprocts densely covered with long setae, without distinct rows of shorter setae along caudal margins.
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF440D09FF28FD8AFE91FC10" blockId="9.[188,1399,607,915]" pageId="9" pageNumber="39">
|
||
MALE SEXUAL CHARACTERS. Leg-pair 1 (
|
||
<figureCitation id="5AA2D8EACF440D09FDEAFDB5FD63FDFA" box="[639,731,607,633]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[243,1336,266,1776]" captionTargetId="figure-12@10.[234,1353,262,1782]" captionTargetPageId="10" captionText="Fig. 3. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., holotype ♂ (A, B) (MHNG), ♂ (C) (IZB) and ♀ (D) (IBER) from Foran di Landri Cave. A. Leg-pair 1, caudal view. B. Penis, oral/caudal view. C. Left gonopods, mesal view. D. Left vulva, caudal, somewhat mesal view. Abbreviations: ab = anterior branch of solenomere; bs = basal spine; fc = flagellum channel; lt = lateral tube; m = mesomere; mt = median tube; op = operculum; p = promere; pb = posterior branch of solenomere; ph = posterior hump; v = velum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6323014" httpUri="https://zenodo.org/record/6323014/files/figure.png" pageId="9" pageNumber="39">Fig. 3A</figureCitation>
|
||
) rather slender hooks oriented towards one another; with three complete and another one or two faintly indicated segments; with apically densely microdentate tibial outgrowths, and without or with barely visible tarsal remnants. Legs of more anterior body rings with an adhesive pad on tibia, most pronounced in leg-pair 2, gradually diminishing in caudal direction, completely disappearing around mid-body; tibia and femora without modifications. Pleurotergum 7 (
|
||
<figureCitation id="5AA2D8EACF440D09FF75FCE5FE84FCAA" box="[224,316,783,809]" captionStart="Fig" captionStartId="6.[189,232,1857,1883]" captionTargetBox="[189,1398,340,1798]" captionTargetId="figure-11@6.[189,1398,340,1798]" captionTargetPageId="6" captionText="Fig. 1. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., ♂♂ (A–H) and ♀ (I) from Foran di Landri Cave (IZB). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Labrum, frontodorsal view. D. Labrum, dorsal view. E. Pleurotergum 7, lateral view. F. Mid-body rings, lateral view. G. Telson, lateral view. H. Posterior node of hypopharynx, fronto-ventral view. I. Lobe of body ring 3, lateral view. Scale bars: A = 1 mm; B–G, I = 0.5 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6323004" httpUri="https://zenodo.org/record/6323004/files/figure.png" pageId="9" pageNumber="39">Fig. 1E</figureCitation>
|
||
) caudo-ventrally forming small rounded lobes directed caudad. Penis (
|
||
<figureCitation id="5AA2D8EACF440D09FB33FCE5FABCFCAA" box="[1190,1284,783,809]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[243,1336,266,1776]" captionTargetId="figure-12@10.[234,1353,262,1782]" captionTargetPageId="10" captionText="Fig. 3. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., holotype ♂ (A, B) (MHNG), ♂ (C) (IZB) and ♀ (D) (IBER) from Foran di Landri Cave. A. Leg-pair 1, caudal view. B. Penis, oral/caudal view. C. Left gonopods, mesal view. D. Left vulva, caudal, somewhat mesal view. Abbreviations: ab = anterior branch of solenomere; bs = basal spine; fc = flagellum channel; lt = lateral tube; m = mesomere; mt = median tube; op = operculum; p = promere; pb = posterior branch of solenomere; ph = posterior hump; v = velum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6323014" httpUri="https://zenodo.org/record/6323014/files/figure.png" pageId="9" pageNumber="39">Fig. 3B</figureCitation>
|
||
) long, in situ visible behind coxae 2, slender hourglass shaped: broadest at base, narrowing until the middle, then slightly widening again, ending with short, diverging, apical lobes bearing fine, blunt, terminal lamellae.
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF440D09FF28FC50FE50FAF8" blockId="9.[187,1400,954,1403]" pageId="9" pageNumber="39">
|
||
GONOPODS (
|
||
<figureCitation id="5AA2D8EACF440D09FED1FC50FE27FC56" box="[324,415,954,981]" captionStart="Fig" captionStartId="6.[189,232,1857,1883]" captionTargetBox="[189,1398,340,1798]" captionTargetId="figure-11@6.[189,1398,340,1798]" captionTargetPageId="6" captionText="Fig. 1. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., ♂♂ (A–H) and ♀ (I) from Foran di Landri Cave (IZB). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Labrum, frontodorsal view. D. Labrum, dorsal view. E. Pleurotergum 7, lateral view. F. Mid-body rings, lateral view. G. Telson, lateral view. H. Posterior node of hypopharynx, fronto-ventral view. I. Lobe of body ring 3, lateral view. Scale bars: A = 1 mm; B–G, I = 0.5 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6323004" httpUri="https://zenodo.org/record/6323004/files/figure.png" pageId="9" pageNumber="39">Figs 1B</figureCitation>
|
||
,
|
||
<figureCitation id="5AA2D8EACF440D09FE39FC50FE57FC56" box="[428,495,954,981]" captionStart="Fig" captionStartId="8.[189,232,1787,1813]" captionTargetBox="[209,1381,264,1720]" captionTargetId="figure-12@8.[209,1381,264,1720]" captionTargetPageId="8" captionText="Fig. 2. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., ♂ from from Foran di Landri Cave (IBER). A. Gnathochilarium, ventral view. B. Gnathal lobe of left mandible, ventral view. C. Left promere, caudal view. D. Left mesomere, oral view. E. Distal part of left opisthomere, mesal view. Abbreviations: ab = anterior branch of solenomere; el = external lobe; et = external tooth; il = internal lobe; it = internal tooth; mp = mollar plate; pb = posterior branch of solenomere; ph = posterior hump; pl = pectinate lamellae; pm = promentum; sf = setiform filaments; v = velum. Scale bars: A = 0.2 mm; B = 0.1 mm; C–D = 0.05 mm; E = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.6323008" httpUri="https://zenodo.org/record/6323008/files/figure.png" pageId="9" pageNumber="39">2C–E</figureCitation>
|
||
,
|
||
<figureCitation id="5AA2D8EACF440D09FE69FC50FDA7FC57" box="[508,543,954,980]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[243,1336,266,1776]" captionTargetId="figure-12@10.[234,1353,262,1782]" captionTargetPageId="10" captionText="Fig. 3. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., holotype ♂ (A, B) (MHNG), ♂ (C) (IZB) and ♀ (D) (IBER) from Foran di Landri Cave. A. Leg-pair 1, caudal view. B. Penis, oral/caudal view. C. Left gonopods, mesal view. D. Left vulva, caudal, somewhat mesal view. Abbreviations: ab = anterior branch of solenomere; bs = basal spine; fc = flagellum channel; lt = lateral tube; m = mesomere; mt = median tube; op = operculum; p = promere; pb = posterior branch of solenomere; ph = posterior hump; v = velum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6323014" httpUri="https://zenodo.org/record/6323014/files/figure.png" pageId="9" pageNumber="39">3C</figureCitation>
|
||
). In situ obliquely protruding from gonopodal sinus with their distal parts (
|
||
<figureCitation id="5AA2D8EACF440D09FF51FC37FE99FC74" box="[196,289,989,1015]" captionStart="Fig" captionStartId="6.[189,232,1857,1883]" captionTargetBox="[189,1398,340,1798]" captionTargetId="figure-11@6.[189,1398,340,1798]" captionTargetPageId="6" captionText="Fig. 1. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., ♂♂ (A–H) and ♀ (I) from Foran di Landri Cave (IZB). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Labrum, frontodorsal view. D. Labrum, dorsal view. E. Pleurotergum 7, lateral view. F. Mid-body rings, lateral view. G. Telson, lateral view. H. Posterior node of hypopharynx, fronto-ventral view. I. Lobe of body ring 3, lateral view. Scale bars: A = 1 mm; B–G, I = 0.5 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6323004" httpUri="https://zenodo.org/record/6323004/files/figure.png" pageId="9" pageNumber="39">Fig. 1B</figureCitation>
|
||
). Promere (
|
||
<figureCitation id="5AA2D8EACF440D09FE24FC37FD94FC7B" box="[433,556,989,1016]" captionStart="Fig" captionStartId="8.[189,232,1787,1813]" captionTargetBox="[209,1381,264,1720]" captionTargetId="figure-12@8.[209,1381,264,1720]" captionTargetPageId="8" captionText="Fig. 2. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., ♂ from from Foran di Landri Cave (IBER). A. Gnathochilarium, ventral view. B. Gnathal lobe of left mandible, ventral view. C. Left promere, caudal view. D. Left mesomere, oral view. E. Distal part of left opisthomere, mesal view. Abbreviations: ab = anterior branch of solenomere; el = external lobe; et = external tooth; il = internal lobe; it = internal tooth; mp = mollar plate; pb = posterior branch of solenomere; ph = posterior hump; pl = pectinate lamellae; pm = promentum; sf = setiform filaments; v = velum. Scale bars: A = 0.2 mm; B = 0.1 mm; C–D = 0.05 mm; E = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.6323008" httpUri="https://zenodo.org/record/6323008/files/figure.png" pageId="9" pageNumber="39">Fig. 2C, p</figureCitation>
|
||
in
|
||
<figureCitation id="5AA2D8EACF440D09FDC1FC37FD16FC74" box="[596,686,989,1015]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[243,1336,266,1776]" captionTargetId="figure-12@10.[234,1353,262,1782]" captionTargetPageId="10" captionText="Fig. 3. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., holotype ♂ (A, B) (MHNG), ♂ (C) (IZB) and ♀ (D) (IBER) from Foran di Landri Cave. A. Leg-pair 1, caudal view. B. Penis, oral/caudal view. C. Left gonopods, mesal view. D. Left vulva, caudal, somewhat mesal view. Abbreviations: ab = anterior branch of solenomere; bs = basal spine; fc = flagellum channel; lt = lateral tube; m = mesomere; mt = median tube; op = operculum; p = promere; pb = posterior branch of solenomere; ph = posterior hump; v = velum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6323014" httpUri="https://zenodo.org/record/6323014/files/figure.png" pageId="9" pageNumber="39">Fig. 3C</figureCitation>
|
||
) somewhat higher than mesomere, both being significantly surpassed by the opisthomere. Promere stout, oar-shaped (broadest apically), forming a near rightangled meso-apical corner and a broadly rounded latero-apical corner; apical margin bent somewhat frontad; caudal face distally densely microsquamose, basally with an internal and an external lobe, both being short and robust, partly fused to one another. Mesomere (
|
||
<figureCitation id="5AA2D8EACF440D09FC01FB80FBA9FB06" box="[916,1041,1130,1157]" captionStart="Fig" captionStartId="8.[189,232,1787,1813]" captionTargetBox="[209,1381,264,1720]" captionTargetId="figure-12@8.[209,1381,264,1720]" captionTargetPageId="8" captionText="Fig. 2. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., ♂ from from Foran di Landri Cave (IBER). A. Gnathochilarium, ventral view. B. Gnathal lobe of left mandible, ventral view. C. Left promere, caudal view. D. Left mesomere, oral view. E. Distal part of left opisthomere, mesal view. Abbreviations: ab = anterior branch of solenomere; el = external lobe; et = external tooth; il = internal lobe; it = internal tooth; mp = mollar plate; pb = posterior branch of solenomere; ph = posterior hump; pl = pectinate lamellae; pm = promentum; sf = setiform filaments; v = velum. Scale bars: A = 0.2 mm; B = 0.1 mm; C–D = 0.05 mm; E = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.6323008" httpUri="https://zenodo.org/record/6323008/files/figure.png" pageId="9" pageNumber="39">Fig. 2D, m</figureCitation>
|
||
in
|
||
<figureCitation id="5AA2D8EACF440D09FBA7FB80FB31FB07" box="[1074,1161,1130,1156]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[243,1336,266,1776]" captionTargetId="figure-12@10.[234,1353,262,1782]" captionTargetPageId="10" captionText="Fig. 3. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., holotype ♂ (A, B) (MHNG), ♂ (C) (IZB) and ♀ (D) (IBER) from Foran di Landri Cave. A. Leg-pair 1, caudal view. B. Penis, oral/caudal view. C. Left gonopods, mesal view. D. Left vulva, caudal, somewhat mesal view. Abbreviations: ab = anterior branch of solenomere; bs = basal spine; fc = flagellum channel; lt = lateral tube; m = mesomere; mt = median tube; op = operculum; p = promere; pb = posterior branch of solenomere; ph = posterior hump; v = velum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6323014" httpUri="https://zenodo.org/record/6323014/files/figure.png" pageId="9" pageNumber="39">Fig. 3C</figureCitation>
|
||
) narrow spoon-like, with the apex turned meso-frontad; caudal face distally deeply concave and sparsely microsquamose. Opisthomere (
|
||
<figureCitation id="5AA2D8EACF440D09FEFDFB5AFE72FB48" box="[360,458,1200,1227]" captionStart="Fig" captionStartId="8.[189,232,1787,1813]" captionTargetBox="[209,1381,264,1720]" captionTargetId="figure-12@8.[209,1381,264,1720]" captionTargetPageId="8" captionText="Fig. 2. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., ♂ from from Foran di Landri Cave (IBER). A. Gnathochilarium, ventral view. B. Gnathal lobe of left mandible, ventral view. C. Left promere, caudal view. D. Left mesomere, oral view. E. Distal part of left opisthomere, mesal view. Abbreviations: ab = anterior branch of solenomere; el = external lobe; et = external tooth; il = internal lobe; it = internal tooth; mp = mollar plate; pb = posterior branch of solenomere; ph = posterior hump; pl = pectinate lamellae; pm = promentum; sf = setiform filaments; v = velum. Scale bars: A = 0.2 mm; B = 0.1 mm; C–D = 0.05 mm; E = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.6323008" httpUri="https://zenodo.org/record/6323008/files/figure.png" pageId="9" pageNumber="39">Figs 2E</figureCitation>
|
||
,
|
||
<figureCitation id="5AA2D8EACF440D09FE4DFB5BFE43FB48" box="[472,507,1201,1227]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[243,1336,266,1776]" captionTargetId="figure-12@10.[234,1353,262,1782]" captionTargetPageId="10" captionText="Fig. 3. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., holotype ♂ (A, B) (MHNG), ♂ (C) (IZB) and ♀ (D) (IBER) from Foran di Landri Cave. A. Leg-pair 1, caudal view. B. Penis, oral/caudal view. C. Left gonopods, mesal view. D. Left vulva, caudal, somewhat mesal view. Abbreviations: ab = anterior branch of solenomere; bs = basal spine; fc = flagellum channel; lt = lateral tube; m = mesomere; mt = median tube; op = operculum; p = promere; pb = posterior branch of solenomere; ph = posterior hump; v = velum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6323014" httpUri="https://zenodo.org/record/6323014/files/figure.png" pageId="9" pageNumber="39">3C</figureCitation>
|
||
) markedly slender, with a distinct, obtuse posterior hump (ph); velum (v) pointing distad, both its frontal and caudal margin apically deeply serrated; solenomere with a slender posterior branch (pb) bent caudad, ending with several long ciliae, anterior branch (ab) vestigial, barely protruding, mostly concealed between the velum and the posterior solenomeral branch, densely ciliate; with 2 or 3 fine distal setiform filaments (sf), and sometimes with a variously developed basal spine (bs) at flagellum channel (fc).
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF440D09FF28FA48FD30F931" blockId="9.[187,1400,1442,1715]" pageId="9" pageNumber="39">
|
||
FEMALE SEXUAL CHARACTERS. Leg-pairs 1 and 2 visibly shorter, 1 also thicker, than following legs. Ventral margin of body ring 3 with roundish anterior lobe (
|
||
<figureCitation id="5AA2D8EACF440D09FCBBFA2FFC3CFA5C" box="[814,900,1477,1503]" captionStart="Fig" captionStartId="6.[189,232,1857,1883]" captionTargetBox="[189,1398,340,1798]" captionTargetId="figure-11@6.[189,1398,340,1798]" captionTargetPageId="6" captionText="Fig. 1. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., ♂♂ (A–H) and ♀ (I) from Foran di Landri Cave (IZB). A. Habitus, lateral view. B. Anterior part of body, lateral view. C. Labrum, frontodorsal view. D. Labrum, dorsal view. E. Pleurotergum 7, lateral view. F. Mid-body rings, lateral view. G. Telson, lateral view. H. Posterior node of hypopharynx, fronto-ventral view. I. Lobe of body ring 3, lateral view. Scale bars: A = 1 mm; B–G, I = 0.5 mm; H = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6323004" httpUri="https://zenodo.org/record/6323004/files/figure.png" pageId="9" pageNumber="39">Fig. 1I</figureCitation>
|
||
). Vulva (
|
||
<figureCitation id="5AA2D8EACF440D09FC62FA2FFBEDFA5C" box="[1015,1109,1477,1503]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[243,1336,266,1776]" captionTargetId="figure-12@10.[234,1353,262,1782]" captionTargetPageId="10" captionText="Fig. 3. Stygiiulus fimbriatus (Strasser, 1971) comb. et stat. nov., holotype ♂ (A, B) (MHNG), ♂ (C) (IZB) and ♀ (D) (IBER) from Foran di Landri Cave. A. Leg-pair 1, caudal view. B. Penis, oral/caudal view. C. Left gonopods, mesal view. D. Left vulva, caudal, somewhat mesal view. Abbreviations: ab = anterior branch of solenomere; bs = basal spine; fc = flagellum channel; lt = lateral tube; m = mesomere; mt = median tube; op = operculum; p = promere; pb = posterior branch of solenomere; ph = posterior hump; v = velum. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6323014" httpUri="https://zenodo.org/record/6323014/files/figure.png" pageId="9" pageNumber="39">Fig. 3D</figureCitation>
|
||
) somewhat compressed in the sagittal plane, mostly symmetric: lateral side of operculum higher and apically more pointed than mesal one; each valve of bursa with one vertical row of setae; operculum (op) broad and thick, with a concave apical margin, exceeding bursa by ca ¼ of total height of vulva, medio-laterally with two longitudinal rows of setae each side. Receptaculum seminis consisting of two short and narrow, somewhat folded tubes – a lateral (lt) and a mesal one (mt), both insignificantly widening towards bottom, not forming distinct ampullae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8A8397E4CF440D09FF28F933FEA4F8C5" pageId="9" pageNumber="39" type="distribution">
|
||
<paragraph id="C226C46FCF440D09FF28F933FEE0F970" blockId="9.[189,344,1753,1779]" box="[189,344,1753,1779]" pageId="9" pageNumber="39">
|
||
<heading id="996E7303CF440D09FF28F933FEE0F970" bold="true" box="[189,344,1753,1779]" fontSize="11" level="3" pageId="9" pageNumber="39" reason="3">
|
||
<emphasis id="F0ED187DCF440D09FF28F933FEE0F970" bold="true" box="[189,344,1753,1779]" pageId="9" pageNumber="39">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF440D09FF28F8E3FEA4F8C5" blockId="9.[189,1398,1800,1862]" pageId="9" pageNumber="39">
|
||
Known only from two caves in the Julian Alps in
|
||
<collectingCountry id="BA8E84FFCF440D09FC98F8E2FCFDF8A1" box="[781,837,1800,1826]" name="Italy" pageId="9" pageNumber="39">Italy</collectingCountry>
|
||
, near the border with
|
||
<collectingCountry id="BA8E84FFCF440D09FBDCF8E2FB09F8A1" box="[1097,1201,1800,1826]" name="Slovenia" pageId="9" pageNumber="39">Slovenia</collectingCountry>
|
||
(
|
||
<figureCitation id="5AA2D8EACF440D09FB54F8E2FAA1F8A1" box="[1217,1305,1800,1826]" captionStart="Fig" captionStartId="33.[189,232,1998,2024]" captionTargetBox="[189,1398,1099,1960]" captionTargetId="figure-332@33.[189,1398,1099,1960]" captionTargetPageId="33" captionText="Fig. 13. General distribution of the species of the genus Stygiiulus stat. nov." figureDoi="http://doi.org/10.5281/zenodo.6323034" httpUri="https://zenodo.org/record/6323034/files/figure.png" pageId="9" pageNumber="39">Fig. 13</figureCitation>
|
||
, yellow circles).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8A8397E4CF440D0BFF28F887FCFEFDBB" lastPageId="11" lastPageNumber="41" pageId="9" pageNumber="39" type="discussion">
|
||
<paragraph id="C226C46FCF440D09FF28F887FE9BF804" blockId="9.[189,291,1901,1927]" box="[189,291,1901,1927]" pageId="9" pageNumber="39">
|
||
<heading id="996E7303CF440D09FF28F887FE9BF804" bold="true" box="[189,291,1901,1927]" fontSize="11" level="3" pageId="9" pageNumber="39" reason="3">
|
||
<emphasis id="F0ED187DCF440D09FF28F887FE9BF804" bold="true" box="[189,291,1901,1927]" pageId="9" pageNumber="39">Remark</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF440D09FF28F871FACEF85A" blockId="9.[187,1398,1947,2009]" pageId="9" pageNumber="39">
|
||
The new locality of
|
||
<taxonomicName id="0599BFECCF440D09FE35F876FD8DF836" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Strasser" baseAuthorityYear="1971" box="[416,565,1947,1974]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="9" pageNumber="39" phylum="Arthropoda" rank="species" species="fimbriatus" status="comb. et stat. nov.">
|
||
<emphasis id="F0ED187DCF440D09FE35F876FD8DF836" box="[416,565,1947,1974]" italics="true" pageId="9" pageNumber="39">S. fimbriatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBDEA506CF440D09FDAFF871FCB5F835" box="[570,781,1947,1974]" pageId="9" pageNumber="39">comb. et stat. nov.</taxonomicNameLabel>
|
||
, Cave Foran di Landri, is an active cave with a stream running through it year-round, flowing out of the entrance, after passing through a series of syphons
|
||
</paragraph>
|
||
<caption id="96E694E7CF470D0AFF28F8F4FD37F86A" ID-DOI="http://doi.org/10.5281/zenodo.6323014" ID-Zenodo-Dep="6323014" httpUri="https://zenodo.org/record/6323014/files/figure.png" pageId="10" pageNumber="40" startId="10.[189,232,1822,1848]" targetBox="[243,1336,266,1776]" targetPageId="10">
|
||
<paragraph id="C226C46FCF470D0AFF28F8F4FD37F86A" blockId="10.[189,1399,1822,2025]" pageId="10" pageNumber="40">
|
||
<emphasis id="F0ED187DCF470D0AFF28F8F4FEB5F8BB" bold="true" box="[189,269,1822,1848]" pageId="10" pageNumber="40">Fig. 3.</emphasis>
|
||
<taxonomicName id="0599BFECCF470D0AFE82F8F4FD69F8BB" authority="(Strasser, 1971)" authorityName="Vagalinski & Borissov & Bobeva & Canciani & Antić" authorityYear="2022" baseAuthorityName="Strasser" baseAuthorityYear="1971" box="[279,721,1822,1848]" class="Diplopoda" family="Julidae" genus="Stygiiulus" kingdom="Animalia" order="Julida" pageId="10" pageNumber="40" phylum="Arthropoda" rank="species" species="fimbriatus" status="comb. et stat. nov.">
|
||
<emphasis id="F0ED187DCF470D0AFE82F8F4FDB4F8BB" box="[279,524,1822,1848]" italics="true" pageId="10" pageNumber="40">Stygiiulus fimbriatus</emphasis>
|
||
(Strasser, 1971)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EBDEA506CF470D0AFD4EF8F4FC03F8BA" box="[731,955,1822,1849]" pageId="10" pageNumber="40">comb. et stat. nov.</taxonomicNameLabel>
|
||
, holotype ♂ (A, B) (MHNG), ♂ (C) (IZB) and ♀ (D) (IBER) from Foran di Landri Cave.
|
||
<emphasis id="F0ED187DCF470D0AFCA2F8ABFCF4F8D8" bold="true" box="[823,844,1857,1883]" pageId="10" pageNumber="40">A</emphasis>
|
||
. Leg-pair 1, caudal view.
|
||
<emphasis id="F0ED187DCF470D0AFB16F8A8FB2EF8DF" bold="true" box="[1155,1174,1858,1884]" pageId="10" pageNumber="40">B</emphasis>
|
||
. Penis, oral/caudal view.
|
||
<emphasis id="F0ED187DCF470D0AFE96F88EFEA0F8FD" bold="true" box="[259,280,1892,1918]" pageId="10" pageNumber="40">C</emphasis>
|
||
. Left gonopods, mesal view.
|
||
<emphasis id="F0ED187DCF470D0AFDE1F88FFD31F8FC" bold="true" box="[628,649,1893,1919]" pageId="10" pageNumber="40">D</emphasis>
|
||
. Left vulva, caudal, somewhat mesal view. Abbreviations: ab = anterior branch of solenomere; bs = basal spine; fc = flagellum channel; lt = lateral tube; m = mesomere; mt = median tube; op = operculum; p = promere; pb = posterior branch of solenomere; ph = posterior hump; v = velum. Scale bars = 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C226C46FCF460D0BFF28FEE3FD72FE50" blockId="11.[189,1399,265,467]" pageId="11" pageNumber="41">
|
||
(four discovered so far). All examined specimens were found right beyond the first two syphons, ca 40 meters from the entrance. This part of the cave includes a number of ponds, and it probably becomes completely flooded during rainy periods (
|
||
<bibRefCitation id="A608B99ECF460D0BFD33FEA5FCE9FEEA" author="Canciani G." box="[678,849,335,361]" pageId="11" pageNumber="41" pagination="65 - 79" refId="ref20585" refString="Canciani G. 2019. La fauna cavernicola del Foran di Landri. In: A. S. Forum Julii Speleo (ed.) Foran di Landri. Il Landri svelato: ricerche e approfondimenti su una grotta tra storia e folklore: 65 - 79." type="book chapter" year="2019">Canciani 2019</bibRefCitation>
|
||
). Part of the individuals was observed moving on the surface of the ponds – a behaviour most likely associated with a filtrating rather than chewing feeding mode, in which small organic particles adhere to the fine and densely set teeth of the pectinate lamellae and to the cilia of the hypopharynx.
|
||
</paragraph>
|
||
<paragraph id="C226C46FCF460D0BFF28FE10FCFEFDBB" blockId="11.[189,1398,506,568]" pageId="11" pageNumber="41">
|
||
The cave of San Giovanni d'Antro (the
|
||
<typeStatus id="1D227ACDCF460D0BFD39FE11FD67FD96" box="[684,735,507,533]" pageId="11" pageNumber="41">type</typeStatus>
|
||
locality) is of the same
|
||
<typeStatus id="1D227ACDCF460D0BFB81FE11FBFFFD96" box="[1044,1095,507,533]" pageId="11" pageNumber="41">type</typeStatus>
|
||
as Foran di Landri, and
|
||
<bibRefCitation id="A608B99ECF460D0BFF28FDF4FEC5FDBB" author="Strasser K." box="[189,381,542,568]" pageId="11" pageNumber="41" pagination="1 - 21" refId="ref22340" refString="Strasser K. 1971 a. Uber italienische, besonders kavernikole Diplopoden. Memorie del Museo civico di Storia naturale di Verona 19: 1 - 21." type="journal article" year="1971">Strasser (1971a)</bibRefCitation>
|
||
noted that it was occasionally flooded.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |