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<document id="BCA71120E14F8A00809B37655D0184E4" ID-DOI="10.1111/j.1096-3642.2003.00099.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5428901" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630419746950" checkinUser="diego" docAuthor="Lozouet, Pierre" docDate="2004" docId="4873053EFF8CFFCF50DCFD49A482F8CA" docLanguage="en" docName="j.1096-3642.2003.00099.x.pdf" docOrigin="Zoological Journal of the Linnean Society 140 (4)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2003.00099.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Bourdieria Lozouet 2004, GEN. NOV." docType="treatment" docVersion="3" lastPageNumber="454" masterDocId="B44A7D46FF88FFC85313FFE8A538FFBB" masterDocTitle="The European Tertiary Neritiliidae (Mollusca, Gastropoda, Neritopsina): indicators of tropical submarine cave environments and freshwater faunas" masterLastPageNumber="467" masterPageNumber="447" pageNumber="451" updateTime="1699263735479" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="367332003DCDCBB0533C48A425BF7663">The European Tertiary Neritiliidae (Mollusca, Gastropoda, Neritopsina): indicators of tropical submarine cave environments and freshwater faunas</mods:title>
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<mods:namePart id="E4C41FD6848155516BA5BC33733F319F">Lozouet, Pierre</mods:namePart>
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<mods:date id="E2C58B23CF1DEBC6A1F553DA643BC869">2004</mods:date>
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<mods:number id="9263AD5DF008043F8E05D6DECC6F429B">2004-04-30</mods:number>
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<mods:number id="27643AB29D00B2C7A067F65B1CBA855E">140</mods:number>
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<mods:identifier id="3639959673C539DD1A8FE8563959199F" type="DOI">10.1111/j.1096-3642.2003.00099.x</mods:identifier>
<mods:identifier id="631C0D92DF61FCE0D31D58ECFD29F47C" type="ISSN">0024-4082</mods:identifier>
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<treatment id="4873053EFF8CFFCF50DCFD49A482F8CA" LSID="urn:lsid:plazi:treatment:4873053EFF8CFFCF50DCFD49A482F8CA" httpUri="http://treatment.plazi.org/id/4873053EFF8CFFCF50DCFD49A482F8CA" lastPageId="7" lastPageNumber="454" pageId="4" pageNumber="451">
<subSubSection id="88C0E7A3FF8CFFCC50DCFD49A1DDFD02" box="[975,1253,673,697]" pageId="4" pageNumber="451" type="nomenclature">
<paragraph id="C065B428FF8CFFCC50DCFD49A1DDFD02" blockId="4.[975,1253,673,697]" box="[975,1253,673,697]" pageId="4" pageNumber="451">
<heading id="9B2D0344FF8CFFCC50DCFD49A1DDFD02" box="[975,1253,673,697]" centered="true" fontSize="9" level="2" pageId="4" pageNumber="451" reason="2">
<taxonomicName id="07DACFABFF8CFFCC50DCFD49A15DFD03" ID-CoL="7NTPC" authority="Lozouet, 2004" authorityName="Lozouet" authorityYear="2004" box="[975,1125,673,697]" class="Gastropoda" family="Neritiliidae" genus="Bourdieria" kingdom="Animalia" order="Cycloneritida" pageId="4" pageNumber="451" phylum="Mollusca" rank="genus" status="gen. nov.">BOURDIERIA</taxonomicName>
<emphasis id="F2AE683AFF8CFFCC577EFD4DA1DDFD02" bold="true" box="[1133,1253,673,697]" pageId="4" pageNumber="451">
<taxonomicNameLabel id="E99DD541FF8CFFCC577EFD4DA1DDFD02" box="[1133,1253,673,697]" pageId="4" pageNumber="451" rank="genus">GEN. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="88C0E7A3FF8CFFCC5035FD3BA6BDF994" pageId="4" pageNumber="451" type="materials_examined">
<paragraph id="C065B428FF8CFFCC5035FD3BA1DFFD5C" blockId="4.[806,1255,722,744]" box="[806,1255,722,744]" pageId="4" pageNumber="451">
<emphasis id="F2AE683AFF8CFFCC5035FD3BA1B2FD5C" box="[806,1162,722,744]" italics="true" pageId="4" pageNumber="451">
<typeStatus id="1F610A8AFF8CFFCC5035FD3BA663FD53" box="[806,859,723,744]" pageId="4" pageNumber="451">Type</typeStatus>
species:
<taxonomicName id="07DACFABFF8CFFCC50D7FD3AA1B2FD5C" authority="Lozouet, 2004" authorityName="Lozouet" authorityYear="2004" box="[964,1162,722,743]" class="Gastropoda" family="Neritiliidae" genus="Bourdieria" kingdom="Animalia" order="Cycloneritida" pageId="4" pageNumber="451" phylum="Mollusca" rank="species" species="faviai" status="sp. nov.">Bourdieria faviai</taxonomicName>
</emphasis>
<taxonomicNameLabel id="E99DD541FF8CFFCC5780FD3AA1DFFD5C" box="[1171,1255,722,743]" pageId="4" pageNumber="451" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="C065B428FF8CFFCC5035FCE8A6C7FCE8" blockId="4.[806,1422,768,851]" pageId="4" pageNumber="451">
<emphasis id="F2AE683AFF8CFFCC5035FCE8A69CFCAE" box="[806,932,768,789]" italics="true" pageId="4" pageNumber="451">Etymology:</emphasis>
Dedicated to the late Franck Bourdier, one of the most original French Quaternary geologists of the last century.
</paragraph>
<paragraph id="C065B428FF8CFFCC5035FC84A140FBE3" blockId="4.[806,1422,875,1112]" pageId="4" pageNumber="451">
<emphasis id="F2AE683AFF8CFFCC5035FC84A696FC3A" box="[806,942,876,897]" italics="true" pageId="4" pageNumber="451">Description:</emphasis>
The shell is small, typically about
<quantity id="072219CDFF8CFFCC564EFC83A643FC24" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="4.4" metricValueMin="3.2" pageId="4" pageNumber="451" unit="mm" value="3.8" valueMax="4.4" valueMin="3.2">3.2- 4.4 mm</quantity>
maximum adult width, very solid, with a flat spire. The protoconch is multispiral, tilted relative to the teleoconch coiling axis and ornamented with spiral ridges. The teleoconch is ornamented with strong spiral cords. The inner lip bears five to six teeth and the outer lip bears four strong teeth. The semicircular aperture is highly prosocline.
</paragraph>
<paragraph id="C065B428FF8CFFCC5035FB98A0B7FAE6" blockId="4.[806,1423,1136,1373]" pageId="4" pageNumber="451">
<emphasis id="F2AE683AFF8CFFCC5035FB98A6A9FB3E" box="[806,913,1136,1157]" italics="true" pageId="4" pageNumber="451">Remarks:</emphasis>
Superficially,
<taxonomicName id="07DACFABFF8CFFCC5753FB99A198FB3E" authorityName="Lozouet" authorityYear="2004" box="[1088,1184,1136,1158]" class="Gastropoda" family="Neritiliidae" genus="Bourdieria" kingdom="Animalia" order="Cycloneritida" pageId="4" pageNumber="451" phylum="Mollusca" rank="species" species="faviai">
<emphasis id="F2AE683AFF8CFFCC5753FB99A198FB3E" box="[1088,1184,1136,1158]" italics="true" pageId="4" pageNumber="451">B. faviai</emphasis>
</taxonomicName>
is more similar to members of
<taxonomicName id="07DACFABFF8CFFCC50ABFB67A11DFB1E" authorityName="Rafinesque" authorityYear="1815" box="[952,1061,1167,1189]" class="Gastropoda" family="Neritidae" kingdom="Animalia" order="Cycloneritida" pageId="4" pageNumber="451" phylum="Mollusca" rank="family">Neritidae</taxonomicName>
than to any genus of
<taxonomicName id="07DACFABFF8CFFCC5623FB67A660FB78" authorityName="SCHEPMAN" authorityYear="1908" class="Gastropoda" family="Neritiliidae" kingdom="Animalia" order="Cycloneritida" pageId="4" pageNumber="451" phylum="Mollusca" rank="family">Neritiliidae</taxonomicName>
. In fact,
<taxonomicName id="07DACFABFF8CFFCC50AFFB45A100FB79" authorityName="Lozouet" authorityYear="2004" box="[956,1080,1197,1218]" class="Gastropoda" family="Neritiliidae" genus="Bourdieria" kingdom="Animalia" order="Cycloneritida" pageId="4" pageNumber="451" phylum="Mollusca" rank="genus">
<emphasis id="F2AE683AFF8CFFCC50AFFB45A100FB79" box="[956,1080,1197,1218]" italics="true" pageId="4" pageNumber="451">Bourdieria</emphasis>
</taxonomicName>
is unique among the
<taxonomicName id="07DACFABFF8CFFCC5622FB45A66EFB59" authorityName="SCHEPMAN" authorityYear="1908" class="Gastropoda" family="Neritiliidae" kingdom="Animalia" order="Cycloneritida" pageId="4" pageNumber="451" phylum="Mollusca" rank="family">Neritiliidae</taxonomicName>
because of the presence of strong spiral cords on the teleoconch. Compared to other
<taxonomicName id="07DACFABFF8CFFCC57C2FB03A060FABA" authorityName="SCHEPMAN" authorityYear="1908" box="[1233,1368,1259,1281]" class="Gastropoda" family="Neritiliidae" kingdom="Animalia" order="Cycloneritida" pageId="4" pageNumber="451" phylum="Mollusca" rank="family">Neritiliidae</taxonomicName>
, the spire is also especially flat. The protoconch is tilted relative to the teleoconch (
<figureCitation id="58E1A8ADFF8CFFCC5747FAC0A1A5FA85" box="[1108,1181,1320,1342]" captionStart="Figure 4" captionStartId="6.[142,220,1705,1724]" captionTargetBox="[195,1368,200,1672]" captionTargetId="figure-138@6.[195,1368,200,1672]" captionTargetPageId="6" captionText="Figure 4. Bourdieria faviai sp. nov. from Peyrère (Upper Oligocene). A, broken specimen (apical whorls removed) showing (arrowed) the ridge inside the aperture (MNHN-PL1634E). B &amp; C, juvenile specimen of 0.8 adult whorl (MNHN- PL1634C). B, apical view of the protoconch showing the position of three weak ridges (arrowed). D-F, views of the holotype (MNHN-PL1634A); D, apertural view, E, right lateral view, F, dorsal view. G, view of the apical part showing the ridges of the protoconch (arrow). H, broken specimen showing internal view of the columellar area. I &amp; J, apical view of the protoconch; arrow indicates the embryonic shell. I, detail of the embryonic shell." figureDoi="http://doi.org/10.5281/zenodo.5428909" httpUri="https://zenodo.org/record/5428909/files/figure.png" pageId="4" pageNumber="451">Fig. 4I</figureCitation>
) with spiral ridges (
<figureCitation id="58E1A8ADFF8CFFCC503DFAAFA6B9FAE6" box="[814,897,1351,1373]" captionStart="Figure 4" captionStartId="6.[142,220,1705,1724]" captionTargetBox="[195,1368,200,1672]" captionTargetId="figure-138@6.[195,1368,200,1672]" captionTargetPageId="6" captionText="Figure 4. Bourdieria faviai sp. nov. from Peyrère (Upper Oligocene). A, broken specimen (apical whorls removed) showing (arrowed) the ridge inside the aperture (MNHN-PL1634E). B &amp; C, juvenile specimen of 0.8 adult whorl (MNHN- PL1634C). B, apical view of the protoconch showing the position of three weak ridges (arrowed). D-F, views of the holotype (MNHN-PL1634A); D, apertural view, E, right lateral view, F, dorsal view. G, view of the apical part showing the ridges of the protoconch (arrow). H, broken specimen showing internal view of the columellar area. I &amp; J, apical view of the protoconch; arrow indicates the embryonic shell. I, detail of the embryonic shell." figureDoi="http://doi.org/10.5281/zenodo.5428909" httpUri="https://zenodo.org/record/5428909/files/figure.png" pageId="4" pageNumber="451">Fig. 4G</figureCitation>
) clearly indicating its position in
<taxonomicName id="07DACFABFF8CFFCC5611FAAFA0B3FAE6" authorityName="SCHEPMAN" authorityYear="1908" box="[1282,1419,1351,1373]" class="Gastropoda" family="Neritiliidae" kingdom="Animalia" order="Cycloneritida" pageId="4" pageNumber="451" phylum="Mollusca" rank="family">Neritiliidae</taxonomicName>
.
</paragraph>
<paragraph id="C065B428FF8CFFCC50BDFA7CA03EFA17" blockId="4.[942,1286,1428,1484]" box="[942,1286,1428,1452]" pageId="4" pageNumber="451">
<taxonomicName id="07DACFABFF8CFFCC50BDFA7CA1A1FA11" authority="Lozouet, 2004" authorityName="Lozouet" authorityYear="2004" box="[942,1177,1428,1452]" class="Gastropoda" family="Neritiliidae" genus="Bourdieria" kingdom="Animalia" order="Cycloneritida" pageId="4" pageNumber="451" phylum="Mollusca" rank="species" species="faviai" status="sp. nov.">BOURDIERIA FAVIAI</taxonomicName>
<emphasis id="F2AE683AFF8CFFCC57B1FA7FA03EFA17" bold="true" box="[1186,1286,1428,1452]" pageId="4" pageNumber="451">
<taxonomicNameLabel id="E99DD541FF8CFFCC57B1FA7FA03EFA17" box="[1186,1286,1428,1452]" pageId="4" pageNumber="451" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="C065B428FF8CFFCC5722FA5CA1BBFA77" blockId="4.[942,1286,1428,1484]" box="[1073,1155,1460,1484]" pageId="4" pageNumber="451">
(
<figureCitation id="58E1A8ADFF8CFFCC572AFA5FA143FA77" box="[1081,1147,1460,1484]" captionStart="Figure 4" captionStartId="6.[142,220,1705,1724]" captionTargetBox="[195,1368,200,1672]" captionTargetId="figure-138@6.[195,1368,200,1672]" captionTargetPageId="6" captionText="Figure 4. Bourdieria faviai sp. nov. from Peyrère (Upper Oligocene). A, broken specimen (apical whorls removed) showing (arrowed) the ridge inside the aperture (MNHN-PL1634E). B &amp; C, juvenile specimen of 0.8 adult whorl (MNHN- PL1634C). B, apical view of the protoconch showing the position of three weak ridges (arrowed). D-F, views of the holotype (MNHN-PL1634A); D, apertural view, E, right lateral view, F, dorsal view. G, view of the apical part showing the ridges of the protoconch (arrow). H, broken specimen showing internal view of the columellar area. I &amp; J, apical view of the protoconch; arrow indicates the embryonic shell. I, detail of the embryonic shell." figureDoi="http://doi.org/10.5281/zenodo.5428909" httpUri="https://zenodo.org/record/5428909/files/figure.png" pageId="4" pageNumber="451">FIG. 4</figureCitation>
)
</paragraph>
<paragraph id="C065B428FF8CFFCC5035FA35A6BDF994" blockId="4.[806,1422,1500,1645]" pageId="4" pageNumber="451">
1917
<taxonomicName id="07DACFABFF8CFFCC5076FA35A1C1FA49" authority="Recluz, 1850" authorityName="Recluz" authorityYear="1850" box="[869,1273,1500,1523]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritida" pageId="4" pageNumber="451" phylum="Mollusca" rank="species" species="basteroti" subGenus="Pila">
<emphasis id="F2AE683AFF8CFFCC5076FA35A696FA49" box="[869,942,1501,1522]" italics="true" pageId="4" pageNumber="451">Nerita</emphasis>
(
<emphasis id="F2AE683AFF8CFFCC50ADFA35A6D5FA49" box="[958,1005,1501,1522]" italics="true" pageId="4" pageNumber="451">Pila</emphasis>
)
<emphasis id="F2AE683AFF8CFFCC50EDFA35A159FA49" box="[1022,1121,1501,1522]" italics="true" pageId="4" pageNumber="451">basteroti</emphasis>
Recluz, 1850
</taxonomicName>
– Cossmann &amp; Peyrot, n∞144, pl. 10, figs 38, 39 [not Recluz, 1850].
</paragraph>
</subSubSection>
<subSubSection id="88C0E7A3FF8CFFCF5035F9D1A482F8CA" lastPageId="7" lastPageNumber="454" pageId="4" pageNumber="451" type="description">
<paragraph id="C065B428FF8CFFCC5035F9D1A046F9D6" blockId="4.[806,1422,1500,1645]" pageId="4" pageNumber="451">
1974
<taxonomicName id="07DACFABFF8CFFCC507AF9D1A1C3F9F5" authority="d'Orbigny, 1852" authorityName="d'Orbigny" authorityYear="1852" box="[873,1275,1592,1615]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritida" pageId="4" pageNumber="451" phylum="Mollusca" rank="species" species="subcaronis">
<emphasis id="F2AE683AFF8CFFCC507AF9D1A100F9F5" box="[873,1080,1593,1614]" italics="true" pageId="4" pageNumber="451">Nerita subcaronis</emphasis>
d’Orbigny, 1852
</taxonomicName>
– Magne &amp; Vergneau-Saubade: 5 (pars) [not d’Orbigny, 1852].
</paragraph>
<paragraph id="C065B428FF8CFFCC5035F96CA10FF902" blockId="4.[806,1421,1668,1721]" pageId="4" pageNumber="451">
<emphasis id="F2AE683AFF8CFFCC5035F96CA69CF922" box="[806,932,1668,1689]" italics="true" pageId="4" pageNumber="451">Etymology:</emphasis>
Dedicated to Raymond Favia for his assistance in field collection.
</paragraph>
<paragraph id="C065B428FF8CFFCC5035F939A1AFF8BE" blockId="4.[806,1422,1745,1797]" pageId="4" pageNumber="451">
<emphasis id="F2AE683AFF8CFFCC5035F939A6D3F95D" box="[806,1003,1745,1766]" italics="true" pageId="4" pageNumber="451">Type specimens:</emphasis>
<materialsCitation id="70B2BE75FF8CFFCC5715F939A1AFF8BE" pageId="4" pageNumber="451" specimenCode="MNHN-PL1634" specimenCount="4" typeStatus="holotype">
<typeStatus id="1F610A8AFF8CFFCC5715F939A154F95C" box="[1030,1132,1745,1767]" pageId="4" pageNumber="451" type="holotype">Holotype</typeStatus>
(
<specimenCode id="907C1C53FF8CFFCC579CF939A07EF95C" box="[1167,1350,1745,1767]" collectionCode="MNHN-PL" pageId="4" pageNumber="451">MNHN-PL1634</specimenCode>
A),
<specimenCount id="D6DC7FA1FF8CFFCC5693F939A6AFF8BE" pageId="4" pageNumber="451" type="generic" typeStatus="paratypes">4 paratypes</specimenCount>
(
<specimenCode id="907C1C53FF8CFFCC50B4F907A158F8BE" box="[935,1120,1775,1797]" collectionCode="MNHN-PL" pageId="4" pageNumber="451">MNHN-PL1634</specimenCode>
B-E).
</materialsCitation>
</paragraph>
<paragraph id="C065B428FF8CFFCC5035F8F6A694F8CB" blockId="4.[806,1422,1821,1904]" pageId="4" pageNumber="451">
<materialsCitation id="70B2BE75FF8CFFCC5035F8F6A690F8CB" country="France" county="Upper Oligocene" location="Landes" municipality="Peyrehorade" pageId="4" pageNumber="451" specimenCount="1" typeStatus="holotype">
<emphasis id="F2AE683AFF8CFFCC5035F8F6A177F889" box="[806,1103,1821,1843]" italics="true" pageId="4" pageNumber="451">
<typeStatus id="1F610A8AFF8CFFCC5035F8F6A663F888" box="[806,859,1822,1843]" pageId="4" pageNumber="451">Type</typeStatus>
locality and horizon:
</emphasis>
<collectingCountry id="B8CDF4B8FF8CFFCC574EF8F6A196F888" box="[1117,1198,1822,1843]" name="France" pageId="4" pageNumber="451">France</collectingCountry>
,
<location id="C505E2F3FF8CFFCC57D2F8F5A02EF888" LSID="urn:lsid:plazi:treatment:4873053EFF8CFFCF50DCFD49A482F8CA:C505E2F3FF8CFFCC57D2F8F5A02EF888" box="[1217,1302,1821,1843]" country="France" county="Upper Oligocene" municipality="Peyrehorade" name="Landes" pageId="4" pageNumber="451">Landes</location>
,
<collectingMunicipality id="20012E52FF8CFFCC5639F8F5A661F8E9" pageId="4" pageNumber="451">Peyrehorade</collectingMunicipality>
(Peyrère),
<collectingCounty id="2904CCA4FF8CFFCC5710F8D4A1E1F8E9" box="[1027,1241,1852,1874]" pageId="4" pageNumber="451">Upper Oligocene</collectingCounty>
, marls with
<taxonomicName id="07DACFABFF8CFFCC5035F8B3A690F8CB" authorityName="H. Douville" authorityYear="1911" box="[806,936,1883,1904]" class="Globothalamea" family="Lepidocyclinidae" genus="Eulepidina" kingdom="Chromista" order="Rotaliida" pageId="4" pageNumber="451" phylum="Foraminifera" rank="genus">
<emphasis id="F2AE683AFF8CFFCC5035F8B3A690F8CB" box="[806,936,1883,1904]" italics="true" pageId="4" pageNumber="451">Eulepidina</emphasis>
</taxonomicName>
</materialsCitation>
<emphasis id="F2AE683AFF8CFFCC50BBF8B3A694F8CB" box="[936,940,1883,1904]" italics="true" pageId="4" pageNumber="451">.</emphasis>
</paragraph>
<caption id="94A5E4A0FF8DFFCD5388F90DA165F8D5" ID-DOI="http://doi.org/10.5281/zenodo.5428907" ID-Zenodo-Dep="5428907" httpUri="https://zenodo.org/record/5428907/files/figure.png" pageId="5" pageNumber="452" startId="5.[155,233,1765,1784]" targetBox="[187,1403,197,1733]" targetPageId="5">
<paragraph id="C065B428FF8DFFCD5388F90DA165F8D5" blockId="5.[155,1434,1765,1902]" pageId="5" pageNumber="452">
<emphasis id="F2AE683AFF8DFFCD5388F90DA43AF943" bold="true" box="[155,258,1765,1784]" pageId="5" pageNumber="452">Figure 3.</emphasis>
A- J,
<taxonomicName id="07DACFABFF8DFFCD522CF90DA67EF943" authority="(Cossmann &amp; Peyrot, 1917)" baseAuthorityName="Cossmann &amp; Peyrot" baseAuthorityYear="1917" box="[319,838,1765,1785]" class="Gastropoda" family="Neritiliidae" genus="Neritilia" kingdom="Animalia" order="Cycloneritida" pageId="5" pageNumber="452" phylum="Mollusca" rank="species" species="neritinoides">
<emphasis id="F2AE683AFF8DFFCD522CF90DA724F943" box="[319,540,1765,1784]" italics="true" pageId="5" pageNumber="452">Neritilia neritinoides</emphasis>
(
<bibRefCitation id="A44BC9D9FF8DFFCD5139F90EA607F943" author="Cossmann M &amp; Peyrot A" box="[554,831,1765,1785]" pageId="5" pageNumber="452" pagination="1 - 384" refId="ref11987" refString="Cossmann M, Peyrot A. 1917. Conchologie Neogenique de l'Aquitaine. Actes de la Societe Linneenne de Bordeaux 3 (1): 1 - 384." type="journal article" year="1917">Cossmann &amp; Peyrot, 1917</bibRefCitation>
)
</taxonomicName>
. K &amp; L,
<taxonomicName id="07DACFABFF8DFFCD50B3F90EA1FBF943" authority="(Ferussac, 1825)" baseAuthorityName="Ferussac" baseAuthorityYear="1825" box="[928,1219,1765,1784]" class="Gastropoda" family="Neritidae" genus="Vitta" kingdom="Animalia" order="Cycloneritida" pageId="5" pageNumber="452" phylum="Mollusca" rank="species" species="picta">
<emphasis id="F2AE683AFF8DFFCD50B3F90EA136F943" box="[928,1038,1766,1784]" italics="true" pageId="5" pageNumber="452">Vitta picta</emphasis>
(Férussac, 1825)
</taxonomicName>
. A- D, operculum. E, apertural view (MNHN-PL15281). F, apical view of the protoconch. G, enlarged portion of the protoconch showing minute pits (MNHN-PL4185). H, apertural view. I, right lateral view. J, dorsal view (MNHN-PL14078). K, apical view of the protoconch. L, juvenile specimen of 0.8 whorl with operculum showing a tooth (arrowed) on the inner lip (MNHN-PL15385).
<emphasis id="F2AE683AFF8DFFCD5388F8B3A445F8D6" box="[155,381,1883,1902]" italics="true" pageId="5" pageNumber="452">Sources of specimens:</emphasis>
A- E, Lucbardez; F &amp; G, Pessac; H- L, Mimbaste (all Lower Miocene).
</paragraph>
</caption>
<caption id="94A5E4A0FF8EFFCE539DF941A692F8F5" ID-DOI="http://doi.org/10.5281/zenodo.5428909" ID-Zenodo-Dep="5428909" httpUri="https://zenodo.org/record/5428909/files/figure.png" pageId="6" pageNumber="453" startId="6.[142,220,1705,1724]" targetBox="[195,1368,200,1672]" targetPageId="6">
<paragraph id="C065B428FF8EFFCE539DF941A692F8F5" blockId="6.[142,1422,1704,1871]" pageId="6" pageNumber="453">
<emphasis id="F2AE683AFF8EFFCE539DF941A711F907" bold="true" box="[142,553,1704,1724]" pageId="6" pageNumber="453">
Figure 4.
<taxonomicName id="07DACFABFF8EFFCE5211F940A4F5F900" authority="Lozouet, 2004" authorityName="Lozouet" authorityYear="2004" box="[258,461,1704,1723]" class="Gastropoda" family="Neritiliidae" genus="Bourdieria" kingdom="Animalia" order="Cycloneritida" pageId="6" pageNumber="453" phylum="Mollusca" rank="species" species="faviai" status="sp. nov.">
<emphasis id="F2AE683AFF8EFFCE5211F940A4F5F900" bold="true" box="[258,461,1704,1723]" italics="true" pageId="6" pageNumber="453">Bourdieria faviai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E99DD541FF8EFFCE52C0F941A711F907" box="[467,553,1705,1724]" pageId="6" pageNumber="453" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Peyrère (Upper Oligocene). A, broken specimen (apical whorls removed) showing (arrowed) the ridge inside the aperture (MNHN-PL1634E). B &amp; C, juvenile specimen of 0.8 adult whorl (MNHN- PL1634C). B, apical view of the protoconch showing the position of three weak ridges (arrowed). D- F, views of the holotype (MNHN-PL1634A); D, apertural view, E, right lateral view, F, dorsal view. G, view of the apical part showing the ridges of the protoconch (arrow). H, broken specimen showing internal view of the columellar area. I &amp; J, apical view of the protoconch; arrow indicates the embryonic shell. I, detail of the embryonic shell.
</paragraph>
</caption>
<paragraph id="C065B428FF8FFFCF5388FF2CA42AFEAC" blockId="7.[155,770,196,279]" pageId="7" pageNumber="454">
<emphasis id="F2AE683AFF8FFFCF5388FF2CA4F9FF62" box="[155,449,196,217]" italics="true" pageId="7" pageNumber="454">Other material examined:</emphasis>
<materialsCitation id="70B2BE75FF8FFFCF52DDFF2CA437FEAC" collectionCode="PL" country="France" county="Peyrehorade" location="Upper Oligocene" municipality="Landes" pageId="7" pageNumber="454" specimenCode="PL1634" specimenCount="50">
<location id="C505E2F3FF8FFFCF52DDFF2CA7A0FF61" LSID="urn:lsid:plazi:treatment:4873053EFF8CFFCF50DCFD49A482F8CA:C505E2F3FF8FFFCF52DDFF2CA7A0FF61" box="[462,664,196,218]" country="France" county="Peyrehorade" municipality="Landes" name="Upper Oligocene" pageId="7" pageNumber="454">Upper Oligocene</location>
,
<collectingCountry id="B8CDF4B8FF8FFFCF51BEFF2CA7C7FF62" box="[685,767,196,217]" name="France" pageId="7" pageNumber="454">France</collectingCountry>
,
<collectingMunicipality id="20012E52FF8FFFCF5388FF0BA5C8FF42" box="[155,240,227,249]" pageId="7" pageNumber="454">Landes</collectingMunicipality>
,
<collectingCounty id="2904CCA4FF8FFFCF5219FF0BA4A1FF42" box="[266,409,227,249]" pageId="7" pageNumber="454">Peyrehorade</collectingCounty>
(Peyrère),
<specimenCount id="D6DC7FA1FF8FFFCF5126FF0AA7B0FF43" box="[565,648,226,248]" pageId="7" pageNumber="454" type="generic">50 ex.</specimenCount>
(MNHN-
<specimenCode id="907C1C53FF8FFFCF5388FEE9A5CFFEAC" box="[155,247,257,279]" collectionCode="PL" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="7" pageNumber="454" type="Museum">PL1634</specimenCode>
F)
</materialsCitation>
.
</paragraph>
<paragraph id="C065B428FF8FFFCF5388FEDDA5CFFED2" blockId="7.[155,770,309,361]" pageId="7" pageNumber="454">
<emphasis id="F2AE683AFF8FFFCF5388FEDDA407FEF1" box="[155,319,309,330]" italics="true" pageId="7" pageNumber="454">Measurements</emphasis>
(
<typeStatus id="1F610A8AFF8FFFCF525CFEDDA48CFEF0" box="[335,436,309,331]" pageId="7" pageNumber="454" type="holotype">holotype</typeStatus>
): Height 3.0 mm; max. width 4.0 mm.
</paragraph>
<paragraph id="C065B428FF8FFFCF5388FE6FA404FB20" blockId="7.[155,771,391,1179]" pageId="7" pageNumber="454">
<emphasis id="F2AE683AFF8FFFCF5388FE6FA41BFE27" box="[155,291,391,412]" italics="true" pageId="7" pageNumber="454">Description:</emphasis>
The shell is small, very solid, obliquely ovate in shape, with a very flat spire. The teleoconch consists of more than 2.5 rounded whorls, increasing very rapidly, with a depressed suture line that is difficult to observe due to the development of spiral cords. The globular protoconch (poor state of preservation in available material) is partially covered by the first teleoconch whorl; the boundary between protoconch and teleoconch is very clearly differentiated. Three or four spiral ridges are present close to the apertural line; the suture separating embryonic and larval shells is not clearly visible on the available specimens. The teleoconch is sculptured with 14–15 spiral cords and axial growth lines. The semicircular aperture is highly prosocline (about 49∞ relative to the shell axis) with a thick septum (inner lip callus) and a very blunt outer line; the arcuate septum is slightly convex and smooth. The columellar area bears 5–6 blunt teeth deeply situated inside the aperture, the adapical three being smaller. The blunt outer lip is thickened by a well-developed inductura, bearing four regularly spaced strong teeth. Inside the aperture there is a relatively long ridge near the base probably representing the innermost limit of the opercular retraction.
</paragraph>
<paragraph id="C065B428FF8FFFCF5388FB50A482F8CA" blockId="7.[155,771,1208,1905]" pageId="7" pageNumber="454">
<emphasis id="F2AE683AFF8FFFCF5388FB50A4E1FB76" box="[155,473,1208,1229]" italics="true" pageId="7" pageNumber="454">
Remarks:
<taxonomicName id="07DACFABFF8FFFCF5200FB50A4E1FB76" authorityName="Lozouet" authorityYear="2004" box="[275,473,1208,1229]" class="Gastropoda" family="Neritiliidae" genus="Bourdieria" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="454" phylum="Mollusca" rank="species" species="faviai">Bourdieria faviai</taxonomicName>
</emphasis>
has been variously interpreted.
<bibRefCitation id="A44BC9D9FF8FFFCF53E0FB3FA713FB56" author="Cossmann M &amp; Peyrot A" box="[243,555,1239,1261]" pageId="7" pageNumber="454" pagination="1 - 384" refId="ref11987" refString="Cossmann M, Peyrot A. 1917. Conchologie Neogenique de l'Aquitaine. Actes de la Societe Linneenne de Bordeaux 3 (1): 1 - 384." type="journal article" year="1917">Cossmann &amp; Peyrot (1917)</bibRefCitation>
incorrectly identified it as a young specimen of one of the most common neritids (
<taxonomicName id="07DACFABFF8FFFCF5292FAFDA778FA92" authorityName="Recluz" authorityYear="1850" box="[385,576,1300,1322]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="454" phylum="Mollusca" rank="species" species="basteroti">
<emphasis id="F2AE683AFF8FFFCF5292FAFDA778FA92" box="[385,576,1300,1322]" italics="true" pageId="7" pageNumber="454">Nerita basteroti</emphasis>
</taxonomicName>
) of the Lower Miocene.
<bibRefCitation id="A44BC9D9FF8FFFCF5219FADBA794FAF2" author="Magne A &amp; Vergneau-Saubade AM" box="[266,684,1331,1353]" pageId="7" pageNumber="454" refId="ref13368" refString="Magne A, Vergneau-Saubade AM. 1974. Les Neritidae tertiaires de l'Aquitaine occidentale. Bulletin de la Societe de Borda." type="book" year="1974">Magne &amp; Vergneau-Saubade (1974)</bibRefCitation>
identified young specimens of
<taxonomicName id="07DACFABFF8FFFCF52CDFABAA46AFA3E" authority="sensu Cossmann &amp; Peyrot" authorityName="Cossmann &amp; Peyrot" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="454" phylum="Mollusca" rank="species" species="basteroti">
<emphasis id="F2AE683AFF8FFFCF52CDFABAA78AFADC" box="[478,690,1361,1383]" italics="true" pageId="7" pageNumber="454">N. basteroti sensu</emphasis>
Cossmann &amp; Peyrot
</taxonomicName>
with
<taxonomicName id="07DACFABFF8FFFCF528CFA99A7C6FA3D" authority="d'Orbigny, 1852" authorityName="d'Orbigny" authorityYear="1852" box="[415,766,1392,1414]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="454" phylum="Mollusca" rank="species" species="subcaronis">
<emphasis id="F2AE683AFF8FFFCF528CFA99A700FA3E" box="[415,568,1392,1414]" italics="true" pageId="7" pageNumber="454">N. subcaronis</emphasis>
d’Orbigny, 1852
</taxonomicName>
, a problematic Lower Miocene species initially described by
<bibRefCitation id="A44BC9D9FF8FFFCF525FFA45A724FA78" author="Grateloup S de" box="[332,540,1453,1475]" pageId="7" pageNumber="454" refId="ref12220" refString="Grateloup S de. 1847. Conchyliologie Fossile des Terrains Tertiaires du Bassin de l'Adour (Environ de Dax). Tome I. Univalves. Atlas. Bordeaux: Lafargue." type="book" year="1847">Grateloup (1847)</bibRefCitation>
under the name
<taxonomicName id="07DACFABFF8FFFCF5388FA25A435FA59" baseAuthorityName="Brongniart" baseAuthorityYear="1823" box="[155,269,1485,1506]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="454" phylum="Mollusca" rank="species" species="caronis">
<emphasis id="F2AE683AFF8FFFCF5388FA25A435FA59" box="[155,269,1485,1506]" italics="true" pageId="7" pageNumber="454">N. caronis</emphasis>
</taxonomicName>
. Despite several visits to the collections of the University of Bordeaux-Talence, where the Grateloup collection is housed, I have been unable to locate the figured and only known specimen of
<taxonomicName id="07DACFABFF8FFFCF5388F9AFA40CF9E7" authorityName="d'Orbigny" authorityYear="1852" box="[155,308,1607,1628]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="454" phylum="Mollusca" rank="species" species="subcaronis">
<emphasis id="F2AE683AFF8FFFCF5388F9AFA40CF9E7" box="[155,308,1607,1628]" italics="true" pageId="7" pageNumber="454">N. subcaronis</emphasis>
</taxonomicName>
. According to the figures of
<bibRefCitation id="A44BC9D9FF8FFFCF519CF9AFA5E5F9C1" author="Grateloup S de" pageId="7" pageNumber="454" refId="ref12220" refString="Grateloup S de. 1847. Conchyliologie Fossile des Terrains Tertiaires du Bassin de l'Adour (Environ de Dax). Tome I. Univalves. Atlas. Bordeaux: Lafargue." type="book" year="1847">Grateloup (1847</bibRefCitation>
: pl. 5, fig. 45) and
<bibRefCitation id="A44BC9D9FF8FFFCF52C4F98EA41BF921" author="Magne A &amp; Vergneau-Saubade AM" pageId="7" pageNumber="454" refId="ref13368" refString="Magne A, Vergneau-Saubade AM. 1974. Les Neritidae tertiaires de l'Aquitaine occidentale. Bulletin de la Societe de Borda." type="book" year="1974">Magne &amp; Vergneau-Saubade (1974)</bibRefCitation>
, the septal lip of
<taxonomicName id="07DACFABFF8FFFCF52FCF96DA783F922" authorityName="d'Orbigny" authorityYear="1852" box="[495,699,1668,1690]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="454" phylum="Mollusca" rank="species" species="subcaronis">
<emphasis id="F2AE683AFF8FFFCF52FCF96DA783F922" box="[495,699,1668,1690]" italics="true" pageId="7" pageNumber="454">Nerita subcaronis</emphasis>
</taxonomicName>
bears many small denticles which
<taxonomicName id="07DACFABFF8FFFCF52FDF94BA776F903" authorityName="Lozouet" authorityYear="2004" box="[494,590,1699,1720]" class="Gastropoda" family="Neritiliidae" genus="Bourdieria" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="454" phylum="Mollusca" rank="species" species="faviai">
<emphasis id="F2AE683AFF8FFFCF52FDF94BA776F903" box="[494,590,1699,1720]" italics="true" pageId="7" pageNumber="454">B. faviai</emphasis>
</taxonomicName>
lacks.
<taxonomicName id="07DACFABFF8FFFCF51B0F94BA63BF903" authorityName="Lozouet" authorityYear="2004" box="[675,771,1699,1720]" class="Gastropoda" family="Neritiliidae" genus="Bourdieria" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="454" phylum="Mollusca" rank="species" species="faviai">
<emphasis id="F2AE683AFF8FFFCF51B0F94BA63BF903" box="[675,771,1699,1720]" italics="true" pageId="7" pageNumber="454">B. faviai</emphasis>
</taxonomicName>
is therefore easily differentiated from the
<taxonomicName id="07DACFABFF8FFFCF5187F929A639F96C" authorityName="Rafinesque" authorityYear="1815" box="[660,769,1729,1751]" class="Gastropoda" family="Neritidae" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="454" phylum="Mollusca" rank="family">Neritidae</taxonomicName>
by its small size, the heavy, smooth, convex callus, the large regular teeth on the outer lip and the tilted protoconch typical of the
<taxonomicName id="07DACFABFF8FFFCF511DF8F5A7ADF888" authorityName="SCHEPMAN" authorityYear="1908" box="[526,661,1821,1843]" class="Gastropoda" family="Neritiliidae" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="454" phylum="Mollusca" rank="family">Neritiliidae</taxonomicName>
.
<taxonomicName id="07DACFABFF8FFFCF51B0F8F6A63BF889" authorityName="Lozouet" authorityYear="2004" box="[675,771,1821,1843]" class="Gastropoda" family="Neritiliidae" genus="Bourdieria" kingdom="Animalia" order="Cycloneritida" pageId="7" pageNumber="454" phylum="Mollusca" rank="species" species="faviai">
<emphasis id="F2AE683AFF8FFFCF51B0F8F6A63BF889" box="[675,771,1821,1843]" italics="true" pageId="7" pageNumber="454">B. faviai</emphasis>
</taxonomicName>
occurs only in the Saubrigues palaeocanyon and is only common in one site.
</paragraph>
</subSubSection>
</treatment>
</document>
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