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<mods:title id="F5E9C703B0920D652EE882FB66475908">Taxonomy and distribution of African chiggers (Acariformes, Trombiculidae)</mods:title>
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<heading id="9B33BD29FFACFFB28B79FC90DB68FCB9" box="[552,1035,838,865]" centered="true" fontSize="11" level="2" pageId="54" pageNumber="55" reason="2">
<taxonomicName id="07C471C6FFACFFB28B79FC90DB68FCB9" authority="(Sambon, 1928)" baseAuthorityName="Sambon" baseAuthorityYear="1928" box="[552,1035,838,865]" class="Arachnida" family="Trombiculidae" genus="Endotrombicula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="pillersi">
<emphasis id="F2B0D657FFACFFB28B79FC90DC24FCB8" bold="true" box="[552,839,838,865]" italics="true" pageId="54" pageNumber="55">Endotrombicula pillersi</emphasis>
(
<bibRefCitation id="A45577B4FFACFFB28A06FC90DB60FCB9" author="Sambon L. W." box="[855,1027,838,864]" pageId="54" pageNumber="55" pagination="67 - 132" refId="ref65761" refString="Sambon L. W. 1928. The parasitic acariens of animals and the part they play in the causation of the eruptive fevers and other diseases of man. Annals of Tropical Medicine and Parasitology 22: 67 - 132. https: // doi. org / 10.1080 / 00034983.1928.11684565" type="journal article" year="1928">Sambon, 1928</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFFACFFB289ECFC5EDA14FB89" pageId="54" pageNumber="55" type="reference_group">
<paragraph id="C07B0A45FFACFFB289ECFC5EDD90FC7B" blockId="54.[189,755,904,931]" box="[189,755,904,931]" pageId="54" pageNumber="55">
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<treatmentCitation id="41652C54FFACFFB289ECFC5EDDC0FC7B" author="Sambon L. W." box="[189,675,904,931]" page="122" pageId="54" pageNumber="55" year="1928">
<taxonomicName id="07C471C6FFACFFB289ECFC5EDDC0FC7B" authority="Sambon, 1928: 122" authorityName="Sambon" authorityPageNumber="122" authorityYear="1928" box="[189,675,904,931]" class="Arachnida" family="Trombiculidae" genus="Schoengastia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="pillersi">
<emphasis id="F2B0D657FFACFFB289ECFC5EDED7FC7B" box="[189,436,904,930]" italics="true" pageId="54" pageNumber="55">Schoengastia pillersi</emphasis>
<bibRefCitation id="A45577B4FFACFFB288EAFC5EDDC0FC7B" author="Sambon L. W." box="[443,675,904,931]" pageId="54" pageNumber="55" pagination="67 - 132" refId="ref65761" refString="Sambon L. W. 1928. The parasitic acariens of animals and the part they play in the causation of the eruptive fevers and other diseases of man. Annals of Tropical Medicine and Parasitology 22: 67 - 132. https: // doi. org / 10.1080 / 00034983.1928.11684565" type="journal article" year="1928">Sambon, 1928: 122</bibRefCitation>
</taxonomicName>
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, fig. 9.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C07B0A45FFACFFB289ECFC1CDA14FC3C" blockId="54.[189,1399,970,1176]" box="[189,1399,970,997]" pageId="54" pageNumber="55">
<treatmentCitationGroup id="E0D42D6BFFACFFB289ECFC1CDA14FC3C" box="[189,1399,970,997]" pageId="54" pageNumber="55">
<taxonomicName id="07C471C6FFACFFB289ECFC1CDED1FC3D" authorityName="Sambon" authorityYear="1928" box="[189,434,970,996]" class="Arachnida" family="Trombiculidae" genus="Schoengastia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="pillersi">
<emphasis id="F2B0D657FFACFFB289ECFC1CDED1FC3D" box="[189,434,970,996]" italics="true" pageId="54" pageNumber="55">Schoengastia pillersi</emphasis>
</taxonomicName>
<treatmentCitation id="41652C54FFACFFB2889CFC1CDDFEFC3D" author="Radford C. D." box="[461,669,970,997]" page="67" pageId="54" pageNumber="55" year="1942">
<bibRefCitation id="A45577B4FFACFFB2889CFC1CDDFEFC3D" author="Radford C. D." box="[461,669,970,997]" pageId="54" pageNumber="55" pagination="55 - 81" refId="ref65343" refString="Radford C. D. 1942. The larval Trombiculinae (Acarina: Trombidiidae) with descriptions of twelve new species. Parasitology 34 (1): 55 - 81. https: // doi. org / 10.1017 / S 0031182000015997" type="journal article" year="1942">Radford 1942: 67</bibRefCitation>
</treatmentCitation>
, fig. 51. —
<treatmentCitation id="41652C54FFACFFB28A70FC1CDB0CFC3D" author="Thor S. &amp; Willmann C." box="[801,1135,970,997]" page="306" pageId="54" pageNumber="55" year="1947">
<bibRefCitation id="A45577B4FFACFFB28A70FC1CDB0CFC3D" author="Thor S. &amp; Willmann C." box="[801,1135,970,997]" pageId="54" pageNumber="55" pagination="71" refId="ref66977" refString="Thor S. &amp; Willmann C. 1947. Acarina. In: Schulze F. E., Kukenthal W. &amp; Heider K. (eds) Das Tierreich 71. Walter de Gruyter &amp; Co., Berlin." type="book chapter" year="1947">Thor &amp; Willmann 1947: 306</bibRefCitation>
</treatmentCitation>
, fig. 365. —
<bibRefCitation id="A45577B4FFACFFB28C52FC1DDA14FC3C" author="Lawrence R. F." box="[1283,1399,971,997]" pageId="54" pageNumber="55" pagination="405 - 486" refId="ref64030" refString="Lawrence R. F. 1949. The larval trombiculid mites of South African vertebrates. Annals of the Natal Museum 11 (3): 405 - 486." type="journal article" year="1949">Lawrence</bibRefCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="C07B0A45FFACFFB289BBFC38DA14FB89" blockId="54.[189,1399,970,1176]" pageId="54" pageNumber="55">
1949: 463.
<taxonomicName id="07C471C6FFACFFB289ECFBC4DDCEFBF5" baseAuthorityName="Sambon" baseAuthorityYear="1928" box="[189,685,1042,1068]" class="Arachnida" family="Trombiculidae" genus="Endotrombicula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="pillersi" subGenus="Endotrombicula">
<emphasis id="F2B0D657FFACFFB289ECFBC4DDCEFBF5" box="[189,685,1042,1068]" italics="true" pageId="54" pageNumber="55">Endotrombicula (Endotrombicula) pillersi</emphasis>
</taxonomicName>
<bibRefCitation id="A45577B4FFACFFB28B9BFBC4DB6CFBF5" author="Wharton G. W. &amp; Fuller H. S." box="[714,1039,1042,1069]" pageId="54" pageNumber="55" pagination="1 - 185" refId="ref69481" refString="Wharton G. W. &amp; Fuller H. S. 1952. A manual of the chiggers. Memoirs of the Entomological Society of Washington 4: 1 - 185." type="journal article" year="1952">Wharton &amp; Fuller 1952: 72</bibRefCitation>
.
<taxonomicName id="07C471C6FFACFFB289ECFBE3DDECFB96" authorityName="Sambon" authorityYear="1928" box="[189,655,1077,1103]" class="Arachnida" family="Trombiculidae" genus="Schoengastia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="pillersi" subGenus="Endotrombicula">
<emphasis id="F2B0D657FFACFFB289ECFBE3DDECFB96" box="[189,655,1077,1103]" italics="true" pageId="54" pageNumber="55">Schoengastia (Endotrombicula) pillersi</emphasis>
</taxonomicName>
<bibRefCitation id="A45577B4FFACFFB28BFFFBE3DCC8FB89" author="Taufflieb R." box="[686,939,1077,1104]" pageId="54" pageNumber="55" pagination="224 - 237" refId="ref66356" refString="Taufflieb R. 1960 b. Etude d'une collection de Trombiculidae d'Afrique Occidentale. Acarologia 2 (2): 224 - 237." type="journal article" year="1960">Taufflieb 1960b: 224</bibRefCitation>
. —
<bibRefCitation id="A45577B4FFACFFB28A8CFBE0DBCDFB89" author="Zumpt F." box="[989,1198,1077,1104]" pageId="54" pageNumber="55" pagination="1 - 457" refId="ref69765" refString="Zumpt F. (ed.) 1961. The arthropod parasites of vertebrates in Africa south of the Sahara (Ethiopian Region). Vol. 1: Chelicerata. Publications of the South African Institute for Medical Research 11 (50): 1 - 457." type="journal article" year="1961">Zumpt 1961: 159</bibRefCitation>
. — Vercammen-
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFFACFFB289BBFB8FDA15FB4E" pageId="54" pageNumber="55" type="description">
<paragraph id="C07B0A45FFACFFB289BBFB8FDA15FB4E" blockId="54.[189,1399,970,1176]" pageId="54" pageNumber="55">
Grandjean 1965c: 83.
<taxonomicName id="07C471C6FFACFFB289ECFBABDEB7FB4E" baseAuthorityName="Sambon" baseAuthorityYear="1928" box="[189,468,1149,1175]" class="Arachnida" family="Trombiculidae" genus="Endotrombicula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="pillersi">
<emphasis id="F2B0D657FFACFFB289ECFBABDEB7FB4E" box="[189,468,1149,1175]" italics="true" pageId="54" pageNumber="55">Endotrombicula pillersi</emphasis>
</taxonomicName>
<bibRefCitation id="A45577B4FFACFFB288A1FBABDC08FB41" author="Spieler M. &amp; Linsenmair K. E." box="[496,875,1149,1176]" pageId="54" pageNumber="55" pagination="152 - 161" refId="ref65871" refString="Spieler M. &amp; Linsenmair K. E. 1999. The larval mite Endotrombicula pillersi as a species-specific parasite of a West African savannah frog (Phrynobatrachus francisci). The American Midland Naturalist 142 (1): 152 - 161. https: // doi. org / 10.1674 / 0003 - 0031 (1999) 142 [0152: TLMEPA] 2.0. CO; 2" type="journal article" year="1999">Spieler &amp; Linsenmair 1999: 152</bibRefCitation>
, fig. 1. —
<bibRefCitation id="A45577B4FFACFFB28AB3FBABDA4AFB4E" author="Wohltmann A. &amp; du Preez L. &amp; Rodel M. O. &amp; Kohler J. &amp; Vences M." box="[994,1321,1149,1176]" pageId="54" pageNumber="55" pagination="225 - 235" refId="ref69625" refString="Wohltmann A., du Preez L., Rodel M. O., Kohler J. &amp; Vences M. 2007. Endoparasitic mites of the genus Endotrombicula Ewing, 1931 (Acari: Prostigmata: Parasitengona: Trombiculidae) from African and Madagascaran anurans, with description of a new species. Folia Parasitologica 54: 225 - 235. https: // doi. org / 10.14411 / fp. 2007.031" type="journal article" year="2007">
Wohltmann
<emphasis id="F2B0D657FFACFFB28D21FBA8DBC9FB4E" box="[1136,1194,1149,1175]" italics="true" pageId="54" pageNumber="55">et al.</emphasis>
2007: 226
</bibRefCitation>
, fig. 1.
</paragraph>
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<typeStatus id="1F7FB4E7FFACFFB289ECFB69DE4CFB00" box="[189,303,1215,1241]" pageId="54" pageNumber="55" type="holotype">Holotype</typeStatus>
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<paragraph id="C07B0A45FFACFFB289ECFB39DD28FAD0" blockId="54.[189,587,1262,1289]" box="[189,587,1262,1289]" pageId="54" pageNumber="55">
<collectionCode id="A6D59280FFACFFB289ECFB39DE76FAD0" box="[189,277,1263,1289]" pageId="54" pageNumber="55">BMNH</collectionCode>
(
<bibRefCitation id="A45577B4FFACFFB28877FB38DD58FAD0" author="Wharton G. W. &amp; Fuller H. S." box="[294,571,1262,1289]" pageId="54" pageNumber="55" pagination="1 - 185" refId="ref69481" refString="Wharton G. W. &amp; Fuller H. S. 1952. A manual of the chiggers. Memoirs of the Entomological Society of Washington 4: 1 - 185." type="journal article" year="1952">Wharton &amp; Fuller 1952</bibRefCitation>
).
</paragraph>
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<heading id="9B33BD29FFACFFB289ECFAE6DEE9FA93" bold="true" box="[189,394,1328,1354]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="F2B0D657FFACFFB289ECFAE6DEE9FA93" bold="true" box="[189,394,1328,1354]" pageId="54" pageNumber="55">Material revised</emphasis>
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<paragraph id="C07B0A45FFACFFB289ECFA89DEF8FA18" blockId="54.[189,1399,1375,1473]" pageId="54" pageNumber="55">
One specimen (obviously
<typeStatus id="1F7FB4E7FFACFFB28B50FA89DD09FAA3" box="[513,618,1375,1402]" pageId="54" pageNumber="55" type="holotype">holotype</typeStatus>
) from
<collectionCode id="A6D59280FFACFFB28B94FAB6DC43FAA3" box="[709,800,1376,1402]" pageId="54" pageNumber="55">BMNH</collectionCode>
, without accession number, ex
<taxonomicName id="07C471C6FFACFFB28DFAFA89DE57FA44" baseAuthorityName="Zumpt" baseAuthorityYear="1961" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" kingdom="Animalia" order="Anura" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="natalensis">
<emphasis id="F2B0D657FFACFFB28DFAFA89DE57FA44" italics="true" pageId="54" pageNumber="55">Phrynobatrachus natalensis</emphasis>
</taxonomicName>
, White
<collectingRegion id="0200C4A7FFACFFB288CAFA55DEBBFA44" box="[411,472,1411,1437]" country="Ghana" name="Volta" pageId="54" pageNumber="55">Volta</collectingRegion>
valley,
<collectingRegion id="0200C4A7FFACFFB28B6DFA55DDF9FA44" box="[572,666,1411,1437]" country="Ghana" name="Ashanti" pageId="54" pageNumber="55">Ashanti</collectingRegion>
, labeled “
<taxonomicName id="07C471C6FFACFFB28A48FA55DBB1FA44" authority="Sambon, 1923" authorityName="Sambon" authorityYear="1923" box="[793,1234,1411,1437]" class="Arachnida" family="Trombiculidae" genus="Schoengastia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="pillersi">
<emphasis id="F2B0D657FFACFFB28A48FA55DB76FA44" box="[793,1045,1411,1437]" italics="true" pageId="54" pageNumber="55">Schoengastia pillersi</emphasis>
Sambon, 1923
</taxonomicName>
, larva”, slide marked by red ink.
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFFACFFB289ECFA3EDCE1F98F" pageId="54" pageNumber="55" type="distribution">
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<emphasis id="F2B0D657FFACFFB289ECFA3EDE35F9DB" bold="true" box="[189,342,1512,1538]" pageId="54" pageNumber="55">Distribution</emphasis>
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</paragraph>
<paragraph id="C07B0A45FFACFFB289ECF9CEDCE1F98F" blockId="54.[189,1398,1560,1622]" pageId="54" pageNumber="55">
<collectingCountry id="B8D34AD5FFACFFB289ECF9CEDE69F9EB" box="[189,266,1560,1586]" name="Ghana" pageId="54" pageNumber="55">Ghana</collectingCountry>
(Kumasi, Black
<collectingRegion id="0200C4A7FFACFFB2889DF9CEDD6DF9EA" box="[460,526,1560,1587]" country="Ghana" name="Volta" pageId="54" pageNumber="55">Volta</collectingRegion>
),
<collectingCountry id="B8D34AD5FFACFFB28B70F9CEDDCAF9EB" box="[545,681,1560,1586]" name="Ivory Coast" pageId="54" pageNumber="55">Ivory Coast</collectingCountry>
(
<collectingRegion id="0200C4A7FFACFFB28BE6F9CEDC6DF9EB" box="[695,782,1560,1586]" country="Ivory Coast" name="Comoe" pageId="54" pageNumber="55">Comoé</collectingRegion>
National Park Research Station, Lamto, Taï National Park),
<collectingCountry id="B8D34AD5FFACFFB2885AF9EDDE37F98C" box="[267,340,1595,1621]" name="Benin" pageId="54" pageNumber="55">Benin</collectingCountry>
,
<collectingCountry id="B8D34AD5FFACFFB28831F9EDDED6F98C" box="[352,437,1595,1621]" name="Guinea-Bissau" pageId="54" pageNumber="55">Guinea</collectingCountry>
(Pic de Fon, Diécké Classified Forest).
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFFACFFB289ECF9A8DA41F933" pageId="54" pageNumber="55" type="biology_ecology">
<paragraph id="C07B0A45FFACFFB289ECF9A8DE60F941" blockId="54.[189,259,1662,1688]" box="[189,259,1662,1688]" pageId="54" pageNumber="55">
<heading id="9B33BD29FFACFFB289ECF9A8DE60F941" bold="true" box="[189,259,1662,1688]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="F2B0D657FFACFFB289ECF9A8DE60F941" bold="true" box="[189,259,1662,1688]" pageId="54" pageNumber="55">Hosts</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FFACFFB289ECF97BDA41F933" blockId="54.[189,1398,1708,1771]" pageId="54" pageNumber="55">
<taxonomicName id="07C471C6FFACFFB289ECF97BDE8DF91E" box="[189,494,1709,1735]" class="Amphibia" family="Bufonidae" genus="Amietophrynus" kingdom="Animalia" order="Anura" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="maculatus">
<emphasis id="F2B0D657FFACFFB289ECF97BDE8DF91E" box="[189,494,1709,1735]" italics="true" pageId="54" pageNumber="55">Amietophrynus maculatus</emphasis>
</taxonomicName>
,
<taxonomicName id="07C471C6FFACFFB288AAF97BDD8CF91F" authorityName="Boulenger" authorityYear="1905" box="[507,751,1709,1735]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="natator">
<emphasis id="F2B0D657FFACFFB288AAF97BDD8CF91F" box="[507,751,1709,1735]" italics="true" pageId="54" pageNumber="55">Petropedetes natator</emphasis>
</taxonomicName>
,
<taxonomicName id="07C471C6FFACFFB28BACF97BDB25F91E" baseAuthorityName="Zumpt" baseAuthorityYear="1961" box="[765,1094,1709,1735]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" kingdom="Animalia" order="Anura" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="natalensis">
<emphasis id="F2B0D657FFACFFB28BACF97BDB25F91E" box="[765,1094,1709,1735]" italics="true" pageId="54" pageNumber="55">Phrynobatrachus natalensis</emphasis>
</taxonomicName>
(
<typeStatus id="1F7FB4E7FFACFFB28D04F97BDBEAF91E" box="[1109,1161,1709,1735]" pageId="54" pageNumber="55">type</typeStatus>
host),
<taxonomicName id="07C471C6FFACFFB28D86F97BDA0CF91E" baseAuthorityName="Cope" baseAuthorityYear="1867" box="[1239,1391,1709,1735]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" kingdom="Animalia" order="Anura" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="acridoides">
<emphasis id="F2B0D657FFACFFB28D86F97BDA0CF91E" box="[1239,1391,1709,1735]" italics="true" pageId="54" pageNumber="55">P. acridoides</emphasis>
</taxonomicName>
,
<taxonomicName id="07C471C6FFACFFB289ECF907DE7EF933" authorityName="Parker" authorityYear="1936" box="[189,285,1744,1770]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" kingdom="Animalia" order="Anura" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="alleni">
<emphasis id="F2B0D657FFACFFB289ECF907DE7EF933" box="[189,285,1744,1770]" italics="true" pageId="54" pageNumber="55">P. alleni</emphasis>
</taxonomicName>
,
<taxonomicName id="07C471C6FFACFFB2887AF907DEA7F933" baseAuthorityName="Peters" baseAuthorityYear="1863" box="[299,452,1744,1770]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" kingdom="Animalia" order="Anura" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="calcaratus">
<emphasis id="F2B0D657FFACFFB2887AF907DEA7F933" box="[299,452,1744,1770]" italics="true" pageId="54" pageNumber="55">P. calcaratus</emphasis>
</taxonomicName>
,
<taxonomicName id="07C471C6FFACFFB28882F907DD35F933" authorityName="Boulenger" authorityYear="1912" box="[467,598,1744,1770]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" kingdom="Animalia" order="Anura" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="francisci">
<emphasis id="F2B0D657FFACFFB28882F907DD35F933" box="[467,598,1744,1770]" italics="true" pageId="54" pageNumber="55">P. francisci</emphasis>
</taxonomicName>
,
<taxonomicName id="07C471C6FFACFFB28B34F907DD80F933" authorityName="Ahl" authorityYear="1924" box="[613,739,1744,1770]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" kingdom="Animalia" order="Anura" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="latifrons">
<emphasis id="F2B0D657FFACFFB28B34F907DD80F933" box="[613,739,1744,1770]" italics="true" pageId="54" pageNumber="55">P. latifrons</emphasis>
</taxonomicName>
,
<taxonomicName id="07C471C6FFACFFB28BA3F907DCC3F933" authorityName="Rodel &amp; Ernst" authorityYear="2002" box="[754,928,1744,1770]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" kingdom="Animalia" order="Anura" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="phyllophilus">
<emphasis id="F2B0D657FFACFFB28BA3F907DCC3F933" box="[754,928,1744,1770]" italics="true" pageId="54" pageNumber="55">P. phyllophilus</emphasis>
</taxonomicName>
,
<taxonomicName id="07C471C6FFACFFB28AFEF907DB4AF933" box="[943,1065,1744,1770]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" kingdom="Animalia" order="Anura" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="plicatus">
<emphasis id="F2B0D657FFACFFB28AFEF907DB4AF933" box="[943,1065,1744,1770]" italics="true" pageId="54" pageNumber="55">P. plicatus</emphasis>
</taxonomicName>
,
<taxonomicName id="07C471C6FFACFFB28D66F907DBF6F933" baseAuthorityName="Chabanaud" baseAuthorityYear="1921" box="[1079,1173,1744,1770]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" kingdom="Animalia" order="Anura" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="tokba">
<emphasis id="F2B0D657FFACFFB28D66F907DBF6F933" box="[1079,1173,1744,1770]" italics="true" pageId="54" pageNumber="55">P. tokba</emphasis>
</taxonomicName>
,
<emphasis id="F2B0D657FFACFFB28DF5F907DA41F933" box="[1188,1314,1744,1770]" italics="true" pageId="54" pageNumber="55">
<taxonomicName id="07C471C6FFACFFB28DF5F907DA7DF933" authorityName="Guibe" authorityYear="1959" box="[1188,1310,1744,1770]" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" kingdom="Animalia" order="Anura" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="villiersi">P. villiersi</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>