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<document id="D8EC71B861A7E0A7B34F7D146CD0B6C0" ID-DOI="10.5852/ejt.2018.395" ID-GBIF-Dataset="10dbb315-5662-4069-a1ae-9692fdf99499" ID-ISSN="2118-9773" ID-Zenodo-Dep="3829724" ID-ZooBank="A244F97F-8CEB-40CE-A182-7E5440337CAA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="juliana" checkinTime="1589571932603" checkinUser="valdenar" docAuthor="Stekolnikov, Alexandr A." docDate="2018" docId="486DBB53FFA2FFBD88ABFA8FDEFFFE85" docLanguage="en" docName="ejt-395_stekolnikov.pdf.imf" docOrigin="European Journal of Taxonomy 395" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Helenicula pilosa" docType="treatment" docVersion="19" lastPageNumber="58" masterDocId="B454C32BFF9AFF848951FFD6DF63FFD9" masterDocTitle="Taxonomy and distribution of African chiggers (Acariformes, Trombiculidae)" masterLastPageNumber="233" masterPageNumber="1" pageNumber="57" updateTime="1740581643606" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5BE17398ED30DA9E96E6FC44E62DD0FA">Taxonomy and distribution of African chiggers (Acariformes, Trombiculidae)</mods:title>
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<mods:namePart id="9A2912003C6ACD93E1497FAA67A5A6C1">Stekolnikov, Alexandr A.</mods:namePart>
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<mods:affiliation id="500D45FFABE9830FD943F118467926A7">Zoological Institute, Russian Academy of Sciences, Universitetskaya embankment 1, St. Petersburg 199034, Russia.</mods:affiliation>
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<mods:title id="51D37A17AADC63599567C921E0735D3A">European Journal of Taxonomy</mods:title>
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<mods:date id="38F7B5C7ABC9F25F5AFE25798F4A2CFE">2018</mods:date>
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<treatment id="486DBB53FFA2FFBD88ABFA8FDEFFFE85" ID-DOI="http://doi.org/10.5281/zenodo.3849288" ID-GBIF-Taxon="164160737" ID-Zenodo-Dep="3849288" LSID="urn:lsid:plazi:treatment:486DBB53FFA2FFBD88ABFA8FDEFFFE85" httpUri="http://treatment.plazi.org/id/486DBB53FFA2FFBD88ABFA8FDEFFFE85" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57" scope_family="Trombiculidae" scope_subFamily="Trombiculinae" scope_superOrder="Acariformes" scope_tribe="Schoengastiini">
<subSubSection id="88DE59CEFFA2FFBC88ABFA8FDB5AFAAD" box="[506,1081,1369,1396]" pageId="56" pageNumber="57" type="nomenclature">
<paragraph id="C07B0A45FFA2FFBC88ABFA8FDB5AFAAD" blockId="56.[506,1081,1369,1396]" box="[506,1081,1369,1396]" pageId="56" pageNumber="57">
<heading id="9B33BD29FFA2FFBC88ABFA8FDB5AFAAD" box="[506,1081,1369,1396]" centered="true" fontSize="11" level="2" pageId="56" pageNumber="57" reason="2">
<taxonomicName id="07C471C6FFA2FFBC88ABFA8FDB5AFAAD" ID-CoL="8ZJHV" authority="(Abonnenc &amp; Taufflieb, 1957)" baseAuthorityName="Abonnenc &amp; Taufflieb" baseAuthorityYear="1957" box="[506,1081,1369,1396]" class="Arachnida" family="Trombiculidae" genus="Helenicula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="F2B0D657FFA2FFBC88ABFA8FDDACFAAD" bold="true" box="[506,719,1369,1396]" italics="true" pageId="56" pageNumber="57">Helenicula pilosa</emphasis>
(
<bibRefCitation id="A45577B4FFA2FFBC8B8FFA8FDB52FAAD" author="Abonnenc &amp; Taufflieb" box="[734,1073,1369,1396]" firstAuthor="Abonnenc" pageId="56" pageNumber="57" pagination="80 - 85" refId="ref66702" refString="Taufflieb R. &amp; Abonnenc E. 1957. Sur deux Trombiculides larvaires parasites du rat de Gambie: Euschoengastia mailloti n. sp. et Elianella anomaluri Vercammen-Grandjean, 1956. Bulletin de la Societe de pathologie exotique 50 (1): 80 - 85." type="journal article" year="1957">Abonnenc &amp; Taufflieb, 1957</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFFA2FFBC89ECFA4ADC85F9C6" pageId="56" pageNumber="57" type="reference_group">
<paragraph id="C07B0A45FFA2FFBC89ECFA4ADB17FA6E" blockId="56.[189,1140,1436,1463]" box="[189,1140,1436,1463]" pageId="56" pageNumber="57">
<treatmentCitationGroup id="E0D42D6BFFA2FFBC89ECFA4ADB17FA6E" box="[189,1140,1436,1463]" pageId="56" pageNumber="57">
<treatmentCitation id="41652C54FFA2FFBC89ECFA4ADB60FA6E" author="Abonnenc E. &amp; Taufflieb R." box="[189,1027,1436,1463]" page="556" pageId="56" pageNumber="57" year="1957">
<taxonomicName id="07C471C6FFA2FFBC89ECFA4ADB60FA6E" authority="Abonnenc &amp; Taufflieb, 1957 a: 556" authorityName="Abonnenc &amp; Taufflieb" authorityPageNumber="556" authorityYear="1957" box="[189,1027,1436,1463]" class="Arachnida" family="Trombiculidae" genus="Euschoengastia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="pilosa" subGenus="Helenicula">
<emphasis id="F2B0D657FFA2FFBC89ECFA4ADD01FA6F" box="[189,610,1436,1462]" italics="true" pageId="56" pageNumber="57">Euschoengastia (Helenicula) pilosa</emphasis>
<bibRefCitation id="A45577B4FFA2FFBC8B36FA4ADB60FA6E" author="Abonnenc &amp; Taufflieb" box="[615,1027,1436,1463]" firstAuthor="Abonnenc" page="556" pageId="56" pageNumber="57" pagination="556 - 567" refId="ref59818" refString="Abonnenc E. &amp; Taufflieb R. 1957 a. Deux Trombiculides nouveaux de la region du Tchad, Euschoengastia pilosa n. sp. et Trombicula youhensis n. sp. Presence de Schoengastia radfordi Jadin et Vercammen- Grandjean, 1954. Bulletin de la Societe de pathologie exotique 50 (4): 556 - 567." type="journal article" year="1957" yearSuffix="a">Abonnenc &amp; Taufflieb, 1957a: 556</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 23.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C07B0A45FFA2FFBC89ECFA36DB44FA22" blockId="56.[189,1063,1504,1568]" box="[189,1063,1504,1531]" pageId="56" pageNumber="57">
<treatmentCitationGroup id="E0D42D6BFFA2FFBC89ECFA36DB44FA22" box="[189,1063,1504,1531]" pageId="56" pageNumber="57">
<taxonomicName id="07C471C6FFA2FFBC89ECFA36DEEEFA23" ID-CoL="8ZJHV" baseAuthorityName="Abonnenc &amp; Taufflieb" baseAuthorityYear="1957" box="[189,397,1504,1530]" class="Arachnida" family="Trombiculidae" genus="Helenicula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="F2B0D657FFA2FFBC89ECFA36DEEEFA23" box="[189,397,1504,1530]" italics="true" pageId="56" pageNumber="57">Helenicula pilosa</emphasis>
</taxonomicName>
<treatmentCitation id="41652C54FFA2FFBC88FBFA36DD1AFA22" author="Zumpt F." box="[426,633,1504,1531]" page="153" pageId="56" pageNumber="57" year="1961">
<bibRefCitation id="A45577B4FFA2FFBC88FBFA36DD1AFA22" author="Zumpt" box="[426,633,1504,1531]" firstAuthor="Zumpt" page="153" pageId="56" pageNumber="57" pagination="1 - 457" refId="ref69765" refString="Zumpt F. (ed.) 1961. The arthropod parasites of vertebrates in Africa south of the Sahara (Ethiopian Region). Vol. 1: Chelicerata. Publications of the South African Institute for Medical Research 11 (50): 1 - 457." type="journal article" year="1961">Zumpt 1961: 153</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="41652C54FFA2FFBC8BF8FA36DB40FA22" author="Nadchatram M. &amp; Traub R." box="[681,1059,1504,1531]" page="591" pageId="56" pageNumber="57" year="1971">
<bibRefCitation id="A45577B4FFA2FFBC8BF8FA36DB40FA22" author="Nadchatram &amp; Traub" box="[681,1059,1504,1531]" firstAuthor="Nadchatram" page="591" pageId="56" pageNumber="57" pagination="562 - 597" refId="ref64896" refString="Nadchatram M. &amp; Traub R. 1971. Chiggers of the genus Helenicula of the Old World including descriptions of 9 new species (Acarina: Prostigmata, Trombiculidae). Journal of Medical Entomology 8 (5): 562 - 597. https: // doi. org / 10.1093 / jmedent / 8.5.562" type="journal article" year="1971">Nadchatram &amp; Traub 1971: 591</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C07B0A45FFA2FFBC89ECF9D3DC85F9C6" blockId="56.[189,1063,1504,1568]" box="[189,998,1541,1568]" pageId="56" pageNumber="57">
<treatmentCitationGroup id="E0D42D6BFFA2FFBC89ECF9D3DC85F9C6" box="[189,998,1541,1568]" pageId="56" pageNumber="57">
<taxonomicName id="07C471C6FFA2FFBC89ECF9D3DD4AF9C6" ID-CoL="8ZJHV" baseAuthorityName="Abonnenc &amp; Taufflieb" baseAuthorityYear="1957" box="[189,553,1541,1567]" class="Arachnida" family="Trombiculidae" genus="Helenicula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="pilosa" subGenus="Helenicula">
<emphasis id="F2B0D657FFA2FFBC89ECF9D3DD4AF9C6" box="[189,553,1541,1567]" italics="true" pageId="56" pageNumber="57">Helenicula (Helenicula) pilosa</emphasis>
</taxonomicName>
<treatmentCitation id="41652C54FFA2FFBC8B14F9D3DC81F9C6" author="Vercammen-Grandjean P. H." box="[581,994,1541,1568]" page="112" pageId="56" pageNumber="57" year="1965">
<bibRefCitation id="A45577B4FFA2FFBC8B14F9D3DC81F9C6" author="Vercammen-Grandjean" box="[581,994,1541,1568]" firstAuthor="Vercammen-Grandjean" page="112" pageId="56" pageNumber="57" refId="ref68052" refString="Vercammen-Grandjean P. H. 1965 c. Trombiculinae of the World. Synopsis with Generic, Subgeneric, and Group Diagnoses (Acarina, Trombiculidae). George Williams Hooper Foundation, San Francisco." type="book" year="1965" yearSuffix="c">Vercammen-Grandjean 1965c: 112</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFFA2FFBC89ECF99FDE6AF8F4" pageId="56" pageNumber="57" type="materials_examined">
<materialsCitation id="70AC0018FFA2FFBC89ECF99FDE7DF94D" box="[189,303,1609,1684]" collectionCode="MNHN" pageId="56" pageNumber="57" specimenCount="1" typeStatus="holotype">
<paragraph id="C07B0A45FFA2FFBC89ECF99FDE4CF9BA" blockId="56.[189,303,1609,1635]" box="[189,303,1609,1635]" pageId="56" pageNumber="57">
<heading id="9B33BD29FFA2FFBC89ECF99FDE4CF9BA" bold="true" box="[189,303,1609,1635]" fontSize="11" level="3" pageId="56" pageNumber="57" reason="3">
<emphasis id="F2B0D657FFA2FFBC89ECF99FDE4CF9BA" bold="true" box="[189,303,1609,1635]" pageId="56" pageNumber="57">
<typeStatus id="1F7FB4E7FFA2FFBC89ECF99FDE4CF9BA" box="[189,303,1609,1635]" pageId="56" pageNumber="57" type="holotype">Holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FFA2FFBC89ECF9ACDE7DF94D" blockId="56.[189,286,1658,1684]" box="[189,286,1658,1684]" pageId="56" pageNumber="57">
<collectionCode id="A6D59280FFA2FFBC89ECF9ACDE78F94D" box="[189,283,1658,1684]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="56" pageNumber="57">MNHN</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="C07B0A45FFA2FFBC89ECF96ADEC9F90F" blockId="56.[189,426,1724,1750]" box="[189,426,1724,1750]" pageId="56" pageNumber="57">
<heading id="9B33BD29FFA2FFBC89ECF96ADEC9F90F" bold="true" box="[189,426,1724,1750]" fontSize="11" level="3" pageId="56" pageNumber="57" reason="3">
<emphasis id="F2B0D657FFA2FFBC89ECF96ADEC9F90F" bold="true" box="[189,426,1724,1750]" pageId="56" pageNumber="57">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FFA2FFBC89ECF93BDE6AF8F4" blockId="56.[189,1398,1773,1837]" pageId="56" pageNumber="57">
<materialsCitation id="70AC0018FFA2FFBC89ECF93BDE6AF8F4" collectionCode="RMCA" determinerName="Vercammen-Grandjean" pageId="56" pageNumber="57" specimenCode="86176" specimenCount="1" typeStatus="paratype">
<specimenCount id="D6C2C1CCFFA2FFBC89ECF93BDF8DF8DE" box="[189,238,1773,1799]" pageId="56" pageNumber="57" type="generic">One</specimenCount>
<typeStatus id="1F7FB4E7FFA2FFBC89A4F938DE39F8D1" box="[245,346,1774,1800]" pageId="56" pageNumber="57">paratype</typeStatus>
from
<collectionCode id="A6D59280FFA2FFBC88F3F93BDE94F8DE" box="[418,503,1773,1799]" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="56" pageNumber="57" type="Museum">RMCA</collectionCode>
(No.
<specimenCode id="9062A23EFFA2FFBC8B6BF93BDDE5F8DE" box="[570,646,1773,1799]" pageId="56" pageNumber="57">86176</specimenCode>
, in same box as
<taxonomicName id="07C471C6FFA2FFBC8A16F93BDF9FF8F4" authority="Vercammen-Grandjean, 1958" authorityName="Vercammen-Grandjean" authorityYear="1958" class="Arachnida" family="Trombiculidae" genus="Schoutedenichia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="F2B0D657FFA2FFBC8A16F93BDB3BF8DE" box="[839,1112,1773,1799]" italics="true" pageId="56" pageNumber="57">Schoutedenichia pilosa</emphasis>
<bibRefCitation id="A45577B4FFA2FFBC8D0EF93BDF9FF8F4" author="Vercammen-Grandjean" firstAuthor="Vercammen-Grandjean" pageId="56" pageNumber="57" pagination="1 - 103" refId="ref67569" refString="Vercammen-Grandjean P. H. 1958 a. Revision du genre Schoutedenichia Jad. et Verc. Annales du Musee royal du Congo belge, serie in- 8 65: 1 - 103." type="journal article" year="1958">
<determinerName id="35A7E681FFA2FFBC8D0EF93BDA11F8DE" box="[1119,1394,1773,1799]" pageId="56" pageNumber="57">Vercammen-Grandjean</determinerName>
, 1958
</bibRefCitation>
</taxonomicName>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFFA2FFBC89ECF880DE6AF832" pageId="56" pageNumber="57" type="distribution">
<paragraph id="C07B0A45FFA2FFBC89ECF880DE35F8A9" blockId="56.[189,342,1878,1904]" box="[189,342,1878,1904]" pageId="56" pageNumber="57">
<heading id="9B33BD29FFA2FFBC89ECF880DE35F8A9" bold="true" box="[189,342,1878,1904]" fontSize="11" level="3" pageId="56" pageNumber="57" reason="3">
<emphasis id="F2B0D657FFA2FFBC89ECF880DE35F8A9" bold="true" box="[189,342,1878,1904]" pageId="56" pageNumber="57">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FFA2FFBC89ECF850DE6AF832" blockId="56.[189,1399,1926,2027]" pageId="56" pageNumber="57">
<collectingCountry id="B8D34AD5FFA2FFBC89ECF850DF98F879" box="[189,251,1926,1952]" name="Chad" pageId="56" pageNumber="57">Chad</collectingCountry>
(Léré). This species was also recorded in
<collectingCountry id="B8D34AD5FFA2FFBC8A63F850DC1BF879" box="[818,888,1926,1952]" name="Nepal" pageId="56" pageNumber="57">Nepal</collectingCountry>
from
<taxonomicName id="07C471C6FFA2FFBC8A83F851DA14F879" authority="Temminck, 1844" authorityName="Temminck" authorityYear="1844" box="[978,1399,1926,1952]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="tanezumi">
<emphasis id="F2B0D657FFA2FFBC8A83F851DBF4F879" box="[978,1175,1927,1952]" italics="true" pageId="56" pageNumber="57">Rattus tanezumi</emphasis>
Temminck, 1844
</taxonomicName>
(
<bibRefCitation id="A45577B4FFA2FFBC8996F87DDD65F81F" author="Nadchatram &amp; Traub" box="[199,518,1963,1990]" firstAuthor="Nadchatram" pageId="56" pageNumber="57" pagination="562 - 597" refId="ref64896" refString="Nadchatram M. &amp; Traub R. 1971. Chiggers of the genus Helenicula of the Old World including descriptions of 9 new species (Acarina: Prostigmata, Trombiculidae). Journal of Medical Entomology 8 (5): 562 - 597. https: // doi. org / 10.1093 / jmedent / 8.5.562" type="journal article" year="1971">Nadchatram &amp; Traub 1971</bibRefCitation>
) and in
<collectingCountry id="B8D34AD5FFA2FFBC8B3AF87DDDB0F81C" box="[619,723,1963,1989]" name="Thailand" pageId="56" pageNumber="57">Thailand</collectingCountry>
from
<taxonomicName id="07C471C6FFA2FFBC8A4CF87DDE67F832" authority="(Bechstein, 1800) (Chaisiri et al. 2016)" authorityName="Chaisiri et al." authorityYear="2016" baseAuthorityName="Bechstein" baseAuthorityYear="1800" class="Mammalia" family="Muridae" genus="Bandicota" kingdom="Animalia" order="Rodentia" pageId="56" pageNumber="57" phylum="Chordata" rank="species" species="indica">
<emphasis id="F2B0D657FFA2FFBC8A4CF87DDC84F81C" box="[797,999,1963,1989]" italics="true" pageId="56" pageNumber="57">Bandicota indica</emphasis>
(Bechstein, 1800) (
<bibRefCitation id="A45577B4FFA2FFBC8D84F87DDF9BF832" author="Chaisiri" etAl="et al." firstAuthor="Chaisiri" pageId="56" pageNumber="57" pagination="321 - 342" refId="ref61362" refString="Chaisiri K., Stekolnikov A. A., Makepeace B. L. &amp; Morand S. 2016. A revised checklist of chigger mites (Acari: Trombiculidae) from Thailand, with the description of three new species. Journal of Medical Entomology 53 (2): 321 - 342. https: // doi. org / 10.1093 / jme / tjv 244" type="journal article" year="2016">
Chaisiri
<emphasis id="F2B0D657FFA2FFBC8C6BF87ADA14F81C" box="[1338,1399,1963,1989]" italics="true" pageId="56" pageNumber="57">et al.</emphasis>
2016
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFFA3FFBD89ECFEDDDEFFFE85" box="[189,412,267,348]" pageId="57" pageNumber="58" type="biology_ecology">
<paragraph id="C07B0A45FFA3FFBD89ECFEDDDF9BFEFC" blockId="57.[189,248,267,293]" box="[189,248,267,293]" pageId="57" pageNumber="58">
<heading id="9B33BD29FFA3FFBD89ECFEDDDF9BFEFC" bold="true" box="[189,248,267,293]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
<emphasis id="F2B0D657FFA3FFBD89ECFEDDDF9BFEFC" bold="true" box="[189,248,267,293]" pageId="57" pageNumber="58">Host</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FFA3FFBD89ECFE97DEFFFE85" blockId="57.[189,412,321,348]" box="[189,412,321,348]" pageId="57" pageNumber="58">
<taxonomicName id="07C471C6FFA3FFBD89ECFE97DEF6FE82" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[189,405,321,347]" class="Aves" family="Numididae" genus="Numida" kingdom="Animalia" order="Galliformes" pageId="57" pageNumber="58" phylum="Chordata" rank="species" species="meleagris">
<emphasis id="F2B0D657FFA3FFBD89ECFE97DEF6FE82" box="[189,405,321,347]" italics="true" pageId="57" pageNumber="58">Numida meleagris</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>