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<document id="6FABEDDEA1909A411C1001078C464620" ID-DOI="10.5852/ejt.2018.395" ID-GBIF-Dataset="10dbb315-5662-4069-a1ae-9692fdf99499" ID-ISSN="2118-9773" ID-Zenodo-Dep="3829724" ID-ZooBank="A244F97F-8CEB-40CE-A182-7E5440337CAA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="juliana" checkinTime="1589571932603" checkinUser="valdenar" docAuthor="Stekolnikov, Alexandr A." docDate="2018" docId="486DBB53FF8EFF9188E3F969DB75FAFA" docLanguage="en" docName="ejt-395_stekolnikov.pdf.imf" docOrigin="European Journal of Taxonomy 395" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Gahrliepia brennani" docType="treatment" docVersion="19" lastPageNumber="22" masterDocId="B454C32BFF9AFF848951FFD6DF63FFD9" masterDocTitle="Taxonomy and distribution of African chiggers (Acariformes, Trombiculidae)" masterLastPageNumber="233" masterPageNumber="1" pageNumber="21" updateTime="1740581643606" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="37ED03246B8CAC87DB147CDA7A9C5A7C">Taxonomy and distribution of African chiggers (Acariformes, Trombiculidae)</mods:title>
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<mods:affiliation id="3A3B438D3A18F4B8698F3057025F39AA">Zoological Institute, Russian Academy of Sciences, Universitetskaya embankment 1, St. Petersburg 199034, Russia.</mods:affiliation>
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<treatment id="486DBB53FF8EFF9188E3F969DB75FAFA" ID-DOI="http://doi.org/10.5281/zenodo.3849115" ID-GBIF-Taxon="164160473" ID-Zenodo-Dep="3849115" LSID="urn:lsid:plazi:treatment:486DBB53FF8EFF9188E3F969DB75FAFA" httpUri="http://treatment.plazi.org/id/486DBB53FF8EFF9188E3F969DB75FAFA" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" scope_family="Trombiculidae" scope_subFamily="Gahrliepiinae" scope_superOrder="Acariformes">
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<paragraph id="C07B0A45FF8EFF9088E3F969DBE2F902" blockId="20.[434,1153,1727,1755]" box="[434,1153,1727,1755]" pageId="20" pageNumber="21">
<heading id="9B33BD29FF8EFF9088E3F969DBE2F902" box="[434,1153,1727,1755]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="21" reason="2">
<taxonomicName id="07C471C6FF8EFF9088E3F969DBE2F902" ID-CoL="8ZKK5" authority="(Jadin &amp; Vercammen-Grandjean, 1952)" baseAuthorityName="Jadin &amp; Vercammen-Grandjean" baseAuthorityYear="1952" box="[434,1153,1727,1755]" class="Arachnida" family="Trombiculidae" genus="Gahrliepia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="brennani">
<emphasis id="F2B0D657FF8EFF9088E3F969DDC8F903" bold="true" box="[434,683,1727,1754]" italics="true" pageId="20" pageNumber="21">Gahrliepia brennani</emphasis>
(
<bibRefCitation id="A45577B4FF8EFF908BEBF916DB1AF902" author="Jadin &amp; Vercammen-Grandjean" box="[698,1145,1728,1755]" firstAuthor="Jadin" pageId="20" pageNumber="21" pagination="605 - 641" refId="ref63049" refString="Jadin J. B. &amp; Vercammen-Grandjean P. H. 1952. Douze nouvelles especes de larves de Trombiculides (Acariens Prostigmates) du Ruanda-Urundi (Congo Belge). Annales de la Societe belge de Medecine tropicale 32: 605 - 641." type="journal article" year="1952">Jadin &amp; Vercammen-Grandjean, 1952</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="C07B0A45FF8EFF9089ECF8D7DC8CF8C5" blockId="20.[189,1399,1793,1891]" box="[189,1007,1793,1820]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="E0D42D6BFF8EFF9089ECF8D7DC8CF8C5" box="[189,1007,1793,1820]" pageId="20" pageNumber="21">
<treatmentCitation id="41652C54FF8EFF9089ECF8D7DCF9F8C2" author="Jadin J. B. &amp; Vercammen-Grandjean P. H." box="[189,922,1793,1820]" page="640" pageId="20" pageNumber="21" year="1952">
<taxonomicName id="07C471C6FF8EFF9089ECF8D7DCF9F8C2" authority="Jadin &amp; Vercammen-Grandjean, 1952: 640" authorityName="Jadin &amp; Vercammen-Grandjean" authorityPageNumber="640" authorityYear="1952" box="[189,922,1793,1820]" class="Arachnida" family="Trombiculidae" genus="Giroudia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="brennani">
<emphasis id="F2B0D657FF8EFF9089ECF8D7DEFBF8C2" box="[189,408,1793,1819]" italics="true" pageId="20" pageNumber="21">Giroudia brennani</emphasis>
<bibRefCitation id="A45577B4FF8EFF9088CEF8D7DCF9F8C2" author="Jadin &amp; Vercammen-Grandjean" box="[415,922,1793,1820]" firstAuthor="Jadin" page="640" pageId="20" pageNumber="21" pagination="605 - 641" refId="ref63049" refString="Jadin J. B. &amp; Vercammen-Grandjean P. H. 1952. Douze nouvelles especes de larves de Trombiculides (Acariens Prostigmates) du Ruanda-Urundi (Congo Belge). Annales de la Societe belge de Medecine tropicale 32: 605 - 641." type="journal article" year="1952">Jadin &amp; Vercammen-Grandjean, 1952: 640</bibRefCitation>
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, pl. 14.
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</paragraph>
<paragraph id="C07B0A45FF8EFF9089ECF8F3DB15F8BB" blockId="20.[189,1399,1793,1891]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="E0D42D6BFF8EFF9089ECF8F3DB15F8BB" pageId="20" pageNumber="21">
<treatmentCitation id="41652C54FF8EFF9089ECF8F3DA53F8E6" author="Zumpt F." box="[189,1328,1829,1855]" page="174" pageId="20" pageNumber="21" year="1961">
<taxonomicName id="07C471C6FF8EFF9089ECF8F3DA53F8E6" authority="Audy, Lawrence &amp; Vercammen-Grandjean in Zumpt, 1961: 174" authorityName="Audy, Lawrence &amp; Vercammen-Grandjean in Zumpt" authorityPageNumber="174" authorityYear="1961" box="[189,1328,1829,1855]" class="Arachnida" family="Trombiculidae" genus="Gahrliepia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="traubi" subGenus="Giroudia">
<emphasis id="F2B0D657FF8EFF9089ECF8F3DD78F8E6" box="[189,539,1829,1855]" italics="true" pageId="20" pageNumber="21">Gahrliepia (Giroudia) traubi</emphasis>
Audy, Lawrence &amp; Vercammen-Grandjean in
<bibRefCitation id="A45577B4FF8EFF908D04F8F3DA53F8E6" author="Zumpt" box="[1109,1328,1829,1855]" firstAuthor="Zumpt" page="174" pageId="20" pageNumber="21" pagination="1 - 457" refId="ref69765" refString="Zumpt F. (ed.) 1961. The arthropod parasites of vertebrates in Africa south of the Sahara (Ethiopian Region). Vol. 1: Chelicerata. Publications of the South African Institute for Medical Research 11 (50): 1 - 457." type="journal article" year="1961">Zumpt, 1961: 174</bibRefCitation>
</taxonomicName>
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(new name for
<taxonomicName id="07C471C6FF8EFF90880AF89EDC99F8BA" authority="Jadin &amp; Vercammen-Grandjean, 1952" authorityName="Jadin &amp; Vercammen-Grandjean" authorityYear="1952" box="[347,1018,1864,1891]" class="Arachnida" family="Trombiculidae" genus="Giroudia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="brennani">
<emphasis id="F2B0D657FF8EFF90880AF89EDD55F8BB" box="[347,566,1864,1890]" italics="true" pageId="20" pageNumber="21">Giroudia brennani</emphasis>
<bibRefCitation id="A45577B4FF8EFF908B6CF89EDC99F8BA" author="Jadin &amp; Vercammen-Grandjean" box="[573,1018,1864,1891]" firstAuthor="Jadin" pageId="20" pageNumber="21" pagination="605 - 641" refId="ref63049" refString="Jadin J. B. &amp; Vercammen-Grandjean P. H. 1952. Douze nouvelles especes de larves de Trombiculides (Acariens Prostigmates) du Ruanda-Urundi (Congo Belge). Annales de la Societe belge de Medecine tropicale 32: 605 - 641." type="journal article" year="1952">Jadin &amp; Vercammen-Grandjean, 1952</bibRefCitation>
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)
<emphasis id="F2B0D657FF8EFF908D5BF89EDB15F8BB" bold="true" box="[1034,1142,1864,1890]" pageId="20" pageNumber="21">syn. nov.</emphasis>
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</paragraph>
<paragraph id="C07B0A45FF8EFF9089ECF85CDBEEF87D" blockId="20.[189,1171,1930,2027]" box="[189,1165,1930,1957]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="E0D42D6BFF8EFF9089ECF85CDBEEF87D" box="[189,1165,1930,1957]" pageId="20" pageNumber="21">
<taxonomicName id="07C471C6FF8EFF9089ECF85CDE9CF87D" box="[189,511,1930,1956]" class="Arachnida" family="Trombiculidae" genus="Gateria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="brennani" subGenus="Gateria">
<emphasis id="F2B0D657FF8EFF9089ECF85CDE9CF87D" box="[189,511,1930,1956]" italics="true" pageId="20" pageNumber="21">Gateria (Gateria) brennani</emphasis>
</taxonomicName>
<treatmentCitation id="41652C54FF8EFF908B4AF85CDB7FF87D" author="Vercammen-Grandjean P. H. &amp; Jadin J. B." box="[539,1052,1930,1957]" page="360" pageId="20" pageNumber="21" year="1956">
<bibRefCitation id="A45577B4FF8EFF908B4AF85CDB7FF87D" author="Vercammen-Grandjean &amp; Jadin" box="[539,1052,1930,1957]" firstAuthor="Vercammen-Grandjean" page="360" pageId="20" pageNumber="21" pagination="360 - 364" refId="ref68743" refString="Vercammen-Grandjean P. H. &amp; Jadin J. B. 1956 a. Considerations diverses au sujet de Gateria (Giroudia) brennani (Jad. et Verc. 1952) et description de la nymphe. Revue de zoologie et de botanique africaines 53: 360 - 364." type="journal article" year="1956" yearSuffix="a">Vercammen-Grandjean &amp; Jadin 1956a: 360</bibRefCitation>
</treatmentCitation>
, figs 12.
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</paragraph>
<paragraph id="C07B0A45FF8EFF9089ECF878DCC4F81E" blockId="20.[189,1171,1930,2027]" box="[189,935,1965,1992]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="E0D42D6BFF8EFF9089ECF878DCC4F81E" box="[189,935,1965,1992]" pageId="20" pageNumber="21">
<taxonomicName id="07C471C6FF8EFF9089ECF878DD6DF81E" box="[189,526,1965,1992]" class="Arachnida" family="Trombiculidae" genus="Gateria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="brennani" subGenus="Giroudia">
<emphasis id="F2B0D657FF8EFF9089ECF878DD6DF81E" box="[189,526,1965,1992]" italics="true" pageId="20" pageNumber="21">Gateria (Giroudia) brennani</emphasis>
</taxonomicName>
<treatmentCitation id="41652C54FF8EFF908B7AF87BDCC0F81E" author="Taufflieb R. &amp; Mouchet J." box="[555,931,1965,1992]" page="243" pageId="20" pageNumber="21" year="1959">
<bibRefCitation id="A45577B4FF8EFF908B7AF87BDCC0F81E" author="Taufflieb &amp; Mouchet" box="[555,931,1965,1992]" firstAuthor="Taufflieb" page="243" pageId="20" pageNumber="21" pagination="228 - 245" refId="ref66791" refString="Taufflieb R. &amp; Mouchet J. 1959. Revue des Trombiculidae du Cameroun et description de huit especes et sous-especes nouvelles. Acarologia 1 (2): 228 - 245." type="journal article" year="1959">Taufflieb &amp; Mouchet 1959: 243</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="C07B0A45FF8EFF9089ECF807DBF0F832" blockId="20.[189,1171,1930,2027]" box="[189,1171,2001,2027]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="E0D42D6BFF8EFF9089ECF807DBF0F832" box="[189,1171,2001,2027]" pageId="20" pageNumber="21">
<taxonomicName id="07C471C6FF8EFF9089ECF807DD50F832" ID-CoL="8ZKK5" baseAuthorityName="Jadin &amp; Vercammen-Grandjean" baseAuthorityYear="1952" box="[189,563,2001,2027]" class="Arachnida" family="Trombiculidae" genus="Gahrliepia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="brennani" subGenus="Giroudia">
<emphasis id="F2B0D657FF8EFF9089ECF807DD50F832" box="[189,563,2001,2027]" italics="true" pageId="20" pageNumber="21">Gahrliepia (Giroudia) brennani</emphasis>
</taxonomicName>
<treatmentCitation id="41652C54FF8EFF908B00F807DBECF832" author="Kolebinova M. &amp; Vercammen-Grandjean P. H." box="[593,1167,2001,2027]" page="123" pageId="20" pageNumber="21" year="1978">
<bibRefCitation id="A45577B4FF8EFF908B00F807DBECF832" author="Kolebinova &amp; Vercammen-Grandjean" box="[593,1167,2001,2027]" firstAuthor="Kolebinova" page="123" pageId="20" pageNumber="21" pagination="101 - 131" refId="ref63661" refString="Kolebinova M. &amp; Vercammen-Grandjean P. H. 1978. Five new larval Trombiculidae (Acarina) from Liberia and one from Zaire. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg 6 (101): 101 - 131." type="journal article" year="1978">Kolebinova &amp; Vercammen-Grandjean 1978: 123</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C07B0A45FF8FFF9189ECFEDDDC21FEFF" blockId="21.[189,1138,267,329]" box="[189,834,267,294]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="E0D42D6BFF8FFF9189ECFEDDDC21FEFF" box="[189,834,267,294]" pageId="21" pageNumber="22">
<taxonomicName id="07C471C6FF8FFF9189ECFEDDDD4AFEFC" authorityName="Audy, Lawrence &amp; Vercammen-Grandjean in Zumpt" authorityYear="1961" box="[189,553,267,293]" class="Arachnida" family="Trombiculidae" genus="Gahrliepia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="traubi" subGenus="Gahrliepia">
<emphasis id="F2B0D657FF8FFF9189ECFEDDDD4AFEFC" box="[189,553,267,293]" italics="true" pageId="21" pageNumber="22">Gahrliepia (Gahrliepia) traubi</emphasis>
</taxonomicName>
<treatmentCitation id="41652C54FF8FFF918B14FEDDDC5CFEFF" author="Taufflieb R." box="[581,831,267,294]" page="515" pageId="21" pageNumber="22" year="1965">
<bibRefCitation id="A45577B4FF8FFF918B14FEDDDC5CFEFF" author="Taufflieb" box="[581,831,267,294]" firstAuthor="Taufflieb" page="515" pageId="21" pageNumber="22" pagination="510 - 522" refId="ref66541" refString="Taufflieb R. 1965 b. Le sous-genre Gahrliepia (Acarina, Trombiculidae) en Afrique Subsaharienne. Acarologia 7 (3): 510 - 522." type="journal article" year="1965" yearSuffix="b">Taufflieb 1965b: 515</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="C07B0A45FF8FFF9189ECFEF9DB11FE90" blockId="21.[189,1138,267,329]" box="[189,1138,303,329]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="E0D42D6BFF8FFF9189ECFEF9DB11FE90" box="[189,1138,303,329]" pageId="21" pageNumber="22">
<taxonomicName id="07C471C6FF8FFF9189ECFEF9DD71FE90" authorityName="Audy, Lawrence &amp; Vercammen-Grandjean in Zumpt" authorityYear="1961" box="[189,530,303,329]" class="Arachnida" family="Trombiculidae" genus="Gahrliepia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="traubi" subGenus="Giroudia">
<emphasis id="F2B0D657FF8FFF9189ECFEF9DD71FE90" box="[189,530,303,329]" italics="true" pageId="21" pageNumber="22">Gahrliepia (Giroudia) traubi</emphasis>
</taxonomicName>
<treatmentCitation id="41652C54FF8FFF918B7EFEF9DB0DFE90" author="Kolebinova M. &amp; Vercammen-Grandjean P. H." box="[559,1134,303,329]" page="126" pageId="21" pageNumber="22" year="1978">
<bibRefCitation id="A45577B4FF8FFF918B7EFEF9DB0DFE90" author="Kolebinova &amp; Vercammen-Grandjean" box="[559,1134,303,329]" firstAuthor="Kolebinova" page="126" pageId="21" pageNumber="22" pagination="101 - 131" refId="ref63661" refString="Kolebinova M. &amp; Vercammen-Grandjean P. H. 1978. Five new larval Trombiculidae (Acarina) from Liberia and one from Zaire. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg 6 (101): 101 - 131." type="journal article" year="1978">Kolebinova &amp; Vercammen-Grandjean 1978: 126</bibRefCitation>
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.
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<heading id="9B33BD29FF8FFF9189ECFEA7DE4CFE52" bold="true" box="[189,303,369,395]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="F2B0D657FF8FFF9189ECFEA7DE4CFE52" bold="true" box="[189,303,369,395]" pageId="21" pageNumber="22">
<typeStatus id="1F7FB4E7FF8FFF9189ECFEA7DE4CFE52" box="[189,303,369,395]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FF8FFF9189ECFE76DE09FE63" blockId="21.[189,362,416,442]" box="[189,362,416,442]" pageId="21" pageNumber="22">
<specimenCode id="9062A23EFF8FFF9189ECFE76DE05FE63" box="[189,358,416,442]" collectionCode="RMCA" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="21" pageNumber="22" type="Museum">RMCA 76149</specimenCode>
.
</paragraph>
</materialsCitation>
<paragraph id="C07B0A45FF8FFF9189ECFE37DEE9FE22" blockId="21.[189,394,481,507]" box="[189,394,481,507]" pageId="21" pageNumber="22">
<heading id="9B33BD29FF8FFF9189ECFE37DEE9FE22" bold="true" box="[189,394,481,507]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="F2B0D657FF8FFF9189ECFE37DEE9FE22" bold="true" box="[189,394,481,507]" pageId="21" pageNumber="22">Material revised</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FF8FFF9189ECFDC6DC38FDAB" blockId="21.[189,1398,528,626]" pageId="21" pageNumber="22">
<materialsCitation id="70AC0018FF8FFF9189ECFDC6DDC8FDF2" ID-GBIF-Occurrence="2620304439" box="[189,683,528,555]" collectionCode="RMCA" pageId="21" pageNumber="22" specimenCount="1" typeStatus="holotype">Holotype (not suitable for examination)</materialsCitation>
<materialsCitation id="70AC0018FF8FFF918BE8FDC6DC38FDAB" ID-GBIF-Occurrence="2620304439" collectionCode="RMCA" pageId="21" pageNumber="22" specimenCode="76150" specimenCount="56" typeStatus="paratype">
and
<typeStatus id="1F7FB4E7FF8FFF918BA0FDC7DC35FDF2" box="[753,854,529,555]" pageId="21" pageNumber="22">paratype</typeStatus>
(No.
<specimenCode id="9062A23EFF8FFF918AF9FDC6DC97FDF2" box="[936,1012,528,555]" pageId="21" pageNumber="22">76150</specimenCode>
, not suitable for examination),
<specimenCount id="D6C2C1CCFF8FFF9189ECFDE2DE03FD97" box="[189,352,564,590]" count="47" pageId="21" pageNumber="22" type="generic">47 specimens</specimenCount>
including
<specimenCount id="D6C2C1CCFF8FFF9188B7FDE2DD03FD97" box="[486,608,564,590]" pageId="21" pageNumber="22">9 nymphs</specimenCount>
, not designated as paratypes, and 86 more specimens labeled as “
<taxonomicName id="07C471C6FF8FFF91899BFD8EDD23FDAB" authorityName="Jadin &amp; Vercammen-Grandjean" authorityYear="1952" box="[202,576,600,626]" class="Arachnida" family="Trombiculidae" genus="Gahrliepia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="breunani" subGenus="Giroudia">
<emphasis id="F2B0D657FF8FFF91899BFD8EDD23FDAB" box="[202,576,600,626]" italics="true" pageId="21" pageNumber="22">Gahrliepia (Giroudia) breunani</emphasis>
</taxonomicName>
” [sic], all from
<collectionCode id="A6D59280FF8FFF918BADFD8EDC35FDAB" box="[764,854,600,626]" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="21" pageNumber="22" type="Museum">RMCA</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFF8FFF9189ECFD4FDCC5FD3A" box="[189,934,665,739]" pageId="21" pageNumber="22" type="distribution">
<paragraph id="C07B0A45FF8FFF9189ECFD4FDE35FD6A" blockId="21.[189,342,665,691]" box="[189,342,665,691]" pageId="21" pageNumber="22">
<heading id="9B33BD29FF8FFF9189ECFD4FDE35FD6A" bold="true" box="[189,342,665,691]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="F2B0D657FF8FFF9189ECFD4FDE35FD6A" bold="true" box="[189,342,665,691]" pageId="21" pageNumber="22">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FF8FFF9189ECFD1EDCC5FD3A" blockId="21.[189,934,712,739]" box="[189,934,712,739]" pageId="21" pageNumber="22">
<collectingCountry id="B8D34AD5FF8FFF9189ECFD1EDE7EFD3B" box="[189,285,712,738]" name="Rwanda" pageId="21" pageNumber="22">Rwanda</collectingCountry>
(Butare),
<collectingCountry id="B8D34AD5FF8FFF9188C4FD1FDD70FD3A" box="[405,531,713,739]" name="Democratic Republic of the Congo" pageId="21" pageNumber="22">DR Congo</collectingCountry>
(Beni, Bukavu, Lwiro, Kabunga).
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFF8FFF9189ECFCDDDE08FC42" pageId="21" pageNumber="22" type="biology_ecology">
<paragraph id="C07B0A45FF8FFF9189ECFCDDDE60FCFC" blockId="21.[189,259,779,805]" box="[189,259,779,805]" pageId="21" pageNumber="22">
<heading id="9B33BD29FF8FFF9189ECFCDDDE60FCFC" bold="true" box="[189,259,779,805]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="F2B0D657FF8FFF9189ECFCDDDE60FCFC" bold="true" box="[189,259,779,805]" pageId="21" pageNumber="22">Hosts</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FF8FFF9189ECFCEFDE08FC42" blockId="21.[189,1399,825,923]" pageId="21" pageNumber="22">
<taxonomicName id="07C471C6FF8FFF9189ECFCEFDEF8FC8A" baseAuthorityName="G. [Baron] Cuvier" baseAuthorityYear="1829" box="[189,411,825,851]" class="Mammalia" family="Herpestidae" genus="Atilax" kingdom="Animalia" order="Carnivora" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="paludinosus">
<emphasis id="F2B0D657FF8FFF9189ECFCEFDEF8FC8A" box="[189,411,825,851]" italics="true" pageId="21" pageNumber="22">Atilax paludinosus</emphasis>
</taxonomicName>
,
<taxonomicName id="07C471C6FF8FFF9188FDFCEFDDC1FC8A" baseAuthorityName="Zumpt" baseAuthorityYear="1961" box="[428,674,825,851]" class="Mammalia" family="Muridae" genus="Arvicanthis" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="niloticus">
<emphasis id="F2B0D657FF8FFF9188FDFCEFDDC1FC8A" box="[428,674,825,851]" italics="true" pageId="21" pageNumber="22">Arvicanthis niloticus</emphasis>
</taxonomicName>
,
<taxonomicName id="07C471C6FF8FFF918BE5FCEFDC39FC8D" box="[692,858,825,852]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="F2B0D657FF8FFF918BE5FCEFDC4EFC8A" box="[692,813,825,851]" italics="true" pageId="21" pageNumber="22">Crocidura</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="07C471C6FF8FFF918A3BFCECDB2BFC8A" baseAuthorityName="Sundevall" baseAuthorityYear="1847" box="[874,1096,826,851]" class="Mammalia" family="Muridae" genus="Dasymys" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="incomtus">
<emphasis id="F2B0D657FF8FFF918A3BFCECDB2BFC8A" box="[874,1096,826,851]" italics="true" pageId="21" pageNumber="22">Dasymys incomtus</emphasis>
</taxonomicName>
(
<typeStatus id="1F7FB4E7FF8FFF918D0BFCEFDBEDFC8A" box="[1114,1166,825,851]" pageId="21" pageNumber="22">type</typeStatus>
host),
<taxonomicName id="07C471C6FF8FFF918DB4FCEFDE0BFCAE" baseAuthorityName="Zumpt" baseAuthorityYear="1961" class="Mammalia" family="Muridae" genus="Lophuromys" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="flavopunctatus">
<emphasis id="F2B0D657FF8FFF918DB4FCEFDE0BFCAE" italics="true" pageId="21" pageNumber="22">Lophuromys flavopunctatus</emphasis>
</taxonomicName>
,
<taxonomicName id="07C471C6FF8FFF91882BFC88DD10FCAE" baseAuthorityName="Zumpt" baseAuthorityYear="1961" box="[378,627,861,887]" class="Mammalia" family="Muridae" genus="Mastomys" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="natalensis">
<emphasis id="F2B0D657FF8FFF91882BFC88DD10FCAE" box="[378,627,861,887]" italics="true" pageId="21" pageNumber="22">Mastomys natalensis</emphasis>
</taxonomicName>
,
<taxonomicName id="07C471C6FF8FFF918BD7FC88DC67FCAE" baseAuthorityName="Thomas" baseAuthorityYear="1909" box="[646,772,862,887]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="triton">
<emphasis id="F2B0D657FF8FFF918BD7FC88DC67FCAE" box="[646,772,862,887]" italics="true" pageId="21" pageNumber="22">Mus triton</emphasis>
</taxonomicName>
,
<taxonomicName id="07C471C6FF8FFF918A46FC8BDB40FCAE" baseAuthorityName="Pucheran" baseAuthorityYear="1855" box="[791,1059,861,887]" class="Mammalia" family="Muridae" genus="Oenomys" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="hypoxanthus">
<emphasis id="F2B0D657FF8FFF918A46FC8BDB40FCAE" box="[791,1059,861,887]" italics="true" pageId="21" pageNumber="22">Oenomys hypoxanthus</emphasis>
</taxonomicName>
,
<taxonomicName id="07C471C6FF8FFF918D64FC8BDB84FCAE" baseAuthorityName="Peters" baseAuthorityYear="1852" box="[1077,1255,861,887]" class="Mammalia" family="Muridae" genus="Pelomys" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="fallax">
<emphasis id="F2B0D657FF8FFF918D64FC8BDB84FCAE" box="[1077,1255,861,887]" italics="true" pageId="21" pageNumber="22">Pelomys fallax</emphasis>
</taxonomicName>
,
<emphasis id="F2B0D657FF8FFF918DA8FC8BDE08FC42" italics="true" pageId="21" pageNumber="22">
<taxonomicName id="07C471C6FF8FFF918DA8FC8BDE04FC42" baseAuthorityName="Tullberg" baseAuthorityYear="1893" class="Mammalia" family="Muridae" genus="Stochomys" kingdom="Animalia" order="Rodentia" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="longicaudatus">Stochomys longicaudatus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFF8FFF9189ECFC15DB75FAFA" pageId="21" pageNumber="22" type="discussion">
<paragraph id="C07B0A45FF8FFF9189ECFC15DE4CFC04" blockId="21.[189,303,963,989]" box="[189,303,963,989]" pageId="21" pageNumber="22">
<heading id="9B33BD29FF8FFF9189ECFC15DE4CFC04" bold="true" box="[189,303,963,989]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="F2B0D657FF8FFF9189ECFC15DE4CFC04" bold="true" box="[189,303,963,989]" pageId="21" pageNumber="22">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FF8FFF9189ECFC27DBA3FB38" blockId="21.[189,1399,1009,1250]" pageId="21" pageNumber="22">
The name
<taxonomicName id="07C471C6FF8FFF918810FC27DDFCFBD2" authorityName="Audy, Lawrence &amp; Vercammen-Grandjean in Zumpt" authorityYear="1961" box="[321,671,1009,1035]" class="Arachnida" family="Trombiculidae" genus="Gahrliepia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="traubi" subGenus="Giroudia">
<emphasis id="F2B0D657FF8FFF918810FC27DDFCFBD2" box="[321,671,1009,1035]" italics="true" pageId="21" pageNumber="22">Gahrliepia (Giroudia) traubi</emphasis>
</taxonomicName>
was proposed after 1960 to fix secondary homonymy with
<taxonomicName id="07C471C6FF8FFF9189ECFBC3DC79FBE9" authority="Womersley, 1952" authorityName="Womersley" authorityYear="1952" box="[189,794,1045,1072]" class="Arachnida" family="Trombiculidae" genus="Gahrliepia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="brennani" subGenus="Walchia">
<emphasis id="F2B0D657FF8FFF9189ECFBC3DD54FBF6" box="[189,567,1045,1071]" italics="true" pageId="21" pageNumber="22">Gahrliepia (Walchia) brennani</emphasis>
Womersley, 1952
</taxonomicName>
(
<bibRefCitation id="A45577B4FF8FFF918A63FBC3DCA9FBF6" author="Zumpt" box="[818,970,1045,1071]" firstAuthor="Zumpt" pageId="21" pageNumber="22" pagination="1 - 457" refId="ref69765" refString="Zumpt F. (ed.) 1961. The arthropod parasites of vertebrates in Africa south of the Sahara (Ethiopian Region). Vol. 1: Chelicerata. Publications of the South African Institute for Medical Research 11 (50): 1 - 457." type="journal article" year="1961">Zumpt 1961</bibRefCitation>
;
<bibRefCitation id="A45577B4FF8FFF918AB0FBC3DBC0FBE9" author="Taufflieb" box="[993,1187,1045,1072]" firstAuthor="Taufflieb" pageId="21" pageNumber="22" pagination="510 - 522" refId="ref66541" refString="Taufflieb R. 1965 b. Le sous-genre Gahrliepia (Acarina, Trombiculidae) en Afrique Subsaharienne. Acarologia 7 (3): 510 - 522." type="journal article" year="1965" yearSuffix="b">Taufflieb 1965b</bibRefCitation>
), but in modern systems the genus
<taxonomicName id="07C471C6FF8FFF9188F8FBEFDD6BFB8A" authorityName="Ewing" authorityYear="1931" box="[425,520,1081,1107]" class="Arachnida" family="Trombiculidae" genus="Walchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="F2B0D657FF8FFF9188F8FBEFDD6BFB8A" box="[425,520,1081,1107]" italics="true" pageId="21" pageNumber="22">Walchia</emphasis>
</taxonomicName>
separates from
<taxonomicName id="07C471C6FF8FFF918B85FBEFDC36FB8A" authorityName="Oudemans" authorityYear="1912" box="[724,853,1081,1107]" class="Arachnida" family="Trombiculidae" genus="Gahrliepia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="F2B0D657FF8FFF918B85FBEFDC36FB8A" box="[724,853,1081,1107]" italics="true" pageId="21" pageNumber="22">Gahrliepia</emphasis>
</taxonomicName>
(
<bibRefCitation id="A45577B4FF8FFF918A3AFBEEDB33FB8A" author="Kudryashova" box="[875,1104,1080,1107]" firstAuthor="Kudryashova" pageId="21" pageNumber="22" refId="ref63886" refString="Kudryashova N. I. 1998. Chigger Mites (Acariformes, Trombiculidae) of East Palaearctics. KMK Scientific Press, Moscow." type="book" year="1998">Kudryashova 1998</bibRefCitation>
;
<bibRefCitation id="A45577B4FF8FFF918D34FBEEDF9BFBAF" author="Fernandes &amp; Kulkarni" firstAuthor="Fernandes" pageId="21" pageNumber="22" refId="ref61861" refString="Fernandes S. J. S. &amp; Kulkarni S. M. 2003. Studies on the Trombiculid Mite Fauna of India. Records of the Zoological Survey of India, Occasional Paper 212." type="journal volume" year="2003">Fernandes &amp; Kulkarni 2003</bibRefCitation>
;
<bibRefCitation id="A45577B4FF8FFF91885AFB8ADD38FBAF" author="Stekolnikov &amp; Daniel" box="[267,603,1116,1142]" firstAuthor="Stekolnikov" pageId="21" pageNumber="22" pagination="1 - 104" refId="ref66024" refString="Stekolnikov A. A. &amp; Daniel M. 2012. Chigger mites (Acari: Trombiculidae) of Turkey. Zootaxa 3216: 1 - 104." type="journal article" year="2012">Stekolnikov &amp; Daniel 2012</bibRefCitation>
). Thus,
<taxonomicName id="07C471C6FF8FFF918B92FB8ADCF4FBAF" authorityName="Audy, Lawrence &amp; Vercammen-Grandjean in Zumpt" authorityYear="1961" box="[707,919,1116,1142]" class="Arachnida" family="Trombiculidae" genus="Gahrliepia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="traubi">
<emphasis id="F2B0D657FF8FFF918B92FB8ADCF4FBAF" box="[707,919,1116,1142]" italics="true" pageId="21" pageNumber="22">Gahrliepia traubi</emphasis>
</taxonomicName>
is synonymized here with
<taxonomicName id="07C471C6FF8FFF918DB7FB8BDDEDFB42" authority="(Jadin &amp; Vercammen-Grandjean, 1952)" baseAuthorityName="Jadin &amp; Vercammen-Grandjean" baseAuthorityYear="1952" class="Arachnida" family="Trombiculidae" genus="Gahrliepia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="brennani">
<emphasis id="F2B0D657FF8FFF918DB7FB8BDA14FBAF" box="[1254,1399,1116,1143]" italics="true" pageId="21" pageNumber="22">G. brennani</emphasis>
(
<bibRefCitation id="A45577B4FF8FFF918994FB56DDE5FB42" author="Jadin &amp; Vercammen-Grandjean" box="[197,646,1152,1179]" firstAuthor="Jadin" pageId="21" pageNumber="22" pagination="605 - 641" refId="ref63049" refString="Jadin J. B. &amp; Vercammen-Grandjean P. H. 1952. Douze nouvelles especes de larves de Trombiculides (Acariens Prostigmates) du Ruanda-Urundi (Congo Belge). Annales de la Societe belge de Medecine tropicale 32: 605 - 641." type="journal article" year="1952">Jadin &amp; Vercammen-Grandjean, 1952</bibRefCitation>
)
</taxonomicName>
according to ICZN Code (
<collectingRegion id="0200C4A7FF8FFF918A9BFB56DC95FB43" box="[970,1014,1152,1178]" country="Djibouti" name="Arta" pageId="21" pageNumber="22">Art</collectingRegion>
. 59.4). Both names,
<taxonomicName id="07C471C6FF8FFF918DB8FB56DDEDFB67" authority="(Jadin &amp; Vercammen-Grandjean, 1952)" baseAuthorityName="Jadin &amp; Vercammen-Grandjean" baseAuthorityYear="1952" class="Arachnida" family="Trombiculidae" genus="Gahrliepia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="brennani">
<emphasis id="F2B0D657FF8FFF918DB8FB56DA14FB43" box="[1257,1399,1152,1178]" italics="true" pageId="21" pageNumber="22">G. brennani</emphasis>
(
<bibRefCitation id="A45577B4FF8FFF918994FB75DDE6FB67" author="Jadin &amp; Vercammen-Grandjean" box="[197,645,1187,1214]" firstAuthor="Jadin" pageId="21" pageNumber="22" pagination="605 - 641" refId="ref63049" refString="Jadin J. B. &amp; Vercammen-Grandjean P. H. 1952. Douze nouvelles especes de larves de Trombiculides (Acariens Prostigmates) du Ruanda-Urundi (Congo Belge). Annales de la Societe belge de Medecine tropicale 32: 605 - 641." type="journal article" year="1952">Jadin &amp; Vercammen-Grandjean, 1952</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="07C471C6FF8FFF918B99FB72DB7CFB67" authority="(Audy et al., 1961)" baseAuthorityName="Audy" baseAuthorityYear="1961" box="[712,1055,1187,1214]" class="Arachnida" family="Trombiculidae" genus="Gahrliepia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="traubi">
<emphasis id="F2B0D657FF8FFF918B99FB72DC57FB64" box="[712,820,1187,1214]" italics="true" pageId="21" pageNumber="22">G. traubi</emphasis>
(
<bibRefCitation id="A45577B4FF8FFF918A12FB75DB75FB67" author="Audy" box="[835,1046,1187,1214]" etAl="et al." firstAuthor="Audy" pageId="21" pageNumber="22" pagination="125 - 133" refId="ref60634" refString="Audy J. R. &amp; Vercammen-Grandjean P. H. 1961 a. African Trombiculidae (Acarina). 1. An interim note on the genus Trombicula Berlese and the subgenus Microtrombicula Ewing. Annals of the Natal Museum 15 (12): 125 - 133. Available from https: // hdl. handle. net / 10520 / AJA 03040798 _ 814 [accessed 4 December 2017]." type="journal article" year="1961">
Audy
<emphasis id="F2B0D657FF8FFF918ADFFB72DCAAFB64" box="[910,969,1187,1213]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
, 1961
</bibRefCitation>
)
</taxonomicName>
, were used by
<bibRefCitation id="A45577B4FF8FFF918D9EFB75DD40FB3B" author="Kolebinova &amp; Vercammen-Grandjean" firstAuthor="Kolebinova" pageId="21" pageNumber="22" pagination="101 - 131" refId="ref63661" refString="Kolebinova M. &amp; Vercammen-Grandjean P. H. 1978. Five new larval Trombiculidae (Acarina) from Liberia and one from Zaire. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg 6 (101): 101 - 131." type="journal article" year="1978">Kolebinova &amp; Vercammen-Grandjean (1978)</bibRefCitation>
simultaneously, without any discussion on their relation.
</paragraph>
<paragraph id="C07B0A45FF8FFF9189ECFADFDB75FAFA" blockId="21.[189,1046,1289,1316]" box="[189,1046,1289,1316]" pageId="21" pageNumber="22">
The deutonymph was described (
<bibRefCitation id="A45577B4FF8FFF918B13FADFDB66FAFD" author="Vercammen-Grandjean &amp; Jadin" box="[578,1029,1289,1316]" firstAuthor="Vercammen-Grandjean" pageId="21" pageNumber="22" pagination="360 - 364" refId="ref68743" refString="Vercammen-Grandjean P. H. &amp; Jadin J. B. 1956 a. Considerations diverses au sujet de Gateria (Giroudia) brennani (Jad. et Verc. 1952) et description de la nymphe. Revue de zoologie et de botanique africaines 53: 360 - 364." type="journal article" year="1956" yearSuffix="a">Vercammen-Grandjean &amp; Jadin 1956a</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>