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<document id="7CE1F8212C873245F51D10A0850A29EE" ID-DOI="10.5852/ejt.2018.395" ID-GBIF-Dataset="10dbb315-5662-4069-a1ae-9692fdf99499" ID-ISSN="2118-9773" ID-Zenodo-Dep="3829724" ID-ZooBank="A244F97F-8CEB-40CE-A182-7E5440337CAA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="juliana" checkinTime="1589571932603" checkinUser="valdenar" docAuthor="Stekolnikov, Alexandr A." docDate="2018" docId="486DBB53FF38FF2788E2F906DCB3FDE0" docLanguage="en" docName="ejt-395_stekolnikov.pdf.imf" docOrigin="European Journal of Taxonomy 395" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Microtrombicula pachydactyli Stekolnikov 2018, comb. nov." docType="treatment" docVersion="21" lastPageNumber="164" masterDocId="B454C32BFF9AFF848951FFD6DF63FFD9" masterDocTitle="Taxonomy and distribution of African chiggers (Acariformes, Trombiculidae)" masterLastPageNumber="233" masterPageNumber="1" pageNumber="163" updateTime="1740599393548" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="70A2D5EED0F7203BC214D79546EF639B">Taxonomy and distribution of African chiggers (Acariformes, Trombiculidae)</mods:title>
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<mods:namePart id="4AF21C9A8EC334EEB239DFC4C5B7A03A">Stekolnikov, Alexandr A.</mods:namePart>
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<mods:affiliation id="53C6D8D47203F8DDC837C28D1A171F95">Zoological Institute, Russian Academy of Sciences, Universitetskaya embankment 1, St. Petersburg 199034, Russia.</mods:affiliation>
<mods:nameIdentifier id="61E15C95E5E61AD8215E46CAC782D943" type="email">Alexandr.Stekolnikov@zin.ru</mods:nameIdentifier>
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<mods:title id="E1584381A7AD7914FEC3AEDAE3E315C9">European Journal of Taxonomy</mods:title>
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<mods:date id="3DF1CD5328CCA9B3881A059D4BB2E585">2018</mods:date>
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<mods:number id="16E1DEB2ECBE2C3D38FCF753D3459F1C">2018-01-16</mods:number>
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<treatment id="486DBB53FF38FF2788E2F906DCB3FDE0" ID-DOI="http://doi.org/10.5281/zenodo.3849123" ID-GBIF-Taxon="164160984" ID-Zenodo-Dep="3849123" LSID="urn:lsid:plazi:treatment:486DBB53FF38FF2788E2F906DCB3FDE0" httpUri="http://treatment.plazi.org/id/486DBB53FF38FF2788E2F906DCB3FDE0" lastPageId="163" lastPageNumber="164" pageId="162" pageNumber="163" scope_family="Trombiculidae" scope_subFamily="Trombiculinae" scope_superOrder="Acariformes" scope_tribe="Trombiculini">
<subSubSection id="88DE59CEFF38FF2688E2F906DBE3F932" box="[435,1152,1744,1771]" pageId="162" pageNumber="163" type="nomenclature">
<paragraph id="C07B0A45FF38FF2688E2F906DBE3F932" blockId="162.[435,1152,1744,1771]" box="[435,1152,1744,1771]" pageId="162" pageNumber="163">
<heading id="9B33BD29FF38FF2688E2F906DBE3F932" box="[435,1152,1744,1771]" centered="true" fontSize="11" level="2" pageId="162" pageNumber="163" reason="2">
<taxonomicName id="07C471C6FF38FF2688E2F906DC94F932" ID-CoL="8ZLHP" authority="(Lawrence, 1949)" authorityName="Stekolnikov" authorityYear="2018" baseAuthorityName="Lawrence" baseAuthorityYear="1949" box="[435,1015,1744,1771]" class="Arachnida" family="Trombiculidae" genus="Microtrombicula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="pachydactyli" status="comb. nov.">
<emphasis id="F2B0D657FF38FF2688E2F906DC43F932" bold="true" box="[435,800,1744,1771]" italics="true" pageId="162" pageNumber="163">Microtrombicula pachydactyli</emphasis>
(
<bibRefCitation id="A45577B4FF38FF268A7EF907DC8CF932" author="Lawrence" box="[815,1007,1745,1771]" firstAuthor="Lawrence" pageId="162" pageNumber="163" pagination="405 - 486" refId="ref64030" refString="Lawrence R. F. 1949. The larval trombiculid mites of South African vertebrates. Annals of the Natal Museum 11 (3): 405 - 486." type="journal article" year="1949">Lawrence, 1949</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="E9836B2CFF38FF268AAEF906DBE3F932" box="[1023,1152,1744,1771]" pageId="162" pageNumber="163" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFF38FF2689ECF8C0DBBBF832" pageId="162" pageNumber="163" type="reference_group">
<paragraph id="C07B0A45FF38FF2689ECF8C0DC38F8E9" blockId="162.[189,859,1814,1840]" box="[189,859,1814,1840]" pageId="162" pageNumber="163">
<treatmentCitationGroup id="E0D42D6BFF38FF2689ECF8C0DC38F8E9" box="[189,859,1814,1840]" pageId="162" pageNumber="163">
<treatmentCitation id="41652C54FF38FF2689ECF8C0DD9EF8E9" author="Lawrence R. F." box="[189,765,1814,1840]" page="443" pageId="162" pageNumber="163" year="1949">
<taxonomicName id="07C471C6FF38FF2689ECF8C0DD9EF8E9" authority="Lawrence, 1949: 443" authorityName="Lawrence" authorityPageNumber="443" authorityYear="1949" box="[189,765,1814,1840]" class="Arachnida" family="Trombiculidae" genus="Eutrombicula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="pachydactyli">
<emphasis id="F2B0D657FF38FF2689ECF8C0DE98F8E9" box="[189,507,1814,1840]" italics="true" pageId="162" pageNumber="163">Eutrombicula pachydactyli</emphasis>
<bibRefCitation id="A45577B4FF38FF268B53F8C0DD9EF8E9" author="Lawrence" box="[514,765,1814,1840]" firstAuthor="Lawrence" page="443" pageId="162" pageNumber="163" pagination="405 - 486" refId="ref64030" refString="Lawrence R. F. 1949. The larval trombiculid mites of South African vertebrates. Annals of the Natal Museum 11 (3): 405 - 486." type="journal article" year="1949">Lawrence, 1949: 443</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 29.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C07B0A45FF38FF2689ECF88DDC8AF8AC" blockId="162.[189,1398,1883,2027]" box="[189,1001,1883,1910]" pageId="162" pageNumber="163">
<treatmentCitationGroup id="E0D42D6BFF38FF2689ECF88DDC8AF8AC" box="[189,1001,1883,1910]" pageId="162" pageNumber="163">
<taxonomicName id="07C471C6FF38FF2689ECF88DDDE7F8AC" box="[189,644,1883,1909]" class="Arachnida" family="Trombiculidae" genus="Trombicula" kingdom="Animalia" order="Trombidiformes" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="pachydactyli" subGenus="Trombicula">
<emphasis id="F2B0D657FF38FF2689ECF88DDDE7F8AC" box="[189,644,1883,1909]" italics="true" pageId="162" pageNumber="163">Trombicula (Trombicula) pachydactyli</emphasis>
</taxonomicName>
<treatmentCitation id="41652C54FF38FF268BF1F88DDC86F8AC" author="Wharton G. W. &amp; Fuller H. S." box="[672,997,1883,1910]" page="68" pageId="162" pageNumber="163" year="1952">
<bibRefCitation id="A45577B4FF38FF268BF1F88DDC86F8AC" author="Wharton &amp; Fuller" box="[672,997,1883,1910]" firstAuthor="Wharton" page="68" pageId="162" pageNumber="163" pagination="1 - 185" refId="ref69481" refString="Wharton G. W. &amp; Fuller H. S. 1952. A manual of the chiggers. Memoirs of the Entomological Society of Washington 4: 1 - 185." type="journal article" year="1952">Wharton &amp; Fuller 1952: 68</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C07B0A45FF38FF2689ECF854DE89F81D" blockId="162.[189,1398,1883,2027]" pageId="162" pageNumber="163">
<treatmentCitationGroup id="E0D42D6BFF38FF2689ECF854DE89F81D" pageId="162" pageNumber="163">
<taxonomicName id="07C471C6FF38FF2689ECF854DDC8F845" authorityName="Lawrence" authorityYear="1949" box="[189,683,1922,1948]" class="Arachnida" family="Trombiculidae" genus="Eutrombicula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="pachydactyli" subGenus="Squamicola">
<emphasis id="F2B0D657FF38FF2689ECF854DDC8F845" box="[189,683,1922,1948]" italics="true" pageId="162" pageNumber="163">Eutrombicula (Squamicola) pachydactyli</emphasis>
</taxonomicName>
<treatmentCitation id="41652C54FF38FF268B83F854DB8EF844" author="Audy J. R. &amp; Vercammen-Grandjean P. H." box="[722,1261,1922,1949]" page="138" pageId="162" pageNumber="163" year="1961">
<bibRefCitation id="A45577B4FF38FF268B83F854DB8EF844" author="Audy &amp; Vercammen-Grandjean" box="[722,1261,1922,1949]" firstAuthor="Audy" page="138" pageId="162" pageNumber="163" pagination="135 - 140" refId="ref60708" refString="Audy J. R. &amp; Vercammen-Grandjean P. H. 1961 b. African Trombiculidae (Acarina). 2. The genera Eutrombicula Ew. and Sauriscus Lawr., with description of a new subgenus, Squamicola. Annals of the Natal Museum 15 (13): 135 - 140. Available from https: // hdl. handle. net / 10520 / AJA 03040798 _ 808 [accessed 4 December 2017]." type="journal article" year="1961" yearSuffix="b">Audy &amp; Vercammen-Grandjean 1961b: 138</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="41652C54FF38FF268C79F855DE00F81D" author="Zumpt F." page="149" pageId="162" pageNumber="163" year="1961">
<bibRefCitation id="A45577B4FF38FF268C79F855DE00F81D" author="Zumpt" firstAuthor="Zumpt" page="149" pageId="162" pageNumber="163" pagination="1 - 457" refId="ref69765" refString="Zumpt F. (ed.) 1961. The arthropod parasites of vertebrates in Africa south of the Sahara (Ethiopian Region). Vol. 1: Chelicerata. Publications of the South African Institute for Medical Research 11 (50): 1 - 457." type="journal article" year="1961">Zumpt 1961: 149</bibRefCitation>
</treatmentCitation>
, fig. 88ac.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C07B0A45FF38FF2689ECF807DBBBF832" blockId="162.[189,1398,1883,2027]" box="[189,1240,2000,2027]" pageId="162" pageNumber="163">
<taxonomicName id="07C471C6FF38FF2689ECF807DD2FF832" box="[189,588,2001,2027]" class="Arachnida" family="Trombiculidae" genus="Eltonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="162" pageNumber="163" phylum="Arthropoda" rank="species" species="pachydactyli" subGenus="Eltonella">
<emphasis id="F2B0D657FF38FF2689ECF807DD2FF832" box="[189,588,2001,2027]" italics="true" pageId="162" pageNumber="163">Eltonella (Eltonella) pachydactyli</emphasis>
</taxonomicName>
<bibRefCitation id="A45577B4FF38FF268B38F807DC94F832" author="Vercammen-Grandjean P. H." box="[617,1015,2000,2027]" pageId="162" pageNumber="163" pagination="34 - 257" refId="ref67955" refString="Vercammen-Grandjean P. H. 1965 a. Revision of the genera: Eltonella Audy, 1956 and Microtrombicula Ewing, 1950, with descriptions of fifty new species and transferal of subgenus Chiroptella to genus Leptotrombidium (Acarina, Trombiculidae). Acarologia 7 (suppl.): 34 - 257." type="journal article" year="1965">Vercammen-Grandjean 1965a: 82</bibRefCitation>
, pl. EE; 1965c: 42.
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFF39FF2789ECFEDDDA69FE8C" box="[189,1290,267,341]" pageId="163" pageNumber="164" type="materials_examined">
<materialsCitation id="70AC0018FF39FF2789ECFEDDDE8BFE8C" box="[189,488,267,341]" collectionCode="TMSA" pageId="163" pageNumber="164" specimenCode="TMSA 13" specimenCount="1" typeStatus="syntype">
<paragraph id="C07B0A45FF39FF2789ECFEDDDE4EFEFC" blockId="163.[189,301,267,293]" box="[189,301,267,293]" pageId="163" pageNumber="164">
<heading id="9B33BD29FF39FF2789ECFEDDDE4EFEFC" bold="true" box="[189,301,267,293]" fontSize="11" level="3" pageId="163" pageNumber="164" reason="3">
<emphasis id="F2B0D657FF39FF2789ECFEDDDE4EFEFC" bold="true" box="[189,301,267,293]" pageId="163" pageNumber="164">
<typeStatus id="1F7FB4E7FF39FF2789ECFEDDDE4EFEFC" box="[189,301,267,293]" pageId="163" pageNumber="164" type="syntype">Syntypes</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FF39FF2789ECFEECDA69FE8C" blockId="163.[189,1290,314,341]" box="[189,1290,314,341]" pageId="163" pageNumber="164">
<specimenCode id="9062A23EFF39FF2789ECFEECDE52FE8D" box="[189,305,314,340]" pageId="163" pageNumber="164">TMSA 13</specimenCode>
(original data),
<materialsCitation id="70AC0018FF39FF2788BEFEECDA69FE8C" box="[495,1290,314,341]" collectionCode="NMSA" determinerName="Wharton &amp; Fuller &amp; Vercammen-Grandjean" pageId="163" pageNumber="164" specimenCode="NMSA 13" specimenCount="1">
<specimenCode id="9062A23EFF39FF2788BEFEECDD05FE8D" box="[495,614,314,340]" pageId="163" pageNumber="164">NMSA 13</specimenCode>
(
<bibRefCitation id="A45577B4FF39FF278B26FEECDCEFFE8C" author="Wharton &amp; Fuller" box="[631,908,314,341]" firstAuthor="Wharton" pageId="163" pageNumber="164" pagination="1 - 185" refId="ref69481" refString="Wharton G. W. &amp; Fuller H. S. 1952. A manual of the chiggers. Memoirs of the Entomological Society of Washington 4: 1 - 185." type="journal article" year="1952">
<determinerName id="35A7E681FF39FF278B26FEECDC29FE8D" box="[631,842,314,340]" pageId="163" pageNumber="164">Wharton &amp; Fuller</determinerName>
1952
</bibRefCitation>
;
<bibRefCitation id="A45577B4FF39FF278ACAFEECDB99FE8C" author="Vercammen-Grandjean P. H." box="[923,1274,314,341]" pageId="163" pageNumber="164" pagination="34 - 257" refId="ref67955" refString="Vercammen-Grandjean P. H. 1965 a. Revision of the genera: Eltonella Audy, 1956 and Microtrombicula Ewing, 1950, with descriptions of fifty new species and transferal of subgenus Chiroptella to genus Leptotrombidium (Acarina, Trombiculidae). Acarologia 7 (suppl.): 34 - 257." type="journal article" year="1965">
<determinerName id="35A7E681FF39FF278ACAFEECDBC8FE8D" box="[923,1195,314,340]" pageId="163" pageNumber="164">Vercammen-Grandjean</determinerName>
1965a
</bibRefCitation>
).
</materialsCitation>
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="88DE59CEFF39FF2789ECFEABDD5DFE1E" box="[189,574,381,455]" pageId="163" pageNumber="164" type="distribution">
<paragraph id="C07B0A45FF39FF2789ECFEABDE35FE4E" blockId="163.[189,342,381,407]" box="[189,342,381,407]" pageId="163" pageNumber="164">
<heading id="9B33BD29FF39FF2789ECFEABDE35FE4E" bold="true" box="[189,342,381,407]" fontSize="11" level="3" pageId="163" pageNumber="164" reason="3">
<emphasis id="F2B0D657FF39FF2789ECFEABDE35FE4E" bold="true" box="[189,342,381,407]" pageId="163" pageNumber="164">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FF39FF2789ECFE7ADD5DFE1E" blockId="163.[189,574,428,455]" box="[189,574,428,455]" pageId="163" pageNumber="164">
<collectingCountry id="B8D34AD5FF39FF2789ECFE7ADE41FE1F" box="[189,290,428,454]" name="Namibia" pageId="163" pageNumber="164">Namibia</collectingCountry>
(Rehoboth, Kamanjab).
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFF39FF2789ECFE39DCB3FDE0" box="[189,976,495,569]" pageId="163" pageNumber="164" type="biology_ecology">
<paragraph id="C07B0A45FF39FF2789ECFE39DE60FDD0" blockId="163.[189,259,495,521]" box="[189,259,495,521]" pageId="163" pageNumber="164">
<heading id="9B33BD29FF39FF2789ECFE39DE60FDD0" bold="true" box="[189,259,495,521]" fontSize="11" level="3" pageId="163" pageNumber="164" reason="3">
<emphasis id="F2B0D657FF39FF2789ECFE39DE60FDD0" bold="true" box="[189,259,495,521]" pageId="163" pageNumber="164">Hosts</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FF39FF2789ECFDC8DCB3FDE0" blockId="163.[189,976,542,569]" box="[189,976,542,569]" pageId="163" pageNumber="164">
<taxonomicName id="07C471C6FF39FF2789ECFDC8DD6AFDE1" baseAuthorityName="Loveridge" baseAuthorityYear="1947" box="[189,521,542,568]" class="Squamata" family="Gekkonidae" genus="Chondrodactylus" kingdom="Animalia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="fitzsimonsi">
<emphasis id="F2B0D657FF39FF2789ECFDC8DD6AFDE1" box="[189,521,542,568]" italics="true" pageId="163" pageNumber="164">Chondrodactylus fitzsimonsi</emphasis>
</taxonomicName>
,
<taxonomicName id="07C471C6FF39FF278B49FDC8DC20FDE1" authorityName="Fischer" authorityYear="1888" box="[536,835,542,568]" class="Squamata" family="Gekkonidae" genus="Pachydactylus" kingdom="Animalia" pageId="163" pageNumber="164" phylum="Chordata" rank="species" species="laevigatus">
<emphasis id="F2B0D657FF39FF278B49FDC8DC20FDE1" box="[536,835,542,568]" italics="true" pageId="163" pageNumber="164">Pachydactylus laevigatus</emphasis>
</taxonomicName>
(
<typeStatus id="1F7FB4E7FF39FF278A03FDC8DCE5FDE1" box="[850,902,542,568]" pageId="163" pageNumber="164">type</typeStatus>
host).
</paragraph>
</subSubSection>
</treatment>
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