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<mods:title id="45D25089FF27305E880C71AAA9DD7301">Taxonomy and distribution of African chiggers (Acariformes, Trombiculidae)</mods:title>
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<heading id="9B33BD29FF36FF288890FAF3DB10FA99" box="[449,1139,1317,1344]" centered="true" fontSize="11" level="2" pageId="172" pageNumber="173" reason="2">
<taxonomicName id="07C471C6FF36FF288890FAF3DC8AFA99" authority="(Oudemans, 1910)" authorityName="Stekolnikov" authorityYear="2018" baseAuthorityName="Oudemans" baseAuthorityYear="1910" box="[449,1001,1317,1344]" class="Arachnida" family="Trombiculidae" genus="Microtrombicula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="172" pageNumber="173" phylum="Arthropoda" rank="species" species="tragardhi" status="comb. nov.">
<emphasis id="F2B0D657FF36FF288890FAF3DC6BFA99" bold="true" box="[449,776,1317,1344]" italics="true" pageId="172" pageNumber="173">Microtrombicula tragardhi</emphasis>
(Oudemans, 1910)
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<treatmentCitation id="41652C54FF36FF2889ECFABEDC7BFA58" author="Oudemans A. C." box="[189,792,1383,1410]" page="86" pageId="172" pageNumber="173" year="1910">
<taxonomicName id="07C471C6FF36FF2889ECFABEDC7BFA58" authority="Oudemans, 1910 a: 86" authorityName="Oudemans" authorityPageNumber="86" authorityYear="1910" box="[189,792,1383,1410]" class="Arachnida" family="Microtrombidiidae" genus="Microthrombidium" kingdom="Animalia" order="Prostigmata" pageId="172" pageNumber="173" phylum="Arthropoda" rank="species" species="tragardhi">
<emphasis id="F2B0D657FF36FF2889ECFABEDD73FA5B" box="[189,528,1384,1410]" italics="true" pageId="172" pageNumber="173">Microthrombidium tragardhi</emphasis>
<bibRefCitation id="A45577B4FF36FF288B49FAB1DC7BFA58" author="Oudemans A. C." box="[536,792,1383,1410]" pageId="172" pageNumber="173" pagination="83 - 90" refId="ref65038" refString="Oudemans A. C. 1910 a. Acarologische aanteekeningen XXXIII. Entomologische Berichten 3 (54): 83 - 90. Available from https: // biodiversitylibrary. org / page / 8983129 [accessed 4 December 2017]." type="journal article" year="1910">Oudemans, 1910a: 86</bibRefCitation>
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<paragraph id="C07B0A45FF36FF2889ECFA7FDC10FA1D" blockId="172.[189,1009,1449,1654]" box="[189,883,1449,1476]" pageId="172" pageNumber="173">
<treatmentCitationGroup id="E0D42D6BFF36FF2889ECFA7FDC10FA1D" box="[189,883,1449,1476]" pageId="172" pageNumber="173">
<taxonomicName id="07C471C6FF36FF2889ECFA7FDD73FA1A" authorityName="Oudemans" authorityYear="1910" box="[189,528,1449,1475]" class="Arachnida" family="Microtrombidiidae" genus="Microthrombidium" kingdom="Animalia" order="Prostigmata" pageId="172" pageNumber="173" phylum="Arthropoda" rank="species" species="tragardhi">
<emphasis id="F2B0D657FF36FF2889ECFA7FDD73FA1A" box="[189,528,1449,1475]" italics="true" pageId="172" pageNumber="173">Microthrombidium tragardhi</emphasis>
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<treatmentCitation id="41652C54FF36FF288B7FFA7FDC7EFA1A" author="Oudemans A. C." box="[558,797,1449,1476]" page="37" pageId="172" pageNumber="173" year="1912">
<bibRefCitation id="A45577B4FF36FF288B7FFA7FDC7EFA1A" author="Oudemans A. C." box="[558,797,1449,1476]" pageId="172" pageNumber="173" pagination="1 - 230" refId="ref65158" refString="Oudemans A. C. 1912. Die bis jetzt bekannten Larven von Thrombidiidae und Erythraeidae. Zoologische Jahrbucher suppl. 14 (1): 1 - 230." type="journal article" year="1912">Oudemans 1912: 37</bibRefCitation>
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, fig. K.
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<treatmentCitationGroup id="E0D42D6BFF36FF2889ECFA1BDC92FA3E" box="[189,1009,1485,1512]" pageId="172" pageNumber="173">
<taxonomicName id="07C471C6FF36FF2889ECFA1BDED8FA3E" authorityName="Oudemans" authorityYear="1910" box="[189,443,1485,1511]" class="Arachnida" family="Trombiculidae" genus="Trombicula" kingdom="Animalia" order="Prostigmata" pageId="172" pageNumber="173" phylum="Arthropoda" rank="species" species="tragardhi">
<emphasis id="F2B0D657FF36FF2889ECFA1BDED8FA3E" box="[189,443,1485,1511]" italics="true" pageId="172" pageNumber="173">Trombicula tragardhi</emphasis>
</taxonomicName>
<treatmentCitation id="41652C54FF36FF288889FA1BDDC9FA3E" author="Radford C. D." box="[472,682,1485,1511]" page="60" pageId="172" pageNumber="173" year="1942">
<bibRefCitation id="A45577B4FF36FF288889FA1BDDC9FA3E" author="Radford C. D." box="[472,682,1485,1511]" pageId="172" pageNumber="173" pagination="55 - 81" refId="ref65343" refString="Radford C. D. 1942. The larval Trombiculinae (Acarina: Trombidiidae) with descriptions of twelve new species. Parasitology 34 (1): 55 - 81. https: // doi. org / 10.1017 / S 0031182000015997" type="journal article" year="1942">Radford 1942: 60</bibRefCitation>
</treatmentCitation>
, fig. 17. —
<treatmentCitation id="41652C54FF36FF288A65FA1BDC8EFA3E" author="Fuller H. S." box="[820,1005,1485,1512]" page="72" pageId="172" pageNumber="173" year="1952">
<bibRefCitation id="A45577B4FF36FF288A65FA1BDC8EFA3E" author="Fuller H. S." box="[820,1005,1485,1512]" pageId="172" pageNumber="173" pagination="1 - 261" refId="ref61953" refString="Fuller H. S. 1952. The mite-larvae of the family Trombiculidae in the Oudemans collection: taxonomy and medical importance. Zoologische Verhandelingen 18: 1 - 261." type="journal article" year="1952">Fuller 1952: 72</bibRefCitation>
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.
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</paragraph>
<paragraph id="C07B0A45FF36FF2889ECFA26DCCFF9D2" blockId="172.[189,1009,1449,1654]" box="[189,940,1520,1547]" pageId="172" pageNumber="173">
<treatmentCitationGroup id="E0D42D6BFF36FF2889ECFA26DCCFF9D2" box="[189,940,1520,1547]" pageId="172" pageNumber="173">
<taxonomicName id="07C471C6FF36FF2889ECFA26DEB3F9D3" box="[189,464,1520,1546]" class="Arachnida" family="Trombiculidae" genus="Pentagonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="172" pageNumber="173" phylum="Arthropoda" rank="species" species="tragardhi">
<emphasis id="F2B0D657FF36FF2889ECFA26DEB3F9D3" box="[189,464,1520,1546]" italics="true" pageId="172" pageNumber="173">Pentagonella tragardhi</emphasis>
</taxonomicName>
<treatmentCitation id="41652C54FF36FF2888BCFA26DC5CF9D2" author="Thor S. &amp; Willmann C." box="[493,831,1520,1547]" page="294" pageId="172" pageNumber="173" year="1947">
<bibRefCitation id="A45577B4FF36FF2888BCFA26DC5CF9D2" author="Thor S. &amp; Willmann C." box="[493,831,1520,1547]" pageId="172" pageNumber="173" pagination="71" refId="ref66977" refString="Thor S. &amp; Willmann C. 1947. Acarina. In: Schulze F. E., Kukenthal W. &amp; Heider K. (eds) Das Tierreich 71. Walter de Gruyter &amp; Co., Berlin." type="book chapter" year="1947">Thor &amp; Willmann 1947: 294</bibRefCitation>
</treatmentCitation>
, fig. 351.
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</paragraph>
<paragraph id="C07B0A45FF36FF2889ECF9C2DC89F9F7" blockId="172.[189,1009,1449,1654]" box="[189,1002,1556,1583]" pageId="172" pageNumber="173">
<treatmentCitationGroup id="E0D42D6BFF36FF2889ECF9C2DC89F9F7" box="[189,1002,1556,1583]" pageId="172" pageNumber="173">
<taxonomicName id="07C471C6FF36FF2889ECF9C2DDE7F9F7" authorityName="Oudemans" authorityYear="1910" box="[189,644,1556,1582]" class="Arachnida" family="Trombiculidae" genus="Trombicula" kingdom="Animalia" order="Prostigmata" pageId="172" pageNumber="173" phylum="Arthropoda" rank="species" species="tragardhi" subGenus="Neotrombicula">
<emphasis id="F2B0D657FF36FF2889ECF9C2DDE7F9F7" box="[189,644,1556,1582]" italics="true" pageId="172" pageNumber="173">Trombicula (Neotrombicula) tragardhi</emphasis>
</taxonomicName>
<treatmentCitation id="41652C54FF36FF288BF3F9C2DC85F9F7" author="Wharton G. W. &amp; Fuller H. S." box="[674,998,1556,1583]" page="60" pageId="172" pageNumber="173" year="1952">
<bibRefCitation id="A45577B4FF36FF288BF3F9C2DC85F9F7" author="Wharton G. W. &amp; Fuller H. S." box="[674,998,1556,1583]" pageId="172" pageNumber="173" pagination="1 - 185" refId="ref69481" refString="Wharton G. W. &amp; Fuller H. S. 1952. A manual of the chiggers. Memoirs of the Entomological Society of Washington 4: 1 - 185." type="journal article" year="1952">Wharton &amp; Fuller 1952: 60</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C07B0A45FF36FF2889ECF9EEDCCFF98B" blockId="172.[189,1009,1449,1654]" box="[189,940,1591,1618]" pageId="172" pageNumber="173">
<treatmentCitationGroup id="E0D42D6BFF36FF2889ECF9EEDCCFF98B" box="[189,940,1591,1618]" pageId="172" pageNumber="173">
<taxonomicName id="07C471C6FF36FF2889ECF9EEDE81F98B" box="[189,482,1592,1618]" class="Arachnida" family="Trombiculidae" genus="Neotrombicula" kingdom="Animalia" order="Prostigmata" pageId="172" pageNumber="173" phylum="Arthropoda" rank="species" species="tragardhi">
<emphasis id="F2B0D657FF36FF2889ECF9EEDE81F98B" box="[189,482,1592,1618]" italics="true" pageId="172" pageNumber="173">Neotrombicula tragardhi</emphasis>
</taxonomicName>
<treatmentCitation id="41652C54FF36FF288B51F9EEDDACF988" author="Zumpt F." box="[512,719,1591,1618]" page="146" pageId="172" pageNumber="173" year="1961">
<bibRefCitation id="A45577B4FF36FF288B51F9EEDDACF988" author="Zumpt F." box="[512,719,1591,1618]" pageId="172" pageNumber="173" pagination="1 - 457" refId="ref69765" refString="Zumpt F. (ed.) 1961. The arthropod parasites of vertebrates in Africa south of the Sahara (Ethiopian Region). Vol. 1: Chelicerata. Publications of the South African Institute for Medical Research 11 (50): 1 - 457." type="journal article" year="1961">Zumpt 1961: 146</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="41652C54FF36FF288BAEF9E1DCC4F98B" author="Goff M. L." box="[767,935,1591,1618]" page="12" pageId="172" pageNumber="173" year="1995">
<bibRefCitation id="A45577B4FF36FF288BAEF9E1DCC4F98B" author="Goff M. L." box="[767,935,1591,1618]" pageId="172" pageNumber="173" pagination="12 - 15" refId="ref62704" refString="Goff M. L. 1995. New species of Neotrombicula (Acari: Trombiculidae) from African primates (Galagidae and Cercopithecidae). Journal of Medical Entomology 32 (1): 12 - 15. https: // doi. org / 10.1093 / jmedent / 32.1.12" type="journal article" year="1995">Goff 1995: 12</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="C07B0A45FF36FF2889ECF98DDCB5F9AC" blockId="172.[189,1009,1449,1654]" box="[189,982,1627,1654]" pageId="172" pageNumber="173">
<treatmentCitationGroup id="E0D42D6BFF36FF2889ECF98DDCB5F9AC" box="[189,982,1627,1654]" pageId="172" pageNumber="173">
<taxonomicName id="07C471C6FF36FF2889ECF98DDD46F9AC" box="[189,549,1627,1653]" class="Arachnida" family="Trombiculidae" genus="Eltonella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="172" pageNumber="173" phylum="Arthropoda" rank="species" species="tragardhi" subGenus="Eltonella">
<emphasis id="F2B0D657FF36FF2889ECF98DDD46F9AC" box="[189,549,1627,1653]" italics="true" pageId="172" pageNumber="173">Eltonella (Eltonella) tragardhi</emphasis>
</taxonomicName>
<treatmentCitation id="41652C54FF36FF288B13F98DDCB1F9AC" author="Vercammen-Grandjean P. H." box="[578,978,1627,1654]" page="43" pageId="172" pageNumber="173" year="1965">
<bibRefCitation id="A45577B4FF36FF288B13F98DDCB1F9AC" author="Vercammen-Grandjean P. H." box="[578,978,1627,1654]" pageId="172" pageNumber="173" refId="ref68052" refString="Vercammen-Grandjean P. H. 1965 c. Trombiculinae of the World. Synopsis with Generic, Subgeneric, and Group Diagnoses (Acarina, Trombiculidae). George Williams Hooper Foundation, San Francisco." type="book" year="1965">Vercammen-Grandjean 1965c: 43</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<heading id="9B33BD29FF36FF2889ECF94BDE4CF96E" bold="true" box="[189,303,1693,1719]" fontSize="11" level="3" pageId="172" pageNumber="173" reason="3">
<emphasis id="F2B0D657FF36FF2889ECF94BDE4CF96E" bold="true" box="[189,303,1693,1719]" pageId="172" pageNumber="173">
<typeStatus id="1F7FB4E7FF36FF2889ECF94BDE4CF96E" box="[189,303,1693,1719]" pageId="172" pageNumber="173" type="holotype">Holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FF36FF2889ECF91ADE35F8D3" blockId="172.[189,1399,1739,1802]" pageId="172" pageNumber="173">
Private collection of Trägårdh, Stockholm (
<bibRefCitation id="A45577B4FF36FF288B97F91ADCEBF93F" author="Oudemans A. C." box="[710,904,1740,1766]" pageId="172" pageNumber="173" pagination="1 - 230" refId="ref65158" refString="Oudemans A. C. 1912. Die bis jetzt bekannten Larven von Thrombidiidae und Erythraeidae. Zoologische Jahrbucher suppl. 14 (1): 1 - 230." type="journal article" year="1912">Oudemans 1912</bibRefCitation>
), lost (
<bibRefCitation id="A45577B4FF36FF288A8EF91ADB09F93E" author="Fuller H. S." box="[991,1130,1740,1767]" pageId="172" pageNumber="173" pagination="1 - 261" refId="ref61953" refString="Fuller H. S. 1952. The mite-larvae of the family Trombiculidae in the Oudemans collection: taxonomy and medical importance. Zoologische Verhandelingen 18: 1 - 261." type="journal article" year="1952">Fuller 1952</bibRefCitation>
),
<collectionCode id="A6D59280FF36FF288DD5F91ADBBFF93F" box="[1156,1244,1740,1766]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="172" pageNumber="173" type="Museum">RMNH</collectionCode>
(
<bibRefCitation id="A45577B4FF36FF288DA1F91ADE26F8D3" author="Wharton G. W. &amp; Fuller H. S." pageId="172" pageNumber="173" pagination="1 - 185" refId="ref69481" refString="Wharton G. W. &amp; Fuller H. S. 1952. A manual of the chiggers. Memoirs of the Entomological Society of Washington 4: 1 - 185." type="journal article" year="1952">Wharton &amp; Fuller 1952</bibRefCitation>
).
</paragraph>
</subSubSection>
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<emphasis id="F2B0D657FF36FF2889ECF8E7DE35F892" bold="true" box="[189,342,1841,1867]" pageId="172" pageNumber="173">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FF36FF2889ECF8B6DEC9F8A2" blockId="172.[189,426,1888,1915]" box="[189,426,1888,1915]" pageId="172" pageNumber="173">
<collectingCountry id="B8D34AD5FF36FF2889ECF8B6DE65F8A3" box="[189,262,1888,1914]" name="Sudan" pageId="172" pageNumber="173">Sudan</collectingCountry>
(
<collectingRegion id="0200C4A7FF36FF288844F8B6DEFEF8A2" box="[277,413,1888,1915]" country="Sudan" name="An Nil al Abyad" pageId="172" pageNumber="173">White Nile</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFF36FF2889ECF874DCB9F832" box="[189,986,1954,2028]" pageId="172" pageNumber="173" type="biology_ecology">
<paragraph id="C07B0A45FF36FF2889ECF874DE60F865" blockId="172.[189,259,1954,1980]" box="[189,259,1954,1980]" pageId="172" pageNumber="173">
<heading id="9B33BD29FF36FF2889ECF874DE60F865" bold="true" box="[189,259,1954,1980]" fontSize="11" level="3" pageId="172" pageNumber="173" reason="3">
<emphasis id="F2B0D657FF36FF2889ECF874DE60F865" bold="true" box="[189,259,1954,1980]" pageId="172" pageNumber="173">Hosts</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FF36FF2889ECF807DCB9F832" blockId="172.[189,986,2001,2028]" box="[189,986,2001,2028]" pageId="172" pageNumber="173">
<taxonomicName id="07C471C6FF36FF2889ECF807DEDFF832" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[189,444,2001,2027]" class="Mammalia" family="Cercopithecidae" genus="Chlorocebus" kingdom="Animalia" order="Primates" pageId="172" pageNumber="173" phylum="Chordata" rank="species" species="aethiops">
<emphasis id="F2B0D657FF36FF2889ECF807DEDFF832" box="[189,444,2001,2027]" italics="true" pageId="172" pageNumber="173">Chlorocebus aethiops</emphasis>
</taxonomicName>
(original data),
<taxonomicName id="07C471C6FF36FF288B2BF807DCB6F832" authority="(Zumpt 1961)" baseAuthorityName="Zumpt" baseAuthorityYear="1961" box="[634,981,2001,2027]" class="Mammalia" family="Cercopithecidae" genus="Chlorocebus" kingdom="Animalia" order="Primates" pageId="172" pageNumber="173" phylum="Chordata" rank="species" species="pygerythrus">
<emphasis id="F2B0D657FF36FF288B2BF807DC4BF832" box="[634,808,2001,2027]" italics="true" pageId="172" pageNumber="173">C. pygerythrus</emphasis>
(
<bibRefCitation id="A45577B4FF36FF288A68F807DCA9F832" author="Zumpt F." box="[825,970,2001,2027]" pageId="172" pageNumber="173" pagination="1 - 457" refId="ref69765" refString="Zumpt F. (ed.) 1961. The arthropod parasites of vertebrates in Africa south of the Sahara (Ethiopian Region). Vol. 1: Chelicerata. Publications of the South African Institute for Medical Research 11 (50): 1 - 457." type="journal article" year="1961">Zumpt 1961</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>