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<document id="4472830911862D6D096209F821EF0CD9" ID-DOI="10.5852/ejt.2018.395" ID-GBIF-Dataset="10dbb315-5662-4069-a1ae-9692fdf99499" ID-ISSN="2118-9773" ID-Zenodo-Dep="3829724" ID-ZooBank="A244F97F-8CEB-40CE-A182-7E5440337CAA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="juliana" checkinTime="1589571932603" checkinUser="valdenar" docAuthor="Stekolnikov, Alexandr A." docDate="2018" docId="486DBB53FF26FF388888FE63DCB4FB82" docLanguage="en" docName="ejt-395_stekolnikov.pdf.imf" docOrigin="European Journal of Taxonomy 395" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Willmannium natalense Stekolnikov 2018, comb. nov." docType="treatment" docVersion="20" lastPageNumber="189" masterDocId="B454C32BFF9AFF848951FFD6DF63FFD9" masterDocTitle="Taxonomy and distribution of African chiggers (Acariformes, Trombiculidae)" masterLastPageNumber="233" masterPageNumber="1" pageNumber="189" updateTime="1740599393548" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="77B8612A4D0110DAD0C9ADCF4AC05B92">Taxonomy and distribution of African chiggers (Acariformes, Trombiculidae)</mods:title>
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<mods:name id="3EAC3529F1AFB943F30FC2020CC9D2CA" type="personal">
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<mods:namePart id="5BD3DCEDF940EB5F2B1DF36D83D08E11">Stekolnikov, Alexandr A.</mods:namePart>
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<mods:affiliation id="C8785E8958F1DCDFCEA75DDA30A28C8F">Zoological Institute, Russian Academy of Sciences, Universitetskaya embankment 1, St. Petersburg 199034, Russia.</mods:affiliation>
<mods:nameIdentifier id="226C078DE45D33879218936D5D397941" type="email">Alexandr.Stekolnikov@zin.ru</mods:nameIdentifier>
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<mods:title id="94657F2435BE462F84B07C451F1FB477">European Journal of Taxonomy</mods:title>
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<mods:part id="C233C3661519BF38646E08DCC74FB9E4">
<mods:date id="0D8D80B312E1012E0ADB2ECD1167AE11">2018</mods:date>
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<mods:number id="C6C13BE003DC1BE9AEA10EB0A1F1D098">2018-01-16</mods:number>
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<mods:number id="C65544EAE428E32CC86DF850BE63F3F4">395</mods:number>
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<treatment id="486DBB53FF26FF388888FE63DCB4FB82" ID-DOI="http://doi.org/10.5281/zenodo.3849429" ID-GBIF-Taxon="164160885" ID-Zenodo-Dep="3849429" LSID="urn:lsid:plazi:treatment:486DBB53FF26FF388888FE63DCB4FB82" httpUri="http://treatment.plazi.org/id/486DBB53FF26FF388888FE63DCB4FB82" lastPageNumber="189" pageId="188" pageNumber="189" scope_family="Trombiculidae" scope_subFamily="Trombiculinae" scope_superOrder="Acariformes" scope_tribe="Trombiculini">
<subSubSection id="88DE59CEFF26FF388888FE63DB38FE09" box="[473,1115,437,464]" pageId="188" pageNumber="189" type="nomenclature">
<paragraph id="C07B0A45FF26FF388888FE63DB38FE09" blockId="188.[473,1115,437,464]" box="[473,1115,437,464]" pageId="188" pageNumber="189">
<heading id="9B33BD29FF26FF388888FE63DB38FE09" box="[473,1115,437,464]" centered="true" fontSize="11" level="2" pageId="188" pageNumber="189" reason="2">
<taxonomicName id="07C471C6FF26FF388888FE63DCB1FE09" authority="(Lawrence, 1949)" authorityName="Stekolnikov" authorityYear="2018" baseAuthorityName="Lawrence" baseAuthorityYear="1949" box="[473,978,437,464]" class="Arachnida" family="Trombiculidae" genus="Willmannium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="natalense" status="comb. nov.">
<emphasis id="F2B0D657FF26FF388888FE63DD98FE09" bold="true" box="[473,763,437,464]" italics="true" pageId="188" pageNumber="189">Willmannium natalense</emphasis>
(
<bibRefCitation id="A45577B4FF26FF388A5BFE60DCA9FE09" author="Lawrence" box="[778,970,438,464]" firstAuthor="Lawrence" pageId="188" pageNumber="189" pagination="405 - 486" refId="ref64030" refString="Lawrence R. F. 1949. The larval trombiculid mites of South African vertebrates. Annals of the Natal Museum 11 (3): 405 - 486." type="journal article" year="1949">Lawrence, 1949</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="E9836B2CFF26FF388A8BFE60DB38FE09" box="[986,1115,438,464]" pageId="188" pageNumber="189" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFF26FF3889ECFE2DDA7FFD21" pageId="188" pageNumber="189" type="reference_group">
<paragraph id="C07B0A45FF26FF3889ECFE2DDC76FDCC" blockId="188.[189,789,507,534]" box="[189,789,507,534]" pageId="188" pageNumber="189">
<treatmentCitationGroup id="E0D42D6BFF26FF3889ECFE2DDC76FDCC" box="[189,789,507,534]" pageId="188" pageNumber="189">
<treatmentCitation id="41652C54FF26FF3889ECFE2DDDA6FDCC" author="Lawrence R. F." box="[189,709,507,534]" page="413" pageId="188" pageNumber="189" year="1949">
<taxonomicName id="07C471C6FF26FF3889ECFE2DDDA6FDCC" authority="Lawrence, 1949: 413" authorityName="Lawrence" authorityPageNumber="413" authorityYear="1949" box="[189,709,507,534]" class="Arachnida" family="Trombiculidae" genus="Trombicula" kingdom="Animalia" order="Trombidiformes" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="natalensis">
<emphasis id="F2B0D657FF26FF3889ECFE2DDEA0FDCC" box="[189,451,507,533]" italics="true" pageId="188" pageNumber="189">Trombicula natalensis</emphasis>
<bibRefCitation id="A45577B4FF26FF38889BFE2ADDA6FDCC" author="Lawrence" box="[458,709,507,534]" firstAuthor="Lawrence" page="413" pageId="188" pageNumber="189" pagination="405 - 486" refId="ref64030" refString="Lawrence R. F. 1949. The larval trombiculid mites of South African vertebrates. Annals of the Natal Museum 11 (3): 405 - 486." type="journal article" year="1949">Lawrence, 1949: 413</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 3.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C07B0A45FF26FF3889ECFD97DD87FD82" blockId="188.[189,1308,576,760]" box="[189,740,576,603]" pageId="188" pageNumber="189">
<treatmentCitationGroup id="E0D42D6BFF26FF3889ECFD97DD87FD82" box="[189,740,576,603]" pageId="188" pageNumber="189">
<taxonomicName id="07C471C6FF26FF3889ECFD97DEA0FD82" authorityName="Lawrence" authorityYear="1949" box="[189,451,577,603]" class="Arachnida" family="Trombiculidae" genus="Trombicula" kingdom="Animalia" order="Trombidiformes" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="natalensis">
<emphasis id="F2B0D657FF26FF3889ECFD97DEA0FD82" box="[189,451,577,603]" italics="true" pageId="188" pageNumber="189">Trombicula natalensis</emphasis>
</taxonomicName>
<treatmentCitation id="41652C54FF26FF3888B1FD97DD83FD82" author="Lawrence R. F." box="[480,736,576,603]" page="114" pageId="188" pageNumber="189" year="1951">
<bibRefCitation id="A45577B4FF26FF3888B1FD97DD83FD82" author="Lawrence" box="[480,736,576,603]" firstAuthor="Lawrence" page="114" pageId="188" pageNumber="189" pagination="91 - 133" refId="ref64060" refString="Lawrence R. F. 1951 a. New fur-mites from South African mammals. Annals of the Natal Museum 12 (1): 91 - 133." type="journal article" year="1951" yearSuffix="a">Lawrence 1951a: 114</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C07B0A45FF26FF3889ECFDBEDCA9FD5B" blockId="188.[189,1308,576,760]" box="[189,970,616,643]" pageId="188" pageNumber="189">
<treatmentCitationGroup id="E0D42D6BFF26FF3889ECFDBEDCA9FD5B" box="[189,970,616,643]" pageId="188" pageNumber="189">
<taxonomicName id="07C471C6FF26FF3889ECFDBEDD06FD5B" authorityName="Lawrence" authorityYear="1949" box="[189,613,616,642]" class="Arachnida" family="Trombiculidae" genus="Trombicula" kingdom="Animalia" order="Trombidiformes" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="natalensis" subGenus="Trombicula">
<emphasis id="F2B0D657FF26FF3889ECFDBEDD06FD5B" box="[189,613,616,642]" italics="true" pageId="188" pageNumber="189">Trombicula (Trombicula) natalensis</emphasis>
</taxonomicName>
<treatmentCitation id="41652C54FF26FF388BD3FDBEDCA5FD5B" author="Wharton G. W. &amp; Fuller H. S." box="[642,966,616,643]" page="67" pageId="188" pageNumber="189" year="1952">
<bibRefCitation id="A45577B4FF26FF388BD3FDBEDCA5FD5B" author="Wharton &amp; Fuller" box="[642,966,616,643]" firstAuthor="Wharton" page="67" pageId="188" pageNumber="189" pagination="1 - 185" refId="ref69481" refString="Wharton G. W. &amp; Fuller H. S. 1952. A manual of the chiggers. Memoirs of the Entomological Society of Washington 4: 1 - 185." type="journal article" year="1952">Wharton &amp; Fuller 1952: 67</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C07B0A45FF26FF3889ECFD59DC1BFD70" blockId="188.[189,1308,576,760]" box="[189,888,655,681]" pageId="188" pageNumber="189">
<treatmentCitationGroup id="E0D42D6BFF26FF3889ECFD59DC1BFD70" box="[189,888,655,681]" pageId="188" pageNumber="189">
<taxonomicName id="07C471C6FF26FF3889ECFD59DD65FD70" box="[189,518,655,681]" class="Arachnida" family="Trombiculidae" genus="Leptotrombidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="natalensis">
<emphasis id="F2B0D657FF26FF3889ECFD59DD65FD70" box="[189,518,655,681]" italics="true" pageId="188" pageNumber="189">Leptotrombidium natalensis</emphasis>
</taxonomicName>
<treatmentCitation id="41652C54FF26FF388B72FD59DD92FD70" author="Zumpt F." box="[547,753,655,681]" page="141" pageId="188" pageNumber="189" year="1961">
<bibRefCitation id="A45577B4FF26FF388B72FD59DD92FD70" author="Zumpt" box="[547,753,655,681]" firstAuthor="Zumpt" page="141" pageId="188" pageNumber="189" pagination="1 - 457" refId="ref69765" refString="Zumpt F. (ed.) 1961. The arthropod parasites of vertebrates in Africa south of the Sahara (Ethiopian Region). Vol. 1: Chelicerata. Publications of the South African Institute for Medical Research 11 (50): 1 - 457." type="journal article" year="1961">Zumpt 1961: 141</bibRefCitation>
</treatmentCitation>
, fig. 84 (a).
</treatmentCitationGroup>
</paragraph>
<paragraph id="C07B0A45FF26FF3889ECFD60DBF6FD08" blockId="188.[189,1308,576,760]" box="[189,1173,694,721]" pageId="188" pageNumber="189">
<treatmentCitationGroup id="E0D42D6BFF26FF3889ECFD60DBF6FD08" box="[189,1173,694,721]" pageId="188" pageNumber="189">
<taxonomicName id="07C471C6FF26FF3889ECFD60DD87FD09" authorityName="Vercammen-Grandjean &amp; Langston" authorityYear="1976" box="[189,740,694,720]" class="Arachnida" family="Trombiculidae" genus="Leptotrombidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="natalense" subGenus="Leptotrombidium">
<emphasis id="F2B0D657FF26FF3889ECFD60DD87FD09" box="[189,740,694,720]" italics="true" pageId="188" pageNumber="189">Leptotrombidium (Leptotrombidium) natalense</emphasis>
</taxonomicName>
<treatmentCitation id="41652C54FF26FF388A53FD60DBF2FD08" author="Vercammen-Grandjean P. H." box="[770,1169,694,721]" page="54" pageId="188" pageNumber="189" year="1965">
<bibRefCitation id="A45577B4FF26FF388A53FD60DBF2FD08" author="Vercammen-Grandjean" box="[770,1169,694,721]" firstAuthor="Vercammen-Grandjean" page="54" pageId="188" pageNumber="189" refId="ref68052" refString="Vercammen-Grandjean P. H. 1965 c. Trombiculinae of the World. Synopsis with Generic, Subgeneric, and Group Diagnoses (Acarina, Trombiculidae). George Williams Hooper Foundation, San Francisco." type="book" year="1965" yearSuffix="c">Vercammen-Grandjean 1965c: 54</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C07B0A45FF26FF3889ECFD0BDA7FFD21" blockId="188.[189,1308,576,760]" box="[189,1308,733,760]" pageId="188" pageNumber="189">
<treatmentCitationGroup id="E0D42D6BFF26FF3889ECFD0BDA7FFD21" box="[189,1308,733,760]" pageId="188" pageNumber="189">
<taxonomicName id="07C471C6FF26FF3889ECFD0BDD1AFD2E" box="[189,633,733,759]" class="Arachnida" family="Trombiculidae" genus="Chiroptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="natalensis" subGenus="Willmannium">
<emphasis id="F2B0D657FF26FF3889ECFD0BDD1AFD2E" box="[189,633,733,759]" italics="true" pageId="188" pageNumber="189">Chiroptella (Willmannium) natalensis</emphasis>
</taxonomicName>
<treatmentCitation id="41652C54FF26FF388BC7FD0BDBDBFD2E" author="Vercammen-Grandjean P. H. &amp; Langston R. L." box="[662,1208,733,760]" page="911" pageId="188" pageNumber="189" year="1976">
<bibRefCitation id="A45577B4FF26FF388BC7FD0BDBDBFD2E" author="Vercammen-Grandjean &amp; Langston" box="[662,1208,733,760]" firstAuthor="Vercammen-Grandjean" page="911" pageId="188" pageNumber="189" refId="ref68993" refString="Vercammen-Grandjean P. H. &amp; Langston R. L. 1976. The Chigger Mites of the World (Acarina: Trombiculidae et Leeuwenhoekiidae). Vol. III. Leptotrombidium Complex. George Williams Hooper Foundation, University of California, San Francisco, CA." type="book" year="1976">Vercammen-Grandjean &amp; Langston 1976: 911</bibRefCitation>
</treatmentCitation>
, pl. 256.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFF26FF3889ECFCF4DE39FCB6" box="[189,346,802,879]" pageId="188" pageNumber="189" type="materials_examined">
<materialsCitation id="70AC0018FF26FF3889ECFCF4DE39FCB6" box="[189,346,802,879]" collectionCode="NMSA" pageId="188" pageNumber="189" specimenCode="NMSA 4789" specimenCount="1" typeStatus="syntype">
<paragraph id="C07B0A45FF26FF3889ECFCF4DE4EFCE5" blockId="188.[189,301,802,828]" box="[189,301,802,828]" pageId="188" pageNumber="189">
<heading id="9B33BD29FF26FF3889ECFCF4DE4EFCE5" bold="true" box="[189,301,802,828]" fontSize="11" level="3" pageId="188" pageNumber="189" reason="3">
<emphasis id="F2B0D657FF26FF3889ECFCF4DE4EFCE5" bold="true" box="[189,301,802,828]" pageId="188" pageNumber="189">
<typeStatus id="1F7FB4E7FF26FF3889ECFCF4DE4EFCE5" box="[189,301,802,828]" pageId="188" pageNumber="189" type="syntype">Syntypes</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FF26FF3889ECFC83DE39FCB6" blockId="188.[189,346,853,879]" box="[189,346,853,879]" pageId="188" pageNumber="189">
<specimenCode id="9062A23EFF26FF3889ECFC83DE36FCB6" box="[189,341,853,879]" pageId="188" pageNumber="189">NMSA 4789</specimenCode>
.
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="88DE59CEFF26FF3889ECFC4FDB43FC3F" box="[189,1056,921,998]" pageId="188" pageNumber="189" type="distribution">
<paragraph id="C07B0A45FF26FF3889ECFC4FDE35FC6A" blockId="188.[189,342,921,947]" box="[189,342,921,947]" pageId="188" pageNumber="189">
<heading id="9B33BD29FF26FF3889ECFC4FDE35FC6A" bold="true" box="[189,342,921,947]" fontSize="11" level="3" pageId="188" pageNumber="189" reason="3">
<emphasis id="F2B0D657FF26FF3889ECFC4FDE35FC6A" bold="true" box="[189,342,921,947]" pageId="188" pageNumber="189">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FF26FF3889ECFC1DDB43FC3F" blockId="188.[189,1056,971,998]" box="[189,1056,971,998]" pageId="188" pageNumber="189">
<collectingCountry id="B8D34AD5FF26FF3889ECFC1DDE31FC3C" box="[189,338,971,997]" name="South Africa" pageId="188" pageNumber="189">South Africa</collectingCountry>
(Noodsberg, Sterkfontein Caves, Town Bush cave, Howick).
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFF26FF3889ECFBD9DCB4FB82" box="[189,983,1039,1115]" pageId="188" pageNumber="189" type="biology_ecology">
<paragraph id="C07B0A45FF26FF3889ECFBD9DE60FBF0" blockId="188.[189,259,1039,1065]" box="[189,259,1039,1065]" pageId="188" pageNumber="189">
<heading id="9B33BD29FF26FF3889ECFBD9DE60FBF0" bold="true" box="[189,259,1039,1065]" fontSize="11" level="3" pageId="188" pageNumber="189" reason="3">
<emphasis id="F2B0D657FF26FF3889ECFBD9DE60FBF0" bold="true" box="[189,259,1039,1065]" pageId="188" pageNumber="189">Hosts</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FF26FF3889ECFB94DCB4FB82" blockId="188.[189,983,1089,1115]" box="[189,983,1089,1115]" pageId="188" pageNumber="189">
<taxonomicName id="07C471C6FF26FF3889ECFB94DE0DFB82" baseAuthorityName="Temminck" baseAuthorityYear="1832" box="[189,366,1089,1115]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="188" pageNumber="189" phylum="Chordata" rank="species" species="tricolor">
<emphasis id="F2B0D657FF26FF3889ECFB94DE0DFB82" box="[189,366,1089,1115]" italics="true" pageId="188" pageNumber="189">Myotis tricolor</emphasis>
</taxonomicName>
,
<taxonomicName id="07C471C6FF26FF38882DFB94DD30FB82" baseAuthorityName="Pallas" baseAuthorityYear="1766" box="[380,595,1090,1115]" class="Mammalia" family="Procaviidae" genus="Procavia" kingdom="Animalia" order="Hyracoidea" pageId="188" pageNumber="189" phylum="Chordata" rank="species" species="capensis">
<emphasis id="F2B0D657FF26FF38882DFB94DD30FB82" box="[380,595,1090,1115]" italics="true" pageId="188" pageNumber="189">Procavia capensis</emphasis>
</taxonomicName>
,
<emphasis id="F2B0D657FF26FF388B33FB97DCB4FB82" box="[610,983,1089,1115]" italics="true" pageId="188" pageNumber="189">
<taxonomicName id="07C471C6FF26FF388B33FB97DCB1FB82" authorityName="K. Andersen" authorityYear="1904" box="[610,978,1089,1115]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="188" pageNumber="189" phylum="Chordata" rank="subSpecies" species="clivosus" subSpecies="zuluensis">Rhinolophus clivosus zuluensis</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>