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<document id="BEAA907EB82D6669FE4ACA956ED6ADAF" ID-CLB-Dataset="6214" ID-DOI="10.11646/zootaxa.4974.3.2" ID-GBIF-Dataset="5b4eaeb3-5722-4021-84ad-125ccd80855f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4777839" ID-ZooBank="1BD6220B-7540-4A12-B3D2-5EDC1A3B7DD8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621588618725" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="4772526A005AFFDEFF1829F5108F3229" docLanguage="en" docName="zootaxa.4974.3.2.pdf" docOrigin="Zootaxa 4974 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus foveatus Haldeman" docType="treatment" docVersion="6" lastPageNumber="476" masterDocId="BB4B2A12004AFFCFFF8F2D18151D3030" masterDocTitle="Taxonomy of Central American water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="503" masterPageNumber="459" pageNumber="475" updateTime="1698940277359" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="ECD82815B8F7BC4912891FE33BF094A6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="85F524C4B996CFF1CA1EF7348183D1C8">
<mods:title id="97B4F257E16EEB7D66B657962F220C9C">Taxonomy of Central American water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)</mods:title>
</mods:titleInfo>
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<mods:roleTerm id="3237AD9A4797CAFD804EA61E993CB188">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="51217DA96AB10EBBF52F72C5A9E15F6A">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="8C665560B3D942C10CCE1E8FDC2E3BDA">text</mods:typeOfResource>
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<mods:title id="8D46CEAB0D424981E4F5BCDB52D5D03D">Zootaxa</mods:title>
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<mods:part id="BE4374BE77905FFF1D8039EF5C96C38A">
<mods:date id="EF6239FD32C04B5FE9A990C7A389893A">2021</mods:date>
<mods:detail id="9191B620190A00CD370FBD691AB4B827" type="pubDate">
<mods:number id="AAAC18456781839CE572039FB9CDC5E0">2021-05-21</mods:number>
</mods:detail>
<mods:detail id="C6D7974B26069E3B268B6F8FD133DFCE" type="volume">
<mods:number id="66FACA906841EEEF54959494BA4BC80E">4974</mods:number>
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<mods:number id="B9448568608B6D6F9274E55EB8E90060">3</mods:number>
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<mods:classification id="1CED69D836DC138F75550BD192D193B7">journal article</mods:classification>
<mods:identifier id="EE34F502C955C2EAD69226039591D886" type="CLB-Dataset">6214</mods:identifier>
<mods:identifier id="A0724A3B959DBEECCFED68484D5571F2" type="DOI">10.11646/zootaxa.4974.3.2</mods:identifier>
<mods:identifier id="F755EEA961C4DDF4FD23F30E71B23965" type="GBIF-Dataset">5b4eaeb3-5722-4021-84ad-125ccd80855f</mods:identifier>
<mods:identifier id="3D04FBEAE3E5A34CE3237FFDB5F12385" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="06D4FD3ED852D61B1BDA51CCDB412D95" type="Zenodo-Dep">4777839</mods:identifier>
<mods:identifier id="D62BF4AD948171604F4303714684516B" type="ZooBank">1BD6220B-7540-4A12-B3D2-5EDC1A3B7DD8</mods:identifier>
</mods:mods>
<treatment id="4772526A005AFFDEFF1829F5108F3229" ID-DOI="http://doi.org/10.5281/zenodo.4777926" ID-GBIF-Taxon="182642559" ID-Zenodo-Dep="4777926" LSID="urn:lsid:plazi:treatment:4772526A005AFFDEFF1829F5108F3229" httpUri="http://treatment.plazi.org/id/4772526A005AFFDEFF1829F5108F3229" lastPageId="17" lastPageNumber="476" pageId="16" pageNumber="475">
<subSubSection id="87C1B0F7005AFFDFFF1829F5170E3537" box="[151,531,1261,1288]" pageId="16" pageNumber="475" type="nomenclature">
<paragraph id="CF64E37C005AFFDFFF1829F5170E3537" blockId="16.[151,531,1261,1324]" box="[151,531,1261,1288]" pageId="16" pageNumber="475">
<heading id="942C5410005AFFDFFF1829F5170E3537" bold="true" box="[151,531,1261,1288]" fontSize="11" level="1" pageId="16" pageNumber="475" reason="1">
<taxonomicName id="08DB98FF005AFFDFFF1829F5170E3537" ID-CoL="6MK9D" authority="Haldeman" authorityName="Haldeman" box="[151,531,1261,1288]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="475" phylum="Arthropoda" rank="species" species="foveatus">
<emphasis id="FDAF3F6E005AFFDFFF1829F5170E3537" bold="true" box="[151,531,1261,1288]" pageId="16" pageNumber="475">
<emphasis id="FDAF3F6E005AFFDFFF1829F514943537" bold="true" box="[151,393,1261,1288]" italics="true" pageId="16" pageNumber="475">Hydrochus foveatus</emphasis>
Haldeman
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="87C1B0F7005AFFDFFF182809147C351B" box="[151,353,1297,1324]" pageId="16" pageNumber="475" type="description">
<paragraph id="CF64E37C005AFFDFFF182809147C351B" blockId="16.[151,531,1261,1324]" box="[151,353,1297,1324]" pageId="16" pageNumber="475">
<figureCitation id="57E0FFF9005AFFDFFF18280915F8351B" box="[151,229,1297,1323]" captionStart="FIGURE 2" captionStartId="5.[151,250,1989,2013]" captionTargetBox="[181,1393,181,1953]" captionTargetId="figure-15@5.[158,1429,181,1965]" captionTargetPageId="5" captionText="FIGURE 2. Dorsal habitus of Central American Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.4777847" httpUri="https://zenodo.org/record/4777847/files/figure.png" pageId="16" pageNumber="475">Figs. 2</figureCitation>
,
<figureCitation id="57E0FFF9005AFFDFFF7E28091424351B" box="[241,313,1297,1324]" captionStart="FIGURES 15–16" captionStartId="29.[151,264,1944,1968]" captionTargetBox="[181,1432,185,1916]" captionTargetId="figure-15@29.[151,1435,181,1920]" captionTargetPageId="29" captionText="FIGURES 15–16. Habitus and male genitalia. 15. Hydrochus foveatus Haldeman, non-type. 16. Hydrochus foveatus Haldeman, lectotype." figureDoi="http://doi.org/10.5281/zenodo.4777875" httpUri="https://zenodo.org/record/4777875/files/figure.png" pageId="16" pageNumber="475">15, 16</figureCitation>
,
<figureCitation id="57E0FFF9005AFFDFFECA2809147C351B" box="[325,353,1297,1323]" captionStart="FIGURES 34–36" captionStartId="37.[151,264,1943,1967]" captionTargetBox="[166,1421,181,1920]" captionTargetId="figure-15@37.[166,1421,181,1920]" captionTargetPageId="37" captionText="FIGURES 34–36. Geographical distribution maps." figureDoi="http://doi.org/10.5281/zenodo.4777912" httpUri="https://zenodo.org/record/4777912/files/figure.png" pageId="16" pageNumber="475">36</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="87C1B0F7005AFFDFFF18284E163335BC" pageId="16" pageNumber="475" type="reference_group">
<paragraph id="CF64E37C005AFFDFFF18284E1138355E" blockId="16.[151,1061,1366,1421]" box="[151,1061,1366,1390]" pageId="16" pageNumber="475">
<taxonomicName id="08DB98FF005AFFDFFF18284E1759355E" ID-CoL="6MK9D" authority="Haldeman, 1852: 375" authorityName="Haldeman" authorityPageNumber="375" authorityYear="1852" box="[151,580,1366,1390]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="475" phylum="Arthropoda" rank="species" species="foveatus">
<emphasis id="FDAF3F6E005AFFDFFF18284E147D355E" box="[151,352,1366,1390]" italics="true" pageId="16" pageNumber="475">Hydrochus foveatus</emphasis>
Haldeman, 1852: 375
</taxonomicName>
;
<typeStatus id="10605DDE005AFFDFFDC0284E17B1355E" box="[591,684,1366,1390]" pageId="16" pageNumber="475" type="lectotype">lectotype</typeStatus>
designation
<bibRefCitation id="AB4A9E8D005AFFDFFCA2284E113C355E" author="Worthington, R. J. &amp; Hellman, J. L. &amp; Lago, P. K." box="[813,1057,1366,1390]" pageId="16" pageNumber="475" pagination="167 - 213" refId="ref25748" refString="Worthington, R. J., Hellman, J. L. &amp; Lago, P. K. (2016) Hydrochidae (Coleoptera) of Mississippi. Transactions of the American Entomological Society, 142, 167 - 213. https: // doi. org / 10.3157 / 061.142.0206" type="journal article" year="2016">
Worthington
<emphasis id="FDAF3F6E005AFFDFFC3D284F16FB355E" box="[946,998,1366,1390]" italics="true" pageId="16" pageNumber="475">et al.</emphasis>
2016
</bibRefCitation>
.
</paragraph>
<paragraph id="CF64E37C005AFFDFFF18286C163335BC" blockId="16.[151,1061,1366,1421]" box="[151,814,1396,1421]" pageId="16" pageNumber="475">
<taxonomicName id="08DB98FF005AFFDFFF18286C14E535BC" ID-CoL="3N69P" authority="Makhan, 1995" authorityName="Makhan" authorityYear="1995" box="[151,504,1396,1420]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="475" phylum="Arthropoda" rank="species" species="monishi">
<emphasis id="FDAF3F6E005AFFDFFF18286C144135BC" box="[151,348,1396,1420]" italics="true" pageId="16" pageNumber="475">Hydrochus monishi</emphasis>
Makhan, 1995
</taxonomicName>
. Syn.
<bibRefCitation id="AB4A9E8D005AFFDFFDBA286C163735BC" author="Worthington, R. J. &amp; Hellman, J. L. &amp; Lago, P. K." box="[565,810,1396,1420]" pageId="16" pageNumber="475" pagination="167 - 213" refId="ref25748" refString="Worthington, R. J., Hellman, J. L. &amp; Lago, P. K. (2016) Hydrochidae (Coleoptera) of Mississippi. Transactions of the American Entomological Society, 142, 167 - 213. https: // doi. org / 10.3157 / 061.142.0206" type="journal article" year="2016">
Worthington
<emphasis id="FDAF3F6E005AFFDFFD34286D17F335BC" box="[699,750,1396,1420]" italics="true" pageId="16" pageNumber="475">et al.</emphasis>
2016
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="87C1B0F7005AFFDFFF1828A3114635E5" box="[151,1115,1467,1493]" pageId="16" pageNumber="475" type="description">
<paragraph id="CF64E37C005AFFDFFF1828A3114635E5" blockId="16.[151,1437,1467,2034]" box="[151,1115,1467,1493]" pageId="16" pageNumber="475">
Originally described from Texas; the following are new locality records from
<collectingCountry id="B7CCA3EC005AFFDFFC7128A3114A35E5" box="[1022,1111,1467,1493]" name="Mexico" pageId="16" pageNumber="475">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="87C1B0F7005AFFDFFF4828C6117936C5" pageId="16" pageNumber="475" type="materials_examined">
<paragraph id="CF64E37C005AFFDFFF4828C6117936C5" blockId="16.[151,1437,1467,2034]" pageId="16" pageNumber="475">
<emphasis id="FDAF3F6E005AFFDFFF4828C614F435C8" bold="true" box="[199,489,1502,1528]" pageId="16" pageNumber="475">Material examined (17)</emphasis>
:
<materialsCitation id="7FB3E921005AFFDFFE7128C617D7362D" ID-GBIF-Occurrence="3128194343" collectingDate="1991-03-22" collectionCode="SEMC" collectorName="R. Brooks &amp; R. Leschen" country="Mexico" elevation="430" location="10 km N Linares" pageId="16" pageNumber="475" specimenCount="1" stateProvince="Nuevo Leon">
<emphasis id="FDAF3F6E005AFFDFFE7128C6174835C8" bold="true" box="[510,597,1502,1528]" pageId="16" pageNumber="475">
<collectingCountry id="B7CCA3EC005AFFDFFE7128C6174835C8" box="[510,597,1502,1528]" name="Mexico" pageId="16" pageNumber="475">Mexico</collectingCountry>
</emphasis>
:
<collectingRegion id="0D1F2D9E005AFFDFFDE628C717E135C9" box="[617,764,1503,1529]" country="Mexico" name="Nuevo Leon" pageId="16" pageNumber="475">Nuevo Leon</collectingRegion>
,
<location id="CA04B5A7005AFFDFFC8228C716C335C9" LSID="urn:lsid:plazi:treatment:4772526A005AFFDEFF1829F5108F3229:CA04B5A7005AFFDFFC8228C716C335C9" box="[781,990,1503,1529]" country="Mexico" name="10 km N Linares" pageId="16" pageNumber="475" stateProvince="Nuevo Leon">
<quantity id="08234E99005AFFDFFC8228C7164435C9" box="[781,857,1503,1529]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="475" unit="km" value="10.0">10 km</quantity>
N Linares
</location>
; ex. mercury vapor lite, elev.
<quantity id="08234E99005AFFDFFAC628C7108A35C9" box="[1353,1431,1503,1529]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="16" pageNumber="475" unit="m" value="430.0">
<elevation id="44F6044F005AFFDFFAC628C7108A35C9" box="[1353,1431,1503,1529]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="16" pageNumber="475" unit="m" value="430.0">430 m</elevation>
</quantity>
,
<date id="BB65C5BC005AFFDFFF182B1B140E362D" box="[151,275,1539,1565]" pageId="16" pageNumber="475" value="1991-03-22">
<collectingDate id="AB213C54005AFFDFFF182B1B140E362D" box="[151,275,1539,1565]" pageId="16" pageNumber="475" value="1991-03-22">22.iii.1991</collectingDate>
</date>
, leg.
<collectorName id="622E86AA005AFFDFFEC02B1B14DE362D" box="[335,451,1539,1565]" pageId="16" pageNumber="475">R. Brooks</collectorName>
,
<collectorName id="622E86AA005AFFDFFE412B1B1751362D" box="[462,588,1539,1565]" pageId="16" pageNumber="475">R. Leschen</collectorName>
(3
<collectionCode id="A9CA7BB9005AFFDFFDFE2B1B17DF362D" box="[625,706,1539,1565]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="16" pageNumber="475" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921005AFFDFFD5A2B1B14073671" ID-GBIF-Occurrence="3128194365" collectingDate="1968-07-19" collectionCode="NMNH" collectorName="F. N. Young" country="Mexico" location="Rio Ramos" pageId="16" pageNumber="475" specimenCount="1" stateProvince="Nuevo Leon">
<collectingRegion id="0D1F2D9E005AFFDFFD5A2B1B167E362D" box="[725,867,1539,1565]" country="Mexico" name="Nuevo Leon" pageId="16" pageNumber="475">Nuevo Leon</collectingRegion>
,
<location id="CA04B5A7005AFFDFFCE12B1B16F2362D" LSID="urn:lsid:plazi:treatment:4772526A005AFFDEFF1829F5108F3229:CA04B5A7005AFFDFFCE12B1B16F2362D" box="[878,1007,1539,1565]" country="Mexico" name="Rio Ramos" pageId="16" pageNumber="475" stateProvince="Nuevo Leon">Rio Ramos</location>
,
<date id="BB65C5BC005AFFDFFC762B1B1166362D" box="[1017,1147,1539,1565]" pageId="16" pageNumber="475" value="1968-07-19">
<collectingDate id="AB213C54005AFFDFFC762B1B1166362D" box="[1017,1147,1539,1565]" pageId="16" pageNumber="475" value="1968-07-19">19.vii.1968</collectingDate>
</date>
, leg.
<collectorName id="622E86AA005AFFDFFB382B1B1021362D" box="[1207,1340,1539,1565]" pageId="16" pageNumber="475">F. N. Young</collectorName>
(#2304) (1
<collectionCode id="A9CA7BB9005AFFDFFF382B3F140F3671" box="[183,274,1575,1601]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="475" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921005AFFDFFEA92B3F11643671" ID-GBIF-Occurrence="3128194594" box="[294,1145,1574,1602]" collectingDate="1954-08-18" collectionCode="NMNH" collectorName="F. N. Young" country="Mexico" location="Allende" municipality="Rio Ramos" pageId="16" pageNumber="475" specimenCount="1">
<collectingMunicipality id="2F007906005AFFDFFEA92B3F14B83671" box="[294,421,1575,1601]" pageId="16" pageNumber="475">Rio Ramos</collectingMunicipality>
near
<location id="CA04B5A7005AFFDFFE6B2B3F175D3671" LSID="urn:lsid:plazi:treatment:4772526A005AFFDEFF1829F5108F3229:CA04B5A7005AFFDFFE6B2B3F175D3671" box="[484,576,1575,1601]" country="Mexico" municipality="Rio Ramos" name="Allende" pageId="16" pageNumber="475">Allende</location>
,
<date id="BB65C5BC005AFFDFFDC32B3E17C83671" box="[588,725,1574,1601]" pageId="16" pageNumber="475" value="1954-08-18">
<collectingDate id="AB213C54005AFFDFFDC32B3E17C83671" box="[588,725,1574,1601]" pageId="16" pageNumber="475" value="1954-08-18">18.viii.1954</collectingDate>
</date>
, leg.
<collectorName id="622E86AA005AFFDFFC9B2B3F16813671" box="[788,924,1575,1601]" pageId="16" pageNumber="475">F. N. Young</collectorName>
(1106) (1
<collectionCode id="A9CA7BB9005AFFDFFB992B3F116C3671" box="[1046,1137,1575,1601]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="475" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921005AFFDFFB0A2B3F16103655" ID-GBIF-Occurrence="3128194319" collectingDate="1966-06-28" collectionCode="NMNH" collectorName="R. E. Woodruff" country="Mexico" location="Monterrey" municipality="Siesta Motel" pageId="16" pageNumber="475" specimenCount="1">
<collectingMunicipality id="2F007906005AFFDFFB0A2B3F100B3671" box="[1157,1302,1575,1601]" pageId="16" pageNumber="475">Siesta Motel</collectingMunicipality>
,
<quantity id="08234E99005AFFDFFAAC2B3F10433671" box="[1315,1374,1575,1601]" metricMagnitude="3" metricUnit="m" metricValue="9.656064" pageId="16" pageNumber="475" unit="mi" value="6.0">6 mi.</quantity>
S. of
<location id="CA04B5A7005AFFDFFF182B5314133655" LSID="urn:lsid:plazi:treatment:4772526A005AFFDEFF1829F5108F3229:CA04B5A7005AFFDFFF182B5314133655" box="[151,270,1611,1637]" country="Mexico" municipality="Siesta Motel" name="Monterrey" pageId="16" pageNumber="475">Monterrey</location>
,
<date id="BB65C5BC005AFFDFFE942B5314883655" box="[283,405,1611,1637]" pageId="16" pageNumber="475" value="1966-06-28">
<collectingDate id="AB213C54005AFFDFFE942B5314883655" box="[283,405,1611,1637]" pageId="16" pageNumber="475" value="1966-06-28">28.vi.1966</collectingDate>
</date>
, leg.
<collectorName id="622E86AA005AFFDFFE5B2B53179F3655" box="[468,642,1611,1637]" pageId="16" pageNumber="475">R. E. Woodruff</collectorName>
(2
<collectionCode id="A9CA7BB9005AFFDFFD252B5316183655" box="[682,773,1611,1637]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="475" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921005AFFDFFC962B53149E36B9" ID-GBIF-Occurrence="3128194391" collectingDate="1964-08-22" collectionCode="NMNH" collectorName="Paul J. Spangler" country="Mexico" location="Ciudad Valles" pageId="16" pageNumber="475" specimenCount="1" stateProvince="San Luis Potosi">
<collectingRegion id="0D1F2D9E005AFFDFFC962B5316D23655" box="[793,975,1611,1637]" country="Mexico" name="San Luis Potosi" pageId="16" pageNumber="475">San Luis Potosi</collectingRegion>
,
<location id="CA04B5A7005AFFDFFC542B5311663655" LSID="urn:lsid:plazi:treatment:4772526A005AFFDEFF1829F5108F3229:CA04B5A7005AFFDFFC542B5311663655" box="[987,1147,1611,1637]" country="Mexico" name="Ciudad Valles" pageId="16" pageNumber="475" stateProvince="San Luis Potosi">Ciudad Valles</location>
,
<date id="BB65C5BC005AFFDFFB072B53100F3655" box="[1160,1298,1611,1637]" pageId="16" pageNumber="475" value="1964-08-22">
<collectingDate id="AB213C54005AFFDFFB072B53100F3655" box="[1160,1298,1611,1637]" pageId="16" pageNumber="475" value="1964-08-22">22.viii.1964</collectingDate>
</date>
, leg.
<collectorName id="622E86AA005AFFDFFADF2B5315E636B9" pageId="16" pageNumber="475">Paul J. Spangler</collectorName>
(1
<collectionCode id="A9CA7BB9005AFFDFFEAF2B77146636B9" box="[288,379,1647,1673]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="475" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921005AFFDFFE012B77108A36B9" ID-GBIF-Occurrence="3128194345" box="[398,1431,1646,1673]" collectingDate="1948-12-22" collectionCode="CAS" collectorName="Hugh B. Leech" country="Mexico" county="Clear" location="Tamazunchale" municipality="Palitla" pageId="16" pageNumber="475" specimenCount="1">
<collectingCounty id="26059BF0005AFFDFFE012B7714D636B9" box="[398,459,1647,1673]" pageId="16" pageNumber="475">Clear</collectingCounty>
stream at
<collectingMunicipality id="2F007906005AFFDFFDB02B77179436B9" box="[575,649,1647,1673]" pageId="16" pageNumber="475">Palitla</collectingMunicipality>
,
<quantity id="08234E99005AFFDFFD1A2B7617D236B9" box="[661,719,1646,1673]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="16" pageNumber="475" unit="mi" value="5.0">5 mi.</quantity>
N. of
<location id="CA04B5A7005AFFDFFC982B7716A636B9" LSID="urn:lsid:plazi:treatment:4772526A005AFFDEFF1829F5108F3229:CA04B5A7005AFFDFFC982B7716A636B9" box="[791,955,1647,1673]" country="Mexico" county="Clear" municipality="Palitla" name="Tamazunchale" pageId="16" pageNumber="475">Tamazunchale</location>
,
<date id="BB65C5BC005AFFDFFC492B77115536B9" box="[966,1096,1647,1673]" pageId="16" pageNumber="475" value="1948-12-22">
<collectingDate id="AB213C54005AFFDFFC492B77115536B9" box="[966,1096,1647,1673]" pageId="16" pageNumber="475" value="1948-12-22">22.xii.1948</collectingDate>
</date>
, leg.
<collectorName id="622E86AA005AFFDFFB0B2B77103336B9" box="[1156,1326,1647,1673]" pageId="16" pageNumber="475">Hugh B. Leech</collectorName>
(1
<collectionCode id="A9CA7BB9005AFFDFFADA2B77108D36B9" box="[1365,1424,1647,1673]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="16" pageNumber="475" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921005AFFDFFF182B8B163C369D" ID-GBIF-Occurrence="3128194393" box="[151,801,1682,1709]" collectingDate="1938-09-25" collectionCode="SEMC" collectorName="L. J. Lipovsky" country="Mexico" location="Huichihuayan" pageId="16" pageNumber="475" specimenCount="1">
<location id="CA04B5A7005AFFDFFF182B8B142A369D" LSID="urn:lsid:plazi:treatment:4772526A005AFFDEFF1829F5108F3229:CA04B5A7005AFFDFFF182B8B142A369D" box="[151,311,1683,1709]" country="Mexico" name="Huichihuayan" pageId="16" pageNumber="475">Huichihuayan</location>
,
<date id="BB65C5BC005AFFDFFECC2B8A14A3369D" box="[323,446,1682,1709]" pageId="16" pageNumber="475" value="1938-09-25">
<collectingDate id="AB213C54005AFFDFFECC2B8A14A3369D" box="[323,446,1682,1709]" pageId="16" pageNumber="475" value="1938-09-25">25.ix.1938</collectingDate>
</date>
, leg.
<collectorName id="622E86AA005AFFDFFE732B8B17BD369D" box="[508,672,1683,1709]" pageId="16" pageNumber="475">L. J. Lipovsky</collectorName>
(1
<collectionCode id="A9CA7BB9005AFFDFFD472B8B1604369D" box="[712,793,1683,1709]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="16" pageNumber="475" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921005AFFDFFCA22B8B17BA36E1" ID-GBIF-Occurrence="3128194323" collectingDate="1970-07-19" collectionCode="NMNH" collectorName="Shaffner &amp; Hart &amp; Murray &amp; Phelps" country="Mexico" location="Gomez Farias" pageId="16" pageNumber="475" specimenCount="1" stateProvince="Tamaulipas">
<collectingRegion id="0D1F2D9E005AFFDFFCA22B8B16AC369D" box="[813,945,1683,1709]" country="Mexico" name="Tamaulipas" pageId="16" pageNumber="475">Tamaulipas</collectingRegion>
,
<quantity id="08234E99005AFFDFFC322B8A110D369D" box="[957,1040,1682,1709]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="16" pageNumber="475" unit="mi" value="5.0">5 miles</quantity>
sse. of
<location id="CA04B5A7005AFFDFFBE62B8B1014369D" LSID="urn:lsid:plazi:treatment:4772526A005AFFDEFF1829F5108F3229:CA04B5A7005AFFDFFBE62B8B1014369D" box="[1129,1289,1683,1709]" country="Mexico" name="Gomez Farias" pageId="16" pageNumber="475" stateProvince="Tamaulipas">Gomez Farias</location>
,
<date id="BB65C5BC005AFFDFFA992B8B1085369D" box="[1302,1432,1683,1709]" pageId="16" pageNumber="475" value="1970-07-19">
<collectingDate id="AB213C54005AFFDFFA992B8B1085369D" box="[1302,1432,1683,1709]" pageId="16" pageNumber="475" value="1970-07-19">19.vii.1970</collectingDate>
</date>
, leg.
<collectorName id="622E86AA005AFFDFFF482BAF143436E1" box="[199,297,1719,1745]" pageId="16" pageNumber="475">Shaffner</collectorName>
,
<collectorName id="622E86AA005AFFDFFEBB2BAF147436E1" box="[308,361,1719,1745]" pageId="16" pageNumber="475">Hart</collectorName>
,
<collectorName id="622E86AA005AFFDFFEFB2BAF14D436E1" box="[372,457,1719,1745]" pageId="16" pageNumber="475">Murray</collectorName>
,
<collectorName id="622E86AA005AFFDFFE5B2BAF170236E1" box="[468,543,1719,1745]" pageId="16" pageNumber="475">Phelps</collectorName>
(1
<collectionCode id="A9CA7BB9005AFFDFFDCA2BAF17BD36E1" box="[581,672,1719,1745]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="475" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921005AFFDFFD3D2BAF15E036C5" ID-GBIF-Occurrence="3128194510" collectingDate="1983-03-16" collectionCode="UMRM" collectorName="W. El Encino &amp; M. S. Davis" country="Mexico" location="Gomez Farias" pageId="16" pageNumber="475" specimenCount="1">
<location id="CA04B5A7005AFFDFFD3D2BAF164C36E1" LSID="urn:lsid:plazi:treatment:4772526A005AFFDEFF1829F5108F3229:CA04B5A7005AFFDFFD3D2BAF164C36E1" box="[690,849,1719,1745]" country="Mexico" name="Gomez Farias" pageId="16" pageNumber="475">Gomez Farias</location>
,
<quantity id="08234E99005AFFDFFCD22BAF168A36E1" box="[861,919,1719,1745]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="16" pageNumber="475" unit="mi" value="3.0">3 mi.</quantity>
<collectorName id="622E86AA005AFFDFFC122BAF112836E1" box="[925,1077,1719,1745]" pageId="16" pageNumber="475">W. El Encino</collectorName>
,
<date id="BB65C5BC005AFFDFFBCF2BAF11A136E1" box="[1088,1212,1719,1745]" pageId="16" pageNumber="475" value="1983-03-16">
<collectingDate id="AB213C54005AFFDFFBCF2BAF11A136E1" box="[1088,1212,1719,1745]" pageId="16" pageNumber="475" value="1983-03-16">16.iii.1983</collectingDate>
</date>
, leg.
<collectorName id="622E86AA005AFFDFFB772BAF106036E1" box="[1272,1405,1719,1745]" pageId="16" pageNumber="475">M. S. Davis</collectorName>
(2
<collectionCode id="A9CA7BB9005AFFDFFF182BC315E836C5" box="[151,245,1755,1781]" country="USA" httpUri="http://grbio.org/cool/w4qt-0wzz" name="W.R. Enns Entomology Museum" pageId="16" pageNumber="475">UMRM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7FB3E921005AFFDFFE882BC3117C36C5" ID-GBIF-Occurrence="3128194397" box="[263,1121,1754,1781]" collectingDate="1965-08-27" collectionCode="NMNH" collectorName="W. Poza Rica &amp; Paul J. Spangler" country="Mexico" location="Veracruz" pageId="16" pageNumber="475" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
<collectingRegion id="0D1F2D9E005AFFDFFE882BC3147336C5" box="[263,366,1755,1781]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="16" pageNumber="475">Veracruz</collectingRegion>
,
<quantity id="08234E99005AFFDFFEF62BC314AE36C5" box="[377,435,1755,1781]" metricMagnitude="4" metricUnit="m" metricValue="1.4484096" pageId="16" pageNumber="475" unit="mi" value="9.0">9 mi.</quantity>
<collectorName id="622E86AA005AFFDFFE362BC3174F36C5" box="[441,594,1755,1781]" pageId="16" pageNumber="475">W. Poza Rica</collectorName>
,
<date id="BB65C5BC005AFFDFFDD12BC217FA36C5" box="[606,743,1754,1781]" pageId="16" pageNumber="475" value="1965-08-27">
<collectingDate id="AB213C54005AFFDFFDD12BC217FA36C5" box="[606,743,1754,1781]" pageId="16" pageNumber="475" value="1965-08-27">27.viii.1965</collectingDate>
</date>
, leg.
<collectorName id="622E86AA005AFFDFFCAC2BC316C436C5" box="[803,985,1755,1781]" pageId="16" pageNumber="475">Paul J. Spangler</collectorName>
(4
<collectionCode id="A9CA7BB9005AFFDFFC712BC3114436C5" box="[1022,1113,1755,1781]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="475" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="87C1B0F7005AFFDEFF482BE611A5319D" lastPageId="17" lastPageNumber="476" pageId="16" pageNumber="475" type="diagnosis">
<paragraph id="CF64E37C005AFFDFFF482BE614BD37FD" blockId="16.[151,1437,1467,2034]" pageId="16" pageNumber="475">
<emphasis id="FDAF3F6E005AFFDFFF482BE614D03728" bold="true" box="[199,461,1790,1816]" pageId="16" pageNumber="475">Differential diagnosis.</emphasis>
Among Central American species, recognized by the combination of moderate size (ca.
<quantity id="08234E99005AFFDFFF182A3A1423370D" box="[151,318,1826,1853]" metricMagnitude="-3" metricUnit="m" metricValue="3.53" metricValueMax="3.56" metricValueMin="3.5" pageId="16" pageNumber="475" unit="mm" value="3.53" valueMax="3.56" valueMin="3.5">3.50–3.56 mm</quantity>
), the deep pronotal foveae, the large elytral tubercle of the 5
<superScript id="38AE4E34005AFFDFFC542A3A16F53700" attach="left" box="[987,1000,1826,1840]" fontSize="6" pageId="16" pageNumber="475">th</superScript>
interstria, and the male genitalia (
<figureCitation id="57E0FFF9005AFFDFFAEB2A3B15FF3751" captionStart="FIGURES 15–16" captionStartId="29.[151,264,1944,1968]" captionTargetBox="[181,1432,185,1916]" captionTargetId="figure-15@29.[151,1435,181,1920]" captionTargetPageId="29" captionText="FIGURES 15–16. Habitus and male genitalia. 15. Hydrochus foveatus Haldeman, non-type. 16. Hydrochus foveatus Haldeman, lectotype." figureDoi="http://doi.org/10.5281/zenodo.4777875" httpUri="https://zenodo.org/record/4777875/files/figure.png" pageId="16" pageNumber="475">Figs. 15, 16</figureCitation>
; described below). This species has a generalized
<taxonomicName id="08DB98FF005AFFDFFCA92A5F16BD3751" authorityName="Leach" authorityYear="1817" box="[806,928,1863,1889]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="FDAF3F6E005AFFDFFCA92A5F16BD3751" box="[806,928,1863,1889]" italics="true" pageId="16" pageNumber="475">Hydrochus</emphasis>
</taxonomicName>
habitus; reliable determinations will require dissection of males. The male genitalia are distinct from other Central American species by virtue of the following combinations of characters: genitalia large, paramere tips sagittate, adtl very near apex of aedeagus, aedeagus wide, pseudoparameres large.
</paragraph>
<paragraph id="CF64E37C005AFFDEFF482ACF11EB3171" blockId="16.[151,1437,1467,2034]" lastBlockId="17.[151,1436,151,537]" lastPageId="17" lastPageNumber="476" pageId="16" pageNumber="475">
Male genitalia (
<figureCitation id="57E0FFF9005AFFDFFEF62ACF171E37C2" box="[377,515,2006,2034]" captionStart="FIGURES 15–16" captionStartId="29.[151,264,1944,1968]" captionTargetBox="[181,1432,185,1916]" captionTargetId="figure-15@29.[151,1435,181,1920]" captionTargetPageId="29" captionText="FIGURES 15–16. Habitus and male genitalia. 15. Hydrochus foveatus Haldeman, non-type. 16. Hydrochus foveatus Haldeman, lectotype." figureDoi="http://doi.org/10.5281/zenodo.4777875" httpUri="https://zenodo.org/record/4777875/files/figure.png" pageId="16" pageNumber="475">Figs. 15, 16</figureCitation>
). General characters: paramere tips sagittate, plsa prominent, narrower at very tip end in lateral view than in dorsal/ventral views, in dorsal/ventral views with medial margins at apex parallel or nearly so, then arcuate away from midline; aedeagus in dorsal/ventral views lanceolate, very wide at agw, at ca. midlength; in lateral view slightly narrower, without emargination where adbl and shaft meet; lateral margins of aedeagus sclerotized; basal piece slightly shorter than parameres, in dorsal/ventral views tapering from distal to proximal, in lateral view dorsal surface arcuate, ventral surface straight, orifice strongly sclerotized.
</paragraph>
<paragraph id="CF64E37C005BFFDEFF482C53142731B9" blockId="17.[151,1436,151,537]" pageId="17" pageNumber="476">Dorsal surface: pdmm moderately widely separated, slightly sinuate, overlapping aedeagus; adbl very small; adtl subapical.</paragraph>
<paragraph id="CF64E37C005BFFDEFF482C8B11A5319D" blockId="17.[151,1436,151,537]" box="[199,1208,403,429]" pageId="17" pageNumber="476">Ventral surface: pvmm approximating lateral margin of aedeagus; avtl at ca. proximal 1/3.</paragraph>
</subSubSection>
<subSubSection id="87C1B0F7005BFFDEFF482CAF108F3229" pageId="17" pageNumber="476" type="discussion">
<paragraph id="CF64E37C005BFFDEFF482CAF108F3229" blockId="17.[151,1436,151,537]" pageId="17" pageNumber="476">
<emphasis id="FDAF3F6E005BFFDEFF482CAF142631E1" bold="true" box="[199,315,439,465]" pageId="17" pageNumber="476">Remarks.</emphasis>
The
<specimenCount id="D9DD28F5005BFFDEFEF52CAF170B31E1" box="[378,534,439,465]" pageId="17" pageNumber="476" type="generic">one specimen</specimenCount>
in the LeConte collection at the MCZ, a female, was designated
<typeStatus id="10605DDE005BFFDEFA9F2CAF106A31E1" box="[1296,1399,439,465]" pageId="17" pageNumber="476" type="lectotype">lectotype</typeStatus>
by
<bibRefCitation id="AB4A9E8D005BFFDEFF182CC314A731C6" author="Worthington, R. J. &amp; Hellman, J. L. &amp; Lago, P. K." box="[151,442,475,502]" pageId="17" pageNumber="476" pagination="167 - 213" refId="ref25748" refString="Worthington, R. J., Hellman, J. L. &amp; Lago, P. K. (2016) Hydrochidae (Coleoptera) of Mississippi. Transactions of the American Entomological Society, 142, 167 - 213. https: // doi. org / 10.3157 / 061.142.0206" type="journal article" year="2016">
Worthington
<emphasis id="FDAF3F6E005BFFDEFEA32CC3147831C5" box="[300,357,475,501]" italics="true" pageId="17" pageNumber="476">et al.</emphasis>
(2016)
</bibRefCitation>
. Images of the habitus and labels of this specimen are given herein (
<figureCitation id="57E0FFF9005BFFDEFB312CC3100F31C6" box="[1214,1298,475,502]" captionStart="FIGURES 15–16" captionStartId="29.[151,264,1944,1968]" captionTargetBox="[181,1432,185,1916]" captionTargetId="figure-15@29.[151,1435,181,1920]" captionTargetPageId="29" captionText="FIGURES 15–16. Habitus and male genitalia. 15. Hydrochus foveatus Haldeman, non-type. 16. Hydrochus foveatus Haldeman, lectotype." figureDoi="http://doi.org/10.5281/zenodo.4777875" httpUri="https://zenodo.org/record/4777875/files/figure.png" pageId="17" pageNumber="476">Fig. 16</figureCitation>
). The specimen imaged (
<figureCitation id="57E0FFF9005BFFDEFEBF2CE7149F3229" box="[304,386,510,537]" captionStart="FIGURES 15–16" captionStartId="29.[151,264,1944,1968]" captionTargetBox="[181,1432,185,1916]" captionTargetId="figure-15@29.[151,1435,181,1920]" captionTargetPageId="29" captionText="FIGURES 15–16. Habitus and male genitalia. 15. Hydrochus foveatus Haldeman, non-type. 16. Hydrochus foveatus Haldeman, lectotype." figureDoi="http://doi.org/10.5281/zenodo.4777875" httpUri="https://zenodo.org/record/4777875/files/figure.png" pageId="17" pageNumber="476">Fig. 15</figureCitation>
) has this locality data:
<collectingRegion id="0D1F2D9E005BFFDEFD0A2CE7160E3229" box="[645,787,511,537]" country="Mexico" name="Nuevo Leon" pageId="17" pageNumber="476">Nuevo Leon</collectingRegion>
, Rio Ramos near Allende,
<date id="BB65C5BC005BFFDEFBB02CE611D53229" box="[1087,1224,510,537]" pageId="17" pageNumber="476" value="1954-08-18">18.viii.1954</date>
, leg. F. N. Young.
</paragraph>
</subSubSection>
</treatment>
</document>
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