172 lines
20 KiB
XML
172 lines
20 KiB
XML
<document id="1416706ED1352BE5F831A0F34D2CCE25" ID-CLB-Dataset="6214" ID-DOI="10.11646/zootaxa.4974.3.2" ID-GBIF-Dataset="5b4eaeb3-5722-4021-84ad-125ccd80855f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4777839" ID-ZooBank="1BD6220B-7540-4A12-B3D2-5EDC1A3B7DD8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621588618725" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="4772526A004CFFC9FF182FCE16DD3719" docLanguage="en" docName="zootaxa.4974.3.2.pdf" docOrigin="Zootaxa 4974 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus argutoides Perkins 2021, new species" docType="treatment" docVersion="6" lastPageNumber="465" masterDocId="BB4B2A12004AFFCFFF8F2D18151D3030" masterDocTitle="Taxonomy of Central American water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="503" masterPageNumber="459" pageNumber="465" updateTime="1698940277359" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="0B072C34BFA857EE10DFAE6AE0F01175">
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<mods:title id="AAD8A49306B1843F8E09CA10E1D744DB">Taxonomy of Central American water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)</mods:title>
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<mods:name id="A837AB8DD639DC872ACE54B025CC89F4" type="personal">
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<mods:namePart id="F0473D9C9305E22777B30BBF20C073A0">Perkins, Philip D.</mods:namePart>
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<mods:title id="1175DB9117E0B1BC39DEC62E84DAE1A5">Zootaxa</mods:title>
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<mods:part id="F04124E52FD10A42CFE18960E53F1308">
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<mods:date id="4EAEB3168CADAC43B0087C4DD4AEF7B0">2021</mods:date>
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<mods:number id="3EF1DA4C18CED228CB6E7DAC2FE7B493">2021-05-21</mods:number>
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<mods:number id="EB7941BDDBAC8EDB3B313042B1E35DF6">4974</mods:number>
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<mods:identifier id="4A89E84437D9A9EE906736F680F6850D" type="DOI">10.11646/zootaxa.4974.3.2</mods:identifier>
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<treatment id="4772526A004CFFC9FF182FCE16DD3719" ID-DOI="http://doi.org/10.5281/zenodo.4777849" ID-GBIF-Taxon="182642565" ID-Zenodo-Dep="4777849" LSID="urn:lsid:plazi:treatment:4772526A004CFFC9FF182FCE16DD3719" httpUri="http://treatment.plazi.org/id/4772526A004CFFC9FF182FCE16DD3719" lastPageNumber="465" pageId="6" pageNumber="465">
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<subSubSection id="87C1B0F7004CFFC9FF182FCE175F32C0" box="[151,578,726,753]" pageId="6" pageNumber="465" type="nomenclature">
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<paragraph id="CF64E37C004CFFC9FF182FCE175F32C0" blockId="6.[151,578,726,789]" box="[151,578,726,753]" pageId="6" pageNumber="465">
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<heading id="942C5410004CFFC9FF182FCE175F32C0" bold="true" box="[151,578,726,753]" fontSize="11" level="1" pageId="6" pageNumber="465" reason="1">
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<emphasis id="FDAF3F6E004CFFC9FF182FCE175F32C0" bold="true" box="[151,578,726,753]" pageId="6" pageNumber="465">
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<emphasis id="FDAF3F6E004CFFC9FF182FCE14B732C1" bold="true" box="[151,426,726,753]" italics="true" pageId="6" pageNumber="465">
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<taxonomicName id="08DB98FF004CFFC9FF182FCE14B932C1" ID-CoL="8Y4P4" authority="Perkins, 2021" authorityName="Perkins" authorityYear="2021" box="[151,420,726,753]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="465" phylum="Arthropoda" rank="species" species="argutoides" status="sp. nov.">Hydrochus argutoides</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="E69C8215004CFFC9FE3D2FCF175F32C0" box="[434,578,726,753]" pageId="6" pageNumber="465" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="87C1B0F7004CFFC9FF182FE314343325" box="[151,297,763,789]" pageId="6" pageNumber="465" type="description">
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<paragraph id="CF64E37C004CFFC9FF182FE314343325" blockId="6.[151,578,726,789]" box="[151,297,763,789]" pageId="6" pageNumber="465">
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<figureCitation id="57E0FFF9004CFFC9FF182FE315F83325" box="[151,229,763,789]" captionStart="FIGURE 2" captionStartId="5.[151,250,1989,2013]" captionTargetBox="[181,1393,181,1953]" captionTargetId="figure-15@5.[158,1429,181,1965]" captionTargetPageId="5" captionText="FIGURE 2. Dorsal habitus of Central American Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.4777847" httpUri="https://zenodo.org/record/4777847/files/figure.png" pageId="6" pageNumber="465">Figs. 2</figureCitation>
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,
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<figureCitation id="57E0FFF9004CFFC9FF7E2FE3141C3325" box="[241,257,763,789]" captionStart="FIGURES 3–4" captionStartId="23.[151,264,1938,1962]" captionTargetBox="[169,1433,200,1895]" captionTargetId="figure-15@23.[151,1435,181,1914]" captionTargetPageId="23" captionText="FIGURES 3–4. Habitus and male genitalia of holotypes. 3. Hydrochus lubricus n. sp. 4. Hydrochus argutoides n. sp." figureDoi="http://doi.org/10.5281/zenodo.4777851" httpUri="https://zenodo.org/record/4777851/files/figure.png" pageId="6" pageNumber="465">4</figureCitation>
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,
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<figureCitation id="57E0FFF9004CFFC9FE822FE314343325" box="[269,297,763,789]" captionStart="FIGURES 37–39" captionStartId="38.[151,264,1943,1967]" captionTargetBox="[159,1428,181,1919]" captionTargetId="figure-22@38.[159,1428,181,1919]" captionTargetPageId="38" captionText="FIGURES 37–39. Geographical distribution maps." figureDoi="http://doi.org/10.5281/zenodo.4777916" httpUri="https://zenodo.org/record/4777916/files/figure.png" pageId="6" pageNumber="465">38</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="87C1B0F7004CFFC9FF182E5B16F733B1" pageId="6" pageNumber="465" type="materials_examined">
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<paragraph id="CF64E37C004CFFC9FF182E5B16F733B1" blockId="6.[151,1437,834,1833]" pageId="6" pageNumber="465">
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<emphasis id="FDAF3F6E004CFFC9FF182E5B1458336C" bold="true" box="[151,325,834,861]" pageId="6" pageNumber="465">Type material.</emphasis>
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<materialsCitation id="7FB3E921004CFFC9FEC32E5B16F733B1" ID-GBIF-Occurrence="3128194417" collectingDate="2004-01-13" collectionCode="SEMC" collectorName="A. E. Z. Short & D. J. Lebbin" country="Costa Rica" elevation="210" location="Rio Animas" pageId="6" pageNumber="465" specimenCount="1" stateProvince="Guanacaste" typeStatus="holotype">
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<emphasis id="FDAF3F6E004CFFC9FEC32E5B1708336C" bold="true" box="[332,533,834,861]" pageId="6" pageNumber="465">
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<typeStatus id="10605DDE004CFFC9FEC32E5B14A4336D" box="[332,441,835,861]" pageId="6" pageNumber="465" type="holotype">Holotype</typeStatus>
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(male):
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</emphasis>
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“
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<collectingCountry id="B7CCA3EC004CFFC9FDAA2E5B17D2336D" box="[549,719,835,861]" name="Costa Rica" pageId="6" pageNumber="465">COSTA RICA</collectingCountry>
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:
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<collectingRegion id="0D1F2D9E004CFFC9FD562E5B1647336D" box="[729,858,835,861]" country="Costa Rica" name="Guanacaste" pageId="6" pageNumber="465">Guanacaste</collectingRegion>
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Prv.,
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<location id="CA04B5A7004CFFC9FC142E5B113D336D" LSID="urn:lsid:plazi:treatment:4772526A004CFFC9FF182FCE16DD3719:CA04B5A7004CFFC9FC142E5B113D336D" box="[923,1056,835,861]" country="Costa Rica" name="Rio Animas" pageId="6" pageNumber="465" stateProvince="Guanacaste">Rio Animas</location>
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@ Rt. 4;
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<quantity id="08234E99004CFFC9FB022E5B11C9336D" box="[1165,1236,835,861]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="465" unit="m" value="210.0">
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<elevation id="44F6044F004CFFC9FB022E5B11C9336D" box="[1165,1236,835,861]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="465" unit="m" value="210.0">210 m</elevation>
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</quantity>
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elv, detrital pools & backwater,
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<date id="BB65C5BC004CFFC9FEBB2E7F14BD33B1" box="[308,416,871,897]" pageId="6" pageNumber="465" value="2004-01-13">
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<collectingDate id="AB213C54004CFFC9FEBB2E7F14BD33B1" box="[308,416,871,897]" pageId="6" pageNumber="465" value="2004-01-13">13.i.2004</collectingDate>
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</date>
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, AS-04-040,
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<collectorName id="622E86AA004CFFC9FDBC2E7F17DF33B1" box="[563,706,871,897]" pageId="6" pageNumber="465">A.E.Z. Short</collectorName>
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&
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<collectorName id="622E86AA004CFFC9FD692E7F166E33B1" box="[742,883,871,897]" pageId="6" pageNumber="465">D. J. Lebbin</collectorName>
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” (
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<collectionCode id="A9CA7BB9004CFFC9FC032E7F16C333B1" box="[908,990,871,897]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="6" pageNumber="465" type="Museum">SEMC</collectionCode>
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).
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||
</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="87C1B0F7004CFFC9FF482E9211B33491" pageId="6" pageNumber="465" type="diagnosis">
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<paragraph id="CF64E37C004CFFC9FF482E9211B33491" blockId="6.[151,1437,834,1833]" pageId="6" pageNumber="465">
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<emphasis id="FDAF3F6E004CFFC9FF482E9214CD3394" bold="true" box="[199,464,906,932]" pageId="6" pageNumber="465">Differential diagnosis.</emphasis>
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Among Central American species, recognized by the combination of small size (ca.
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<quantity id="08234E99004CFFC9FF182EB6141D33F9" box="[151,256,942,969]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" pageId="6" pageNumber="465" unit="mm" value="2.65">2.65 mm</quantity>
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), the pronotum with shallow, densely punctate depressions, the pronotal shape, with strongly denticulate lateral margins, the elytral disc with very large and deep punctures, the large punctures of the elytral apices, and the male genitalia (
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<figureCitation id="57E0FFF9004CFFC9FEE12EEF14AE3421" box="[366,435,1015,1041]" captionStart="FIGURES 3–4" captionStartId="23.[151,264,1938,1962]" captionTargetBox="[169,1433,200,1895]" captionTargetId="figure-15@23.[151,1435,181,1914]" captionTargetPageId="23" captionText="FIGURES 3–4. Habitus and male genitalia of holotypes. 3. Hydrochus lubricus n. sp. 4. Hydrochus argutoides n. sp." figureDoi="http://doi.org/10.5281/zenodo.4777851" httpUri="https://zenodo.org/record/4777851/files/figure.png" pageId="6" pageNumber="465">Fig. 4</figureCitation>
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; described below). Differentiation from other similarly small sized species with similarly shaped pronotum should include dissection of males. This species is a member of the
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<taxonomicName id="08DB98FF004CFFC9FBF8290311EC3404" box="[1143,1265,1051,1076]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="465" phylum="Arthropoda" rank="species" species="argutus">
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<emphasis id="FDAF3F6E004CFFC9FBF8290311EC3404" box="[1143,1265,1051,1076]" italics="true" pageId="6" pageNumber="465">H. argutus</emphasis>
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</taxonomicName>
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species group. Other members of the group include
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<taxonomicName id="08DB98FF004CFFC9FDB62927161A3469" authority="Knisch" authorityName="Knisch" box="[569,775,1087,1113]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="465" phylum="Arthropoda" rank="species" species="argutus">
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<emphasis id="FDAF3F6E004CFFC9FDB6292717AC3468" box="[569,689,1087,1112]" italics="true" pageId="6" pageNumber="465">H. argutus</emphasis>
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Knisch
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</taxonomicName>
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(
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<collectingCountry id="B7CCA3EC004CFFC9FC98292716733469" box="[791,878,1087,1113]" name="Bolivia" pageId="6" pageNumber="465">Bolivia</collectingCountry>
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, southern
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<collectingCountry id="B7CCA3EC004CFFC9FC6D292711343469" box="[994,1065,1087,1113]" name="Brazil" pageId="6" pageNumber="465">Brazil</collectingCountry>
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,
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<collectingCountry id="B7CCA3EC004CFFC9FBBA292711BF3469" box="[1077,1186,1087,1113]" name="Paraguay" pageId="6" pageNumber="465">Paraguay</collectingCountry>
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),
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<taxonomicName id="08DB98FF004CFFC9FB3A2927144B344E" authority="Perkins" authorityName="Perkins" authorityYear="2019" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="465" phylum="Arthropoda" rank="species" species="argutissimus">
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<emphasis id="FDAF3F6E004CFFC9FB3A2927107B3468" box="[1205,1382,1087,1112]" italics="true" pageId="6" pageNumber="465">H. argutissimus</emphasis>
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Perkins (
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<collectingCountry id="B7CCA3EC004CFFC9FF5A297B1453344E" box="[213,334,1123,1150]" name="Venezuela" pageId="6" pageNumber="465">Venezuela</collectingCountry>
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)
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</taxonomicName>
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, and an undescribed species from
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<collectingCountry id="B7CCA3EC004CFFC9FD58297B1634344D" box="[727,809,1123,1149]" name="Bolivia" pageId="6" pageNumber="465">Bolivia</collectingCountry>
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(Perkins, in prep.). All members have the characteristic general shape of the male genitalia, which has a markedly arcuate basal piece, in lateral view.
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</paragraph>
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</subSubSection>
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<subSubSection id="87C1B0F7004CFFC9FF4829B216233735" pageId="6" pageNumber="465" type="description">
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||
<paragraph id="CF64E37C004CFFC9FF4829B216E93565" blockId="6.[151,1437,834,1833]" pageId="6" pageNumber="465">
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||
<emphasis id="FDAF3F6E004CFFC9FF4829B2144C34F4" bold="true" box="[199,337,1194,1220]" pageId="6" pageNumber="465">Description</emphasis>
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. Habitus as illustrated (
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<figureCitation id="57E0FFF9004CFFC9FDD229B317BE34F5" box="[605,675,1195,1221]" captionStart="FIGURES 3–4" captionStartId="23.[151,264,1938,1962]" captionTargetBox="[169,1433,200,1895]" captionTargetId="figure-15@23.[151,1435,181,1914]" captionTargetPageId="23" captionText="FIGURES 3–4. Habitus and male genitalia of holotypes. 3. Hydrochus lubricus n. sp. 4. Hydrochus argutoides n. sp." figureDoi="http://doi.org/10.5281/zenodo.4777851" httpUri="https://zenodo.org/record/4777851/files/figure.png" pageId="6" pageNumber="465">Fig. 4</figureCitation>
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). Size:
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<typeStatus id="10605DDE004CFFC9FD7829B3164434F5" box="[759,857,1195,1221]" pageId="6" pageNumber="465" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 2.67 /1.01; head width 0.67; pronotum l/w 0.63/0.70; PA 0.66; PB 0.50; elytra 1.68 /1.01. Body size small (ca.
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||
<quantity id="08234E99004CFFC9FAE429D715DA353D" metricMagnitude="-3" metricUnit="m" metricValue="2.67" pageId="6" pageNumber="465" unit="mm" value="2.67">2.67 mm</quantity>
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). Dorsum dark brown to reddish brown, with slight iridescence, elytra without black spots. Punctation on elytra deep, ca. 2–4x that of pronotum. Elytra interstriae ca. 0.5–1× strial puncture diameter. Interstria 9
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<superScript id="38AE4E34004CFFC9FB7C280E101D3514" attach="left" box="[1267,1280,1302,1316]" fontSize="6" pageId="6" pageNumber="465">th</superScript>
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more convex than others, overhanging 10
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<superScript id="38AE4E34004CFFC9FE5F282214C03578" attach="left" box="[464,477,1338,1352]" fontSize="6" pageId="6" pageNumber="465">th</superScript>
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||
interstria. Usual area of callus on 5
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<superScript id="38AE4E34004CFFC9FCE4282216653578" attach="left" box="[875,888,1338,1352]" fontSize="6" pageId="6" pageNumber="465">th</superScript>
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not raised.
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</paragraph>
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<paragraph id="CF64E37C004CFFC9FF48284714C135F1" blockId="6.[151,1437,834,1833]" pageId="6" pageNumber="465">Pronotum wider than long (ratio ca. 70/63), widest slightly behind anterior margin, markedly narrowed at base, sides straight, strongly denticulate; anterior margin bisinuate; depressions very shallow, densely punctate; reliefs very low, generally punctate.</paragraph>
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<paragraph id="CF64E37C004CFFC9FF4828D314C8361D" blockId="6.[151,1437,834,1833]" pageId="6" pageNumber="465">Elytra with punctures large and deep, spaces between strial punctures narrow ridges to ca. 0.5 puncture diameter; apices slightly widened, with submarginal groove having large translucent punctures and, apically, each elytron one large elongate puncture.</paragraph>
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<paragraph id="CF64E37C004CFFC9FF482B2F11EA3661" blockId="6.[151,1437,834,1833]" box="[199,1271,1591,1617]" pageId="6" pageNumber="465">Ventral characters: Mentum densely punctate, with median fovea. Submentum with two foveae.</paragraph>
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<paragraph id="CF64E37C004CFFC9FF482B431078368D" blockId="6.[151,1437,834,1833]" pageId="6" pageNumber="465">
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Male genitalia (
|
||
<figureCitation id="57E0FFF9004CFFC9FEF82B4314A73645" box="[375,442,1627,1653]" captionStart="FIGURES 3–4" captionStartId="23.[151,264,1938,1962]" captionTargetBox="[169,1433,200,1895]" captionTargetId="figure-15@23.[151,1435,181,1914]" captionTargetPageId="23" captionText="FIGURES 3–4. Habitus and male genitalia of holotypes. 3. Hydrochus lubricus n. sp. 4. Hydrochus argutoides n. sp." figureDoi="http://doi.org/10.5281/zenodo.4777851" httpUri="https://zenodo.org/record/4777851/files/figure.png" pageId="6" pageNumber="465">Fig. 4</figureCitation>
|
||
) general characters: paramere tips appearing sagittate,
|
||
<emphasis id="FDAF3F6E004CFFC9FBAD2B4310393644" box="[1058,1316,1627,1653]" italics="true" pageId="6" pageNumber="465">but only in lateral view</emphasis>
|
||
; in dorsal/ ventral views tips have medial margins separated, except meeting at distal end, tips extend well beyond distal end of aedeagus; basal piece long, in lateral view proximal 1/2 strongly arcuate ventrad, with terminal sclerotization.
|
||
</paragraph>
|
||
<paragraph id="CF64E37C004CFFC9FF482BDF16C436D1" blockId="6.[151,1437,834,1833]" box="[199,985,1735,1761]" pageId="6" pageNumber="465">Dorsal surface: adtl and adbl lacking; pdmm slightly overlapping alm.</paragraph>
|
||
<paragraph id="CF64E37C004CFFC9FF182BF316233735" blockId="6.[151,1437,834,1833]" box="[151,830,1771,1797]" pageId="6" pageNumber="465">Ventral surface: pvmm closer to one another than are pdmm.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="87C1B0F7004CFFC9FF482A1716DD3719" box="[199,960,1807,1833]" pageId="6" pageNumber="465" type="etymology">
|
||
<paragraph id="CF64E37C004CFFC9FF482A1716DD3719" blockId="6.[151,1437,834,1833]" box="[199,960,1807,1833]" pageId="6" pageNumber="465">
|
||
<emphasis id="FDAF3F6E004CFFC9FF482A1714533719" bold="true" box="[199,334,1807,1833]" pageId="6" pageNumber="465">Etymology.</emphasis>
|
||
Named in reference to the relationship with
|
||
<taxonomicName id="08DB98FF004CFFC9FCCD2A1716A73718" box="[834,954,1807,1832]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="465" phylum="Arthropoda" rank="species" species="argutus">
|
||
<emphasis id="FDAF3F6E004CFFC9FCCD2A1716A73718" box="[834,954,1807,1832]" italics="true" pageId="6" pageNumber="465">H. argutus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |