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<document id="47AE2EA8D582B06D0013BEE9A8805FD4" ID-CLB-Dataset="24395" ID-DOI="10.11646/zootaxa.4722.1.3" ID-GBIF-Dataset="c7d7ba85-5c89-4c67-958d-877b72cca225" ID-ISSN="1175-5326" ID-Zenodo-Dep="3603656" ID-ZooBank="206172F4-B350-4AE0-940E-B21CE2D7AD1C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1578644045100" checkinUser="plazi" docAuthor="Liu, Weixin &amp; Huo, Qingbo" docDate="2020" docId="4728879D1A10280A05FF55AA91543427" docLanguage="en" docName="zootaxa.4722.1.3.pdf" docOrigin="Zootaxa 4722 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Niponia nodulosa Verhoeff 1931" docType="treatment" docVersion="7" lastPageNumber="42" masterDocId="BB11FFE51A11280B0568554B951A370C" masterDocTitle="Two new species and a new record of polydesmoid millipedes from Jiangsu and Zhejiang provinces, eastern China (Diplopoda, Polydesmida)" masterLastPageNumber="49" masterPageNumber="41" pageNumber="42" updateTime="1698777207143" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="04360CFA85E31ECAD7D870378B772D49">Two new species and a new record of polydesmoid millipedes from Jiangsu and Zhejiang provinces, eastern China (Diplopoda, Polydesmida)</mods:title>
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<taxonomicName id="08814D081A10280A05FF55AA973737F7" ID-CoL="73ZHB" authority="Verhoeff, 1931" authorityName="Verhoeff" authorityYear="1931" box="[151,557,225,252]" class="Diplopoda" family="Cryptodesmidae" genus="Niponia" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="42" phylum="Arthropoda" rank="species" species="nodulosa">
<emphasis id="FDF5EA991A10280A05FF55AA947537F0" bold="true" box="[151,367,225,252]" italics="true" pageId="1" pageNumber="42">Niponia nodulosa</emphasis>
<bibRefCitation id="AB104B7A1A10280A041F55AA973737F7" author="Verhoeff, K. W." box="[375,557,225,251]" pageId="1" pageNumber="42" pagination="397 - 522" refId="ref3493" refString="Verhoeff, K. W. (1931) Chilognathen aus den Bergamasker Alpen und Nachbargebieten; auch uber zwei neue Gattungen der Polydesmidea aus Spanien und Japan. Zoologische Jahrbucher, Systematik, 61, 397 - 522." type="journal article" year="1931">Verhoeff, 1931</bibRefCitation>
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, new record from continental
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<figureCitation id="57BA2A0E1A10280A05FF544E95E23613" box="[151,248,261,287]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,1355,1380]" captionStartId-1="2.[151,250,1919,1944]" captionStartId-2="3.[151,250,1180,1205]" captionTargetBox-0="[155,1432,826,1329]" captionTargetBox-1="[151,1406,190,1855]" captionTargetBox-2="[155,1429,415,1153]" captionTargetId-0="figure@1.[151,1437,823,1333]" captionTargetId-1="figure@2.[151,1410,174,1874]" captionTargetId-2="figure@3.[151,1435,409,1160]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Niponia nodulosa in nature. A: female B: a mating pair." captionText-1="FIGURE 2. Niponia nodulosa, male from Yangzhou, Zhejiang. A, B: anterior part of body, ventral and dorsal views, respectively C, D: midbody segments, ventral and dorsal views, respectively E, F:posterior part of body, ventral and dorsal views, respectively." captionText-2="FIGURE 3. Niponia nodulosa, male from Yangzhou, Zhejiang. A, B: left gonopod, mesal view C: gonopods in situ, ventral view D, E: left gonopod, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.3603658" figureDoi-1="http://doi.org/10.5281/zenodo.3603660" figureDoi-2="http://doi.org/10.5281/zenodo.3603662" httpUri-0="https://zenodo.org/record/3603658/files/figure.png" httpUri-1="https://zenodo.org/record/3603660/files/figure.png" httpUri-2="https://zenodo.org/record/3603662/files/figure.png" pageId="1" pageNumber="42">Figs 13</figureCitation>
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<taxonomicName id="08814D081A10280A05FF54019705366E" ID-CoL="73ZHB" authority="Verhoeff, 1931: 442" authorityName="Verhoeff" authorityPageNumber="442" authorityYear="1931" box="[151,543,329,354]" class="Diplopoda" family="Cryptodesmidae" genus="Niponia" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="42" phylum="Arthropoda" rank="species" species="nodulosa">
<emphasis id="FDF5EA991A10280A05FF54019456366E" box="[151,332,330,354]" italics="true" pageId="1" pageNumber="42">Niponia nodulosa</emphasis>
<treatmentCitation id="4E20109A1A10280A043A54029705366E" author="Verhoeff, K. W." box="[338,543,329,354]" page="442" pageId="1" pageNumber="42" year="1931">
<bibRefCitation id="AB104B7A1A10280A043A54029705366E" author="Verhoeff, K. W." box="[338,543,329,354]" pageId="1" pageNumber="42" pagination="397 - 522" refId="ref3493" refString="Verhoeff, K. W. (1931) Chilognathen aus den Bergamasker Alpen und Nachbargebieten; auch uber zwei neue Gattungen der Polydesmidea aus Spanien und Japan. Zoologische Jahrbucher, Systematik, 61, 397 - 522." type="journal article" year="1931">Verhoeff, 1931: 442</bibRefCitation>
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, figs 4655.
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</paragraph>
<paragraph id="CF3E368B1A10280A05FF542297C6368C" blockId="1.[151,732,329,415]" box="[151,732,360,385]" pageId="1" pageNumber="42">
<treatmentCitationGroup id="EF9111A51A10280A05FF542297C6368C" box="[151,732,360,385]" pageId="1" pageNumber="42">
<emphasis id="FDF5EA991A10280A05FF54229478368C" box="[151,354,360,384]" italics="true" pageId="1" pageNumber="42">
<taxonomicName id="08814D081A10280A05FF542295F3368C" ID-CoL="7PQCY" authorityName="Verhoeff" authorityYear="1931" box="[151,233,361,384]" class="Diplopoda" family="Cryptodesmidae" genus="Niponia" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus">Niponia</taxonomicName>
nodulosa—
</emphasis>
<treatmentCitation id="4E20109A1A10280A040B54239763368C" author="Golovatch, S. I. &amp; Mikhalkova, E. V. &amp; Chang, H. W." box="[355,633,360,385]" page="120" pageId="1" pageNumber="42" year="2011">
<bibRefCitation id="AB104B7A1A10280A040B54239763368C" author="Golovatch, S. I. &amp; Mikhalkova, E. V. &amp; Chang, H. W." box="[355,633,360,385]" pageId="1" pageNumber="42" pagination="119 - 134" refId="ref3372" refString="Golovatch, S. I., Mikhalkova, E. V. &amp; Chang, H. W. (2011) The millipede families Cryptodesmidae, Haplodesmidae, Pyrgodesmidae, Opisotretidae and Xystodesmidae in Taiwan (Diplopoda, Polydesmida). Tropical Natural History, 11 (2), 119 - 134." type="journal article" year="2011">
Golovatch
<emphasis id="FDF5EA991A10280A04BA5422971C368C" box="[466,518,360,384]" italics="true" pageId="1" pageNumber="42">et al.</emphasis>
, 2011: 120
</bibRefCitation>
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, figs 13.
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</paragraph>
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<treatmentCitationGroup id="EF9111A51A10280A05FF54C397233693" box="[151,569,390,415]" pageId="1" pageNumber="42">
<emphasis id="FDF5EA991A10280A05FF54C394783693" box="[151,354,391,415]" italics="true" pageId="1" pageNumber="42">
<taxonomicName id="08814D081A10280A05FF54C395F33693" ID-CoL="7PQCY" authorityName="Verhoeff" authorityYear="1931" box="[151,233,392,415]" class="Diplopoda" family="Cryptodesmidae" genus="Niponia" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus">Niponia</taxonomicName>
nodulosa—
</emphasis>
<treatmentCitation id="4E20109A1A10280A040B54CD972F3693" author="Golovatch, S. I." box="[355,565,390,415]" page="34" pageId="1" pageNumber="42" year="2016">
<bibRefCitation id="AB104B7A1A10280A040B54CD972F3693" author="Golovatch, S. I." box="[355,565,390,415]" pageId="1" pageNumber="42" pagination="33 - 43" refId="ref3331" refString="Golovatch, S. I. (2016) The millipede family Cryptodesmidae in Indochina (Diplopoda, Polydesmida). ZooKeys, 578, 33 - 43. https: // doi. org / 10.3897 / zookeys. 578.7994" type="journal article" year="2016">Golovatch, 2016: 34</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<emphasis id="FDF5EA991A10280A05FF5486946136E4" bold="true" box="[151,379,461,488]" pageId="1" pageNumber="42">Material examined</emphasis>
:
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,
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,
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,
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a.s.l.
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,
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.
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(
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,
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,
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,
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a.s.l.
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,
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<collectingDate id="AB7BE9A31A10280A05B157729452355F" box="[217,328,569,595]" pageId="1" pageNumber="42" value="2018-10-09">2018-X-9</collectingDate>
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, all Qing-Bo
<collectorName id="6274535D1A10280A04885771970A3558" box="[480,528,570,596]" pageId="1" pageNumber="42">Huo</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="879B65001A10280A05AF571591543427" pageId="1" pageNumber="42" type="discussion">
<paragraph id="CF3E368B1A10280A05AF571597FF35EF" blockId="1.[151,1437,461,812]" pageId="1" pageNumber="42">
<emphasis id="FDF5EA991A10280A05AF5715942E3574" bold="true" box="[199,308,606,632]" pageId="1" pageNumber="42">Remarks</emphasis>
: This species is very common in southern
<collectingCountry id="B796761B1A10280A0674571596453574" box="[796,863,606,632]" name="Japan" pageId="1" pageNumber="42">Japan</collectingCountry>
,
<collectingCountry id="B796761B1A10280A0602571696A6357B" box="[874,956,605,631]" name="Taiwan" pageId="1" pageNumber="42">Taiwan</collectingCountry>
and also has been recorded from
<collectingCountry id="B796761B1A10280A005557169086357B" box="[1341,1436,605,631]" name="Vietnam" pageId="1" pageNumber="42">Vietnam</collectingCountry>
(
<bibRefCitation id="AB104B7A1A10280A05F757CA948F3597" author="Golovatch, S. I. &amp; Mikhalkova, E. V. &amp; Chang, H. W." box="[159,405,641,667]" pageId="1" pageNumber="42" pagination="119 - 134" refId="ref3372" refString="Golovatch, S. I., Mikhalkova, E. V. &amp; Chang, H. W. (2011) The millipede families Cryptodesmidae, Haplodesmidae, Pyrgodesmidae, Opisotretidae and Xystodesmidae in Taiwan (Diplopoda, Polydesmida). Tropical Natural History, 11 (2), 119 - 134." type="journal article" year="2011">
Golovatch
<emphasis id="FDF5EA991A10280A047457C9944F3597" box="[284,341,641,667]" italics="true" pageId="1" pageNumber="42">et al.</emphasis>
2011
</bibRefCitation>
,
<bibRefCitation id="AB104B7A1A10280A04C957CA97433590" author="Golovatch, S. I." box="[417,601,641,668]" pageId="1" pageNumber="42" pagination="33 - 43" refId="ref3331" refString="Golovatch, S. I. (2016) The millipede family Cryptodesmidae in Indochina (Diplopoda, Polydesmida). ZooKeys, 578, 33 - 43. https: // doi. org / 10.3897 / zookeys. 578.7994" type="journal article" year="2016">Golovatch 2016</bibRefCitation>
). The new samples almost fully agree with the original description and the later detailed redescription (
<bibRefCitation id="AB104B7A1A10280A04B857EE976E35B3" author="Verhoeff, K. W." box="[464,628,677,703]" pageId="1" pageNumber="42" pagination="397 - 522" refId="ref3493" refString="Verhoeff, K. W. (1931) Chilognathen aus den Bergamasker Alpen und Nachbargebieten; auch uber zwei neue Gattungen der Polydesmidea aus Spanien und Japan. Zoologische Jahrbucher, Systematik, 61, 397 - 522." type="journal article" year="1931">Verhoeff 1931</bibRefCitation>
,
<bibRefCitation id="AB104B7A1A10280A07E857EE966D35B3" author="Golovatch, S. I. &amp; Mikhalkova, E. V. &amp; Chang, H. W." box="[640,887,677,703]" pageId="1" pageNumber="42" pagination="119 - 134" refId="ref3372" refString="Golovatch, S. I., Mikhalkova, E. V. &amp; Chang, H. W. (2011) The millipede families Cryptodesmidae, Haplodesmidae, Pyrgodesmidae, Opisotretidae and Xystodesmidae in Taiwan (Diplopoda, Polydesmida). Tropical Natural History, 11 (2), 119 - 134." type="journal article" year="2011">
Golovatch
<emphasis id="FDF5EA991A10280A079457ED962E35B3" box="[764,820,677,703]" italics="true" pageId="1" pageNumber="42">et al.</emphasis>
2011
</bibRefCitation>
). The habitus and gonopodal structure (
<figureCitation id="57BA2A0E1A10280A005857EE908935B3" box="[1328,1427,677,703]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,1355,1380]" captionStartId-1="2.[151,250,1919,1944]" captionStartId-2="3.[151,250,1180,1205]" captionTargetBox-0="[155,1432,826,1329]" captionTargetBox-1="[151,1406,190,1855]" captionTargetBox-2="[155,1429,415,1153]" captionTargetId-0="figure@1.[151,1437,823,1333]" captionTargetId-1="figure@2.[151,1410,174,1874]" captionTargetId-2="figure@3.[151,1435,409,1160]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Niponia nodulosa in nature. A: female B: a mating pair." captionText-1="FIGURE 2. Niponia nodulosa, male from Yangzhou, Zhejiang. A, B: anterior part of body, ventral and dorsal views, respectively C, D: midbody segments, ventral and dorsal views, respectively E, F:posterior part of body, ventral and dorsal views, respectively." captionText-2="FIGURE 3. Niponia nodulosa, male from Yangzhou, Zhejiang. A, B: left gonopod, mesal view C: gonopods in situ, ventral view D, E: left gonopod, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.3603658" figureDoi-1="http://doi.org/10.5281/zenodo.3603660" figureDoi-2="http://doi.org/10.5281/zenodo.3603662" httpUri-0="https://zenodo.org/record/3603658/files/figure.png" httpUri-1="https://zenodo.org/record/3603660/files/figure.png" httpUri-2="https://zenodo.org/record/3603662/files/figure.png" pageId="1" pageNumber="42">Figs 13</figureCitation>
) are illustrated here to document the species identity.
</paragraph>
<paragraph id="CF3E368B1A10280A05AF57A691543427" blockId="1.[151,1437,461,812]" pageId="1" pageNumber="42">
Willows are very common in Yangzhou City,
<collectingRegion id="0D45F8691A10280A07B957A6968E340B" box="[721,916,749,775]" country="China" name="Jiangsu" pageId="1" pageNumber="42">Jiangsu Province</collectingRegion>
. Most samples of
<taxonomicName id="08814D081A10280A010D57A591F4340B" authorityName="Verhoeff" authorityYear="1931" box="[1125,1262,749,775]" class="Diplopoda" family="Cryptodesmidae" genus="Niponia" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="42" phylum="Arthropoda" rank="species" species="nodulosa">
<emphasis id="FDF5EA991A10280A010D57A591F4340B" box="[1125,1262,749,775]" italics="true" pageId="1" pageNumber="42">N. nodulosa</emphasis>
</taxonomicName>
have been collected from dead willows by the second author during a study of termite infestations.
</paragraph>
</subSubSection>
</treatment>
</document>