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<document id="CC488EE5D50844C3BD81DC5DD7472B24" ID-DOI="10.1655/06-001.1" ID-Zenodo-Dep="14384794" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1733861782093" checkinUser="juliana" docAuthor="Torres-Carvajal, Omar" docDate="2007" docId="467D8791FFABFF86FD44FBF41545FAB6" docLanguage="en" docName="HerpetolMonogr.21.1.76-178.pdf.pdf" docOrigin="Herpetological Monographs 21 (1)" docSource="https://doi.org/10.1655/06-001.1" docTitle="Stenocercus boettgeri Boulenger 1911" docType="treatment" docVersion="6" lastPageNumber="92" masterDocId="BA44FFE9FFBBFF97FFBCFFC41632FFF9" masterDocTitle="A TAXONOMIC REVISION OF SOUTH AMERICAN STENOCERCUS (SQUAMATA: IGUANIA) LIZARDS" masterLastPageNumber="178" masterPageNumber="76" pageNumber="91" updateTime="1733923315685" updateUser="juliana">
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<mods:title id="5719263C839EE6231DD4664BF12F235A">A TAXONOMIC REVISION OF SOUTH AMERICAN STENOCERCUS (SQUAMATA: IGUANIA) LIZARDS</mods:title>
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<mods:namePart id="51C6E33C2EC41D790945088F6326AB83">Torres-Carvajal, Omar</mods:namePart>
<mods:affiliation id="1F64526D6034640FE607BED87DB5FDD1">Department of Vertebrate Zoologŋ, National Museum of Natural Historŋ, Smithsonian Institution, Washington, DC 20560, USA &amp; Escuela de Biología, Pontificia UniƲersidad Católica del Ecuador, AƲenida 12 de Octubre ŋ Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
<mods:nameIdentifier id="38585F2339C289DBC659E87FFA883DC3" type="email">Torreso@si.edu</mods:nameIdentifier>
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<mods:title id="61AA5C10B35B39CB23B17988C1A30372">Herpetological Monographs</mods:title>
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<mods:part id="EE87AE89FC886829ED012DFED1981B92">
<mods:date id="6274FFD59127B0DE9A27D313D72D8AE8">2007</mods:date>
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<mods:number id="22794E143BD00D0A66501DCC33408D1A">2007-12-28</mods:number>
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<mods:number id="079053885C1135AEA430A291468D4D58">21</mods:number>
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<mods:classification id="57C40E62AEFBA235D0846B2A30561617">journal article</mods:classification>
<mods:identifier id="298C36D74D116CB5447320CBA3540769" type="DOI">10.1655/06-001.1</mods:identifier>
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<treatment id="467D8791FFABFF86FD44FBF41545FAB6" ID-DOI="http://doi.org/10.5281/zenodo.14372655" ID-Zenodo-Dep="14372655" LSID="urn:lsid:plazi:treatment:467D8791FFABFF86FD44FBF41545FAB6" httpUri="http://treatment.plazi.org/id/467D8791FFABFF86FD44FBF41545FAB6" lastPageId="17" lastPageNumber="92" pageId="16" pageNumber="91">
<subSubSection id="86CE650CFFABFF87FD44FBF415E4FB9F" pageId="16" pageNumber="91" type="nomenclature">
<paragraph id="CE6B3687FFABFF87FD44FBF415E4FB9F" blockId="16.[685,1196,1072,1722]" pageId="16" pageNumber="91">
<heading id="952381EBFFABFF87FD44FBF415E4FB9F" centered="true" fontSize="10" level="3" pageId="16" pageNumber="91" reason="4">
<taxonomicName id="09D44D04FFABFF87FD44FBF41253FBB1" ID-CoL="4ZWJ4" authority="Boulenger" authorityName="Boulenger" authorityYear="1911" box="[760,1121,1072,1096]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="16" pageNumber="91" phylum="Chordata" rank="species" species="boettgeri">
<emphasis id="FCA0EA95FFABFF87FD44FBF415D7FBB1" box="[760,997,1072,1096]" italics="true" pageId="16" pageNumber="91">Stenocercus boettgeri</emphasis>
Boulenger
</taxonomicName>
(
<figureCitation id="56EF2A02FFABFF87FC37FB8A15FCFB9F" box="[907,974,1102,1126]" captionStart="FIG" captionStartId="9.[143,157,1719,1738]" captionTargetBox="[122,1176,189,1701]" captionTargetId="figure-0@9.[122,1176,189,1701]" captionTargetPageId="9" captionText="FIG. 7.—Eight species of Stenocercus. (A) S. angel (LAC); (B) S. angulifer (FAY); (C) S. apurimacus (WED); (D) S. azureus (SCA); (E) S. boettgeri (MLU); (F) S. boliƲarensis (FCA); (G) S. caducus (LSI); (H) S. chlorostictus (MLU)." figureDoi="http://doi.org/10.5281/zenodo.14384814" httpUri="https://zenodo.org/record/14384814/files/figure.png" pageId="16" pageNumber="91">Fig. 7</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFFABFF87FD11FBB11516FA6D" pageId="16" pageNumber="91" type="reference_group">
<paragraph id="CE6B3687FFABFF87FD11FBB11516FA6D" blockId="16.[685,1196,1072,1722]" pageId="16" pageNumber="91">
<taxonomicName id="09D44D04FFABFF87FD11FBB1129AFB75" authority="Boulenger, 1911: 22" authorityName="Boulenger" authorityPageNumber="22" authorityYear="1911" box="[685,1192,1140,1164]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="16" pageNumber="91" phylum="Chordata" rank="species" species="boettgeri">
<emphasis id="FCA0EA95FFABFF87FD11FBB11594FB75" box="[685,934,1140,1164]" italics="true" pageId="16" pageNumber="91">Stenocercus boettgeri</emphasis>
<bibRefCitation id="AA454B76FFABFF87FC03FBB0129AFB75" author="BOULENGER, G. A." box="[959,1192,1140,1164]" journalOrPublisher="Annals and Magazine of Natural History" pageId="16" pageNumber="91" pagination="19 - 25" part="8" refId="ref76316" refString="BOULENGER, G. A. 1911. Descriptions of new reptiles from the Andes of South America, preserved in the British Museum. Annals and Magazine of Natural History 8: 19 - 25." title="Descriptions of new reptiles from the Andes of South America, preserved in the British Museum" type="journal article" year="1911">Boulenger, 1911:22</bibRefCitation>
</taxonomicName>
.
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<typeStatus id="116F8825FFABFF87FD74FB56151EFB53" box="[712,812,1170,1194]" pageId="16" pageNumber="91">Syntypes</typeStatus>
:
<specimenCode id="9E729EFCFFABFF87FCEEFB56126EFB50" box="[850,1116,1169,1193]" pageId="16" pageNumber="91">BM 1911.12.13.25–32</specimenCode>
(
<specimenCode id="9E729EFCFFABFF87FB3AFB561548FB3E" pageId="16" pageNumber="91">RR 1946.8.11.92–99</specimenCode>
)from‘‘
<location id="CB0B605CFFABFF87FC78FB6B1254FB3E" LSID="urn:lsid:plazi:treatment:467D8791FFABFF86FD44FBF41545FAB6:CB0B605CFFABFF87FC78FB6B1254FB3E" box="[964,1126,1199,1223]" country="Peru" latitude="-10.349999" longLatPrecision="21" longitude="-75.53334" name="Huancabamba" pageId="16" pageNumber="91" stateProvince="Departamento Pasco">Huancabamba</location>
,Peru̕̕ (restricted to Huancabamba [
<geoCoordinate id="ABE05040FFABFF87FB90FB081295FB1D" box="[1068,1191,1228,1252]" degrees="10" direction="south" minutes="20" orientation="latitude" pageId="16" pageNumber="91" precision="15" seconds="60" value="-10.349999">
10
<emphasis id="FCA0EA95FFABFF87FBFBFB081263FB1D" bold="true" box="[1095,1105,1228,1252]" pageId="16" pageNumber="91">°</emphasis>
20'60&quot;S
</geoCoordinate>
,
<geoCoordinate id="ABE05040FFABFF87FD74FB2D157FFAF8" box="[712,845,1257,1282]" degrees="75" direction="west" minutes="31" orientation="longitude" pageId="16" pageNumber="91" precision="15" seconds="60" value="-75.53334">
75
<emphasis id="FCA0EA95FFABFF87FD5EFB2D14DEFAF8" bold="true" box="[738,748,1257,1281]" pageId="16" pageNumber="91">°</emphasis>
31'60&quot;W
</geoCoordinate>
,
<quantity id="092C9B62FFABFF87FCE4FB2D159DFAF8" box="[856,943,1257,1281]" metricMagnitude="3" metricUnit="m" metricValue="2.686" pageId="16" pageNumber="91" unit="m" value="2686.0">
<elevation id="45F9D1B4FFABFF87FCE4FB2D159DFAF8" box="[856,943,1257,1281]" metricMagnitude="3" metricUnit="m" metricValue="2.686" pageId="16" pageNumber="91" unit="m" value="2686.0">2686 m</elevation>
</quantity>
],
<collectingRegion id="0C10F865FFABFF87FC7EFB2E129AFAF8" box="[962,1192,1258,1281]" pageId="16" pageNumber="91">Departamento Pasco</collectingRegion>
,
<collectingCountry id="B6C37617FFABFF87FD74FAC314CDFAE7" box="[712,767,1287,1310]" name="Peru" pageId="16" pageNumber="91">Peru</collectingCountry>
, by
<bibRefCitation id="AA454B76FFABFF87FCFCFAC315EBFAE6" author="CADLE, J. E." box="[832,985,1287,1311]" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="16" pageNumber="91" pagination="1 - 96" part="143" refId="ref76540" refString="CADLE, J. E. 1991. Systematics of lizards of the genus Stenocercus (Iguania: Tropiduridae) from northern Peru: new species and comments on relationships and distribution patterns. Proceedings of the Academy of Natural Sciences of Philadelphia 143: 1 - 96." title="Systematics of lizards of the genus Stenocercus (Iguania: Tropiduridae) from northern Peru: new species and comments on relationships and distribution patterns" type="proceedings paper" year="1991">Cadle [1991]</bibRefCitation>
);
</materialsCitation>
<bibRefCitation id="AA454B76FFABFF87FC4BFAC3152DFAC5" author="BURT, C. E. &amp; M. D. BURT" journalOrPublisher="Proceedings of the United States National Museum" pageId="16" pageNumber="91" pagination="1 - 52" part="78" refId="ref76353" refString="BURT, C. E., AND M. D. BURT. 1930. The South American lizards in the collection of the United States National Museum. Proceedings of the United States National Museum 78 (6): 1 - 52." title="The South American lizards in the collection of the United States National Museum" type="journal article" year="1930">Burt and Burt, 1930:22</bibRefCitation>
;
<bibRefCitation id="AA454B76FFABFF87FC90FAE1120EFAC5" author="BURT, C. E. &amp; M. D. BURT" box="[812,1084,1316,1340]" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="16" pageNumber="91" pagination="227 - 395" part="61" refId="ref76399" refString="BURT, C. E., AND M. D. BURT. 1931. South American lizards in the collection of the American Museum of Natural History and Ecology. Bulletin of the American Museum of Natural History 61 (7): 227 - 395." title="South American lizards in the collection of the American Museum of Natural History and Ecology" type="journal article" year="1931">Burt and Burt, 1931:287</bibRefCitation>
;
<bibRefCitation id="AA454B76FFABFF87FBF5FAE1156CFAA3" author="BURT, C. E. &amp; M. D. BURT" journalOrPublisher="Transactions of the Academy of Science of St. Louis" pageId="16" pageNumber="91" pagination="1 - 104" part="28" refId="ref76448" refString="BURT, C. E., AND M. D. BURT. 1933. A preliminary check list of the lizards of South America. Transactions of the Academy of Science of St. Louis 28 (1): 1 - 104." title="A preliminary check list of the lizards of South America" type="journal article" year="1933">Burt and Burt, 1933:42</bibRefCitation>
;
<bibRefCitation id="AA454B76FFABFF87FCD5FA861223FAA0" author="PETERS, J. A." box="[873,1041,1345,1369]" journalOrPublisher="Proceedings of the United States National Museum" pageId="16" pageNumber="91" pagination="1 - 49" part="119" refId="ref79202" refString="PETERS, J. A. 1967. The lizards of Ecuador, a check list and key. Proceedings of the United States National Museum 119: 1 - 49." title="The lizards of Ecuador, a check list and key" type="proceedings paper" year="1967">Peters, 1967:34</bibRefCitation>
; Etheridge, in
<bibRefCitation id="AA454B76FFABFF87FD74FA9B126FFA8E" author="PETERS, J. A. &amp; R. DONOSO-BARROS" box="[712,1117,1375,1399]" journalOrPublisher="United States National Museum Bulletin" pageId="16" pageNumber="91" pagination="1 - 293" part="297" refId="ref79234" refString="PETERS, J. A., AND R. DONOSO-BARROS. 1970. Catalogue of the Neotropical Squamata. Part 2, lizards and amphisbaenians. United States National Museum Bulletin 297: 1 - 293." title="Catalogue of the Neotropical Squamata. Part 2, lizards and amphisbaenians" type="journal article" year="1970">Peters and Donoso-Barros, 1970:255</bibRefCitation>
;
<bibRefCitation id="AA454B76FFABFF87FBD6FA9B1512FA6D" author="FRITTS, T. H." journalOrPublisher="San Diego Society of Natural History Memoir" pageId="16" pageNumber="91" pagination="1 - 89" part="7" refId="ref77735" refString="FRITTS, T. H. 1974. A multivariate and evolutionary analysis of the andean iguanid lizards of the genus Stenocercus. San Diego Society of Natural History Memoir 7: 1 - 89." title="A multivariate and evolutionary analysis of the andean iguanid lizards of the genus Stenocercus" type="journal article" year="1974">Fritts, 1974:39</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFFABFF86FD74FA5F16D7FDC0" lastPageId="17" lastPageNumber="92" pageId="16" pageNumber="91" type="diagnosis">
<paragraph id="CE6B3687FFABFF86FD74FA5F16D7FDC0" blockId="16.[685,1196,1072,1722]" lastBlockId="17.[122,633,189,1722]" lastPageId="17" lastPageNumber="92" pageId="16" pageNumber="91">
<emphasis id="FCA0EA95FFABFF87FD74FA5F1506FA4B" box="[712,820,1435,1458]" italics="true" pageId="16" pageNumber="91">Diagnosis</emphasis>
.—
<taxonomicName id="09D44D04FFABFF87FCE9FA5F1270FA4B" authorityName="Boulenger" authorityYear="1911" box="[853,1090,1434,1458]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="16" pageNumber="91" phylum="Chordata" rank="species" species="boettgeri">
<emphasis id="FCA0EA95FFABFF87FCE9FA5F1270FA4B" box="[853,1090,1434,1458]" italics="true" pageId="16" pageNumber="91">Stenocercus boettgeri</emphasis>
</taxonomicName>
is distinguished from other species of
<taxonomicName id="09D44D04FFABFF87FB95FA7C129EFA36" box="[1065,1196,1464,1487]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="16" pageNumber="91" phylum="Chordata" rank="genus">
<emphasis id="FCA0EA95FFABFF87FB95FA7C129EFA36" box="[1065,1196,1464,1487]" italics="true" pageId="16" pageNumber="91">Stenocercus</emphasis>
</taxonomicName>
except
<taxonomicName id="09D44D04FFABFF87FD47FA111545FA14" baseAuthorityName="Werner" baseAuthorityYear="1901" box="[763,887,1493,1517]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="16" pageNumber="91" phylum="Chordata" rank="species" species="haenschi">
<emphasis id="FCA0EA95FFABFF87FD47FA111545FA14" box="[763,887,1493,1517]" italics="true" pageId="16" pageNumber="91">S. haenschi</emphasis>
</taxonomicName>
,
<taxonomicName id="09D44D04FFABFF87FC39FA11123CFA14" authorityName="Gunther" authorityYear="1859" box="[901,1038,1493,1517]" class="Squamata" family="Tropiduridae" genus="Microphractus" kingdom="Animalia" pageId="16" pageNumber="91" phylum="Chordata" rank="species" species="humeralis">
<emphasis id="FCA0EA95FFABFF87FC39FA11123CFA14" box="[901,1038,1493,1517]" italics="true" pageId="16" pageNumber="91">S. humeralis</emphasis>
</taxonomicName>
, and
<taxonomicName id="09D44D04FFABFF87FBF7FA111299FA15" box="[1099,1195,1493,1516]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="16" pageNumber="91" phylum="Chordata" rank="species" species="varius">
<emphasis id="FCA0EA95FFABFF87FBF7FA111299FA15" box="[1099,1195,1493,1516]" italics="true" pageId="16" pageNumber="91">S. varius</emphasis>
</taxonomicName>
by having granular scales on the posterior surface of thighs, enlarged vertebrals, three caudal whorls per autotomic segment, a medially complete antegular fold, non-spinose caudals, and by males lacking a black transverse band on the ventral surface of neck.
<taxonomicName id="09D44D04FFABFF87FB2AF941153CF943" authorityName="Boulenger" authorityYear="1911" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="16" pageNumber="91" phylum="Chordata" rank="species" species="boettgeri">
<emphasis id="FCA0EA95FFABFF87FB2AF941153CF943" italics="true" pageId="16" pageNumber="91">S. boettgeri</emphasis>
</taxonomicName>
has more scales around midbody (79–104,
<emphasis id="FCA0EA95FFAAFF86FF5EFF7A16C0FF2C" box="[226,242,190,213]" italics="true" pageId="17" pageNumber="92">X</emphasis>
= 88.61) than
<taxonomicName id="09D44D04FFAAFF86FE1CFF7A142CFF2F" baseAuthorityName="Werner" baseAuthorityYear="1901" box="[416,542,190,214]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="17" pageNumber="92" phylum="Chordata" rank="species" species="haenschi">
<emphasis id="FCA0EA95FFAAFF86FE1CFF7A142CFF2F" box="[416,542,190,214]" italics="true" pageId="17" pageNumber="92">S. haenschi</emphasis>
</taxonomicName>
(57–64,
<emphasis id="FCA0EA95FFAAFF86FFC6FF1816B8FF0A" box="[122,138,220,243]" italics="true" pageId="17" pageNumber="92">X</emphasis>
= 60.50) and
<taxonomicName id="09D44D04FFAAFF86FE8CFF1817A0FF0A" box="[304,402,220,243]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="17" pageNumber="92" phylum="Chordata" rank="species" species="varius">
<emphasis id="FCA0EA95FFAAFF86FE8CFF1817A0FF0A" box="[304,402,220,243]" italics="true" pageId="17" pageNumber="92">S. varius</emphasis>
</taxonomicName>
(74–88,
<emphasis id="FCA0EA95FFAAFF86FE49FF181437FF0A" box="[501,517,220,243]" italics="true" pageId="17" pageNumber="92">X</emphasis>
= 82.35). Both males and females of
<taxonomicName id="09D44D04FFAAFF86FE79FF3D147AFEE8" authorityName="Boulenger" authorityYear="1911" box="[453,584,249,273]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="17" pageNumber="92" phylum="Chordata" rank="species" species="boettgeri">
<emphasis id="FCA0EA95FFAAFF86FE79FF3D147AFEE8" box="[453,584,249,273]" italics="true" pageId="17" pageNumber="92">S. boettgeri</emphasis>
</taxonomicName>
are larger (maximum SVL = 108 and 94 mm, respectively) than
<taxonomicName id="09D44D04FFAAFF86FEFAFEF01794FEB2" box="[326,422,308,331]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="17" pageNumber="92" phylum="Chordata" rank="species" species="varius">
<emphasis id="FCA0EA95FFAAFF86FEFAFEF01794FEB2" box="[326,422,308,331]" italics="true" pageId="17" pageNumber="92">S. varius</emphasis>
</taxonomicName>
(maximum SVL = 85 in both sexes).
<taxonomicName id="09D44D04FFAAFF86FEFAFE9617F6FE90" authorityName="Boulenger" authorityYear="1911" box="[326,452,337,361]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="17" pageNumber="92" phylum="Chordata" rank="species" species="boettgeri">
<emphasis id="FCA0EA95FFAAFF86FEFAFE9617F6FE90" box="[326,452,337,361]" italics="true" pageId="17" pageNumber="92">S. boettgeri</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="09D44D04FFAAFF86FF05FEAB1774FE7E" authorityName="Gunther" authorityYear="1859" box="[185,326,367,391]" class="Squamata" family="Tropiduridae" genus="Microphractus" kingdom="Animalia" pageId="17" pageNumber="92" phylum="Chordata" rank="species" species="humeralis">
<emphasis id="FCA0EA95FFAAFF86FF05FEAB1774FE7E" box="[185,326,367,391]" italics="true" pageId="17" pageNumber="92">S. humeralis</emphasis>
</taxonomicName>
(character states in parentheses) by having lateral nuchals less than half the size of dorsal nuchals (lateral and dorsal nuchals similar in size), 79–104 (
<emphasis id="FCA0EA95FFAAFF86FE4BFE0C1435FE26" box="[503,519,456,479]" italics="true" pageId="17" pageNumber="92">X</emphasis>
= 88.61) scales around midbody (98–125,
<emphasis id="FCA0EA95FFAAFF86FE55FE2217CBFE04" box="[489,505,486,509]" italics="true" pageId="17" pageNumber="92">X</emphasis>
= 110.05), and 64–93 (
<emphasis id="FCA0EA95FFAAFF86FEBDFDC01723FDE2" box="[257,273,516,539]" italics="true" pageId="17" pageNumber="92">X</emphasis>
= 76.86) vertebrals (81–112,
<emphasis id="FCA0EA95FFAAFF86FDD4FDC0144AFDE2" box="[616,632,516,539]" italics="true" pageId="17" pageNumber="92">X</emphasis>
= 92.21).
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFFAAFF86FF29FDFB1276FD30" pageId="17" pageNumber="92" type="description">
<paragraph id="CE6B3687FFAAFF86FF29FDFB1213FE02" blockId="17.[122,633,189,1722]" lastBlockId="17.[664,1175,190,1359]" pageId="17" pageNumber="92">
<emphasis id="FCA0EA95FFAAFF86FF29FDFB1724FDAF" box="[149,278,575,598]" italics="true" pageId="17" pageNumber="92">Description</emphasis>
.—(1) Maximum SVL in males 108 mm (
<emphasis id="FCA0EA95FFAAFF86FF4CFD9916CCFD8D" box="[240,254,605,628]" italics="true" pageId="17" pageNumber="92">n</emphasis>
= 20); (2) maximum SVL in females 94 mm (
<emphasis id="FCA0EA95FFAAFF86FE8FFDBF1773FD6B" box="[307,321,635,658]" italics="true" pageId="17" pageNumber="92">n</emphasis>
= 15); (3) vertebrals 64–93; (4) paravertebrals 93–123; (5) scales around midbody 79–104; (6) supraoculars 5–7; (7) internasals 2–6; (8) postrostrals 4–6; (9) loreals 2–5; (10) gulars 41–62; (11) subdigitals on Finger IV 22–30; (12) subdigitals on Toe IV 27–34; (13) posthumeral mite pocket present as one or more vertical folds or ridges; (14) postfemoral mite pocket present as one or more vertical folds or ridges; (15) parietal eye visible through interparietal cornea in 5% of specimens; (16) scales on occipitoparietal region small, smooth, juxtaposed; (17) projecting angulate temporals absent; (18) row of enlarged supraoculars occupying most of supraocular region absent; (19) scales on frontonasal region juxtaposed; (20) preauricular fringe present; (21) antegular (continuous medially), antehumeral, gular, longitudinal, oblique, and postauricular neck folds present; (22) lateral nuchals less than half the size of dorsal nuchals; (23) posterior gulars cycloid, smooth, slightly imbricate, not notched; (24) lateral scales reduced in size, approximately half the size of dorsal body scales; (25) vertebrals larger than adjacent paravertebrals; (26) dorsolateral crest absent; (27) ventrals smooth, imbricate; (28) scales on posterior surfaces of thighs granular; (29) inguinal granular pocket present; (30) inguinal groove present; (31) preanals not projected; (32) tail not compressed laterally in adult males; (33) tail length 55–67% of total length; (34) caudal whorls per autotomic segment three; (35) caudals not spinose; (36) dark brown stripe extending anterodorsally from subocular region to supraciliaries absent; (37) dark patch extensively covering gular region in 13% of adult females; (38) dark patch extensively covering gular region of adult males absent; (39) black patch on ventral surface of neck in adult males absent; (40) dark midventral longitudinal mark such as faint line, conspicuous stripe, or extensive patch in adult males absent; (41) dark patches on ventral surface of thighs in adult males absent; (42) postxiphisternal inscriptional ribs not in contact midventrally, Patterns 1B, 2C, and 4B.
</paragraph>
<paragraph id="CE6B3687FFAAFF86FD0FFDC51276FD30" blockId="17.[664,1175,190,1359]" pageId="17" pageNumber="92">
<emphasis id="FCA0EA95FFAAFF86FD0FFDC51570FDE0" box="[691,834,513,537]" italics="true" pageId="17" pageNumber="92">Color in life</emphasis>
.—Dorsum black with yellow flecks or grayish brown with yellow and black spots; black vertical bar on shoulder delimited by irregular yellow line posteriorly in males; dorsal head scales with black borders; venter slightly pink (based on
<bibRefCitation id="AA454B76FFAAFF86FC79FD571269FD52" author="FRITTS, T. H." box="[965,1115,659,683]" journalOrPublisher="San Diego Society of Natural History Memoir" pageId="17" pageNumber="92" pagination="1 - 89" part="7" refId="ref77735" refString="FRITTS, T. H. 1974. A multivariate and evolutionary analysis of the andean iguanid lizards of the genus Stenocercus. San Diego Society of Natural History Memoir 7: 1 - 89." title="A multivariate and evolutionary analysis of the andean iguanid lizards of the genus Stenocercus" type="journal article" year="1974">Fritts [1974]</bibRefCitation>
and a photograph taken by M. Lundberg.)
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFFAAFF86FD0FFD0B15CEFC4D" pageId="17" pageNumber="92" type="biology_ecology">
<paragraph id="CE6B3687FFAAFF86FD0FFD0B15CEFC4D" blockId="17.[664,1175,190,1359]" pageId="17" pageNumber="92">
<emphasis id="FCA0EA95FFAAFF86FD0FFD0B155AFD1F" box="[691,872,719,743]" italics="true" pageId="17" pageNumber="92">Natural History</emphasis>
.—The microhabitat occupied by
<taxonomicName id="09D44D04FFAAFF86FD46FD281549FCFD" authorityName="Boulenger" authorityYear="1911" box="[762,891,748,772]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="17" pageNumber="92" phylum="Chordata" rank="species" species="boettgeri">
<emphasis id="FCA0EA95FFAAFF86FD46FD281549FCFD" box="[762,891,748,772]" italics="true" pageId="17" pageNumber="92">S. boettgeri</emphasis>
</taxonomicName>
usually consists of large piles and walls made of dark rocks partially covered with yellow lichens, which suggests that this species mimics the color of the substrate;
<taxonomicName id="09D44D04FFAAFF86FCADFCA615A4FC80" authorityName="Boulenger" authorityYear="1911" box="[785,918,865,889]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="17" pageNumber="92" phylum="Chordata" rank="species" species="boettgeri">
<emphasis id="FCA0EA95FFAAFF86FCADFCA615A4FC80" box="[785,918,865,889]" italics="true" pageId="17" pageNumber="92">S. boettgeri</emphasis>
</taxonomicName>
seeks refuge in rock crevices, where it wedges its body tightly after inflating its lungs (
<bibRefCitation id="AA454B76FFAAFF86FCD5FC5815DDFC4D" author="FRITTS, T. H." box="[873,1007,924,948]" journalOrPublisher="San Diego Society of Natural History Memoir" pageId="17" pageNumber="92" pagination="1 - 89" part="7" refId="ref77735" refString="FRITTS, T. H. 1974. A multivariate and evolutionary analysis of the andean iguanid lizards of the genus Stenocercus. San Diego Society of Natural History Memoir 7: 1 - 89." title="A multivariate and evolutionary analysis of the andean iguanid lizards of the genus Stenocercus" type="journal article" year="1974">Fritts, 1974</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFFAAFF86FD0FFC7E1545FAB6" pageId="17" pageNumber="92" type="distribution">
<paragraph id="CE6B3687FFAAFF86FD0FFC7E1545FAB6" blockId="17.[664,1175,190,1359]" pageId="17" pageNumber="92">
<emphasis id="FCA0EA95FFAAFF86FD0FFC7E1509FC2B" box="[691,827,954,978]" italics="true" pageId="17" pageNumber="92">Distribution</emphasis>
.—
<taxonomicName id="09D44D04FFAAFF86FCE0FC7E1250FC2B" authorityName="Boulenger" authorityYear="1911" box="[860,1122,954,978]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="17" pageNumber="92" phylum="Chordata" rank="species" species="boettgeri">
<emphasis id="FCA0EA95FFAAFF86FCE0FC7E1250FC2B" box="[860,1122,954,978]" italics="true" pageId="17" pageNumber="92">Stenocercus boettgeri</emphasis>
</taxonomicName>
is known from the central Andes in the eastern Cordillera in Peru (12
<emphasis id="FCA0EA95FFAAFF86FC24FC311590FBF4" bold="true" box="[920,930,1013,1037]" pageId="17" pageNumber="92">°</emphasis>
S–10
<emphasis id="FCA0EA95FFAAFF86FC65FC3115D1FBF4" bold="true" box="[985,995,1013,1037]" pageId="17" pageNumber="92">°</emphasis>
S). This species occurs in the upper valleys of Río Perene at elevations between 2900–3250 m in Departamentos Huánuco, Junín, and Pasco (
<figureCitation id="56EF2A02FFAAFF86FB8BFB8812B8FB9C" box="[1079,1162,1100,1125]" captionStart="FIG" captionStartId="18.[165,179,1042,1061]" captionTargetBox="[143,1197,189,1025]" captionTargetId="figure-251@18.[143,1197,189,1025]" captionTargetPageId="18" captionText="FIG. 12.—Distribution of six species of Stenocercus in Peru. Black triangle pointed by arrow corresponds to María Teresa (10°42'05&quot;S, 75°27'22&quot;W, 1470 m, Departamento Pasco), where Stenocercus boettgeri, S. formosus, S. scapularis, and S. torquatus occur in sympatry." figureDoi="http://doi.org/10.5281/zenodo.14384828" httpUri="https://zenodo.org/record/14384828/files/figure.png" pageId="17" pageNumber="92">Fig. 12</figureCitation>
).
<taxonomicName id="09D44D04FFAAFF86FD24FBAE152BFB7B" authorityName="Boulenger" authorityYear="1911" box="[664,793,1130,1154]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="17" pageNumber="92" phylum="Chordata" rank="species" species="boettgeri">
<emphasis id="FCA0EA95FFAAFF86FD24FBAE152BFB7B" box="[664,793,1130,1154]" italics="true" pageId="17" pageNumber="92">S. boettgeri</emphasis>
</taxonomicName>
is sympatric with
<taxonomicName id="09D44D04FFAAFF86FC52FBAE1242FB7B" box="[1006,1136,1130,1154]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="17" pageNumber="92" phylum="Chordata" rank="species" species="formosus">
<emphasis id="FCA0EA95FFAAFF86FC52FBAE1242FB7B" box="[1006,1136,1130,1154]" italics="true" pageId="17" pageNumber="92">S. formosus</emphasis>
</taxonomicName>
,
<taxonomicName id="09D44D04FFAAFF86FB3DFBAE1535FB66" baseAuthorityName="Boulenger" baseAuthorityYear="1901" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="17" pageNumber="92" phylum="Chordata" rank="species" species="scapularis">
<emphasis id="FCA0EA95FFAAFF86FB3DFBAE1535FB66" italics="true" pageId="17" pageNumber="92">S. scapularis</emphasis>
</taxonomicName>
, and
<taxonomicName id="09D44D04FFAAFF86FCF6FB4315E1FB66" box="[842,979,1159,1183]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="17" pageNumber="92" phylum="Chordata" rank="species" species="torquatus">
<emphasis id="FCA0EA95FFAAFF86FCF6FB4315E1FB66" box="[842,979,1159,1183]" italics="true" pageId="17" pageNumber="92">S. torquatus</emphasis>
</taxonomicName>
at María Teresa, 10
<emphasis id="FCA0EA95FFAAFF86FD0FFB60148FFB45" bold="true" box="[691,701,1188,1212]" pageId="17" pageNumber="92">°</emphasis>
42'05&quot;S, 75
<emphasis id="FCA0EA95FFAAFF86FCF9FB60157DFB45" bold="true" box="[837,847,1188,1212]" pageId="17" pageNumber="92">°</emphasis>
27'22&quot;W, 1470 m, Departamento Pasco (
<bibRefCitation id="AA454B76FFAAFF86FCFEFB0612B9FB23" author="TORRES-CARVAJAL, O. &amp; E. LEHR &amp; M. LUNDBERG" box="[834,1163,1218,1242]" journalOrPublisher="Herpetologica" pageId="17" pageNumber="92" pagination="440 - 448" part="61" refId="ref80223" refString="TORRES-CARVAJAL, O., E. LEHR, AND M. LUNDBERG. 2005. Resurrection of Stenocercus torquatus Boulenger, a spiny-tailed iguanid lizard (Squamata: Iguania) from Peru. Herpetologica 61 (4): 440 - 448." title="Resurrection of Stenocercus torquatus Boulenger, a spiny-tailed iguanid lizard (Squamata: Iguania) from Peru" type="journal article" year="2005">Torres-Carvajal et al., 2005</bibRefCitation>
). This species has been erroneously reported for northern Peru (
<bibRefCitation id="AA454B76FFAAFF86FCCDFB3815C7FAED" author="FRITTS, T. H." box="[881,1013,1276,1300]" journalOrPublisher="San Diego Society of Natural History Memoir" pageId="17" pageNumber="92" pagination="1 - 89" part="7" refId="ref77735" refString="FRITTS, T. H. 1974. A multivariate and evolutionary analysis of the andean iguanid lizards of the genus Stenocercus. San Diego Society of Natural History Memoir 7: 1 - 89." title="A multivariate and evolutionary analysis of the andean iguanid lizards of the genus Stenocercus" type="journal article" year="1974">Fritts, 1974</bibRefCitation>
) and Ecuador (Etheridge, in
<bibRefCitation id="AA454B76FFAAFF86FCEFFADE14E2FAB6" author="PETERS, J. A. &amp; R. DONOSO-BARROS" journalOrPublisher="United States National Museum Bulletin" pageId="17" pageNumber="92" pagination="1 - 293" part="297" refId="ref79234" refString="PETERS, J. A., AND R. DONOSO-BARROS. 1970. Catalogue of the Neotropical Squamata. Part 2, lizards and amphisbaenians. United States National Museum Bulletin 297: 1 - 293." title="Catalogue of the Neotropical Squamata. Part 2, lizards and amphisbaenians" type="journal article" year="1970">Peters and Donoso-Barros, 1970</bibRefCitation>
;
<bibRefCitation id="AA454B76FFAAFF86FD61FAF31559FAB6" author="PETERS, J. A." box="[733,875,1335,1359]" journalOrPublisher="Proceedings of the United States National Museum" pageId="17" pageNumber="92" pagination="1 - 49" part="119" refId="ref79202" refString="PETERS, J. A. 1967. The lizards of Ecuador, a check list and key. Proceedings of the United States National Museum 119: 1 - 49." title="The lizards of Ecuador, a check list and key" type="proceedings paper" year="1967">Peters, 1967</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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