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<document id="4746A5A1D420525AE7A6CBC89E69B378" ID-DOI="10.1655/06-001.1" ID-Zenodo-Dep="14384794" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1733861782093" checkinUser="juliana" docAuthor="Torres-Carvajal, Omar" docDate="2007" docId="467D8791FF84FFD7FF5EFA541718FCFD" docLanguage="en" docName="HerpetolMonogr.21.1.76-178.pdf.pdf" docOrigin="Herpetological Monographs 21 (1)" docSource="https://doi.org/10.1655/06-001.1" docTitle="Stenocercus nubicola Fritts 1972" docType="treatment" docVersion="6" lastPageNumber="139" masterDocId="BA44FFE9FFBBFF97FFBCFFC41632FFF9" masterDocTitle="A TAXONOMIC REVISION OF SOUTH AMERICAN STENOCERCUS (SQUAMATA: IGUANIA) LIZARDS" masterLastPageNumber="178" masterPageNumber="76" pageNumber="138" updateTime="1733923315685" updateUser="juliana">
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<mods:title id="9CDDAB252046CE0666973A2002C225BF">A TAXONOMIC REVISION OF SOUTH AMERICAN STENOCERCUS (SQUAMATA: IGUANIA) LIZARDS</mods:title>
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<mods:roleTerm id="9018A46CBA9D00E1C20C6A0EDAFE5C9A">Author</mods:roleTerm>
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<mods:namePart id="A326B43F709732C656E8E267ABB5CA2E">Torres-Carvajal, Omar</mods:namePart>
<mods:affiliation id="CBE1AD224BB6CA658F2B9701556837AA">Department of Vertebrate Zoologŋ, National Museum of Natural Historŋ, Smithsonian Institution, Washington, DC 20560, USA &amp; Escuela de Biología, Pontificia UniƲersidad Católica del Ecuador, AƲenida 12 de Octubre ŋ Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
<mods:nameIdentifier id="F4A193A2CD92BFA395CEA36F28C08369" type="email">Torreso@si.edu</mods:nameIdentifier>
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<mods:typeOfResource id="5A551F5AAF965557AB4CCFCD617F6E0C">text</mods:typeOfResource>
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<mods:title id="5185F44559D840CFABD83C30A708D859">Herpetological Monographs</mods:title>
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<mods:part id="D0C4DCB18662EC38A9AA4C12B90020B8">
<mods:date id="392E9EC7495FF5AF26BF0E41EB81A9A1">2007</mods:date>
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<mods:number id="B47FB58CB4735404C3372C11EB509651">2007-12-28</mods:number>
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<mods:number id="B30DF9AAB9800B796F49E610392F6342">21</mods:number>
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<mods:classification id="C751200E01EF26732B2C8A331E922002">journal article</mods:classification>
<mods:identifier id="072C164A258F67A1868D7D04364AAA15" type="DOI">10.1655/06-001.1</mods:identifier>
<mods:identifier id="370B4E221DDEE2E9011A182750C15AB0" type="Zenodo-Dep">14384794</mods:identifier>
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<treatment id="467D8791FF84FFD7FF5EFA541718FCFD" ID-DOI="http://doi.org/10.5281/zenodo.14372743" ID-Zenodo-Dep="14372743" LSID="urn:lsid:plazi:treatment:467D8791FF84FFD7FF5EFA541718FCFD" httpUri="http://treatment.plazi.org/id/467D8791FF84FFD7FF5EFA541718FCFD" lastPageId="64" lastPageNumber="139" pageId="63" pageNumber="138">
<subSubSection id="86CE650CFF84FFA8FF5EFA541423FA51" box="[226,529,1424,1448]" pageId="63" pageNumber="138" type="nomenclature">
<paragraph id="CE6B3687FF84FFA8FF5EFA541423FA51" blockId="63.[122,633,1424,1722]" box="[226,529,1424,1448]" pageId="63" pageNumber="138">
<heading id="952381EBFF84FFA8FF5EFA541423FA51" box="[226,529,1424,1448]" centered="true" fontSize="10" level="3" pageId="63" pageNumber="138" reason="4">
<taxonomicName id="09D44D04FF84FFA8FF5EFA541423FA51" ID-CoL="6ZLRL" authority="Fritts" authorityName="Fritts" authorityYear="1972" box="[226,529,1424,1448]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="63" pageNumber="138" phylum="Chordata" rank="species" species="nubicola">
<emphasis id="FCA0EA95FF84FFA8FF5EFA5417F9FA51" box="[226,459,1424,1448]" italics="true" pageId="63" pageNumber="138">Stenocercus nubicola</emphasis>
Fritts
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFF84FFA8FEF5FA6A1798FA3F" box="[329,426,1454,1478]" pageId="63" pageNumber="138" type="description">
<paragraph id="CE6B3687FF84FFA8FEF5FA6A1798FA3F" blockId="63.[122,633,1424,1722]" box="[329,426,1454,1478]" pageId="63" pageNumber="138">
<heading id="952381EBFF84FFA8FEF5FA6A1798FA3F" box="[329,426,1454,1478]" centered="true" fontSize="10" level="3" pageId="63" pageNumber="138" reason="4">
(
<figureCitation id="56EF2A02FF84FFA8FEEDFA6A1790FA3F" box="[337,418,1454,1478]" captionStart="FIG" captionStartId="59.[143,157,1718,1737]" captionTargetBox="[122,1174,189,1701]" captionTargetId="figure-1@59.[122,1174,189,1701]" captionTargetPageId="59" captionText="FIG. 20.—Eight species of Stenocercus. (A) S. marmoratus (AMU); (B) S. melanopŋgus (PVE); (C) S. modestus (MLU); (D) S. nigromaculatus (WED); (E) S. nubicola (PVE); (F) S. ochoai (WED); (G) S. orientalis (PVE); (H) S. ornatissimus (MLU)." figureDoi="http://doi.org/10.5281/zenodo.14384850" httpUri="https://zenodo.org/record/14384850/files/figure.png" pageId="63" pageNumber="138">Fig. 20</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFF84FFA8FFC6FA11141BF998" pageId="63" pageNumber="138" type="reference_group">
<paragraph id="CE6B3687FF84FFA8FFC6FA11141BF998" blockId="63.[122,633,1424,1722]" pageId="63" pageNumber="138">
<taxonomicName id="09D44D04FF84FFA8FFC6FA111417FA15" authority="Fritts, 1972: 11" authorityName="Fritts" authorityPageNumber="11" authorityYear="1972" box="[122,549,1492,1516]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="63" pageNumber="138" phylum="Chordata" rank="species" species="nubicola">
<emphasis id="FCA0EA95FF84FFA8FFC6FA111758FA15" box="[122,362,1492,1516]" italics="true" pageId="63" pageNumber="138">Stenocercus nubicola</emphasis>
<bibRefCitation id="AA454B76FF84FFA8FEC5FA101417FA15" author="FRITTS, T. H." box="[377,549,1492,1516]" journalOrPublisher="Ocassional Papers University of Kansas Museum of Natural History" pageId="63" pageNumber="138" pagination="1 - 21" part="10" refId="ref77696" refString="FRITTS, T. H. 1972. New species of lizards of the genus Stenocercus from Peru (Sauria: Iguanidae). Ocassional Papers University of Kansas Museum of Natural History 10: 1 - 21." title="New species of lizards of the genus Stenocercus from Peru (Sauria: Iguanidae)" type="journal article" year="1972">Fritts, 1972:11</bibRefCitation>
</taxonomicName>
.
<materialsCitation id="7EBC3CDAFF84FFA8FD84FA10141BF998" collectionCode="KU" country="Peru" elevation="3100" location="Cordillera Huancabamba" municipality="Canchaque" pageId="63" pageNumber="138" specimenCode="KU 134107" specimenCount="1" specimenCount-male="1" stateProvince="Departamento Piura" typeStatus="holotype">
<typeStatus id="116F8825FF84FFA8FD84FA1016FAF9F3" pageId="63" pageNumber="138">Holotype</typeStatus>
:
<specimenCode id="9E729EFCFF84FFA8FF69FA36176AF9F3" box="[213,344,1522,1546]" pageId="63" pageNumber="138">KU 134107</specimenCode>
, a
<specimenCount id="D8D2FD0EFF84FFA8FEC0FA351783F9F0" box="[380,433,1521,1545]" pageId="63" pageNumber="138" type="male">male</specimenCount>
from ‘‘summit of
<location id="CB0B605CFF84FFA8FF29F9CB179FF9DE" LSID="urn:lsid:plazi:treatment:467D8791FF84FFD7FF5EFA541718FCFD:CB0B605CFF84FFA8FF29F9CB179FF9DE" box="[149,429,1551,1575]" country="Peru" municipality="Canchaque" name="Cordillera Huancabamba" pageId="63" pageNumber="138" stateProvince="Departamento Piura">Cordillera Huancabamba</location>
between
<collectingMunicipality id="2E0FACFDFF84FFA8FDA0F9CB168CF9BD" pageId="63" pageNumber="138">Canchaque</collectingMunicipality>
and Huancabamba,
<quantity id="092C9B62FF84FFA8FE13F9E81437F9BD" box="[431,517,1580,1604]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="63" pageNumber="138" unit="m" value="3100.0">
<elevation id="45F9D1B4FF84FFA8FE13F9E81437F9BD" box="[431,517,1580,1604]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="63" pageNumber="138" unit="m" value="3100.0">3100 m</elevation>
</quantity>
,
<collectingRegion id="0C10F865FF84FFA8FDA8F9E91713F998" pageId="63" pageNumber="138">Departamento Piura</collectingRegion>
,
<collectingCountry id="B6C37617FF84FFA8FE93F98D1756F998" box="[303,356,1609,1633]" name="Peru" pageId="63" pageNumber="138">Perú</collectingCountry>
̕̕;
<bibRefCitation id="AA454B76FF84FFA8FEC3F98D1417F998" author="FRITTS, T. H." box="[383,549,1609,1633]" journalOrPublisher="San Diego Society of Natural History Memoir" pageId="63" pageNumber="138" pagination="1 - 89" part="7" refId="ref77735" refString="FRITTS, T. H. 1974. A multivariate and evolutionary analysis of the andean iguanid lizards of the genus Stenocercus. San Diego Society of Natural History Memoir 7: 1 - 89." title="A multivariate and evolutionary analysis of the andean iguanid lizards of the genus Stenocercus" type="journal article" year="1974">Fritts, 1974:59</bibRefCitation>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFF84FFA8FF29F9AC1237FE19" pageId="63" pageNumber="138" type="diagnosis">
<paragraph id="CE6B3687FF84FFA8FF29F9AC1237FE19" blockId="63.[122,633,1424,1722]" lastBlockId="63.[664,1175,190,1722]" pageId="63" pageNumber="138">
<emphasis id="FCA0EA95FF84FFA8FF29F9AC1733F986" box="[149,257,1640,1663]" italics="true" pageId="63" pageNumber="138">Diagnosis</emphasis>
.—
<taxonomicName id="09D44D04FF84FFA8FE9EF9AC1428F979" authorityName="Fritts" authorityYear="1972" box="[290,538,1640,1664]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="63" pageNumber="138" phylum="Chordata" rank="species" species="nubicola">
<emphasis id="FCA0EA95FF84FFA8FE9EF9AC1428F979" box="[290,538,1640,1664]" italics="true" pageId="63" pageNumber="138">Stenocercus nubicola</emphasis>
</taxonomicName>
differs from all other species of
<taxonomicName id="09D44D04FF84FFA8FE1FF9411414F965" box="[419,550,1669,1692]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="63" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="FCA0EA95FF84FFA8FE1FF9411414F965" box="[419,550,1669,1692]" italics="true" pageId="63" pageNumber="138">Stenocercus</emphasis>
</taxonomicName>
except
<taxonomicName id="09D44D04FF84FFA8FFC6F96716E5F943" authorityName="Torres-Carvajal" authorityYear="2005" box="[122,215,1698,1722]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="63" pageNumber="138" phylum="Chordata" rank="species" species="frittsi">
<emphasis id="FCA0EA95FF84FFA8FFC6F96716E5F943" box="[122,215,1698,1722]" italics="true" pageId="63" pageNumber="138">S. frittsi</emphasis>
</taxonomicName>
and
<taxonomicName id="09D44D04FF84FFA8FEAAF96717AEF943" authorityName="Fritts" authorityYear="1974" box="[278,412,1698,1722]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="63" pageNumber="138" phylum="Chordata" rank="species" species="variabilis">
<emphasis id="FCA0EA95FF84FFA8FEAAF96717AEF943" box="[278,412,1698,1722]" italics="true" pageId="63" pageNumber="138">S. variabilis</emphasis>
</taxonomicName>
in having granular scales on the posterior surface of thighs, a distinct vertebral crest, lateral nuchals less than half the size of dorsal nuchals, strongly keeled lateral body scales, and three caudal whorls per autotomic segment. It can be distinguished from
<taxonomicName id="09D44D04FF84FFA8FCCAFE9615E0FE93" authorityName="Torres-Carvajal" authorityYear="2005" box="[886,978,338,362]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="63" pageNumber="138" phylum="Chordata" rank="species" species="frittsi">
<emphasis id="FCA0EA95FF84FFA8FCCAFE9615E0FE93" box="[886,978,338,362]" italics="true" pageId="63" pageNumber="138">S. frittsi</emphasis>
</taxonomicName>
and
<taxonomicName id="09D44D04FF84FFA8FBACFE9612A4FE90" authorityName="Fritts" authorityYear="1974" box="[1040,1174,337,361]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="63" pageNumber="138" phylum="Chordata" rank="species" species="variabilis">
<emphasis id="FCA0EA95FF84FFA8FBACFE9612A4FE90" box="[1040,1174,337,361]" italics="true" pageId="63" pageNumber="138">S. variabilis</emphasis>
</taxonomicName>
by having a distinct mite pocket underneath the oblique neck fold, lateral and dorsal body scales similar in size, and a black longitudinal midventral stripe in adult males.
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFF84FFD7FD0FFE231791FE24" lastPageId="64" lastPageNumber="139" pageId="63" pageNumber="138" type="description">
<paragraph id="CE6B3687FF84FFA8FD0FFE2315BCF943" blockId="63.[664,1175,190,1722]" pageId="63" pageNumber="138">
<emphasis id="FCA0EA95FF84FFA8FD0FFE231506FE07" box="[691,820,487,510]" italics="true" pageId="63" pageNumber="138">Description</emphasis>
.—(1) Maximum SVL in males 71 mm (
<emphasis id="FCA0EA95FF84FFA8FCBAFDC01526FDE2" box="[774,788,516,539]" italics="true" pageId="63" pageNumber="138">n</emphasis>
= 1); (3) vertebrals 67; (4) paravertebrals 97; (5) scales around midbody 93–94; (6) supraoculars 5–6; (7) internasals four; (8) postrostrals four; (9) loreals three; (10) gulars 38–39; (11) subdigitals on Finger IV 17–22; (12) subdigitals on Toe IV 24–28; (13) posthumeral mite pocket present as a shallow depression with a wide opening; (14) postfemoral mite pocket distinct with slit-like opening; (15) parietal eye not visible through interparietal cornea; (16) scales on occipitoparietal region small, smooth, juxtaposed; (17) projecting angulate temporals absent; (18) row of enlarged supraoculars occupying most of supraocular region absent; (19) scales on frontonasal region juxtaposed anteriorly; (20) preauricular fringe present; (21) antehumeral, gular, longitudinal, oblique, and supra-auricular neck folds present; (22) lateral nuchals less than half the size of dorsal nuchals; (23) posterior gulars cycloid, smooth, slightly imbricate, not notched; (24) lateral and dorsal body scales similar in size; (25) vertebrals larger than adjacent paravertebrals; (26) dorsolateral crest absent; (27) ventrals smooth, imbricate; (28) scales on posterior surfaces of thighs granular; (29) inguinal granular pocket absent; (30) inguinal groove absent; (31) preanals not projected; (32) tail not compressed laterally in adult males; (33) tail length 67% of total length; (34) caudal whorls per autotomic segment three; (35) caudals not spinose; (36) dark brown stripe extending anterodorsally from subocular region to supraciliaries absent; (38) dark patch extensively covering gular region of adult males present; (39) black patch on ventral surface of neck in adult males absent; (40) dark midventral longitudinal stripe present; (41) dark patches on ventral surface of thighs in adult males absent.
</paragraph>
<paragraph id="CE6B3687FFFBFFD7FF16FF7A1791FE24" blockId="64.[143,654,190,772]" pageId="64" pageNumber="139">
<emphasis id="FCA0EA95FFFBFFD7FF16FF7A178DFF2F" box="[170,447,190,214]" italics="true" pageId="64" pageNumber="139">Color in life of holotype</emphasis>
.—Dorsum brown with small dark brown or black blotches, bluish-green transverse marks, and beige dorsolateral stripes; antehumeral region with black vertical bar; supralabials and infralabials black; loreal region white; gular and pectoral regions white with black spots; wide, gray midventral stripe; lateral aspect of body and venter light green with reddish-orange spots and flecks (
<bibRefCitation id="AA454B76FFFBFFD7FEADFE0117A5FE24" author="FRITTS, T. H." box="[273,407,453,477]" journalOrPublisher="Ocassional Papers University of Kansas Museum of Natural History" pageId="64" pageNumber="139" pagination="1 - 21" part="10" refId="ref77696" refString="FRITTS, T. H. 1972. New species of lizards of the genus Stenocercus from Peru (Sauria: Iguanidae). Ocassional Papers University of Kansas Museum of Natural History 10: 1 - 21." title="New species of lizards of the genus Stenocercus from Peru (Sauria: Iguanidae)" type="journal article" year="1972">Fritts, 1972</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFFFBFFD7FF16FE2716E1FDCF" pageId="64" pageNumber="139" type="biology_ecology">
<paragraph id="CE6B3687FFFBFFD7FF16FE2716E1FDCF" blockId="64.[143,654,190,772]" pageId="64" pageNumber="139">
<emphasis id="FCA0EA95FFFBFFD7FF16FE27175EFE02" box="[170,364,483,507]" italics="true" pageId="64" pageNumber="139">Natural History</emphasis>
.—The holotype (adult male) was found basking on a fallen tree trunk.
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFFFBFFD7FF16FDF81718FCFD" pageId="64" pageNumber="139" type="distribution">
<paragraph id="CE6B3687FFFBFFD7FF16FDF81718FCFD" blockId="64.[143,654,190,772]" pageId="64" pageNumber="139">
<emphasis id="FCA0EA95FFFBFFD7FF16FDF81703FDAD" box="[170,305,572,596]" italics="true" pageId="64" pageNumber="139">Distribution</emphasis>
.—
<taxonomicName id="09D44D04FFFBFFD7FEEFFDF9140EFDAD" authorityName="Fritts" authorityYear="1972" box="[339,572,572,596]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="64" pageNumber="139" phylum="Chordata" rank="species" species="nubicola">
<emphasis id="FCA0EA95FFFBFFD7FEEFFDF9140EFDAD" box="[339,572,572,596]" italics="true" pageId="64" pageNumber="139">Stenocercus nubicola</emphasis>
</taxonomicName>
is only known from its type locality (5
<emphasis id="FCA0EA95FFFBFFD7FD89FD9D140DFD88" bold="true" box="[565,575,601,625]" pageId="64" pageNumber="139">°</emphasis>
19'0&quot;S, 79
<emphasis id="FCA0EA95FFFBFFD7FF16FDB31686FD76" bold="true" box="[170,180,631,655]" pageId="64" pageNumber="139">°</emphasis>
29'0&quot;W, 3100 m) in Departamento Piura, Peru (
<figureCitation id="56EF2A02FFFBFFD7FF6BFD50171BFD55" box="[215,297,660,684]" captionStart="FIG" captionStartId="42.[333,347,1042,1061]" captionTargetBox="[143,1196,189,1025]" captionTargetId="figure-262@42.[143,1196,189,1025]" captionTargetPageId="42" captionText="FIG. 17.—Distribution of four species of Stenocercus in Ecuador and Peru." figureDoi="http://doi.org/10.5281/zenodo.14384842" httpUri="https://zenodo.org/record/14384842/files/figure.png" pageId="64" pageNumber="139">Fig. 17</figureCitation>
). This locality lies north of the western Cordillera of the central Andes between 6
<emphasis id="FCA0EA95FFFBFFD7FEB4FD0B1720FD1E" bold="true" box="[264,274,719,743]" pageId="64" pageNumber="139">°</emphasis>
S–5
<emphasis id="FCA0EA95FFFBFFD7FE80FD0B1774FD1E" bold="true" box="[316,326,719,743]" pageId="64" pageNumber="139">°</emphasis>
S. This species is sympatric with
<taxonomicName id="09D44D04FFFBFFD7FF74FD281710FCFD" authorityName="Fritts" authorityYear="1972" box="[200,290,748,772]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="64" pageNumber="139" phylum="Chordata" rank="species" species="ivitus">
<emphasis id="FCA0EA95FFFBFFD7FF74FD281710FCFD" box="[200,290,748,772]" italics="true" pageId="64" pageNumber="139">S. ivitus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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