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<document id="E38474F9E3F9DE7F8633D4D14A0CCD22" ID-DOI="10.1655/06-001.1" ID-Zenodo-Dep="14384794" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1733861782093" checkinUser="juliana" docAuthor="Torres-Carvajal, Omar" docDate="2007" docId="467D8791FF81FFABFEBAFC1314BCFB0E" docLanguage="en" docName="HerpetolMonogr.21.1.76-178.pdf.pdf" docOrigin="Herpetological Monographs 21 (1)" docSource="https://doi.org/10.1655/06-001.1" docTitle="Stenocercus marmoratus" docType="treatment" docVersion="6" lastPageNumber="135" masterDocId="BA44FFE9FFBBFF97FFBCFFC41632FFF9" masterDocTitle="A TAXONOMIC REVISION OF SOUTH AMERICAN STENOCERCUS (SQUAMATA: IGUANIA) LIZARDS" masterLastPageNumber="178" masterPageNumber="76" pageNumber="133" updateTime="1733923315685" updateUser="juliana">
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<mods:title id="124D32CD4F2421D734118A9D980B5505">A TAXONOMIC REVISION OF SOUTH AMERICAN STENOCERCUS (SQUAMATA: IGUANIA) LIZARDS</mods:title>
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<mods:namePart id="704C859B5F451044377E805BB8E0C3A7">Torres-Carvajal, Omar</mods:namePart>
<mods:affiliation id="D4711C688289A901F0220415F662A2B4">Department of Vertebrate Zoologŋ, National Museum of Natural Historŋ, Smithsonian Institution, Washington, DC 20560, USA &amp; Escuela de Biología, Pontificia UniƲersidad Católica del Ecuador, AƲenida 12 de Octubre ŋ Roca, Apartado 17 - 01 - 2184, Quito, Ecuador</mods:affiliation>
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<taxonomicName id="09D44D04FF81FFADFEBAFC131438FBF2" ID-CoL="4ZWK9" authority="(Dumeril and Bibron)" baseAuthorityName="Dumeril and Bibron" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="58" pageNumber="133" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="FCA0EA95FF81FFADFEBAFC131425FC17" box="[262,535,983,1006]" italics="true" pageId="58" pageNumber="133">Stenocercus marmoratus</emphasis>
(Duméril and Bibron)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFF81FFADFEE2FBD6178DFBD3" box="[350,447,1042,1066]" pageId="58" pageNumber="133" type="description">
<paragraph id="CE6B3687FF81FFADFEE2FBD6178DFBD3" blockId="58.[143,654,983,1722]" box="[350,447,1042,1066]" pageId="58" pageNumber="133">
<heading id="952381EBFF81FFADFEE2FBD6178DFBD3" box="[350,447,1042,1066]" centered="true" fontSize="10" level="3" pageId="58" pageNumber="133" reason="4">
(
<figureCitation id="56EF2A02FF81FFADFEDAFBD61785FBD3" box="[358,439,1042,1066]" captionStart="FIG" captionStartId="59.[143,157,1718,1737]" captionTargetBox="[122,1174,189,1701]" captionTargetId="figure-1@59.[122,1174,189,1701]" captionTargetPageId="59" captionText="FIG. 20.—Eight species of Stenocercus. (A) S. marmoratus (AMU); (B) S. melanopŋgus (PVE); (C) S. modestus (MLU); (D) S. nigromaculatus (WED); (E) S. nubicola (PVE); (F) S. ochoai (WED); (G) S. orientalis (PVE); (H) S. ornatissimus (MLU)." figureDoi="http://doi.org/10.5281/zenodo.14384850" httpUri="https://zenodo.org/record/14384850/files/figure.png" pageId="58" pageNumber="133">Fig. 20</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFF81FFADFF33FBFC17CCF987" pageId="58" pageNumber="133" type="reference_group">
<paragraph id="CE6B3687FF81FFADFF33FBFC1732FAE4" blockId="58.[143,654,983,1722]" pageId="58" pageNumber="133">
<taxonomicName id="09D44D04FF81FFADFF33FBFC176EFB94" authority="Dumeril and Bibron, 1837: 356" authorityName="Dumeril and Bibron" authorityPageNumber="356" authorityYear="1837" class="Squamata" family="Iguanidae" genus="Trachncnclus" kingdom="Animalia" pageId="58" pageNumber="133" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="FCA0EA95FF81FFADFF33FBFC1785FBA9" box="[143,439,1080,1104]" italics="true" pageId="58" pageNumber="133">Trachycyclus marmoratus</emphasis>
<bibRefCitation id="AA454B76FF81FFADFE7AFBFC176EFB94" author="DUMERIL, A. M. &amp; G. BIBRON" bookContentInfo="Librairie Encyclopedique de Roret, Paris, France." pageId="58" pageNumber="133" refId="ref77458" refString="DUMERIL, A. M., AND G. BIBRON. 1837. Erpetologie Generale ou Histoire Naturelle Complete des Reptiles. Librairie Encyclopedique de Roret, Paris, France." title="Erpetologie Generale ou Histoire Naturelle Complete des Reptiles" type="book" year="1837">Duméril and Bibron, 1837:356</bibRefCitation>
</taxonomicName>
.
<materialsCitation id="7EBC3CDAFF81FFADFED2FB911732FAE4" collectionCode="MNHN" country="Bolivia" location="Rio-Grande" municipality="Provincia de La Laguna" pageId="58" pageNumber="133" specimenCode="MNHN 2513" specimenCount="1" specimenCount-juvenile="1" stateProvince="Departamento Chuquisaca" typeStatus="holotype">
<typeStatus id="116F8825FF81FFADFED2FB9117EAFB94" box="[366,472,1109,1133]" pageId="58" pageNumber="133">Holotype</typeStatus>
:
<specimenCode id="9E729EFCFF81FFADFE56FB9214B8FB94" box="[490,650,1109,1133]" pageId="58" pageNumber="133">MNHN 2513</specimenCode>
, a
<specimenCount id="D8D2FD0EFF81FFADFF7DFBB6172AFB73" box="[193,280,1138,1162]" pageId="58" pageNumber="133" type="juvenile">juvenile</specimenCount>
from province de
<location id="CB0B605CFF81FFADFE43FBB614B8FB72" LSID="urn:lsid:plazi:treatment:467D8791FF81FFABFEBAFC1314BCFB0E:CB0B605CFF81FFADFE43FBB614B8FB72" box="[511,650,1138,1163]" country="Bolivia" municipality="Provincia de La Laguna" name="Rio-Grande" pageId="58" pageNumber="133" stateProvince="Departamento Chuquisaca">Rio-Grande</location>
̕̕ (restricted to
<location id="CB0B605CFF81FFADFEF0FB5417E8FB51" LSID="urn:lsid:plazi:treatment:467D8791FF81FFABFEBAFC1314BCFB0E:CB0B605CFF81FFADFEF0FB5417E8FB51" box="[332,474,1168,1192]" country="Bolivia" municipality="Provincia de La Laguna" name="Pampa Ruiz" pageId="58" pageNumber="133" stateProvince="Departamento Chuquisaca">Pampa Ruiz</location>
, between
<location id="CB0B605CFF81FFADFDEAFB5416CFFB3C" LSID="urn:lsid:plazi:treatment:467D8791FF81FFABFEBAFC1314BCFB0E:CB0B605CFF81FFADFDEAFB5416CFFB3C" country="Bolivia" municipality="Provincia de La Laguna" name="Valle Grande" pageId="58" pageNumber="133" stateProvince="Departamento Chuquisaca">Valle Grande</location>
and
<location id="CB0B605CFF81FFADFEFCFB6917F6FB3C" LSID="urn:lsid:plazi:treatment:467D8791FF81FFABFEBAFC1314BCFB0E:CB0B605CFF81FFADFEFCFB6917F6FB3C" box="[320,452,1197,1221]" country="Bolivia" municipality="Provincia de La Laguna" name="El Pescado" pageId="58" pageNumber="133" stateProvince="Departamento Chuquisaca">El Pescado</location>
,
<collectingMunicipality id="2E0FACFDFF81FFADFE69FB6916CDFB1A" pageId="58" pageNumber="133">Provincia de La Laguna</collectingMunicipality>
,
<locationDeviation id="DFD92FA5FF81FFADFEA5FB0F143AFB1B" box="[281,520,1226,1250]" pageId="58" pageNumber="133">east of Chuquisaca</locationDeviation>
, [
<collectingRegion id="0C10F865FF81FFADFD9BFB0F17B1FB06" pageId="58" pageNumber="133">Departamento Chuquisaca</collectingRegion>
],
<collectingCountry id="B6C37617FF81FFADFE26FB2317DBFB06" box="[410,489,1255,1279]" name="Bolivia" pageId="58" pageNumber="133">Bolivia</collectingCountry>
, by d̕
<bibRefCitation id="AA454B76FF81FFADFD8CFB2216C6FAE4" author="ORBIGNY, A." journalOrPublisher="Reptiles." pageId="58" pageNumber="133" pagination="1 - 12" part="5" refId="ref77328" refString="D ̕ ORBIGNY, A. 1847. Voyage dans L ̕ Amerique Meridionale execute pendant les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833. Reptiles. 5 (1): 1 - 12." title="Voyage dans L ̕ Amerique Meridionale execute pendant les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833" type="journal article" year="1847">Orbigny [1847]</bibRefCitation>
.)
</materialsCitation>
</paragraph>
<paragraph id="CE6B3687FF81FFADFF33FAE01708FAA1" blockId="58.[143,654,983,1722]" pageId="58" pageNumber="133">
<taxonomicName id="09D44D04FF81FFADFF33FAE01705FAA1" authority="Fitzinger, 1843: 71" authorityName="Fitzinger" authorityPageNumber="71" authorityYear="1843" class="Squamata" family="Tropiduridae" genus="Heterotropis" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" pageId="58" pageNumber="133" phylum="Chordata" rank="species" species="marmorata" subGenus="Trachncnlus">
<emphasis id="FCA0EA95FF81FFADFF33FAE01728FAC2" box="[143,282,1316,1339]" italics="true" pageId="58" pageNumber="133">Heterotropis</emphasis>
(
<emphasis id="FCA0EA95FF81FFADFE97FAE71783FAC2" box="[299,433,1315,1339]" italics="true" pageId="58" pageNumber="133">Trachycylus</emphasis>
)
<emphasis id="FCA0EA95FF81FFADFE7DFAE01409FAC2" box="[449,571,1316,1339]" italics="true" pageId="58" pageNumber="133">marmorata</emphasis>
<bibRefCitation id="AA454B76FF81FFADFDF8FAE71705FAA1" author="FITZINGER, L. I." bookContentInfo="Baumuller und Seidel, Wien, Austria" pageId="58" pageNumber="133" refId="ref77645" refString="FITZINGER, L. I. 1843. Systema Reptilium, Fasciculus Primus, Amblyglossae. Baumuller und Seidel, Wien, Austria." title="Systema Reptilium, Fasciculus Primus, Amblyglossae" type="book" year="1843">Fitzinger, 1843:71</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="CE6B3687FF81FFADFF33FA9B144BFA37" blockId="58.[143,654,983,1722]" pageId="58" pageNumber="133">
<taxonomicName id="09D44D04FF81FFADFF33FA9B16E6FA6D" authority="Boulenger, 1885 a: 132" authorityName="Boulenger" authorityPageNumber="132" authorityYear="1885" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="58" pageNumber="133" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="FCA0EA95FF81FFADFF33FA9B179BFA8F" box="[143,425,1375,1398]" italics="true" pageId="58" pageNumber="133">Stenocercus marmoratus</emphasis>
<bibRefCitation id="AA454B76FF81FFADFE06FA9A16E6FA6D" author="BOULENGER, G. A." bookContentInfo="Taylor and Francis, London, U. K." pageId="58" pageNumber="133" refId="ref76038" refString="BOULENGER, G. A. 1885 a. Catalogue of the Lizards in the British Museum (Natural History), Volume 2. Taylor and Francis, London, U. K." title="Catalogue of the Lizards in the British Museum (Natural History), Volume 2" type="book" year="1885">
Boulenger, 1885
<emphasis id="FCA0EA95FF81FFADFDC6FA9B14B5FA8F" box="[634,647,1375,1398]" italics="true" pageId="58" pageNumber="133">a</emphasis>
: 132
</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="AA454B76FF81FFADFF4EFAB8141FFA6D" author="BURT, C. E. &amp; M. D. BURT" box="[242,557,1404,1428]" journalOrPublisher="Transactions of the Academy of Science of St. Louis" pageId="58" pageNumber="133" pagination="1 - 104" part="28" refId="ref76448" refString="BURT, C. E., AND M. D. BURT. 1933. A preliminary check list of the lizards of South America. Transactions of the Academy of Science of St. Louis 28 (1): 1 - 104." title="A preliminary check list of the lizards of South America" type="journal article" year="1933">Burt and Burt, 1933:43</bibRefCitation>
;
<bibRefCitation id="AA454B76FF81FFADFDF6FAB81731FA48" author="FRITTS, T. H." journalOrPublisher="San Diego Society of Natural History Memoir" pageId="58" pageNumber="133" pagination="1 - 89" part="7" refId="ref77735" refString="FRITTS, T. H. 1974. A multivariate and evolutionary analysis of the andean iguanid lizards of the genus Stenocercus. San Diego Society of Natural History Memoir 7: 1 - 89." title="A multivariate and evolutionary analysis of the andean iguanid lizards of the genus Stenocercus" type="journal article" year="1974">Fritts, 1974:57</bibRefCitation>
; Etheridge, in
<bibRefCitation id="AA454B76FF81FFADFE12FA5D175BFA37" author="PETERS, J. A. &amp; R. DONOSO-BARROS" journalOrPublisher="United States National Museum Bulletin" pageId="58" pageNumber="133" pagination="1 - 293" part="297" refId="ref79234" refString="PETERS, J. A., AND R. DONOSO-BARROS. 1970. Catalogue of the Neotropical Squamata. Part 2, lizards and amphisbaenians. United States National Museum Bulletin 297: 1 - 293." title="Catalogue of the Neotropical Squamata. Part 2, lizards and amphisbaenians" type="journal article" year="1970">Peters and Donoso-Barros, 1970:256</bibRefCitation>
;
<bibRefCitation id="AA454B76FF81FFADFECAFA731444FA37" author="TORRES, S. &amp; G. J. SCROCCHI &amp; M. B. HARVEY" box="[374,630,1462,1486]" journalOrPublisher="Journal of Herpetology" pageId="58" pageNumber="133" pagination="129 - 134" part="34" refId="ref80329" refString="TORRES, S., G. J. SCROCCHI, AND M. B. HARVEY. 2000. The South American tropidurid lizard Stenocercus marmoratus: redescription, distribution, and natural history. Journal of Herpetology 34: 129 - 134." title="The South American tropidurid lizard Stenocercus marmoratus: redescription, distribution, and natural history" type="journal article" year="2000">Torres et al., 2000:129</bibRefCitation>
.
</paragraph>
<paragraph id="CE6B3687FF81FFADFF33FA1117CCF987" blockId="58.[143,654,983,1722]" pageId="58" pageNumber="133">
<taxonomicName id="09D44D04FF81FFADFF33FA11140DFA14" authority="Werner, 1910: 23" authorityName="Werner" authorityPageNumber="23" authorityYear="1910" box="[143,575,1492,1517]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="58" pageNumber="133" phylum="Chordata" rank="species" species="difficilis">
<emphasis id="FCA0EA95FF81FFADFF33FA111747FA15" box="[143,373,1492,1516]" italics="true" pageId="58" pageNumber="133">Stenocercus difficilis</emphasis>
<bibRefCitation id="AA454B76FF81FFADFEC3FA11140DFA14" author="WERNER, F." box="[383,575,1493,1517]" pageId="58" pageNumber="133" refId="ref80539" refString="WERNER, F. 1910. Uber neue oder seltene Reptilien des Naturhistorischen Museums in Hamburg. Mitteilungen" title="Uber neue oder seltene Reptilien des Naturhistorischen Museums in Hamburg. Mitteilungen" type="book" year="1910">Werner, 1910:23</bibRefCitation>
</taxonomicName>
. Holotype: ZMH specimen destroyed in WWII, a male from Cochabamba [Departamento Cochabamba], Bolivia̕̕;
<bibRefCitation id="AA454B76FF81FFADFE6DF9E91736F99B" author="BURT, C. E. &amp; M. D. BURT" journalOrPublisher="Transactions of the Academy of Science of St. Louis" pageId="58" pageNumber="133" pagination="1 - 104" part="28" refId="ref76448" refString="BURT, C. E., AND M. D. BURT. 1933. A preliminary check list of the lizards of South America. Transactions of the Academy of Science of St. Louis 28 (1): 1 - 104." title="A preliminary check list of the lizards of South America" type="journal article" year="1933">Burt and Burt, 1933:43</bibRefCitation>
. Synonymy fide Etheridge in
<bibRefCitation id="AA454B76FF81FFADFDF4F98E17C8F987" author="PETERS, J. A. &amp; R. DONOSO-BARROS" journalOrPublisher="United States National Museum Bulletin" pageId="58" pageNumber="133" pagination="1 - 293" part="297" refId="ref79234" refString="PETERS, J. A., AND R. DONOSO-BARROS. 1970. Catalogue of the Neotropical Squamata. Part 2, lizards and amphisbaenians. United States National Museum Bulletin 297: 1 - 293." title="Catalogue of the Neotropical Squamata. Part 2, lizards and amphisbaenians" type="journal article" year="1970">Peters and Donoso-Barros, 1970:256</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFF81FFADFF16F942152BFDAF" pageId="58" pageNumber="133" type="diagnosis">
<paragraph id="CE6B3687FF81FFADFF16F942152BFDAF" blockId="58.[143,654,983,1722]" lastBlockId="58.[685,1196,190,1722]" pageId="58" pageNumber="133">
<emphasis id="FCA0EA95FF81FFADFF16F9421724F964" box="[170,278,1670,1693]" italics="true" pageId="58" pageNumber="133">Diagnosis</emphasis>
.—Among species of
<taxonomicName id="09D44D04FF81FFADFDB7F94114BCF965" box="[523,654,1669,1692]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="58" pageNumber="133" phylum="Chordata" rank="genus">
<emphasis id="FCA0EA95FF81FFADFDB7F94114BCF965" box="[523,654,1669,1692]" italics="true" pageId="58" pageNumber="133">Stenocercus</emphasis>
</taxonomicName>
with imbricate scales on the posterior surface of thighs,
<taxonomicName id="09D44D04FF81FFADFC98FF7A15F9FF2C" authorityName="Boulenger" authorityYear="1885" box="[804,971,190,213]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="58" pageNumber="133" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="FCA0EA95FF81FFADFC98FF7A15F9FF2C" box="[804,971,190,213]" italics="true" pageId="58" pageNumber="133">S. marmoratus</emphasis>
</taxonomicName>
and
<taxonomicName id="09D44D04FF81FFADFBB7FF7A1299FF2C" authorityName="Dumeril and Bibron" authorityYear="1837" box="[1035,1195,190,213]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="58" pageNumber="133" phylum="Chordata" rank="species" species="roseiventris">
<emphasis id="FCA0EA95FF81FFADFBB7FF7A1299FF2C" box="[1035,1195,190,213]" italics="true" pageId="58" pageNumber="133">S. roseiventris</emphasis>
</taxonomicName>
are unique in having caudal scales with strongly projected mucrons.
<taxonomicName id="09D44D04FF81FFADFBBDFF3D1299FEE8" authorityName="Boulenger" authorityYear="1885" box="[1025,1195,249,273]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="58" pageNumber="133" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="FCA0EA95FF81FFADFBBDFF3D1299FEE8" box="[1025,1195,249,273]" italics="true" pageId="58" pageNumber="133">S. marmoratus</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="09D44D04FF81FFADFC26FED31206FED7" authorityName="Dumeril and Bibron" authorityYear="1837" box="[922,1076,279,302]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="58" pageNumber="133" phylum="Chordata" rank="species" species="roseiventris">
<emphasis id="FCA0EA95FF81FFADFC26FED31206FED7" box="[922,1076,279,302]" italics="true" pageId="58" pageNumber="133">S. roseiventris</emphasis>
</taxonomicName>
(character states in parentheses) by having a low and discontinuous vertebral crest (conspicuous and continuous), granular temporals (keeled and imbricate), lateral body scales half the size of dorsal body scales (laterals and dorsals similar in size), a prominent ventrolateral fold (fold inconspicuous or absent), 4459 (
<emphasis id="FCA0EA95FF81FFADFBC4FE2212BAFE04" box="[1144,1160,486,509]" italics="true" pageId="58" pageNumber="133">X</emphasis>
= 52.00) scales around midbody (5582,
<emphasis id="FCA0EA95FF81FFADFBCBFDC012B5FDE2" box="[1143,1159,516,539]" italics="true" pageId="58" pageNumber="133">X</emphasis>
= 69.80), and by lacking a preauricular fringe (present).
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFF81FFABFD74FD991421FC81" lastPageId="60" lastPageNumber="135" pageId="58" pageNumber="133" type="description">
<paragraph id="CE6B3687FF81FFABFD74FD99147FFDE1" blockId="58.[685,1196,190,1722]" lastBlockId="60.[143,654,190,1271]" lastPageId="60" lastPageNumber="135" pageId="58" pageNumber="133">
<emphasis id="FCA0EA95FF81FFADFD74FD99157BFD8D" box="[712,841,605,628]" italics="true" pageId="58" pageNumber="133">Description</emphasis>
.—(1) Maximum SVL in males 83 mm (
<bibRefCitation id="AA454B76FF81FFADFCAEFDBE15C4FD6B" author="TORRES, S. &amp; G. J. SCROCCHI &amp; M. B. HARVEY" box="[786,1014,634,658]" journalOrPublisher="Journal of Herpetology" pageId="58" pageNumber="133" pagination="129 - 134" part="34" refId="ref80329" refString="TORRES, S., G. J. SCROCCHI, AND M. B. HARVEY. 2000. The South American tropidurid lizard Stenocercus marmoratus: redescription, distribution, and natural history. Journal of Herpetology 34: 129 - 134." title="The South American tropidurid lizard Stenocercus marmoratus: redescription, distribution, and natural history" type="journal article" year="2000">Torres et al., 2000</bibRefCitation>
); (2) maximum SVL in females 77 mm (
<emphasis id="FCA0EA95FF81FFADFC03FD5C15FFFD56" box="[959,973,664,687]" italics="true" pageId="58" pageNumber="133">n</emphasis>
= 7); (3) vertebrals 4466; (4) paravertebrals 5168; (5) scales around midbody 4459; (6) supraoculars 48; (7) internasals 24; (8) postrostrals five; (9) loreals 24; (10) gulars 2330; (11) subdigitals on Finger IV 1221; (12) subdigitals on Toe IV 1623; (13) posthumeral mite pocket present as one or more vertical folds or ridges; (14) postfemoral mite pocket distinct with slit-like opening; (15) parietal eye not visible through interparietal cornea; (16) scales on occipitoparietal region small, smooth, juxtaposed; (17) projecting angulate temporals absent; (18) row of enlarged supraoculars occupying most of supraocular region absent; (19) scales on frontonasal region juxtaposed anteriorly; (20) preauricular fringe inconspicuous or absent; (21) antegular, antehumeral, gular, longitudinal, oblique, postauricular, and neck folds present (but see
<bibRefCitation id="AA454B76FF81FFADFBB1FB0E14D4FB06" author="TORRES, S. &amp; G. J. SCROCCHI &amp; M. B. HARVEY" journalOrPublisher="Journal of Herpetology" pageId="58" pageNumber="133" pagination="129 - 134" part="34" refId="ref80329" refString="TORRES, S., G. J. SCROCCHI, AND M. B. HARVEY. 2000. The South American tropidurid lizard Stenocercus marmoratus: redescription, distribution, and natural history. Journal of Herpetology 34: 129 - 134." title="The South American tropidurid lizard Stenocercus marmoratus: redescription, distribution, and natural history" type="journal article" year="2000">Torres et al., 2000</bibRefCitation>
); (22) lateral nuchals less than half the size of dorsal nuchals; (23) posterior gulars cycloid, smooth, slightly imbricate, not notched; (24) lateral and dorsal body scales similar in size; (25) anteriormost and posteriormost vertebrals larger than adjacent paravertebrals; (26) dorsolateral crest absent; (27) ventrals smooth, imbricate; (28) scales on posterior surfaces of thighs keeled, imbricate; (29) inguinal granular pocket absent; (30) inguinal groove present; (31) preanals not projected; (32) tail not compressed laterally in adult males; (33) tail length 5660% of total length; (34) caudal whorls per autotomic segment two; (35) caudals strongly spinose; (36) dark brown stripe extending anterodorsally from subocular region to supraciliaries absent; (37) dark patch extensively covering gular region of females absent; (38) dark patch extensively covering gular region of adult males absent; (39) black patch on ventral surface of neck in adult males absent; (40) dark midventral longitudinal mark such as faint line, conspicuous stripe, or extensive patch in adult males absent; (41) dark patches on ventral surface of thighs in adult males absent; (42) postxiphisternal inscriptional ribs not in contact midventrally, Pattern 1B.
</paragraph>
<caption id="9AAB660FFF80FFACFF33F9721704F901" ID-DOI="http://doi.org/10.5281/zenodo.14384850" ID-Zenodo-Dep="14384850" httpUri="https://zenodo.org/record/14384850/files/figure.png" pageId="59" pageNumber="134" startId="59.[143,157,1718,1737]" targetBox="[122,1174,189,1701]" targetPageId="59" targetType="figure">
<paragraph id="CE6B3687FF80FFACFF33F9721704F901" blockId="59.[122,1175,1718,1784]" pageId="59" pageNumber="134">
<smallCapsWord id="C88DA05BFF80FFACFF33F972169CF931" baselines="1732,1732" box="[143,174,1718,1737]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Fig" pageId="59" pageNumber="134">FIG</smallCapsWord>
. 20.—Eight species of
<taxonomicName id="09D44D04FF80FFACFE36F97217C0F930" box="[394,498,1718,1737]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="59" pageNumber="134" phylum="Chordata" rank="genus">
<emphasis id="FCA0EA95FF80FFACFE36F97217C0F930" box="[394,498,1718,1737]" italics="true" pageId="59" pageNumber="134">Stenocercus</emphasis>
</taxonomicName>
. (A)
<taxonomicName id="09D44D04FF80FFACFD9AF97214C4F930" authority="(AMU)" baseAuthorityName="AMU" box="[550,758,1718,1737]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="59" pageNumber="134" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="FCA0EA95FF80FFACFD9AF9721499F931" box="[550,683,1718,1737]" italics="true" pageId="59" pageNumber="134">S. marmoratus</emphasis>
(AMU)
</taxonomicName>
; (B)
<taxonomicName id="09D44D04FF80FFACFC94F97215C5F930" authority="(PVE)" baseAuthorityName="PVE" box="[808,1015,1718,1737]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="59" pageNumber="134" phylum="Chordata" rank="species" species="melanopngus">
<emphasis id="FCA0EA95FF80FFACFC94F9721584F930" box="[808,950,1718,1737]" italics="true" pageId="59" pageNumber="134">S. melanopygus</emphasis>
(PVE)
</taxonomicName>
; (C)
<taxonomicName id="09D44D04FF80FFACFB97F972168EF918" authority="(MLU)" baseAuthorityName="MLU" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="59" pageNumber="134" phylum="Chordata" rank="species" species="modestus">
<emphasis id="FCA0EA95FF80FFACFB97F97212A4F930" box="[1067,1174,1718,1737]" italics="true" pageId="59" pageNumber="134">S. modestus</emphasis>
(MLU)
</taxonomicName>
; (D)
<taxonomicName id="09D44D04FF80FFACFF52F90A17EBF918" authority="(WED)" baseAuthorityName="WED" box="[238,473,1741,1761]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="59" pageNumber="134" phylum="Chordata" rank="species" species="nigromaculatus">
<emphasis id="FCA0EA95FF80FFACFF52F90A17BCF919" box="[238,398,1741,1761]" italics="true" pageId="59" pageNumber="134">S. nigromaculatus</emphasis>
(WED)
</taxonomicName>
; (E)
<taxonomicName id="09D44D04FF80FFACFDB5F90A149FF918" authority="(PVE)" baseAuthorityName="PVE" box="[521,685,1741,1761]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="59" pageNumber="134" phylum="Chordata" rank="species" species="nubicola">
<emphasis id="FCA0EA95FF80FFACFDB5F90A145EF919" box="[521,620,1741,1761]" italics="true" pageId="59" pageNumber="134">S. nubicola</emphasis>
(PVE)
</taxonomicName>
; (F)
<taxonomicName id="09D44D04FF80FFACFD67F90A154AF918" authority="(WED)" baseAuthorityName="WED" box="[731,888,1741,1761]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="59" pageNumber="134" phylum="Chordata" rank="species" species="ochoai">
<emphasis id="FCA0EA95FF80FFACFD67F90A151EF919" box="[731,812,1741,1761]" italics="true" pageId="59" pageNumber="134">S. ochoai</emphasis>
(WED)
</taxonomicName>
; (G)
<taxonomicName id="09D44D04FF80FFACFC15F90A1266F918" authority="(PVE)" baseAuthorityName="PVE" box="[937,1108,1741,1761]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="59" pageNumber="134" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="FCA0EA95FF80FFACFC15F90A1221F919" box="[937,1043,1741,1761]" italics="true" pageId="59" pageNumber="134">S. orientalis</emphasis>
(PVE)
</taxonomicName>
; (H)
<taxonomicName id="09D44D04FF80FFACFB39F90A1700F901" authority="(MLU)" baseAuthorityName="MLU" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="59" pageNumber="134" phylum="Chordata" rank="species" species="ornatissimus">
<emphasis id="FCA0EA95FF80FFACFB39F90A16DBF901" italics="true" pageId="59" pageNumber="134">S. ornatissimus</emphasis>
(MLU)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="CE6B3687FF87FFABFF16FDDA1421FC81" blockId="60.[143,654,190,1271]" pageId="60" pageNumber="135">
<emphasis id="FCA0EA95FF87FFABFF16FDDA1703FDCF" box="[170,305,542,566]" italics="true" pageId="60" pageNumber="135">Color in life</emphasis>
.—Dorsum grayish brown with dark brown cross bands and white or yellow blotches; white or yellow dorsolateral stripe, continuous or discontinuous, extending from subocular region to approximately midbody; flanks and sides of neck with white or yellow blotches; dark brown or black vertical bar anterior to fore limb in some specimens; throat charcoal with cream spots in some males; venter uniform cream, orange in some males (based on
<bibRefCitation id="AA454B76FF87FFABFED8FC871466FCA2" author="TORRES, S. &amp; G. J. SCROCCHI &amp; M. B. HARVEY" box="[356,596,835,859]" journalOrPublisher="Journal of Herpetology" pageId="60" pageNumber="135" pagination="129 - 134" part="34" refId="ref80329" refString="TORRES, S., G. J. SCROCCHI, AND M. B. HARVEY. 2000. The South American tropidurid lizard Stenocercus marmoratus: redescription, distribution, and natural history. Journal of Herpetology 34: 129 - 134." title="The South American tropidurid lizard Stenocercus marmoratus: redescription, distribution, and natural history" type="journal article" year="2000">Torres et al. [2000]</bibRefCitation>
and a photograph taken by A. Muñoz.)
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFF87FFABFF16FCBA16E7FBF5" pageId="60" pageNumber="135" type="biology_ecology">
<paragraph id="CE6B3687FF87FFABFF16FCBA16E7FBF5" blockId="60.[143,654,190,1271]" pageId="60" pageNumber="135">
<emphasis id="FCA0EA95FF87FFABFF16FCBA176CFC6F" box="[170,350,894,918]" italics="true" pageId="60" pageNumber="135">Natural History</emphasis>
.—Females of
<taxonomicName id="09D44D04FF87FFABFDB7FCBB168EFC4D" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="60" pageNumber="135" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="FCA0EA95FF87FFABFDB7FCBB168EFC4D" italics="true" pageId="60" pageNumber="135">S. marmoratus</emphasis>
</taxonomicName>
probably lay eggs twice a year. This species prefers rock outcrops and hides in crevices or underneath rocks (
<bibRefCitation id="AA454B76FF87FFABFE48FC1216FBFBF5" author="TORRES, S. &amp; G. J. SCROCCHI &amp; M. B. HARVEY" journalOrPublisher="Journal of Herpetology" pageId="60" pageNumber="135" pagination="129 - 134" part="34" refId="ref80329" refString="TORRES, S., G. J. SCROCCHI, AND M. B. HARVEY. 2000. The South American tropidurid lizard Stenocercus marmoratus: redescription, distribution, and natural history. Journal of Herpetology 34: 129 - 134." title="The South American tropidurid lizard Stenocercus marmoratus: redescription, distribution, and natural history" type="journal article" year="2000">Torres et al., 2000</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="86CE650CFF87FFABFF16FBD614BCFB0E" pageId="60" pageNumber="135" type="distribution">
<paragraph id="CE6B3687FF87FFABFF16FBD614BCFB0E" blockId="60.[143,654,190,1271]" pageId="60" pageNumber="135">
<emphasis id="FCA0EA95FF87FFABFF16FBD61703FBD3" box="[170,305,1042,1066]" italics="true" pageId="60" pageNumber="135">Distribution</emphasis>
.—
<taxonomicName id="09D44D04FF87FFABFEEFFBD6145EFBD3" box="[339,620,1042,1066]" class="Squamata" family="Tropiduridae" genus="Stenocercus" kingdom="Animalia" pageId="60" pageNumber="135" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="FCA0EA95FF87FFABFEEFFBD6145EFBD3" box="[339,620,1042,1066]" italics="true" pageId="60" pageNumber="135">Stenocercus marmoratus</emphasis>
</taxonomicName>
is known from the eastern Cordilleras of the central and southern Andes between 24
<emphasis id="FCA0EA95FF87FFABFDD4FB881440FB9D" bold="true" box="[616,626,1100,1124]" pageId="60" pageNumber="135">°</emphasis>
S 16
<emphasis id="FCA0EA95FF87FFABFF16FBAE1686FB7B" bold="true" box="[170,180,1130,1154]" pageId="60" pageNumber="135">°</emphasis>
S (
<figureCitation id="56EF2A02FF87FFABFF6BFBAE171BFB7B" box="[215,297,1130,1154]" captionStart="FIG" captionStartId="21.[143,157,997,1016]" captionTargetBox="[122,1176,189,979]" captionTargetId="figure-283@21.[122,1176,189,979]" captionTargetPageId="21" captionText="FIG. 13.—Distribution of seven species of Stenocercus in South America. Question mark indicates possible distribution of S. tricristatus." figureDoi="http://doi.org/10.5281/zenodo.14384832" httpUri="https://zenodo.org/record/14384832/files/figure.png" pageId="60" pageNumber="135">Fig. 13</figureCitation>
). This species occurs at elevations of 10003350 m in Provincia Salta, Argentina, as well as Departamentos Chuquisaca, Cochabamba, Santa Cruz, and Tarija in Bolivia (
<bibRefCitation id="AA454B76FF87FFABFF57FB1B179AFB0E" author="CRUZ, F. B. &amp; G. J. SCROCCHI &amp; S. A. MORO" box="[235,424,1247,1271]" journalOrPublisher="Herpetological Review" pageId="60" pageNumber="135" pagination="211" part="27" refId="ref77030" refString="CRUZ, F. B., G. J. SCROCCHI, AND S. A. MORO. 1996. Geographic distribution. Stenocercus marmoratus (spotted whorltail iguana). Herpetological Review 27: 211." title="Geographic distribution. Stenocercus marmoratus (spotted whorltail iguana)" type="journal volume" year="1996">Cruz et al., 1996</bibRefCitation>
;
<bibRefCitation id="AA454B76FF87FFABFE0FFB1B14B1FB0E" author="TORRES, S. &amp; G. J. SCROCCHI &amp; M. B. HARVEY" box="[435,643,1247,1271]" journalOrPublisher="Journal of Herpetology" pageId="60" pageNumber="135" pagination="129 - 134" part="34" refId="ref80329" refString="TORRES, S., G. J. SCROCCHI, AND M. B. HARVEY. 2000. The South American tropidurid lizard Stenocercus marmoratus: redescription, distribution, and natural history. Journal of Herpetology 34: 129 - 134." title="The South American tropidurid lizard Stenocercus marmoratus: redescription, distribution, and natural history" type="journal article" year="2000">Torres et al., 2000</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>