153 lines
19 KiB
XML
153 lines
19 KiB
XML
<document id="A47BC7D0643A2375CD342ECB092F4650" ID-DOI="10.5281/zenodo.212283" ID-GBIF-Dataset="87c1b7d7-8ba9-48b9-8069-226863c4818d" ID-ISSN="1175-5326" ID-Zenodo-Dep="212283" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460524491036" checkinUser="plazi" docAuthor="Khramov, Alexander V." docDate="2012" docId="46624403FFF0C050A79F5999FE0AFB64" docLanguage="en" docName="zt03478p308.pdf" docOrigin="Zootaxa 3478" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Grammolingiidae" docType="treatment" docVersion="8" lastPageNumber="305" masterDocId="BA5B3C7BFFF8C058A708590EFFF4FFA6" masterDocTitle="The new fossil lacewings of Grammolingiidae (Neuroptera) from the Jurassic of Central Asia and Mongolia, with notes on biogeography of the family" masterLastPageNumber="308" masterPageNumber="297" pageNumber="305" updateTime="1698314878310" updateUser="plazi">
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<mods:titleInfo id="0F99E1A7E08166D0A1D3FBAF557C870A">
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<mods:title id="0BAF8E9EF49C690BC2B200A2D22A031E">The new fossil lacewings of Grammolingiidae (Neuroptera) from the Jurassic of Central Asia and Mongolia, with notes on biogeography of the family</mods:title>
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</mods:titleInfo>
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<mods:name id="9D62403FE8476E7453FD7A0A5B4A1041" type="personal">
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<mods:roleTerm id="6A2B8C9B39BC34DBEE62A4DD28615563">Author</mods:roleTerm>
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<mods:namePart id="E04D73D85CC2038948CD0119AAAFECA0">Khramov, Alexander V.</mods:namePart>
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<mods:typeOfResource id="57CF65DF379310E118CFF7D554FCDF9E">text</mods:typeOfResource>
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<mods:title id="3F438283197CCC0801F906ECA451C0CD">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="3EBC228288F07B5AB505C3398A6C42D4">
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<mods:date id="F71F9285597DB7FCF6DE9B3F8290EC68">2012</mods:date>
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<mods:detail id="EA944D3774DE56A7484319DD3D9C94AD" type="volume">
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<mods:number id="8092E3F8905999FACC29C673D75119E3">3478</mods:number>
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<mods:extent id="4E9E3BC817DCC286E6A724D2B679A05B" unit="page">
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<mods:start id="AD924B118748D794AD2CBC050211BD69">297</mods:start>
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<mods:end id="0C3C4142FF4A5AB08C7287D5A568008D">308</mods:end>
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<mods:classification id="F1758933667EFA6DDDE90548548A9353">journal article</mods:classification>
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<mods:identifier id="090167AF12ABC2F9E39F1E81218B76B3" type="DOI">10.5281/zenodo.212283</mods:identifier>
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<mods:identifier id="E7E50FBCC95BBD66C6C896B6FA5E3AF1" type="GBIF-Dataset">87c1b7d7-8ba9-48b9-8069-226863c4818d</mods:identifier>
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<mods:identifier id="6DF2E665E22E68CEC2803B39D66AED39" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="CBB65820788DFAAC995F0AB471FF5C80" type="Zenodo-Dep">212283</mods:identifier>
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<treatment id="46624403FFF0C050A79F5999FE0AFB64" ID-DOI="http://doi.org/10.5281/zenodo.6180100" ID-GBIF-Taxon="119526078" ID-Zenodo-Dep="6180100" LSID="urn:lsid:plazi:treatment:46624403FFF0C050A79F5999FE0AFB64" httpUri="http://treatment.plazi.org/id/46624403FFF0C050A79F5999FE0AFB64" lastPageNumber="305" pageId="8" pageNumber="305">
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<subSubSection id="86D1A69EFFF0C050A79F5999FCFCFF17" box="[151,776,151,177]" pageId="8" pageNumber="305" type="nomenclature">
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<paragraph id="CE74F515FFF0C050A79F5999FCFCFF17" blockId="8.[151,776,151,177]" box="[151,776,151,177]" pageId="8" pageNumber="305">
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<heading id="953C4279FFF0C050A79F5999FCFCFF17" bold="true" box="[151,776,151,177]" fontSize="11" level="1" pageId="8" pageNumber="305" reason="1">
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<emphasis id="FCBF2907FFF0C050A79F5999FCFCFF17" bold="true" box="[151,776,151,177]" pageId="8" pageNumber="305">
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<taxonomicName id="09CB8E96FFF0C050A79F5999FE80FF17" ID-CoL="623NB" box="[151,372,151,177]" class="Insecta" family="Grammolingiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="family">Grammolingiidae</taxonomicName>
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gen. et sp. indet. from Sai-Sagul
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="86D1A69EFFF0C050A79F59D0FEE7FD9C" pageId="8" pageNumber="305" type="description">
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<paragraph id="CE74F515FFF0C050A79F59D0FAC0FEBC" blockId="8.[151,1436,222,1218]" pageId="8" pageNumber="305">
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<emphasis id="FCBF2907FFF0C050A79F59D0FEDDFF51" bold="true" box="[151,297,222,247]" pageId="8" pageNumber="305">Description.</emphasis>
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<emphasis id="FCBF2907FFF0C050A63C59EEFE95FF51" box="[308,353,224,247]" italics="true" pageId="8" pageNumber="305">PIN</emphasis>
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№
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<emphasis id="FCBF2907FFF0C050A69B59D0FE08FF50" box="[403,508,222,246]" italics="true" pageId="8" pageNumber="305">2389/523</emphasis>
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. Distal part of hindwing (
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<figureCitation id="56F0E990FFF0C050A43D59D0FC89FF50" box="[821,893,222,246]" captionStart="FIGURE 7" captionStartId="7.[151,250,1892,1915]" captionTargetBox="[231,1368,213,1859]" captionTargetId="figure@7.[166,1420,199,1918]" captionTargetPageId="7" captionText="FIGURE 7. Grammolingiidae incertae sedis, Sai Sagul locality, Kyrgyzstan, upper Lower Jurassic – lower Middle Jurassic. The fragments of the hindwings. Photographs: A, PIN № 3073 / 838; B, PIN № 3073 / 443; C, PIN № 3073 / 839; D, PIN № 2389 / 523. Drawings: E, PIN № 3073 / 838; G, PIN № 3073 / 839; F, PIN № 3073 / 443; H, PIN № 2389 / 523. Scale bar represents 5 mm." httpUri="https://zenodo.org/record/212290/files/figure.png" pageId="8" pageNumber="305">Fig. 7</figureCitation>
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D,H). Length of fragment
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<quantity id="093358F0FFF0C050A3BE59D0FAD5FF50" box="[1206,1313,222,246]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="8" pageNumber="305" unit="mm" value="19.5">19,5 mm</quantity>
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, estimated complete length of wing about
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<quantity id="093358F0FFF0C050A6FC580CFD84FEBC" box="[500,624,258,282]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="8" pageNumber="305" unit="mm" value="47.5" valueMax="50.0" valueMin="45.0">45–50 mm</quantity>
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. Trichosors present. Sc and R1 run parallel to the wing margin.
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</paragraph>
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<paragraph id="CE74F515FFF0C050A7CF5826FE28FEC4" blockId="8.[151,1436,222,1218]" pageId="8" pageNumber="305">
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<emphasis id="FCBF2907FFF0C050A7CF5826FF00FE99" box="[199,244,296,319]" italics="true" pageId="8" pageNumber="305">PIN</emphasis>
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№
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<emphasis id="FCBF2907FFF0C050A62A5828FE78FE98" box="[290,396,294,318]" italics="true" pageId="8" pageNumber="305">3073/838</emphasis>
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. Proximal part of hindwing (
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<figureCitation id="56F0E990FFF0C050A5D75828FCD1FE98" box="[735,805,294,318]" captionStart="FIGURE 7" captionStartId="7.[151,250,1892,1915]" captionTargetBox="[231,1368,213,1859]" captionTargetId="figure@7.[166,1420,199,1918]" captionTargetPageId="7" captionText="FIGURE 7. Grammolingiidae incertae sedis, Sai Sagul locality, Kyrgyzstan, upper Lower Jurassic – lower Middle Jurassic. The fragments of the hindwings. Photographs: A, PIN № 3073 / 838; B, PIN № 3073 / 443; C, PIN № 3073 / 839; D, PIN № 2389 / 523. Drawings: E, PIN № 3073 / 838; G, PIN № 3073 / 839; F, PIN № 3073 / 443; H, PIN № 2389 / 523. Scale bar represents 5 mm." httpUri="https://zenodo.org/record/212290/files/figure.png" pageId="8" pageNumber="305">Fig. 7</figureCitation>
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A,E). Length of fragment
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<quantity id="093358F0FFF0C050A3565828FB46FE98" box="[1118,1202,294,318]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="305" unit="mm" value="27.0">27 mm</quantity>
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, estimated complete length of wing about
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<quantity id="093358F0FFF0C050A68E5844FE2CFEC4" box="[390,472,330,354]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="305" unit="mm" value="45.0">45 mm</quantity>
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.
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</paragraph>
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<paragraph id="CE74F515FFF0C050A7CF587EFDCFFE68" blockId="8.[151,1436,222,1218]" pageId="8" pageNumber="305">
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<emphasis id="FCBF2907FFF0C050A7CF587EFF00FE21" box="[199,244,368,391]" italics="true" pageId="8" pageNumber="305">PIN</emphasis>
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№
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<emphasis id="FCBF2907FFF0C050A62C5860FE79FE20" box="[292,397,366,390]" italics="true" pageId="8" pageNumber="305">3073/443</emphasis>
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. Proximal part of hindwing (
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<figureCitation id="56F0E990FFF0C050A5ED5860FCD8FE20" box="[741,812,366,390]" captionStart="FIGURE 7" captionStartId="7.[151,250,1892,1915]" captionTargetBox="[231,1368,213,1859]" captionTargetId="figure@7.[166,1420,199,1918]" captionTargetPageId="7" captionText="FIGURE 7. Grammolingiidae incertae sedis, Sai Sagul locality, Kyrgyzstan, upper Lower Jurassic – lower Middle Jurassic. The fragments of the hindwings. Photographs: A, PIN № 3073 / 838; B, PIN № 3073 / 443; C, PIN № 3073 / 839; D, PIN № 2389 / 523. Drawings: E, PIN № 3073 / 838; G, PIN № 3073 / 839; F, PIN № 3073 / 443; H, PIN № 2389 / 523. Scale bar represents 5 mm." httpUri="https://zenodo.org/record/212290/files/figure.png" pageId="8" pageNumber="305">Fig. 7</figureCitation>
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B,F). Length of fragment
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<quantity id="093358F0FFF0C050A3535860FB44FE20" box="[1115,1200,366,390]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="305" unit="mm" value="28.0">28 mm</quantity>
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, estimated complete length of wing about
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<quantity id="093358F0FFF0C050A69F589CFE19FE0C" box="[407,493,402,426]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="305" unit="mm" value="45.0">45 mm</quantity>
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. CuA dichotomously branched. CuP pectinately branched, its branches long and connected by 4–5 rows of crossveins.
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</paragraph>
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<paragraph id="CE74F515FFF0C050A7CF58D2FEE7FD9C" blockId="8.[151,1436,222,1218]" pageId="8" pageNumber="305">
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<emphasis id="FCBF2907FFF0C050A7CF58D2FF00FE55" box="[199,244,476,499]" italics="true" pageId="8" pageNumber="305">PIN</emphasis>
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№
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<emphasis id="FCBF2907FFF0C050A62A58D5FE7FFE55" box="[290,395,475,499]" italics="true" pageId="8" pageNumber="305">3073/839</emphasis>
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. Proximal part of hindwing (
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<figureCitation id="56F0E990FFF0C050A5D758D4FCD1FE54" box="[735,805,474,498]" captionStart="FIGURE 7" captionStartId="7.[151,250,1892,1915]" captionTargetBox="[231,1368,213,1859]" captionTargetId="figure@7.[166,1420,199,1918]" captionTargetPageId="7" captionText="FIGURE 7. Grammolingiidae incertae sedis, Sai Sagul locality, Kyrgyzstan, upper Lower Jurassic – lower Middle Jurassic. The fragments of the hindwings. Photographs: A, PIN № 3073 / 838; B, PIN № 3073 / 443; C, PIN № 3073 / 839; D, PIN № 2389 / 523. Drawings: E, PIN № 3073 / 838; G, PIN № 3073 / 839; F, PIN № 3073 / 443; H, PIN № 2389 / 523. Scale bar represents 5 mm." httpUri="https://zenodo.org/record/212290/files/figure.png" pageId="8" pageNumber="305">Fig. 7</figureCitation>
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C,G). Length of fragment
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<quantity id="093358F0FFF0C050A35658D4FB46FE54" box="[1118,1202,474,498]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="305" unit="mm" value="25.0">25 mm</quantity>
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, estimated complete length of wing about
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<quantity id="093358F0FFF0C050A69158F0FDEEFDB0" box="[409,538,510,534]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="8" pageNumber="305" unit="mm" value="37.5" valueMax="40.0" valueMin="35.0">35–40 mm</quantity>
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. CuP pectinately branched, its branches short and connected by one row of crossveins.
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</paragraph>
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</subSubSection>
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<subSubSection id="86D1A69EFFF0C050A7CF5B48FC06FD00" pageId="8" pageNumber="305" type="materials_examined">
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<paragraph id="CE74F515FFF0C050A7CF5B48FC06FD00" blockId="8.[151,1436,222,1218]" pageId="8" pageNumber="305">
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<emphasis id="FCBF2907FFF0C050A7CF5B48FE5DFDF9" bold="true" box="[199,425,582,607]" pageId="8" pageNumber="305">Material examined</emphasis>
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. Specimens
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<collectionCode id="A8DA6DD0FFF0C050A53F5B49FD90FDF8" box="[567,612,583,606]" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="8" pageNumber="305">PIN</collectionCode>
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№ 2389/523, a hindwing apex,
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<collectionCode id="A8DA6DD0FFF0C050A4DB5B49FBF4FDF8" box="[979,1024,583,606]" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="8" pageNumber="305">PIN</collectionCode>
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№ 3073/838,
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<collectionCode id="A8DA6DD0FFF0C050A3A95B49FB3AFDF8" box="[1185,1230,583,606]" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="8" pageNumber="305">PIN</collectionCode>
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№ 3073/443,
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<collectionCode id="A8DA6DD0FFF0C050A2675B49FA68FDF8" box="[1391,1436,583,606]" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="8" pageNumber="305">PIN</collectionCode>
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№ 3073/839, proximal fragments of hindwings.
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<collectingCountry id="B6DCB585FFF0C050A5B65B65FCB7FD25" box="[702,835,619,643]" name="Kyrgyzstan" pageId="8" pageNumber="305">Kyrgyzstan</collectingCountry>
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: Batken Province, Batken district on the border with
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<collectingCountry id="B6DCB585FFF0C050A79F5B80FEFDFD00" box="[151,265,654,678]" name="Tajikistan" pageId="8" pageNumber="305">Tajikistan</collectingCountry>
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; Sai-Sagul locality, upper Lower Jurassic – lower Middle Jurassic.
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</paragraph>
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</subSubSection>
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<subSubSection id="86D1A69EFFF0C050A7CF5BBCFE0AFB64" pageId="8" pageNumber="305" type="discussion">
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<paragraph id="CE74F515FFF0C050A7CF5BBCFCB9FB94" blockId="8.[151,1436,222,1218]" pageId="8" pageNumber="305">
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<emphasis id="FCBF2907FFF0C050A7CF5BBCFEBBFD6D" bold="true" box="[199,335,690,715]" pageId="8" pageNumber="305">Comments.</emphasis>
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Unfortunately, the generic affinity of these specimens is unclear because of their incompleteness, and it is impossible to associate hind wings with forewings from the same locality (some of them probably belong to
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<taxonomicName id="09CB8E96FFF0C050A7BC5BF5FE8DFCB5" box="[180,377,763,787]" class="Insecta" family="Grammolingiidae" genus="Litholingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="species" species="longa">
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<emphasis id="FCBF2907FFF0C050A7BC5BF5FE8DFCB5" box="[180,377,763,787]" italics="true" pageId="8" pageNumber="305">Litholingia longa</emphasis>
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</taxonomicName>
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, which is described only by forewings). In spite of this, these fragments are important for the understanding of the grammolingiid fauna of Sai-Sagul. The apex of the hindwing PIN № 2389/523 demonstrates the venation which is typical for
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<taxonomicName id="09CB8E96FFF0C050A5345A4CFCF6FCFC" box="[572,770,834,858]" class="Insecta" family="Grammolingiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="family">Grammolingiidae</taxonomicName>
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and could be assigned to
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<taxonomicName id="09CB8E96FFF0C050A3505A4DFB09FCFD" box="[1112,1277,835,859]" class="Insecta" family="Grammolingiidae" genus="Grammolingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="genus">
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<emphasis id="FCBF2907FFF0C050A3505A4DFB09FCFD" box="[1112,1277,835,859]" italics="true" pageId="8" pageNumber="305">Grammolingia</emphasis>
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</taxonomicName>
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,
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<emphasis id="FCBF2907FFF0C050A21C5A4DFE35FCD8" italics="true" pageId="8" pageNumber="305">
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<taxonomicName id="09CB8E96FFF0C050A21C5A4DFA6CFCFD" box="[1300,1432,835,859]" class="Insecta" family="Grammolingiidae" genus="Leptolingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="genus">Leptolingia</taxonomicName>
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,
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<taxonomicName id="09CB8E96FFF0C050A79F5A68FEE7FCD8" box="[151,275,870,894]" class="Insecta" family="Grammolingiidae" genus="Litholingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="genus">Litholingia</taxonomicName>
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or Chorilineia
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</emphasis>
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. This specimen shows, that in Sai-Sagul were represented not only
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<taxonomicName id="09CB8E96FFF0C050A3DE5A68FA6FFCD8" box="[1238,1435,870,894]" class="Insecta" family="Grammolingiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="family">Grammolingiidae</taxonomicName>
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with fused Sc and R1 (as
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<taxonomicName id="09CB8E96FFF0C050A6B55A85FDC9FC05" box="[445,573,907,931]" class="Insecta" family="Grammolingiidae" genus="Protolingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="300" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="FCBF2907FFF0C050A6B55A85FDC9FC05" box="[445,573,907,931]" italics="true" pageId="8" pageNumber="305">Protolingia</emphasis>
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</taxonomicName>
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), but also
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<taxonomicName id="09CB8E96FFF0C050A5BF5A84FC88FC04" box="[695,892,906,930]" class="Insecta" family="Grammolingiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="family">Grammolingiidae</taxonomicName>
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with typical configuration of Sc and R1, which are separate distally. Fragments PIN № 3073/838 and PIN № 3073/443 have similar venation and size and most likely belong to the same species. The remarkable peculiarity of the specimen PIN № 3073/443 (and apparently also of PIN № 3073/838) is the pectinate CuP, that is not typical for
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<taxonomicName id="09CB8E96FFF0C050A4AD5AF8FB9FFBA8" box="[933,1131,1014,1038]" class="Insecta" family="Grammolingiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="family">Grammolingiidae</taxonomicName>
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from Daohugou and Shar- Teg, whose hindwings have the dichotomously branched CuP.
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</paragraph>
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<paragraph id="CE74F515FFF0C050A7CF5D30FE0AFB64" blockId="8.[151,1436,222,1218]" pageId="8" pageNumber="305">
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Specimen PIN № 3073/839 probably belongs to other taxon of
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<taxonomicName id="09CB8E96FFF0C050A4835D30FBA5FBF0" box="[907,1105,1086,1110]" class="Insecta" family="Grammolingiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="305" phylum="Arthropoda" rank="family">Grammolingiidae</taxonomicName>
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than previous. It differs from two others by smaller size and by the structure of cubital and anal areas. Branches of CuP are shorter and are connected by only one row of crossveins; A1 is not so long and not so adjacent to the wing margin, as in PIN № 3073/838 and PIN № 3073/443.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |