treatments-xml/data/46/12/CF/4612CF45FF9EFFCEB7CB4613FDCAFB56.xml

<document id="7431517BA8C1051725E50F04B06B7533" ID-CLB-Dataset="5319" ID-DOI="10.11646/zootaxa.3826.3.8" ID-GBIF-Dataset="10a760f3-303c-4196-a3ff-ce29c7654408" ID-ISSN="1175-5326" ID-Zenodo-Dep="4920416" ID-ZooBank="DCEAAE0A-3500-41E2-B3B7-63C4F8D88D69" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623275268148" checkinUser="felipe" docAuthor="P. - H &amp; Ng, Peter K. L." docDate="2014" docId="4612CF45FF9EFFCEB7CB4613FDCAFB56" docLanguage="en" docName="zt03826p590.pdf" docOrigin="Zootaxa 3826 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neoliomera fragraea P.-H &amp; Ng 2014, n. sp." docType="treatment" docVersion="9" lastPageNumber="586" masterDocId="BA2BB73DFF9FFFC9B75C4406FFC0FFA8" masterDocTitle="A new species of Neoliomera Odhner, 1925, from the western Pacific, and the first record of N. demani Forest &amp; Guinot, 1961, from Taiwan (Crustacea: Decapoda: Brachyura: Xanthidae)" masterLastPageNumber="590" masterPageNumber="579" pageNumber="580" updateTime="1699008772129" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2E5F6D735924B6327757CE841A25E4B0">A new species of Neoliomera Odhner, 1925, from the western Pacific, and the first record of N. demani Forest &amp; Guinot, 1961, from Taiwan (Crustacea: Decapoda: Brachyura: Xanthidae)</mods:title>
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<mods:namePart id="08FFE5AF6D65494D572FA9A84C832288">P. - H</mods:namePart>
<mods:affiliation id="6A415F182490750B63D4E2F7477DCDCF">Department of Environment Biology and Fishery Science, National Taiwan Ocean University, 2 Pei-Ning Road, Keelung, Taiwan 202, R. O. C. E-mail: pingho @ mail. ntou. edu. tw</mods:affiliation>
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<mods:namePart id="C28572EF03B54F907297F9C0661AE2B5">Ng, Peter K. L.</mods:namePart>
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<emphasis id="FCCFA241FF9EFFC8B7CB4613FE54FD87" bold="true" box="[151,404,533,559]" italics="true" pageId="1" pageNumber="580">Neoliomera fragraea</emphasis>
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(
<figureCitation id="568062D6FF9EFFC8B7C34631FEE2FDF8" box="[159,290,567,592]" captionStart="FIGURE 1" captionStartId="3.[151,250,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure-95@3.[151,1436,193,1884]" captionTargetPageId="3" captionText="FIGURE 1. Colours in life. A–C, Neoliomera fragraea n. sp., female (25.3 × 14.3 mm) (NTOU), Taiwan; D, Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam [photograph by G. Paulay]; E, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU) [photograph by G. Paulay]; F, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan [photograph by Wang Kuei-Ching]." figureDoi="http://doi.org/10.5281/zenodo.4920418" httpUri="https://zenodo.org/record/4920418/files/figure.png" pageId="1" pageNumber="580">Figs. 1A–E</figureCitation>
, 
<figureCitation id="568062D6FF9EFFC8B6724631FEFEFDF8" box="[302,318,567,592]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="1" pageNumber="580">2</figureCitation>
, 
<figureCitation id="568062D6FF9EFFC8B6164631FE4FFDE7" box="[330,399,567,592]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-79@5.[241,1345,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. A, Neoliomera fragraea n. sp., paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="1" pageNumber="580">3A, B</figureCitation>
, 
<figureCitation id="568062D6FF9EFFC8B6C74631FE3FFDE7" box="[411,511,567,592]" captionStart="FIGURE 4" captionStartId="6.[151,250,1267,1289]" captionTargetBox="[164,1428,193,1248]" captionTargetId="figure-419@6.[164,1429,193,1248]" captionTargetPageId="6" captionText="FIGURE 4. A, B, E, Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam; C, D, F, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, outer views of right chelae; B, D, inner views of right chelae; E, F, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4920424" httpUri="https://zenodo.org/record/4920424/files/figure.png" pageId="1" pageNumber="580">4A, B, E</figureCitation>
, 
<figureCitation id="568062D6FF9EFFC8B5574631FDDAFDF8" box="[523,538,567,592]" captionStart="FIGURE 5" captionStartId="8.[151,250,1727,1749]" captionTargetBox="[151,1436,193,1706]" captionTargetId="figure-48@8.[151,1436,193,1706]" captionTargetPageId="8" captionText="FIGURE 5. Neoliomera fragraea n. sp., female (23.9 × 15.0 mm) (ZRC 2000.0483), Hawai‘i. A, dorsal view; B, thoracic sternum showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.4920426" httpUri="https://zenodo.org/record/4920426/files/figure.png" pageId="1" pageNumber="580">5</figureCitation>
, 
<figureCitation id="568062D6FF9EFFC8B57A463EFDA7FDF8" box="[550,615,567,592]" captionStart="FIGURE 7" captionStartId="10.[151,250,1223,1245]" captionTargetBox="[176,1403,401,1181]" captionTargetId="figure-371@10.[151,1436,393,1200]" captionTargetPageId="10" captionText="FIGURE 7. A, C, Neoliomera pubescens (H. Milne Edwards, 1873), lectotype male (30.5 × 17.5 mm) (MNHN), Mauritius; D, Neoliomera demani Forest &amp; Guinot, 1961, paratype male (23.0 × 13.0 mm) (MNHN), Tahiti; B, E, Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, B, left G1; C–E, distal parts of left G1s. A, C, after Forest &amp; Guinot (1961: fig. 75); D, after Forest &amp; Guinot (1961: fig. 76)." figureDoi="http://doi.org/10.5281/zenodo.4920430" httpUri="https://zenodo.org/record/4920430/files/figure.png" pageId="1" pageNumber="580">7B, E</figureCitation>
)
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<taxonomicName id="09BB05D0FF9EFFC8B7CB4686FE44FD3D" ID-CoL="46L2D" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" box="[151,388,640,661]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="580" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="FCCFA241FF9EFFC8B7CB4686FE44FD3D" box="[151,388,640,661]" italics="true" pageId="1" pageNumber="580">Neoliomera pubescens</emphasis>
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— 
<treatmentCitation id="4F1A5842FF9EFFC8B6C14686FDACFD3E" author="Rathbun, M. J." box="[413,620,640,662]" page="844" pageId="1" pageNumber="580" year="1906">
<bibRefCitation id="AA2A03A2FF9EFFC8B6C14686FDACFD3E" author="Rathbun, M. J." box="[413,620,640,662]" pageId="1" pageNumber="580" pagination="827 - 930" refId="ref6950" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23 (3), 827 - 930, pls. 1 - 24. [1903]" type="journal article" year="1906">Rathbun 1906: 844</bibRefCitation>
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(Molokai, Hawaiian Islands); 
<treatmentCitation id="4F1A5842FF9EFFC8B4E14686FB72FD3E" author="Edmondson, C. H." box="[957,1202,640,662]" page="292" pageId="1" pageNumber="580" year="1946">
<bibRefCitation id="AA2A03A2FF9EFFC8B4E14686FB72FD3E" author="Edmondson, C. H." box="[957,1202,640,662]" pageId="1" pageNumber="580" pagination="1 - 381" refId="ref5765" refString="Edmondson, C. H. (1946) Reef and Shore Fauna of Hawaii. Special Publication of the Bernice P. Bishop Museum, 22, i-iii, 1 - 381, figs. 1 - 223." type="journal article" year="1946">Edmondson 1946: 292</bibRefCitation>
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, fig. 178b (Hawaiian Islands); 
<treatmentCitation id="4F1A5842FF9EFFC8B6784698FDCAFD1C" author="Edmondson, C. H." box="[292,522,670,692]" page="252" pageId="1" pageNumber="580" year="1962">
<bibRefCitation id="AA2A03A2FF9EFFC8B6784698FDCAFD1C" author="Edmondson, C. H." box="[292,522,670,692]" pageId="1" pageNumber="580" pagination="215 - 309" refId="ref5805" refString="Edmondson, C. H. (1962) Xanthidae of Hawaii. Occasional Papers of the Bernice P. Bishop Museum, 22 (13), 215 - 309, figs. 1 - 34." type="journal article" year="1962">Edmondson 1962: 252</bibRefCitation>
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, fig. 9d (Hawaiian Islands); 
<treatmentCitation id="4F1A5842FF9EFFC8B47B4698FC0DFD1C" author="Sakai, T." box="[807,973,670,692]" page="399" pageId="1" pageNumber="580" year="1976">
<bibRefCitation id="AA2A03A2FF9EFFC8B47B4698FC0DFD1C" author="Sakai, T." box="[807,973,670,692]" pageId="1" pageNumber="580" pagination="1 - 251" refId="ref7044" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Vol. 1. - 3. Kodansha, Tokyo, English text: xxxix + 773 pp., Japanese text: 461 pp., plates: 16 pp., pls. 1 - 251. (plates)" type="book chapter" year="1976">Sakai 1976: 399</bibRefCitation>
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, part (O‘ahu, Hawaiian Islands); 
<treatmentCitation id="4F1A5842FF9EFFC8B2474698FF20FD7B" author="Castro, P." page="99" pageId="1" pageNumber="580" year="2011">
<bibRefCitation id="AA2A03A2FF9EFFC8B2474698FF20FD7B" author="Castro, P." pageId="1" pageNumber="580" pagination="1 - 154" refId="ref5729" refString="Castro, P. (2011) Catalog of the anomuran and brachyuran crabs (Crustacea: Decapoda: Anomura, Brachyura) of the Hawaiian Islands. Zootaxa, 2947, 1 - 154." type="journal article" year="2011">Castro 2011: 99</bibRefCitation>
</treatmentCitation>
(list). (not 
<taxonomicName id="09BB05D0FF9EFFC8B60E46B8FCDAFD7B" authority="H. Milne Edwards, 1834" authorityName="H. Milne Edwards" authorityYear="1834" box="[338,794,701,723]" class="Malacostraca" family="Xanthidae" genus="Zozymus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="580" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="FCCFA241FF9EFFC8B60E46B8FDD9FD7A" box="[338,537,701,723]" italics="true" pageId="1" pageNumber="580">Zozymus pubescens</emphasis>
H. 
<bibRefCitation id="AA2A03A2FF9EFFC8B56246BBFCDAFD7B" author="Milne Edwards, H." box="[574,794,701,723]" pageId="1" pageNumber="580" pagination="1 - 638" refId="ref6436" refString="Milne Edwards, H. (1834 - 1837) Histoire Naturelle des Crustaces Comprenant l'Anatomie, la Physiologie et la Classification de ces Animaux. Vol. 1. - 3. Librairie Encyclopedique de Roret, Paris, xxxv + 468 pp., 531 pp. + 1837: Atlas (1 - 32) + pls. 1 - 42., 1840: 1 - 638." type="book chapter" year="1834">Milne Edwards, 1834</bibRefCitation>
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)
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<emphasis id="FCCFA241FF9EFFC8B7CB46F9FEBAFCB0" bold="true" box="[151,378,767,792]" pageId="1" pageNumber="580">Material examined</emphasis>
.
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<typeStatus id="1100C0F1FF9EFFC8B6D546F9FE31FCB0" box="[393,497,767,792]" pageId="1" pageNumber="580" type="holotype">Holotype</typeStatus>
male (18.3 × 
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reef front, 
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water, coll. 
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. 
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<specimenCount id="D8BDB5DAFF9EFFC8B5AC4722FCFCFC95" box="[752,828,804,829]" pageId="1" pageNumber="580" type="male">1 male</specimenCount>
(16.4 × 
<quantity id="0943D3B6FF9EFFC8B4C34722FC38FC95" box="[927,1016,804,829]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="1" pageNumber="580" unit="mm" value="9.6">9.6 mm</quantity>
) (
<specimenCode id="9E1DD628FF9EFFC8B3534722FB0CFC95" box="[1039,1228,804,829]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="580">ZRC 2014.0207</specimenCode>
)
</materialsCitation>
, 
<materialsCitation id="7ED3740EFF9EFFC8B3834722FDC7FCC8" ID-GBIF-Occurrence="3326196305" collectionCode="NTOU" country="Guam" pageId="1" pageNumber="580" specimenCode="NTOU 2014.031401" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="D8BDB5DAFF9EFFC8B3834722FA81FC95" box="[1247,1345,804,829]" pageId="1" pageNumber="580" type="female">1 female</specimenCount>
(16.8 × 
<quantity id="0943D3B6FF9EFFC8B7CB4741FF32FCC8" box="[151,242,839,864]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="1" pageNumber="580" unit="mm" value="9.6">9.6 mm</quantity>
) (
<specimenCode id="9E1DD628FF9EFFC8B6514741FE3EFCC8" box="[269,510,839,864]" collectionCode="NTOU" country="Taiwan" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="1" pageNumber="580">NTOU 2014.031401</specimenCode>
)
</materialsCitation>
, 
<materialsCitation id="7ED3740EFF9EFFC8B54A4741FF3FFC2D" ID-GBIF-Occurrence="3326196303" collectingDate="2000-06-01" collectionCode="NTOU" collectorName="G. Paulay" country="Guam" elevation="2" location="Urunao" pageId="1" pageNumber="580" specimenCode="NTOU 2014.031402" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="D8BDB5DAFF9EFFC8B54A4741FDA5FCC8" box="[534,613,839,864]" pageId="1" pageNumber="580" type="male">1 male</specimenCount>
(18.2 × 
<quantity id="0943D3B6FF9EFFC8B58E4741FCFDFCF7" box="[722,829,839,864]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="1" pageNumber="580" unit="mm" value="10.8">10.8 mm</quantity>
, with bopyrid) (
<specimenCode id="9E1DD628FF9EFFC8B4A14741FB2FFCC8" box="[1021,1263,839,864]" collectionCode="NTOU" country="Taiwan" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="1" pageNumber="580">NTOU 2014.031402</specimenCode>
), same data as holotype
</materialsCitation>
.— 
<materialsCitation id="7ED3740EFF9EFFC8B67D476AFCDEFC00" ID-GBIF-Occurrence="3326196312" collectingDate="2000-10-25" collectionCode="ZRC" collectorName="F. Schroeder" country="Northern Mariana Islands" elevation="5" location="Agat Bay" municipality="Alutom Islands" pageId="1" pageNumber="580" specimenCode="ZRC 2014.0184" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="D8BDB5DAFF9EFFC8B67D476AFEA7FC2D" box="[289,359,876,901]" pageId="1" pageNumber="580" type="male">1 male</specimenCount>
(12.3 × 
<quantity id="0943D3B6FF9EFFC8B699476AFDDCFC2D" box="[453,540,876,901]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="1" pageNumber="580" unit="mm" value="6.8">6.8 mm</quantity>
) (
<specimenCode id="9E1DD628FF9EFFC8B56E476AFD2DFC2D" box="[562,749,876,901]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="580">ZRC 2014.0184</specimenCode>
), 
<location id="CB642888FF9EFFC8B45C476AFCABFC2C" LSID="urn:lsid:plazi:treatment:4612CF45FF9EFFCEB7CB4613FDCAFB56:CB642888FF9EFFC8B45C476AFCABFC2C" box="[768,875,876,901]" country="Northern Mariana Islands" municipality="Alutom Islands" name="Agat Bay" pageId="1" pageNumber="580">Agat Bay</location>
, north of 
<collectingMunicipality id="2E60E429FF9EFFC8B48A476AFB44FC2D" box="[982,1156,876,901]" pageId="1" pageNumber="580">Alutom Islands</collectingMunicipality>
, 
<collectingCountry id="B6AC3EC3FF9EFFC8B3CC476AFA85FC2D" box="[1168,1349,876,901]" name="Northern Mariana Islands" pageId="1" pageNumber="580">Mariana Islands</collectingCountry>
, 
<collectingCountry id="B6AC3EC3FF9EFFC8B20D476AFA58FC2D" box="[1361,1432,876,901]" name="Guam" pageId="1" pageNumber="580">Guam</collectingCountry>
, 
<quantity id="0943D3B6FF9EFFC8B7CB4789FF20FC0F" box="[151,224,911,936]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="1" pageNumber="580" unit="m" value="4.5" valueMax="6.0" valueMin="3.0">
<elevation id="45969960FF9EFFC8B7CB4789FF20FC0F" box="[151,224,911,936]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="1" pageNumber="580" unit="m" value="4.5" valueMax="6.0" valueMin="3.0">3–6 m</elevation>
</quantity>
, among rocks, coll. 
<collectorName id="634E1B85FF9EFFC8B69F4796FD93FC00" box="[451,595,911,936]" pageId="1" pageNumber="580">F. Schroeder</collectorName>
, 
<date id="BA055893FF9EFFC8B5024789FCDEFC00" box="[606,798,911,936]" pageId="1" pageNumber="580" value="2000-10-25">
<collectingDate id="AA41A17BFF9EFFC8B5024789FCDEFC00" box="[606,798,911,936]" pageId="1" pageNumber="580" value="2000-10-25">25 October 2000</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="7ED3740EFF9EFFC8B41C4789FF14FC58" ID-GBIF-Occurrence="3326196315" collectingDate="2000-11-09" collectionCode="ZRC" collectorName="H. Conley" country="Northern Mariana Islands" elevation="5" location="Agat Bay" municipality="Alutom Islands" pageId="1" pageNumber="580" specimenCode="ZRC 2014.0182" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="D8BDB5DAFF9EFFC8B41C4789FC54FC00" box="[832,916,911,936]" pageId="1" pageNumber="580" type="male">3 males</specimenCount>
(9.6 × 5.5–16.5 × 
<quantity id="0943D3B6FF9EFFC8B33B4789FB7FFC00" box="[1127,1215,911,936]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="1" pageNumber="580" unit="mm" value="9.5">9.5 mm</quantity>
) (
<specimenCode id="9E1DD628FF9EFFC8B3894789FA51FC00" box="[1237,1425,911,936]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="580">ZRC 2014.0182</specimenCode>
), 
<location id="CB642888FF9EFFC8B7CB47B2FEC2FC64" LSID="urn:lsid:plazi:treatment:4612CF45FF9EFFCEB7CB4613FDCAFB56:CB642888FF9EFFC8B7CB47B2FEC2FC64" box="[151,258,948,973]" country="Northern Mariana Islands" municipality="Alutom Islands" name="Agat Bay" pageId="1" pageNumber="580">Agat Bay</location>
, north of 
<collectingMunicipality id="2E60E429FF9EFFC8B63747B2FDD9FC65" box="[363,537,948,973]" pageId="1" pageNumber="580">Alutom Islands</collectingMunicipality>
, 
<collectingCountry id="B6AC3EC3FF9EFFC8B57847B2FD19FC65" box="[548,729,948,973]" name="Northern Mariana Islands" pageId="1" pageNumber="580">Mariana Islands</collectingCountry>
, 
<collectingCountry id="B6AC3EC3FF9EFFC8B5B847B2FCEBFC65" box="[740,811,948,973]" name="Guam" pageId="1" pageNumber="580">Guam</collectingCountry>
, 
<quantity id="0943D3B6FF9EFFC8B46A47B2FCBEFC63" box="[822,894,948,973]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="1" pageNumber="580" unit="m" value="4.5" valueMax="6.0" valueMin="3.0">
<elevation id="45969960FF9EFFC8B46A47B2FCBEFC63" box="[822,894,948,973]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="1" pageNumber="580" unit="m" value="4.5" valueMax="6.0" valueMin="3.0">3–6 m</elevation>
</quantity>
, among silty rocks, coll. 
<collectorName id="634E1B85FF9EFFC8B3C847B2FAC7FC65" box="[1172,1287,948,973]" pageId="1" pageNumber="580">H. Conley</collectorName>
, 
<date id="BA055893FF9EFFC8B24E47B2FF14FC58" pageId="1" pageNumber="580" value="2000-11-09">
<collectingDate id="AA41A17BFF9EFFC8B24E47B2FF14FC58" pageId="1" pageNumber="580" value="2000-11-09">9 November 2000</collectingDate>
</date>
</materialsCitation>
.— 
<specimenCount id="D8BDB5DAFF9EFFC8B7AA47D1FEFDFC58" box="[246,317,983,1008]" pageId="1" pageNumber="580" type="male">1 male</specimenCount>
(10.4 × 
<quantity id="0943D3B6FF9EFFC8B6C347D1FE3AFC58" box="[415,506,983,1008]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="1" pageNumber="580" unit="mm" value="6.1">6.1 mm</quantity>
), 
<materialsCitation id="7ED3740EFF9EFFC8B55147D1FBB2FBBD" ID-GBIF-Occurrence="3326196304" collectingDate="2000-11" collectionCode="ZRC" collectorName="H. Conley" country="Northern Mariana Islands" elevation="5" location="Agat Bay" municipality="Alutom Islands" pageId="1" pageNumber="580" specimenCode="ZRC 2014.0183" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="D8BDB5DAFF9EFFC8B55147D1FDAFFC58" box="[525,623,983,1008]" pageId="1" pageNumber="580" type="female">1 female</specimenCount>
(11.9 × 
<quantity id="0943D3B6FF9EFFC8B58C47D1FCE8FC58" box="[720,808,983,1008]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="1" pageNumber="580" unit="mm" value="7.5">7.5 mm</quantity>
) (
<specimenCode id="9E1DD628FF9EFFC8B46347D1FC3BFC58" box="[831,1019,983,1008]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="580">ZRC 2014.0183</specimenCode>
), 
<location id="CB642888FF9EFFC8B35247D1FBBAFC58" LSID="urn:lsid:plazi:treatment:4612CF45FF9EFFCEB7CB4613FDCAFB56:CB642888FF9EFFC8B35247D1FBBAFC58" box="[1038,1146,983,1008]" country="Northern Mariana Islands" municipality="Alutom Islands" name="Agat Bay" pageId="1" pageNumber="580">Agat Bay</location>
, north of 
<collectingMunicipality id="2E60E429FF9EFFC8B3B447D1FA57FC58" box="[1256,1431,983,1008]" pageId="1" pageNumber="580">Alutom Islands</collectingMunicipality>
, 
<collectingCountry id="B6AC3EC3FF9EFFC8B7CB47FAFE8EFBBD" box="[151,334,1020,1045]" name="Northern Mariana Islands" pageId="1" pageNumber="580">Mariana Islands</collectingCountry>
, 
<collectingCountry id="B6AC3EC3FF9EFFC8B60647FAFE61FBBD" box="[346,417,1020,1045]" name="Guam" pageId="1" pageNumber="580">Guam</collectingCountry>
, 
<quantity id="0943D3B6FF9EFFC8B6F147FAFE37FBBB" box="[429,503,1020,1045]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="1" pageNumber="580" unit="m" value="4.5" valueMax="6.0" valueMin="3.0">
<elevation id="45969960FF9EFFC8B6F147FAFE37FBBB" box="[429,503,1020,1045]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="1" pageNumber="580" unit="m" value="4.5" valueMax="6.0" valueMin="3.0">3–6 m</elevation>
</quantity>
, among rocks and silt, coll. 
<collectorName id="634E1B85FF9EFFC8B46A47FAFC6BFBBD" box="[822,939,1020,1045]" pageId="1" pageNumber="580">H. Conley</collectorName>
, 
<date id="BA055893FF9EFFC8B4EB47FAFBB2FBBD" box="[951,1138,1020,1045]" pageId="1" pageNumber="580" value="2000-11">
<collectingDate id="AA41A17BFF9EFFC8B4EB47FAFBB2FBBD" box="[951,1138,1020,1045]" pageId="1" pageNumber="580" value="2000-11">November 2000</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="7ED3740EFF9EFFC8B3C947FAFE17FBF5" ID-GBIF-Occurrence="3326196309" collectingDate="2000-11-15" collectionCode="ZRC" collectorName="H. Conley" country="Northern Mariana Islands" elevation="5" location="Agat Bay" municipality="Alutom Islands" pageId="1" pageNumber="580" specimenCode="ZRC 2014.0185" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="D8BDB5DAFF9EFFC8B3C947FAFB1CFBBD" box="[1173,1244,1020,1045]" pageId="1" pageNumber="580" type="male">1 male</specimenCount>
(11.6 × 
<quantity id="0943D3B6FF9EFFC8B26047FAFA54FBBD" box="[1340,1428,1020,1045]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="1" pageNumber="580" unit="mm" value="7.1">7.1 mm</quantity>
) (
<specimenCode id="9E1DD628FF9EFFC8B7C24019FE9EFB90" box="[158,350,1055,1080]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="580">ZRC 2014.0185</specimenCode>
), 
<location id="CB642888FF9EFFC8B6284019FE24FB90" LSID="urn:lsid:plazi:treatment:4612CF45FF9EFFCEB7CB4613FDCAFB56:CB642888FF9EFFC8B6284019FE24FB90" box="[372,484,1055,1080]" country="Northern Mariana Islands" municipality="Alutom Islands" name="Agat Bay" pageId="1" pageNumber="580">Agat Bay</location>
, north of 
<collectingMunicipality id="2E60E429FF9EFFC8B5014019FCD0FB90" box="[605,784,1055,1080]" pageId="1" pageNumber="580">Alutom Islands</collectingMunicipality>
, 
<collectingCountry id="B6AC3EC3FF9EFFC8B4434019FC1AFB90" box="[799,986,1055,1080]" name="Northern Mariana Islands" pageId="1" pageNumber="580">Mariana Islands</collectingCountry>
, 
<collectingCountry id="B6AC3EC3FF9EFFC8B4B64019FBF1FB90" box="[1002,1073,1055,1080]" name="Guam" pageId="1" pageNumber="580">Guam</collectingCountry>
, 
<quantity id="0943D3B6FF9EFFC8B31D4019FB4EFB9F" box="[1089,1166,1055,1080]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="1" pageNumber="580" unit="m" value="4.5" valueMax="6.0" valueMin="3.0">
<elevation id="45969960FF9EFFC8B31D4019FB4EFB9F" box="[1089,1166,1055,1080]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="1" pageNumber="580" unit="m" value="4.5" valueMax="6.0" valueMin="3.0">3–6 m</elevation>
</quantity>
, among rocks, coll. 
<collectorName id="634E1B85FF9EFFC8B2DD4026FF29FBF5" pageId="1" pageNumber="580">H. Conley</collectorName>
, 
<date id="BA055893FF9EFFC8B7AA4042FE17FBF5" box="[246,471,1092,1117]" pageId="1" pageNumber="580" value="2000-11-15">
<collectingDate id="AA41A17BFF9EFFC8B7AA4042FE17FBF5" box="[246,471,1092,1117]" pageId="1" pageNumber="580" value="2000-11-15">15 November 2000</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="7ED3740EFF9EFFC8B6BF4042FD43FBF5" ID-GBIF-Occurrence="3326196311" box="[483,643,1092,1117]" country="Northern Mariana Islands" location="Other" pageId="1" pageNumber="580" specimenCount="1" typeStatus="paratype">
<location id="CB642888FF9EFFC8B6BF4042FDE3FBF5" LSID="urn:lsid:plazi:treatment:4612CF45FF9EFFCEB7CB4613FDCAFB56:CB642888FF9EFFC8B6BF4042FDE3FBF5" box="[483,547,1092,1117]" country="Northern Mariana Islands" name="Other" pageId="1" pageNumber="580">Other</location>
records
</materialsCitation>
: 
<materialsCitation id="7ED3740EFF9EFFC8B5D34042FB0FFBF5" ID-GBIF-Occurrence="3326196314" box="[655,1231,1092,1117]" collectionCode="NTOU" country="Northern Mariana Islands" pageId="1" pageNumber="580" specimenCode="NTOU 2009.062401" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="D8BDB5DAFF9EFFC8B5D34042FD32FBF5" box="[655,754,1092,1117]" pageId="1" pageNumber="580" type="female">1 female</specimenCount>
(25.3 × 
<quantity id="0943D3B6FF9EFFC8B4044042FC7FFBF5" box="[856,959,1092,1117]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="1" pageNumber="580" unit="mm" value="14.3">14.3 mm</quantity>
) (
<specimenCode id="9E1DD628FF9EFFC8B4844042FB07FBF5" box="[984,1223,1092,1117]" collectionCode="NTOU" country="Taiwan" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="1" pageNumber="580">NTOU 2009.062401</specimenCode>
)
</materialsCitation>
, 
<materialsCitation id="7ED3740EFF9EFFC8B3804042FED3FB0D" ID-GBIF-Occurrence="3326196302" collectingDate="2009-06-24" collectionCode="ZRC" collectorName="Y. - J. Shih" country="Taiwan" county="Pintung County" location="Pintung County" pageId="1" pageNumber="580" specimenCode="ZRC 2014.0186" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="D8BDB5DAFF9EFFC8B3804042FAFFFBF5" box="[1244,1343,1092,1117]" pageId="1" pageNumber="580" type="female">1 female</specimenCount>
(17.7 × 
<quantity id="0943D3B6FF9EFFC8B7CB4061FF3FFB28" box="[151,255,1127,1152]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="1" pageNumber="580" unit="mm" value="10.2">10.2 mm</quantity>
) (
<specimenCode id="9E1DD628FF9EFFC8B6444061FE15FB28" box="[280,469,1127,1152]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="580">ZRC 2014.0186</specimenCode>
), Chia-Le-Sui, 
<collectingCounty id="276506DFFF9EFFC8B5DA4061FCFBFB28" box="[646,827,1127,1152]" pageId="1" pageNumber="580">Pintung County</collectingCounty>
, 
<collectingCountry id="B6AC3EC3FF9EFFC8B4144061FC5DFB28" box="[840,925,1127,1152]" name="Taiwan" pageId="1" pageNumber="580">Taiwan</collectingCountry>
, intertidal, among rocks, coll. 
<collectorName id="634E1B85FF9EFFC8B3A3406EFAB2FB28" box="[1279,1394,1127,1152]" pageId="1" pageNumber="580">Y.-J. Shih</collectorName>
, 
<date id="BA055893FF9EFFC8B2DC4061FED3FB0D" pageId="1" pageNumber="580" value="2009-06-24">
<collectingDate id="AA41A17BFF9EFFC8B2DC4061FED3FB0D" pageId="1" pageNumber="580" value="2009-06-24">24 June 2009</collectingDate>
</date>
</materialsCitation>
.— 
<materialsCitation id="7ED3740EFF9EFFC8B66A408AFDA6FB60" ID-GBIF-Occurrence="3326196308" collectingDate="1997-04-29" collectionCode="ZRC" country="Taiwan" elevation="1" pageId="1" pageNumber="580" specimenCode="ZRC 2000.0483" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="D8BDB5DAFF9EFFC8B66A408AFE57FB0D" box="[310,407,1164,1189]" pageId="1" pageNumber="580" type="female">1 female</specimenCount>
(23.9 × 15.0 mm) (
<specimenCode id="9E1DD628FF9EFFC8B5D0408AFC8CFB0D" box="[652,844,1164,1189]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="580">ZRC 2000.0483</specimenCode>
), ca. 
<quantity id="0943D3B6FF9EFFC8B4CC408AFC02FB0B" box="[912,962,1164,1189]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="580" unit="m" value="1.0">
<elevation id="45969960FF9EFFC8B4CC408AFC02FB0B" box="[912,962,1164,1189]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="580" unit="m" value="1.0">1 m</elevation>
</quantity>
, in coral head, 
<collectorName id="634E1B85FF9EFFC8B3DD408AFAFEFB0B" box="[1153,1342,1164,1189]" pageId="1" pageNumber="580">Chinaman’s Hat</collectorName>
, O‘ahu, 
<collectorName id="634E1B85FF9EFFC8B7CB40A9FF34FB60" box="[151,244,1199,1224]" pageId="1" pageNumber="580">Hawai‘i</collectorName>
, coll. 
<collectorName id="634E1B85FF9EFFC8B66B40B6FE7AFB60" box="[311,442,1199,1224]" pageId="1" pageNumber="580">D. Takaoka</collectorName>
, 
<date id="BA055893FF9EFFC8B69A40A9FDA6FB60" box="[454,614,1199,1224]" pageId="1" pageNumber="580" value="1997-04-29">
<collectingDate id="AA41A17BFF9EFFC8B69A40A9FDA6FB60" box="[454,614,1199,1224]" pageId="1" pageNumber="580" value="1997-04-29">29 April 1997</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="86A12DD8FF9EFFC8B79B40D2FE35F9D0" pageId="1" pageNumber="580" type="diagnosis">
<paragraph id="CE047E53FF9EFFC8B79B40D2FE35F9D0" blockId="1.[151,1437,767,2017]" pageId="1" pageNumber="580">
<emphasis id="FCCFA241FF9EFFC8B79B40D2FE81FB43" box="[199,321,1236,1261]" inLineHeading="true" pageId="1" pageNumber="580" reason="1">
<emphasis id="FCCFA241FF9EFFC8B79B40D2FEFAFB45" bold="true" box="[199,314,1236,1261]" pageId="1" pageNumber="580">Diagnosis</emphasis>
.
</emphasis>
Carapace grooves separating gastric, branchial regions shallow, with scattered short setae that never obscure margins or surfaces; dorsal surface covered with numerous evenly spaced, small rounded granules; pterygostomial, suborbital, sub-branchial regions covered with numerous small rounded granules (
<figureCitation id="568062D6FF9EFFC8B25B411AFA9AFA9D" box="[1287,1370,1308,1333]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="1" pageNumber="580">Figs. 2</figureCitation>
, 
<figureCitation id="568062D6FF9EFFC8B235411AFA4EFA9D" box="[1385,1422,1308,1333]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-79@5.[241,1345,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. A, Neoliomera fragraea n. sp., paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="1" pageNumber="580">3A</figureCitation>
); front bilobed, subtruncate lobes separated by V-shaped cleft (
<figureCitation id="568062D6FF9EFFC8B40A4139FC67FAF0" box="[854,935,1343,1368]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="1" pageNumber="580">Figs. 2</figureCitation>
, 
<figureCitation id="568062D6FF9EFFC8B4E94139FC1BFAF0" box="[949,987,1343,1368]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-79@5.[241,1345,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. A, Neoliomera fragraea n. sp., paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="1" pageNumber="580">3A</figureCitation>
); anterolateral margin strongly convex; posterior one-third with 2 low but distinct granular lobes separated by shallow broad cleft; anterior two-thirds granular, uneven, appearing entire (
<figureCitation id="568062D6FF9EFFC8B5764181FD72FA37" box="[554,690,1415,1440]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="1" pageNumber="580">Figs. 2A, B</figureCitation>
, 
<figureCitation id="568062D6FF9EFFC8B5E34181FD24FA08" box="[703,740,1415,1440]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-79@5.[241,1345,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. A, Neoliomera fragraea n. sp., paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="1" pageNumber="580">3A</figureCitation>
); chelipeds with outer surfaces covered with numerous small granules, fingers shorter than palm; dactylus, pollex completely pigmented, pigmentation on pollex extending slightly into palm (
<figureCitation id="568062D6FF9EFFC8B63641C9FE0DFA40" box="[362,461,1487,1512]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="1" pageNumber="580">Figs. 2A</figureCitation>
, 
<figureCitation id="568062D6FF9EFFC8B68441C9FDDDFA40" box="[472,541,1487,1512]" captionStart="FIGURE 4" captionStartId="6.[151,250,1267,1289]" captionTargetBox="[164,1428,193,1248]" captionTargetId="figure-419@6.[164,1429,193,1248]" captionTargetPageId="6" captionText="FIGURE 4. A, B, E, Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam; C, D, F, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, outer views of right chelae; B, D, inner views of right chelae; E, F, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4920424" httpUri="https://zenodo.org/record/4920424/files/figure.png" pageId="1" pageNumber="580">4A, B</figureCitation>
); ambulatory legs relatively short, margins of all articles with scattered long and short simple setae that do not obscure margins; margins, submarginal outer surfaces not cristate, surfaces covered with distinct granules, merus very short (
<figureCitation id="568062D6FF9EFFC8B53E4211FD06F998" box="[610,710,1559,1584]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="1" pageNumber="580">Figs. 2A</figureCitation>
, 
<figureCitation id="568062D6FF9EFFC8B58E4211FCD8F987" box="[722,792,1559,1584]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-79@5.[241,1345,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. A, Neoliomera fragraea n. sp., paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="1" pageNumber="580">3A, B</figureCitation>
, 
<figureCitation id="568062D6FF9EFFC8B4784211FC86F998" box="[804,838,1559,1584]" captionStart="FIGURE 4" captionStartId="6.[151,250,1267,1289]" captionTargetBox="[164,1428,193,1248]" captionTargetId="figure-419@6.[164,1429,193,1248]" captionTargetPageId="6" captionText="FIGURE 4. A, B, E, Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam; C, D, F, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, outer views of right chelae; B, D, inner views of right chelae; E, F, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4920424" httpUri="https://zenodo.org/record/4920424/files/figure.png" pageId="1" pageNumber="580">4E</figureCitation>
); thoracic sternum broad, entire surface covered with numerous small rounded granules (
<figureCitation id="568062D6FF9EFFC8B57B423AFD41F9FD" box="[551,641,1596,1621]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-79@5.[241,1345,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. A, Neoliomera fragraea n. sp., paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="1" pageNumber="580">Fig. 3B</figureCitation>
); G1 with prominent subovate subdistal lobe, long, extending beyond almost straight tip (
<figureCitation id="568062D6FF9EFFC8B62C4259FE28F9D0" box="[368,488,1631,1656]" captionStart="FIGURE 7" captionStartId="10.[151,250,1223,1245]" captionTargetBox="[176,1403,401,1181]" captionTargetId="figure-371@10.[151,1436,393,1200]" captionTargetPageId="10" captionText="FIGURE 7. A, C, Neoliomera pubescens (H. Milne Edwards, 1873), lectotype male (30.5 × 17.5 mm) (MNHN), Mauritius; D, Neoliomera demani Forest &amp; Guinot, 1961, paratype male (23.0 × 13.0 mm) (MNHN), Tahiti; B, E, Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, B, left G1; C–E, distal parts of left G1s. A, C, after Forest &amp; Guinot (1961: fig. 75); D, after Forest &amp; Guinot (1961: fig. 76)." figureDoi="http://doi.org/10.5281/zenodo.4920430" httpUri="https://zenodo.org/record/4920430/files/figure.png" pageId="1" pageNumber="580">Fig. 7B, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="86A12DD8FF9EFFCBB79B4282FAADFA36" lastPageId="2" lastPageNumber="581" pageId="1" pageNumber="580" type="description">
<paragraph id="CE047E53FF9EFFCBB79B4282FE74FE5D" blockId="1.[151,1437,767,2017]" lastBlockId="2.[151,1437,151,2027]" lastPageId="2" lastPageNumber="581" pageId="1" pageNumber="580">
<emphasis id="FCCFA241FF9EFFC8B79B4282FE98F933" box="[199,344,1668,1693]" inLineHeading="true" pageId="1" pageNumber="580" reason="1">
<emphasis id="FCCFA241FF9EFFC8B79B4282FE91F935" bold="true" box="[199,337,1668,1693]" pageId="1" pageNumber="580">Description</emphasis>
.
</emphasis>
Carapace regions not well defined; grooves separating gastric, branchial regions shallow, just discernible, with scattered short setae which never obscure margins or surfaces; shallow Y-shaped groove separating low epigastric regions; dorsal surface covered with numerous evenly spaced small rounded granules, those on median parts lower, relatively smaller; pterygostomial, suborbital, sub-branchial regions covered numerous small rounded granules (
<figureCitation id="568062D6FF9EFFC8B57F4312FDB2F885" box="[547,626,1812,1837]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="1" pageNumber="580">Figs. 2</figureCitation>
, 
<figureCitation id="568062D6FF9EFFC8B5234312FD63F885" box="[639,675,1812,1837]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-79@5.[241,1345,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. A, Neoliomera fragraea n. sp., paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="1" pageNumber="580">3A</figureCitation>
, 
<figureCitation id="568062D6FF9EFFC8B5EC4312FD16F885" box="[688,726,1812,1837]" captionStart="FIGURE 5" captionStartId="8.[151,250,1727,1749]" captionTargetBox="[151,1436,193,1706]" captionTargetId="figure-48@8.[151,1436,193,1706]" captionTargetPageId="8" captionText="FIGURE 5. Neoliomera fragraea n. sp., female (23.9 × 15.0 mm) (ZRC 2000.0483), Hawai‘i. A, dorsal view; B, thoracic sternum showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.4920426" httpUri="https://zenodo.org/record/4920426/files/figure.png" pageId="1" pageNumber="580">5A</figureCitation>
). Front produced just beyond orbits; bilobed, subtruncate lobes separated by distinct V-shaped cleft; margin convex, gently deflexed downwards (
<figureCitation id="568062D6FF9EFFC8B3294331FB0BF8F8" box="[1141,1227,1847,1872]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="1" pageNumber="580">Figs. 2</figureCitation>
, 
<figureCitation id="568062D6FF9EFFC8B3824331FAC2F8F8" box="[1246,1282,1847,1872]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-79@5.[241,1345,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. A, Neoliomera fragraea n. sp., paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="1" pageNumber="580">3A</figureCitation>
, 
<figureCitation id="568062D6FF9EFFC8B2494331FAFBF8F8" box="[1301,1339,1847,1872]" captionStart="FIGURE 5" captionStartId="8.[151,250,1727,1749]" captionTargetBox="[151,1436,193,1706]" captionTargetId="figure-48@8.[151,1436,193,1706]" captionTargetPageId="8" captionText="FIGURE 5. Neoliomera fragraea n. sp., female (23.9 × 15.0 mm) (ZRC 2000.0483), Hawai‘i. A, dorsal view; B, thoracic sternum showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.4920426" httpUri="https://zenodo.org/record/4920426/files/figure.png" pageId="1" pageNumber="580">5A</figureCitation>
). Orbits subovate; cornea relatively large, pigmented; ocular peduncle short, stout, covered with small granules (
<figureCitation id="568062D6FF9EFFC8B269435AFA50F8DD" box="[1333,1424,1884,1909]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="1" pageNumber="580">Fig. 2C</figureCitation>
). Supraorbital margin granular; external orbital angle marked by low granular lobe or group of granules (
<figureCitation id="568062D6FF9EFFC8B2194379FA57F830" box="[1349,1431,1919,1944]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="1" pageNumber="580">Figs. 2</figureCitation>
, 
<figureCitation id="568062D6FF9EFFC8B7CB43A2FF7CF815" box="[151,188,1956,1981]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-79@5.[241,1345,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. A, Neoliomera fragraea n. sp., paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="1" pageNumber="580">3A</figureCitation>
). Infraorbital margin uneven, granular, otherwise unarmed (
<figureCitation id="568062D6FF9EFFC8B42F43A2FC0FF815" box="[883,975,1956,1981]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="1" pageNumber="580">Fig. 2C</figureCitation>
). Anterolateral margin strongly convex; posterior one-third with 2 low but distinct granular lobes, separated by shallow broad cleft; anterior two-thirds uneven, appears entire, lined with granules of various sizes (
<figureCitation id="568062D6FF9DFFCBB40C4491FC19FF07" box="[848,985,151,176]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="2" pageNumber="581">Figs. 2A, B</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB4BB4491FBCCFF18" box="[999,1036,151,176]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-79@5.[241,1345,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. A, Neoliomera fragraea n. sp., paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="2" pageNumber="581">3A</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB3464491FB80FF18" box="[1050,1088,151,176]" captionStart="FIGURE 5" captionStartId="8.[151,250,1727,1749]" captionTargetBox="[151,1436,193,1706]" captionTargetId="figure-48@8.[151,1436,193,1706]" captionTargetPageId="8" captionText="FIGURE 5. Neoliomera fragraea n. sp., female (23.9 × 15.0 mm) (ZRC 2000.0483), Hawai‘i. A, dorsal view; B, thoracic sternum showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.4920426" httpUri="https://zenodo.org/record/4920426/files/figure.png" pageId="2" pageNumber="581">5A</figureCitation>
). Posterolateral margin almost straight or gently convex, converging sharply to posterior carapace margin (
<figureCitation id="568062D6FF9DFFCBB4BB44BAFBACFF7B" box="[999,1132,188,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="2" pageNumber="581">Figs. 2A, B</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB32444BAFB5CFF7D" box="[1144,1180,188,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-79@5.[241,1345,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. A, Neoliomera fragraea n. sp., paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="2" pageNumber="581">3A</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB3F444BAFB0EFF7D" box="[1192,1230,188,213]" captionStart="FIGURE 5" captionStartId="8.[151,250,1727,1749]" captionTargetBox="[151,1436,193,1706]" captionTargetId="figure-48@8.[151,1436,193,1706]" captionTargetPageId="8" captionText="FIGURE 5. Neoliomera fragraea n. sp., female (23.9 × 15.0 mm) (ZRC 2000.0483), Hawai‘i. A, dorsal view; B, thoracic sternum showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.4920426" httpUri="https://zenodo.org/record/4920426/files/figure.png" pageId="2" pageNumber="581">5A</figureCitation>
). Posterior margin of carapace gently convex, lined with closely-set small submarginal granules (
<figureCitation id="568062D6FF9DFFCBB34444D9FB61FF5F" box="[1048,1185,223,248]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="2" pageNumber="581">Figs. 2A, B</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB3F344D9FB13FF50" box="[1199,1235,223,248]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-79@5.[241,1345,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. A, Neoliomera fragraea n. sp., paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="2" pageNumber="581">3A</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB3BD44D9FAC7FF50" box="[1249,1287,223,248]" captionStart="FIGURE 5" captionStartId="8.[151,250,1727,1749]" captionTargetBox="[151,1436,193,1706]" captionTargetId="figure-48@8.[151,1436,193,1706]" captionTargetPageId="8" captionText="FIGURE 5. Neoliomera fragraea n. sp., female (23.9 × 15.0 mm) (ZRC 2000.0483), Hawai‘i. A, dorsal view; B, thoracic sternum showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.4920426" httpUri="https://zenodo.org/record/4920426/files/figure.png" pageId="2" pageNumber="581">5A</figureCitation>
). Antennules short, folding transversely (
<figureCitation id="568062D6FF9DFFCBB6954502FDE1FEB5" box="[457,545,260,285]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="2" pageNumber="581">Fig. 2C</figureCitation>
). Antennal flagellum short, passes through orbital hiatus, not just reaching outer edge of orbit; basal segment subrectangular, relatively large (
<figureCitation id="568062D6FF9DFFCBB4054521FC74FEE8" box="[857,948,295,320]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="2" pageNumber="581">Fig. 2C</figureCitation>
). Surface of epistome medially depressed; posterior margin gently sinuous from frontal view, median part slightly dilated, with small median cleft (
<figureCitation id="568062D6FF9DFFCBB26A454AFA4FFECD" box="[1334,1423,332,357]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="2" pageNumber="581">Fig. 2C</figureCitation>
). Endostome without discernible ridges. Outer surface of third maxilliped gently granular to smooth. Ischium subrectangular, with distinct median oblique sulcus. Merus quadrate, anteroexternal margin angular but rounded, slightly produced, median part gently depressed. Exopod relatively stout proximally, distal part not to anterior edge of merus; flagellum long.
</paragraph>
<paragraph id="CE047E53FF9DFFCBB79B45F9FD5EFCA3" blockId="2.[151,1437,151,2027]" pageId="2" pageNumber="581">
Chelipeds symmetrical (
<figureCitation id="568062D6FF9DFFCBB68145F9FD83FDB0" box="[477,579,511,536]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="2" pageNumber="581">Figs. 2A</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB50D45F9FDB6FDB0" box="[593,630,511,536]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-79@5.[241,1345,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. A, Neoliomera fragraea n. sp., paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="2" pageNumber="581">3A</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB5D845F9FD6AFDB0" box="[644,682,511,536]" captionStart="FIGURE 5" captionStartId="8.[151,250,1727,1749]" captionTargetBox="[151,1436,193,1706]" captionTargetId="figure-48@8.[151,1436,193,1706]" captionTargetPageId="8" captionText="FIGURE 5. Neoliomera fragraea n. sp., female (23.9 × 15.0 mm) (ZRC 2000.0483), Hawai‘i. A, dorsal view; B, thoracic sternum showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.4920426" httpUri="https://zenodo.org/record/4920426/files/figure.png" pageId="2" pageNumber="581">5A</figureCitation>
). Outer surfaces of merus, carpus, chelae covered with numerous rounded granules of varying sizes, granules on dorsal margins generally larger, sharper (
<figureCitation id="568062D6FF9DFFCBB3C74624FAC2FD93" box="[1179,1282,546,571]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="2" pageNumber="581">Figs. 2A</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB24E4624FAF6FD93" box="[1298,1334,546,571]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-79@5.[241,1345,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. A, Neoliomera fragraea n. sp., paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="2" pageNumber="581">3A</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB2194624FA4EFD93" box="[1349,1422,546,571]" captionStart="FIGURE 4" captionStartId="6.[151,250,1267,1289]" captionTargetBox="[164,1428,193,1248]" captionTargetId="figure-419@6.[164,1429,193,1248]" captionTargetPageId="6" captionText="FIGURE 4. A, B, E, Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam; C, D, F, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, outer views of right chelae; B, D, inner views of right chelae; E, F, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4920424" httpUri="https://zenodo.org/record/4920424/files/figure.png" pageId="2" pageNumber="581">4A, B</figureCitation>
). Carpus subovate, inner distal angle with low sharp tooth (
<figureCitation id="568062D6FF9DFFCBB4774643FC50FDF6" box="[811,912,581,606]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="2" pageNumber="581">Figs. 2A</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB4C24643FC02FDF6" box="[926,962,581,606]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-79@5.[241,1345,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. A, Neoliomera fragraea n. sp., paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="2" pageNumber="581">3A</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB48C4643FBD6FDF6" box="[976,1046,581,606]" captionStart="FIGURE 4" captionStartId="6.[151,250,1267,1289]" captionTargetBox="[164,1428,193,1248]" captionTargetId="figure-419@6.[164,1429,193,1248]" captionTargetPageId="6" captionText="FIGURE 4. A, B, E, Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam; C, D, F, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, outer views of right chelae; B, D, inner views of right chelae; E, F, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4920424" httpUri="https://zenodo.org/record/4920424/files/figure.png" pageId="2" pageNumber="581">4A, B</figureCitation>
). Merus without distinct spines or teeth. Fingers shorter than palm, outer surface of each finger with 2 longitudinal grooves, cutting margins with several well developed teeth with convex margins; distal part curved, tip with inner surface excavated, subspatuliform, inner surface with scattered simple setae (
<figureCitation id="568062D6FF9DFFCBB47046ABFC68FD6E" box="[812,936,685,710]" captionStart="FIGURE 4" captionStartId="6.[151,250,1267,1289]" captionTargetBox="[164,1428,193,1248]" captionTargetId="figure-419@6.[164,1429,193,1248]" captionTargetPageId="6" captionText="FIGURE 4. A, B, E, Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam; C, D, F, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, outer views of right chelae; B, D, inner views of right chelae; E, F, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4920424" httpUri="https://zenodo.org/record/4920424/files/figure.png" pageId="2" pageNumber="581">Fig. 4A, B</figureCitation>
). Dactylus completely black to dark brown, tip lighter coloured (
<figureCitation id="568062D6FF9DFFCBB62146C9FE37FD40" box="[381,503,719,744]" captionStart="FIGURE 4" captionStartId="6.[151,250,1267,1289]" captionTargetBox="[164,1428,193,1248]" captionTargetId="figure-419@6.[164,1429,193,1248]" captionTargetPageId="6" captionText="FIGURE 4. A, B, E, Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam; C, D, F, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, outer views of right chelae; B, D, inner views of right chelae; E, F, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4920424" httpUri="https://zenodo.org/record/4920424/files/figure.png" pageId="2" pageNumber="581">Fig. 4A, B</figureCitation>
). Pollex black to dark brown except for lighter coloured tip, pigmentation reaching slightly into inner, outer surfaces (
<figureCitation id="568062D6FF9DFFCBB54A46F4FD50FCA3" box="[534,656,754,779]" captionStart="FIGURE 4" captionStartId="6.[151,250,1267,1289]" captionTargetBox="[164,1428,193,1248]" captionTargetId="figure-419@6.[164,1429,193,1248]" captionTargetPageId="6" captionText="FIGURE 4. A, B, E, Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam; C, D, F, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, outer views of right chelae; B, D, inner views of right chelae; E, F, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4920424" httpUri="https://zenodo.org/record/4920424/files/figure.png" pageId="2" pageNumber="581">Fig. 4A, B</figureCitation>
).
</paragraph>
<paragraph id="CE047E53FF9DFFCBB79B4713FAF2FC73" blockId="2.[151,1437,151,2027]" pageId="2" pageNumber="581">
Ambulatory legs relatively short; second pair longest (
<figureCitation id="568062D6FF9DFFCBB4174713FC5DFC86" box="[843,925,789,814]" captionStart="FIGURE 1" captionStartId="3.[151,250,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure-95@3.[151,1436,193,1884]" captionTargetPageId="3" captionText="FIGURE 1. Colours in life. A–C, Neoliomera fragraea n. sp., female (25.3 × 14.3 mm) (NTOU), Taiwan; D, Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam [photograph by G. Paulay]; E, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU) [photograph by G. Paulay]; F, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan [photograph by Wang Kuei-Ching]." figureDoi="http://doi.org/10.5281/zenodo.4920418" httpUri="https://zenodo.org/record/4920418/files/figure.png" pageId="2" pageNumber="581">Figs. 1</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB4EC4713FC15FC86" box="[944,981,789,814]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="2" pageNumber="581">2A</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB4BA4713FBCAFC86" box="[998,1034,789,814]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-79@5.[241,1345,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. A, Neoliomera fragraea n. sp., paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="2" pageNumber="581">3A</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB3474713FB81FC86" box="[1051,1089,789,814]" captionStart="FIGURE 5" captionStartId="8.[151,250,1727,1749]" captionTargetBox="[151,1436,193,1706]" captionTargetId="figure-48@8.[151,1436,193,1706]" captionTargetPageId="8" captionText="FIGURE 5. Neoliomera fragraea n. sp., female (23.9 × 15.0 mm) (ZRC 2000.0483), Hawai‘i. A, dorsal view; B, thoracic sternum showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.4920426" httpUri="https://zenodo.org/record/4920426/files/figure.png" pageId="2" pageNumber="581">5A</figureCitation>
). Margins of all articles with scattered long and short simple setae that do not obscure margins (
<figureCitation id="568062D6FF9DFFCBB4D84731FC27FCF8" box="[900,999,823,848]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="2" pageNumber="581">Figs. 2A</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB4A84731FBF9FCE7" box="[1012,1081,823,848]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-79@5.[241,1345,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. A, Neoliomera fragraea n. sp., paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="2" pageNumber="581">3A, B</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB31A4731FBA7FCF8" box="[1094,1127,823,848]" captionStart="FIGURE 4" captionStartId="6.[151,250,1267,1289]" captionTargetBox="[164,1428,193,1248]" captionTargetId="figure-419@6.[164,1429,193,1248]" captionTargetPageId="6" captionText="FIGURE 4. A, B, E, Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam; C, D, F, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, outer views of right chelae; B, D, inner views of right chelae; E, F, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4920424" httpUri="https://zenodo.org/record/4920424/files/figure.png" pageId="2" pageNumber="581">4E</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB32F4731FB59FCF8" box="[1139,1177,823,848]" captionStart="FIGURE 5" captionStartId="8.[151,250,1727,1749]" captionTargetBox="[151,1436,193,1706]" captionTargetId="figure-48@8.[151,1436,193,1706]" captionTargetPageId="8" captionText="FIGURE 5. Neoliomera fragraea n. sp., female (23.9 × 15.0 mm) (ZRC 2000.0483), Hawai‘i. A, dorsal view; B, thoracic sternum showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.4920426" httpUri="https://zenodo.org/record/4920426/files/figure.png" pageId="2" pageNumber="581">5A</figureCitation>
). Merus conspicuously short; margins, submarginal outer surfaces not cristate, surfaces covered with distinct granules; granules on dorsal margin relatively larger, uneven in shape (
<figureCitation id="568062D6FF9DFFCBB533477BFD12FC3E" box="[623,722,893,918]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-79@5.[241,1345,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. A, Neoliomera fragraea n. sp., paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="2" pageNumber="581">Figs. 3B</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB582477BFCC1FC3E" box="[734,769,893,918]" captionStart="FIGURE 4" captionStartId="6.[151,250,1267,1289]" captionTargetBox="[164,1428,193,1248]" captionTargetId="figure-419@6.[164,1429,193,1248]" captionTargetPageId="6" captionText="FIGURE 4. A, B, E, Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam; C, D, F, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, outer views of right chelae; B, D, inner views of right chelae; E, F, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4920424" httpUri="https://zenodo.org/record/4920424/files/figure.png" pageId="2" pageNumber="581">4E</figureCitation>
). Outer surfaces, margins of carpus, propodus covered with numerous small, rounded granules (
<figureCitation id="568062D6FF9DFFCBB5714799FD52FC10" box="[557,658,927,952]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="2" pageNumber="581">Figs. 2A</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB5C34799FD03FC10" box="[671,707,927,952]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-79@5.[241,1345,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. A, Neoliomera fragraea n. sp., paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="2" pageNumber="581">3A</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB58D4799FD33FC10" box="[721,755,927,952]" captionStart="FIGURE 4" captionStartId="6.[151,250,1267,1289]" captionTargetBox="[164,1428,193,1248]" captionTargetId="figure-419@6.[164,1429,193,1248]" captionTargetPageId="6" captionText="FIGURE 4. A, B, E, Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam; C, D, F, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, outer views of right chelae; B, D, inner views of right chelae; E, F, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4920424" httpUri="https://zenodo.org/record/4920424/files/figure.png" pageId="2" pageNumber="581">4E</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB5A34799FCE5FC10" box="[767,805,927,952]" captionStart="FIGURE 5" captionStartId="8.[151,250,1727,1749]" captionTargetBox="[151,1436,193,1706]" captionTargetId="figure-48@8.[151,1436,193,1706]" captionTargetPageId="8" captionText="FIGURE 5. Neoliomera fragraea n. sp., female (23.9 × 15.0 mm) (ZRC 2000.0483), Hawai‘i. A, dorsal view; B, thoracic sternum showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.4920426" httpUri="https://zenodo.org/record/4920426/files/figure.png" pageId="2" pageNumber="581">5A</figureCitation>
). Dactylus relatively short, stout, distinctly shorter than propodus, gently curved with corneous distal tip (
<figureCitation id="568062D6FF9DFFCBB59F47C4FCE6FC73" box="[707,806,962,987]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="2" pageNumber="581">Figs. 2A</figureCitation>
, 
<figureCitation id="568062D6FF9DFFCBB46E47C4FC95FC73" box="[818,853,962,987]" captionStart="FIGURE 4" captionStartId="6.[151,250,1267,1289]" captionTargetBox="[164,1428,193,1248]" captionTargetId="figure-419@6.[164,1429,193,1248]" captionTargetPageId="6" captionText="FIGURE 4. A, B, E, Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam; C, D, F, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, outer views of right chelae; B, D, inner views of right chelae; E, F, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4920424" httpUri="https://zenodo.org/record/4920424/files/figure.png" pageId="2" pageNumber="581">4E</figureCitation>
). Dactylo-propodal lock distinct (
<figureCitation id="568062D6FF9DFFCBB39247C4FAE6FC73" box="[1230,1318,962,987]" captionStart="FIGURE 4" captionStartId="6.[151,250,1267,1289]" captionTargetBox="[164,1428,193,1248]" captionTargetId="figure-419@6.[164,1429,193,1248]" captionTargetPageId="6" captionText="FIGURE 4. A, B, E, Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam; C, D, F, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, outer views of right chelae; B, D, inner views of right chelae; E, F, right fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4920424" httpUri="https://zenodo.org/record/4920424/files/figure.png" pageId="2" pageNumber="581">Fig. 4E</figureCitation>
).
</paragraph>
<paragraph id="CE047E53FF9DFFCBB79B47E3FC74FB20" blockId="2.[151,1437,151,2027]" pageId="2" pageNumber="581">
Thoracic sternum broad, entire surface covered with numerous small rounded granules; sternites 1–3 completely fused; sternites 3, 4 fused, medial suture not clearly discernible with only lateral part distinct (
<figureCitation id="568062D6FF9DFFCBB26B4001FA4EFB88" box="[1335,1422,1031,1056]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-79@5.[241,1345,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. A, Neoliomera fragraea n. sp., paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="2" pageNumber="581">Fig. 3B</figureCitation>
). Gonopore coxal. Abdomen with somites 3–5 completely fused, sutures separating segments not discernible, lateral margins gently concave; somites 1–3 trapezoidal, somite 6 about as broad as long, lateral margins gently concave; telson triangular, lateral margins gently concave, tip rounded (
<figureCitation id="568062D6FF9DFFCBB4134069FC66FB20" box="[847,934,1135,1160]" captionStart="FIGURE 3" captionStartId="5.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-79@5.[241,1345,193,1932]" captionTargetPageId="5" captionText="FIGURE 3. A, Neoliomera fragraea n. sp., paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen." figureDoi="http://doi.org/10.5281/zenodo.4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="2" pageNumber="581">Fig. 3B</figureCitation>
).
</paragraph>
<paragraph id="CE047E53FF9DFFCBB79B4094FF3CFB58" blockId="2.[151,1437,151,2027]" pageId="2" pageNumber="581">
G1 long, relatively stout, gently sinuous; with prominent subovate subdistal lobe, long, extending beyond almost straight tip; dorsal margin with numerous long plumose setae (
<figureCitation id="568062D6FF9DFFCBB48C40B3FB8FFB66" box="[976,1103,1205,1230]" captionStart="FIGURE 7" captionStartId="10.[151,250,1223,1245]" captionTargetBox="[176,1403,401,1181]" captionTargetId="figure-371@10.[151,1436,393,1200]" captionTargetPageId="10" captionText="FIGURE 7. A, C, Neoliomera pubescens (H. Milne Edwards, 1873), lectotype male (30.5 × 17.5 mm) (MNHN), Mauritius; D, Neoliomera demani Forest &amp; Guinot, 1961, paratype male (23.0 × 13.0 mm) (MNHN), Tahiti; B, E, Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, B, left G1; C–E, distal parts of left G1s. A, C, after Forest &amp; Guinot (1961: fig. 75); D, after Forest &amp; Guinot (1961: fig. 76)." figureDoi="http://doi.org/10.5281/zenodo.4920430" httpUri="https://zenodo.org/record/4920430/files/figure.png" pageId="2" pageNumber="581">Fig. 7B, E</figureCitation>
). G2 short, with short distal segment.
</paragraph>
<paragraph id="CE047E53FF9DFFCBB79B40FCFAADFA36" blockId="2.[151,1437,151,2027]" pageId="2" pageNumber="581">
<emphasis id="FCCFA241FF9DFFCBB79B40FCFEFEFABA" box="[199,318,1274,1299]" inLineHeading="true" pageId="2" pageNumber="581" reason="1">
<emphasis id="FCCFA241FF9DFFCBB79B40FCFEF7FABB" bold="true" box="[199,311,1274,1299]" pageId="2" pageNumber="581">Variation</emphasis>
.
</emphasis>
There is no obvious dimorphism between the sexes. The structures of the carapaces as well as proportions of the ambulatory legs are the same in both sexes. The structure of the anterolateral margin varies slightly, with the lobes sometimes lower in smaller specimens but is, however, always distinct. The female abdomen is ovate, covering about half of the thoracic sternum, with all the somites and telson free. The vulvae are large, swollen and are on the anterior half of thoracic sternite 6. There is no obvious opercular cover (
<figureCitation id="568062D6FF9DFFCBB2544183FAA0FA36" box="[1288,1376,1413,1438]" captionStart="FIGURE 5" captionStartId="8.[151,250,1727,1749]" captionTargetBox="[151,1436,193,1706]" captionTargetId="figure-48@8.[151,1436,193,1706]" captionTargetPageId="8" captionText="FIGURE 5. Neoliomera fragraea n. sp., female (23.9 × 15.0 mm) (ZRC 2000.0483), Hawai‘i. A, dorsal view; B, thoracic sternum showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.4920426" httpUri="https://zenodo.org/record/4920426/files/figure.png" pageId="2" pageNumber="581">Fig. 5B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="86A12DD8FF9DFFCBB79B41A1FBA0F980" pageId="2" pageNumber="581" type="etymology">
<paragraph id="CE047E53FF9DFFCBB79B41A1FD39FA4B" blockId="2.[151,1437,151,2027]" pageId="2" pageNumber="581">
<emphasis id="FCCFA241FF9DFFCBB79B41A1FE8DFA17" box="[199,333,1447,1472]" inLineHeading="true" pageId="2" pageNumber="581" reason="1">
<emphasis id="FCCFA241FF9DFFCBB79B41A1FE86FA68" bold="true" box="[199,326,1447,1472]" pageId="2" pageNumber="581">Etymology</emphasis>
.
</emphasis>
The species is named after the genus name for strawberries (
<emphasis id="FCCFA241FF9DFFCBB36441AFFB63FA68" box="[1080,1187,1449,1472]" italics="true" pageId="2" pageNumber="581">Fragraea</emphasis>
), alluding to its live colouration. The name is used as a noun in apposition.
</paragraph>
<paragraph id="CE047E53FF9DFFCBB79B41EBFBA0F980" blockId="2.[151,1437,151,2027]" pageId="2" pageNumber="581">
<emphasis id="FCCFA241FF9DFFCBB79B41EBFEAFF9AD" box="[199,367,1517,1542]" inLineHeading="true" pageId="2" pageNumber="581" reason="1">
<emphasis id="FCCFA241FF9DFFCBB79B41EBFEA8F9AE" bold="true" box="[199,360,1517,1542]" pageId="2" pageNumber="581">Colour in life</emphasis>
.
</emphasis>
The species is an overall bright red to crimson overall with regularly arranged white spots on the dorsal surface of the carapace (
<figureCitation id="568062D6FF9DFFCBB5404209FD9AF980" box="[540,602,1551,1576]" captionStart="FIGURE 1" captionStartId="3.[151,250,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure-95@3.[151,1436,193,1884]" captionTargetPageId="3" captionText="FIGURE 1. Colours in life. A–C, Neoliomera fragraea n. sp., female (25.3 × 14.3 mm) (NTOU), Taiwan; D, Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam [photograph by G. Paulay]; E, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU) [photograph by G. Paulay]; F, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan [photograph by Wang Kuei-Ching]." figureDoi="http://doi.org/10.5281/zenodo.4920418" httpUri="https://zenodo.org/record/4920418/files/figure.png" pageId="2" pageNumber="581">Fig. 1</figureCitation>
A-E). The fingers are black to greyish-brown.
</paragraph>
</subSubSection>
<subSubSection id="86A12DD8FF9DFFCEB79B4234FDCAFB56" lastPageId="7" lastPageNumber="586" pageId="2" pageNumber="581" type="discussion">
<paragraph id="CE047E53FF9DFFCBB79B4234FAE4F843" blockId="2.[151,1437,151,2027]" pageId="2" pageNumber="581">
<emphasis id="FCCFA241FF9DFFCBB79B4234FEFBF9E2" box="[199,315,1586,1611]" inLineHeading="true" pageId="2" pageNumber="581" reason="1">
<emphasis id="FCCFA241FF9DFFCBB79B4234FEF4F9E3" bold="true" box="[199,308,1586,1611]" pageId="2" pageNumber="581">Remarks</emphasis>
.
</emphasis>
<bibRefCitation id="AA2A03A2FF9DFFCBB6194235FD48F9E3" author="Forest, J. &amp; Guinot, D." box="[325,648,1586,1611]" pageId="2" pageNumber="581" pagination="1 - 195" refId="ref5843" refString="Forest, J. &amp; Guinot, D. (1961) Crustaces Decapodes Brachyoures de Tahiti et des Tuamotu. In: Expedition francaise sur les recifs coralliens de la Nouvelle-Caledonie. Volume preliminaire. Editions de la Fondation Singer Polignac, Paris, 9 - 11, 1 - 195, pls. 1 - 18." type="journal article" year="1961">Forest &amp; Guinot (1961: 80)</bibRefCitation>
clarified the identity of 
<taxonomicName id="09BB05D0FF9DFFCBB4F64235FF18F9C6" authority="(H. Milne Edwards, 1834)" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="581" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="FCCFA241FF9DFFCBB4F64235FB6BF9E3" box="[938,1195,1587,1611]" italics="true" pageId="2" pageNumber="581">Neoliomera pubescens</emphasis>
(H. 
<bibRefCitation id="AA2A03A2FF9DFFCBB3B94234FF10F9C6" author="Milne Edwards, H." pageId="2" pageNumber="581" pagination="1 - 638" refId="ref6436" refString="Milne Edwards, H. (1834 - 1837) Histoire Naturelle des Crustaces Comprenant l'Anatomie, la Physiologie et la Classification de ces Animaux. Vol. 1. - 3. Librairie Encyclopedique de Roret, Paris, xxxv + 468 pp., 531 pp. + 1837: Atlas (1 - 32) + pls. 1 - 42., 1840: 1 - 638." type="book chapter" year="1834">Milne Edwards, 1834</bibRefCitation>
)
</taxonomicName>
and noted that the species is known for certain only from 
<collectingCountry id="B6AC3EC3FF9DFFCBB4344253FC18F9C6" box="[872,984,1621,1646]" name="Mauritius" pageId="2" pageNumber="581">Mauritius</collectingCountry>
, the type locality (see also 
<bibRefCitation id="AA2A03A2FF9DFFCBB2554253FF76F938" author="Serene, R." pageId="2" pageNumber="581" pagination="1 - 400" refId="ref7211" refString="Serene, R. (1984) Crustaces Decapodes Brachyoures de l'Ocean Indien occidental et de la Mer Rouge. Xanthoidea: Xanthidae et Trapeziidae. Addendum Carpiliidae et Menippidae par A. Crosnier. Faune Tropicale, 24, 1 - 400, pls. 1 - 48." type="journal article" year="1984">Serène 1984: 71</bibRefCitation>
). Henri 
<bibRefCitation id="AA2A03A2FF9DFFCBB6794271FD6DF938" author="Milne Edwards, H." box="[293,685,1655,1680]" pageId="2" pageNumber="581" pagination="1 - 638" refId="ref6436" refString="Milne Edwards, H. (1834 - 1837) Histoire Naturelle des Crustaces Comprenant l'Anatomie, la Physiologie et la Classification de ces Animaux. Vol. 1. - 3. Librairie Encyclopedique de Roret, Paris, xxxv + 468 pp., 531 pp. + 1837: Atlas (1 - 32) + pls. 1 - 42., 1840: 1 - 638." type="book chapter" year="1834">Milne Edwards (1834: 384-385)</bibRefCitation>
described 
<taxonomicName id="09BB05D0FF9DFFCBB467427FFBE1F938" authorityName="H. Milne Edwards" authorityYear="1834" box="[827,1057,1656,1680]" class="Malacostraca" family="Xanthidae" genus="Zozymus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="581" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="FCCFA241FF9DFFCBB467427FFBE1F938" box="[827,1057,1656,1680]" italics="true" pageId="2" pageNumber="581">Zozymus pubescens</emphasis>
</taxonomicName>
from 
<collectingCountry id="B6AC3EC3FF9DFFCBB3264271FB2AF938" box="[1146,1258,1655,1680]" name="Mauritius" pageId="2" pageNumber="581">Mauritius</collectingCountry>
, and provided measurements (but no figure) of 
<specimenCount id="D8BDB5DAFF9DFFCBB54E429DFD6EF91B" box="[530,686,1690,1715]" pageId="2" pageNumber="581" type="generic">one specimen</specimenCount>
(sex not stated) although he did not say if he had more material. Alphonse 
<bibRefCitation id="AA2A03A2FF9DFFCBB65542BBFDFCF97E" author="Milne-Edwards, A." box="[265,572,1725,1750]" pageId="2" pageNumber="581" pagination="177 - 308" refId="ref6508" refString="Milne-Edwards, A. (1865) Etudes zoologiques sur les Crustaces recents de la famille des Canceriens. Cancerides, Pirimelides, Carpilides, Premiere Partie. Nouvelles Archives du Museum national d'Histoire naturelle, 1, 177 - 308, pls. 11 - 19." type="journal article" year="1865">Milne-Edwards (1865: 223</bibRefCitation>
, pl. 12 fig. 6) redescribed and figured the species (as a 
<taxonomicName id="09BB05D0FF9DFFCBB3FB42B8FAA0F97E" box="[1191,1376,1725,1750]" class="Malacostraca" family="Xanthidae" genus="Liomera" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="581" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="FCCFA241FF9DFFCBB3FB42B8FAC6F97D" box="[1191,1286,1726,1749]" italics="true" pageId="2" pageNumber="581">Liomera</emphasis>
species
</taxonomicName>
) (see 
<figureCitation id="568062D6FF9DFFCBB7CB42D9FED3F950" box="[151,275,1759,1784]" captionStart="FIGURE 6" captionStartId="9.[151,250,1179,1201]" captionTargetBox="[164,1418,205,1147]" captionTargetId="figure-474@9.[151,1436,193,1178]" captionTargetPageId="9" captionText="FIGURE 6. A–D, Neoliomera pubescens (H. Milne Edwards, 1834), dried lectotype male (30.5 × 17.5 mm) (MNHN-B10037), Mauritius; E, F, Neoliomera demani Forest &amp; Guinot, 1961, paratype male (23.0 × 13.0 mm) (MNHN-B10035), Tahiti; G, Neoliomera demani Forest &amp; Guinot, 1961, holotype male (38.0 × 21.0 mm) (MNHN), Hikueru, French Polynesia. A, dorsal view; B, frontal view of carapace; C, E, dorsal views of carapaces; D, F, G, outer views of left chelae. A, B, after A. Milne- Edwards (1865: pl. 12 fig. 6); C, D, after Forest &amp; Guinot (1961: pl. 3 figs. 1, 2); E–G, after Forest &amp; Guinot (1961: pl. 3 figs. 3–5)." figureDoi="http://doi.org/10.5281/zenodo.4920428" httpUri="https://zenodo.org/record/4920428/files/figure.png" pageId="2" pageNumber="581">Fig. 6A, B</figureCitation>
), presumably based on the same specimen but the sex was again not stated. In describing 
<taxonomicName id="09BB05D0FF9DFFCBB27C42E7FE19F8B3" authority=", Forest &amp; Guinot (1961: 80)" authorityName=", Forest &amp; Guinot" authorityPageNumber="80" authorityYear="1961" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="581" phylum="Arthropoda" rank="species" species="demani">
<emphasis id="FCCFA241FF9DFFCBB27C42E7FA55F950" box="[1312,1429,1760,1784]" italics="true" pageId="2" pageNumber="581">N. demani</emphasis>
, 
<bibRefCitation id="AA2A03A2FF9DFFCBB7CB4305FE19F8B3" author="Forest, J. &amp; Guinot, D." box="[151,473,1794,1819]" pageId="2" pageNumber="581" pagination="1 - 195" refId="ref5843" refString="Forest, J. &amp; Guinot, D. (1961) Crustaces Decapodes Brachyoures de Tahiti et des Tuamotu. In: Expedition francaise sur les recifs coralliens de la Nouvelle-Caledonie. Volume preliminaire. Editions de la Fondation Singer Polignac, Paris, 9 - 11, 1 - 195, pls. 1 - 18." type="journal article" year="1961">Forest &amp; Guinot (1961: 80)</bibRefCitation>
</taxonomicName>
noted that they examined the male type of 
<taxonomicName id="09BB05D0FF9DFFCBB4824305FBB5F8B3" authorityName="H. Milne Edwards" authorityYear="1834" box="[990,1141,1795,1819]" class="Malacostraca" family="Xanthidae" genus="Zozymus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="581" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="FCCFA241FF9DFFCBB4824305FBB5F8B3" box="[990,1141,1795,1819]" italics="true" pageId="2" pageNumber="581">N. pubescens</emphasis>
</taxonomicName>
. In their captions for the figures of the G1 (
<bibRefCitation id="AA2A03A2FF9DFFCBB62D4323FD62F896" author="Forest, J. &amp; Guinot, D." box="[369,674,1829,1854]" pageId="2" pageNumber="581" pagination="1 - 195" refId="ref5843" refString="Forest, J. &amp; Guinot, D. (1961) Crustaces Decapodes Brachyoures de Tahiti et des Tuamotu. In: Expedition francaise sur les recifs coralliens de la Nouvelle-Caledonie. Volume preliminaire. Editions de la Fondation Singer Polignac, Paris, 9 - 11, 1 - 195, pls. 1 - 18." type="journal article" year="1961">Forest &amp; Guinot 1961: 83</bibRefCitation>
) and carapace (
<bibRefCitation id="AA2A03A2FF9DFFCBB4074323FBA2F896" author="Forest, J. &amp; Guinot, D." box="[859,1122,1829,1854]" pageId="2" pageNumber="581" pagination="1 - 195" refId="ref5843" refString="Forest, J. &amp; Guinot, D. (1961) Crustaces Decapodes Brachyoures de Tahiti et des Tuamotu. In: Expedition francaise sur les recifs coralliens de la Nouvelle-Caledonie. Volume preliminaire. Editions de la Fondation Singer Polignac, Paris, 9 - 11, 1 - 195, pls. 1 - 18." type="journal article" year="1961">Forest &amp; Guinot 1961</bibRefCitation>
: pl. 3), they stated that the specimen was a 
<typeStatus id="1100C0F1FF9DFFCBB60D4341FE73F8C8" box="[337,435,1863,1888]" pageId="2" pageNumber="581" type="holotype">holotype</typeStatus>
male. This is not strictly correct. Because H. 
<bibRefCitation id="AA2A03A2FF9DFFCBB4E24341FB7EF8C8" author="Milne Edwards, H." box="[958,1214,1863,1888]" pageId="2" pageNumber="581" pagination="1 - 638" refId="ref6436" refString="Milne Edwards, H. (1834 - 1837) Histoire Naturelle des Crustaces Comprenant l'Anatomie, la Physiologie et la Classification de ces Animaux. Vol. 1. - 3. Librairie Encyclopedique de Roret, Paris, xxxv + 468 pp., 531 pp. + 1837: Atlas (1 - 32) + pls. 1 - 42., 1840: 1 - 638." type="book chapter" year="1834">Milne Edwards (1834)</bibRefCitation>
did not designate a 
<typeStatus id="1100C0F1FF9DFFCBB7CB436CFF39F82B" box="[151,249,1898,1923]" pageId="2" pageNumber="581" type="holotype">holotype</typeStatus>
and did not clearly state how many specimens he had, all his material should be treated as 
<typeStatus id="1100C0F1FF9DFFCBB25C436DFAA6F82B" box="[1280,1382,1899,1923]" pageId="2" pageNumber="581" type="syntype">syntypes</typeStatus>
. The only extant male that 
<bibRefCitation id="AA2A03A2FF9DFFCBB6D54388FD51F80E" author="Forest, J. &amp; Guinot, D." box="[393,657,1933,1958]" pageId="2" pageNumber="581" pagination="1 - 195" refId="ref5843" refString="Forest, J. &amp; Guinot, D. (1961) Crustaces Decapodes Brachyoures de Tahiti et des Tuamotu. In: Expedition francaise sur les recifs coralliens de la Nouvelle-Caledonie. Volume preliminaire. Editions de la Fondation Singer Polignac, Paris, 9 - 11, 1 - 195, pls. 1 - 18." type="journal article" year="1961">Forest &amp; Guinot (1961)</bibRefCitation>
studied should now be treated as the 
<typeStatus id="1100C0F1FF9DFFCBB368438BFB5BF80E" box="[1076,1179,1933,1958]" pageId="2" pageNumber="581" type="lectotype">lectotype</typeStatus>
of 
<taxonomicName id="09BB05D0FF9DFFCBB39C4388FE74F861" authority="H. Milne Edwards, 1834" authorityName="H. Milne Edwards" authorityYear="1834" class="Malacostraca" family="Xanthidae" genus="Zozymus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="581" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="FCCFA241FF9DFFCBB39C4388FA5CF80D" box="[1216,1436,1933,1957]" italics="true" pageId="2" pageNumber="581">Zozymus pubescens</emphasis>
H. 
<bibRefCitation id="AA2A03A2FF9DFFCBB7E743B6FE74F861" author="Milne Edwards, H." box="[187,436,1968,1993]" pageId="2" pageNumber="581" pagination="1 - 638" refId="ref6436" refString="Milne Edwards, H. (1834 - 1837) Histoire Naturelle des Crustaces Comprenant l'Anatomie, la Physiologie et la Classification de ces Animaux. Vol. 1. - 3. Librairie Encyclopedique de Roret, Paris, xxxv + 468 pp., 531 pp. + 1837: Atlas (1 - 32) + pls. 1 - 42., 1840: 1 - 638." type="book chapter" year="1834">Milne Edwards, 1834</bibRefCitation>
</taxonomicName>
. 
<taxonomicName id="09BB05D0FF9DFFCBB69E43B6FD02F860" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" box="[450,706,1968,1992]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="581" phylum="Arthropoda" rank="species" sensu="stricto" species="pubescens">
<emphasis id="FCCFA241FF9DFFCBB69E43B6FD02F860" box="[450,706,1968,1992]" italics="true" pageId="2" pageNumber="581">Neoliomera pubescens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E7FC1F3AFF9DFFCBB59743B7FCC8F860" box="[715,776,1969,1992]" pageId="2" pageNumber="581" sensu="stricto">s. str.</taxonomicNameLabel>
has also been listed and discussed by Michel (1964: 28), 
<bibRefCitation id="AA2A03A2FF9DFFCBB7CB43D4FEB1F843" author="Guinot, D." box="[151,369,2002,2027]" pageId="2" pageNumber="581" pagination="237 - 352" refId="ref5972" refString="Guinot, D. (1967) La faune carcinologique (Crustacea Brachyura) de l'Ocean Indien occidental et de la Mer Rouge. Catalogue, remarques biogeographiques et bibliographie. In: Reunion de Specialistes C. S. A. sur les Crustaces, Zanzibar 1964. Memoires I. F. A. N., (77), 1966 (1967), 237 - 352." type="journal article" year="1967">Guinot (1967: 267)</bibRefCitation>
, 
<bibRefCitation id="AA2A03A2FF9DFFCBB62143D4FD88F843" author="Serene, R." box="[381,584,2002,2027]" pageId="2" pageNumber="581" pagination="33 - 112" refId="ref7101" refString="Serene, R. (1968) The Brachyura of the Indo-West Pacific Region. In: Prodromus for a Check List of the (non-planctonic) Marine Fauna of South East Asia. Singapore National Academy of Sciences, Special Publication No. 1 (Fauna IIICc 3), pp. 33 - 112." type="journal article" year="1968">Serène (1968: 73)</bibRefCitation>
, 
<bibRefCitation id="AA2A03A2FF9DFFCBB50843D4FCD5F843" author="Serene, R." box="[596,789,2002,2027]" pageId="2" pageNumber="581" pagination="1 - 400" refId="ref7211" refString="Serene, R. (1984) Crustaces Decapodes Brachyoures de l'Ocean Indien occidental et de la Mer Rouge. Xanthoidea: Xanthidae et Trapeziidae. Addendum Carpiliidae et Menippidae par A. Crosnier. Faune Tropicale, 24, 1 - 400, pls. 1 - 48." type="journal article" year="1984">Serène (1984: 71</bibRefCitation>
, text-fig. 30, pl. 8A) and 
<bibRefCitation id="AA2A03A2FF9DFFCBB36D43D5FAE0F843" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[1073,1312,2002,2027]" pageId="2" pageNumber="581" pagination="1 - 286" refId="ref6649" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, 17 (Supplement), 1 - 286." type="journal article" year="2008">
Ng 
<emphasis id="FCCFA241FF9DFFCBB30643D5FB53F843" box="[1114,1171,2003,2027]" italics="true" pageId="2" pageNumber="581">et al.</emphasis>
(2008: 201)
</bibRefCitation>
.
</paragraph>
<caption id="9AC42EDBFF9CFFCAB7CB4377FC64F84B" ID-DOI="http://doi.org/10.5281/zenodo.4920418" ID-Zenodo-Dep="4920418" httpUri="https://zenodo.org/record/4920418/files/figure.png" pageId="3" pageNumber="582" startId="3.[151,250,1905,1927]" targetBox="[151,1436,193,1884]" targetPageId="3">
<paragraph id="CE047E53FF9CFFCAB7CB4377FC64F84B" blockId="3.[151,1436,1905,2019]" pageId="3" pageNumber="582">
<emphasis id="FCCFA241FF9CFFCAB7CB4377FECFF82F" bold="true" box="[151,271,1905,1927]" pageId="3" pageNumber="582">FIGURE 1</emphasis>
. Colours in life. A–C, 
<taxonomicName id="09BB05D0FF9CFFCAB5584374FD1DF82F" authority="P.-H &amp; Ng, 2014" authorityName="P.-H &amp; Ng" authorityYear="2014" box="[516,733,1906,1927]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="582" phylum="Arthropoda" rank="species" species="fragraea" status="sp. nov.">
<emphasis id="FCCFA241FF9CFFCAB5584374FD1DF82F" box="[516,733,1906,1927]" italics="true" pageId="3" pageNumber="582">Neoliomera fragraea</emphasis>
</taxonomicName>
<emphasis id="FCCFA241FF9CFFCAB5BA4377FCE2F82F" bold="true" box="[742,802,1905,1927]" pageId="3" pageNumber="582">
<taxonomicNameLabel id="E7FC1F3AFF9CFFCAB5BA4377FCE2F82F" box="[742,802,1905,1927]" pageId="3" pageNumber="582" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female (25.3 × 14.3 mm) (NTOU), Taiwan; D, 
<taxonomicName id="09BB05D0FF9CFFCAB2784374FF30F80D" authority="P.-H &amp; Ng, 2014" authorityName="P.-H &amp; Ng" authorityYear="2014" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="582" phylum="Arthropoda" rank="species" species="fragraea" status="sp. nov.">
<emphasis id="FCCFA241FF9CFFCAB2784374FF30F80D" italics="true" pageId="3" pageNumber="582">Neoliomera fragraea</emphasis>
</taxonomicName>
<emphasis id="FCCFA241FF9CFFCAB7AA4396FEEFF80E" bold="true" box="[246,303,1936,1958]" pageId="3" pageNumber="582">
<taxonomicNameLabel id="E7FC1F3AFF9CFFCAB7AA4396FEEFF80E" box="[246,303,1936,1958]" pageId="3" pageNumber="582" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam [photograph by G. Paulay]; E, 
<taxonomicName id="09BB05D0FF9CFFCAB3994396FA5CF80D" authority="P.-H &amp; Ng, 2014" authorityName="P.-H &amp; Ng" authorityYear="2014" box="[1221,1436,1936,1957]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="582" phylum="Arthropoda" rank="species" species="fragraea" status="sp. nov.">
<emphasis id="FCCFA241FF9CFFCAB3994396FA5CF80D" box="[1221,1436,1936,1957]" italics="true" pageId="3" pageNumber="582">Neoliomera fragraea</emphasis>
</taxonomicName>
<emphasis id="FCCFA241FF9CFFCAB7CB43A8FF12F86C" bold="true" box="[151,210,1966,1988]" pageId="3" pageNumber="582">
<taxonomicNameLabel id="E7FC1F3AFF9CFFCAB7CB43A8FF12F86C" box="[151,210,1966,1988]" pageId="3" pageNumber="582" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, paratype male (18.2 × 10.8 mm) (NTOU) [photograph by G. Paulay]; F, 
<taxonomicName id="09BB05D0FF9CFFCAB48B43A9FA58F86C" authority="Forest &amp; Guinot, 1961" authorityName=", Forest &amp; Guinot" authorityYear="1961" box="[983,1432,1966,1988]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="582" phylum="Arthropoda" rank="species" species="demani">
<emphasis id="FCCFA241FF9CFFCAB48B43A9FB62F86C" box="[983,1186,1967,1988]" italics="true" pageId="3" pageNumber="582">Neoliomera demani</emphasis>
<bibRefCitation id="AA2A03A2FF9CFFCAB3F643A9FA58F86C" author="Forest, J. &amp; Guinot, D." box="[1194,1432,1966,1988]" pageId="3" pageNumber="582" pagination="1 - 195" refId="ref5843" refString="Forest, J. &amp; Guinot, D. (1961) Crustaces Decapodes Brachyoures de Tahiti et des Tuamotu. In: Expedition francaise sur les recifs coralliens de la Nouvelle-Caledonie. Volume preliminaire. Editions de la Fondation Singer Polignac, Paris, 9 - 11, 1 - 195, pls. 1 - 18." type="journal article" year="1961">Forest &amp; Guinot, 1961</bibRefCitation>
</taxonomicName>
, male (35.7 × 18.7 mm) (NTOU), Taiwan [photograph by Wang Kuei-Ching].
</paragraph>
</caption>
<caption id="9AC42EDBFF9BFFCDB7CB43A7FDA9F87D" ID-DOI="http://doi.org/10.5281/zenodo.4920420" ID-Zenodo-Dep="4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="4" pageNumber="583" startId="4.[151,250,1953,1975]" targetBox="[241,1345,193,1932]" targetPageId="4">
<paragraph id="CE047E53FF9BFFCDB7CB43A7FDA9F87D" blockId="4.[151,1436,1953,2006]" pageId="4" pageNumber="583">
<emphasis id="FCCFA241FF9BFFCDB7CB43A7FECDF81F" bold="true" box="[151,269,1953,1975]" pageId="4" pageNumber="583">FIGURE 2</emphasis>
. 
<taxonomicName id="09BB05D0FF9BFFCDB64643A7FE31F81E" authority="P.-H &amp; Ng, 2014" authorityName="P.-H &amp; Ng" authorityYear="2014" box="[282,497,1953,1974]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="583" phylum="Arthropoda" rank="species" species="fragraea" status="sp. nov.">
<emphasis id="FCCFA241FF9BFFCDB64643A7FE31F81E" box="[282,497,1953,1974]" italics="true" pageId="4" pageNumber="583">Neoliomera fragraea</emphasis>
</taxonomicName>
<emphasis id="FCCFA241FF9BFFCDB6A443A7FDF1F81F" bold="true" box="[504,561,1953,1975]" pageId="4" pageNumber="583">
<taxonomicNameLabel id="E7FC1F3AFF9BFFCDB6A443A7FDF1F81F" box="[504,561,1953,1975]" pageId="4" pageNumber="583" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace.
</paragraph>
</caption>
<caption id="9AC42EDBFF9AFFCCB7CB43A7FB40F85C" ID-DOI="http://doi.org/10.5281/zenodo.4920422" ID-Zenodo-Dep="4920422" httpUri="https://zenodo.org/record/4920422/files/figure.png" pageId="5" pageNumber="584" startId="5.[151,250,1953,1975]" targetBox="[241,1345,193,1932]" targetPageId="5">
<paragraph id="CE047E53FF9AFFCCB7CB43A7FB40F85C" blockId="5.[151,1436,1953,2036]" pageId="5" pageNumber="584">
<emphasis id="FCCFA241FF9AFFCCB7CB43A7FED1F81F" bold="true" box="[151,273,1953,1975]" pageId="5" pageNumber="584">FIGURE 3</emphasis>
. A, 
<taxonomicName id="09BB05D0FF9AFFCCB61943A7FDE1F81E" authority="P.-H &amp; Ng, 2014" authorityName="P.-H &amp; Ng" authorityYear="2014" box="[325,545,1953,1974]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="584" phylum="Arthropoda" rank="species" species="fragraea" status="sp. nov.">
<emphasis id="FCCFA241FF9AFFCCB61943A7FDE1F81E" box="[325,545,1953,1974]" italics="true" pageId="5" pageNumber="584">Neoliomera fragraea</emphasis>
</taxonomicName>
<emphasis id="FCCFA241FF9AFFCCB57743A7FDA9F81F" bold="true" box="[555,617,1953,1975]" pageId="5" pageNumber="584">
<taxonomicNameLabel id="E7FC1F3AFF9AFFCCB57743A7FDA9F81F" box="[555,617,1953,1975]" pageId="5" pageNumber="584" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, paratype female (11.9 × 7.5 mm) (ZRC 2014.0207), Guam; B, 
<taxonomicName id="09BB05D0FF9AFFCCB27843A7FF30F87D" authority="P.-H &amp; Ng, 2014" authorityName="P.-H &amp; Ng" authorityYear="2014" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="584" phylum="Arthropoda" rank="species" species="fragraea" status="sp. nov.">
<emphasis id="FCCFA241FF9AFFCCB27843A7FF30F87D" italics="true" pageId="5" pageNumber="584">Neoliomera fragraea</emphasis>
</taxonomicName>
<emphasis id="FCCFA241FF9AFFCCB7AA43C6FEEFF87E" bold="true" box="[246,303,1984,2006]" pageId="5" pageNumber="584">
<taxonomicNameLabel id="E7FC1F3AFF9AFFCCB7AA43C6FEEFF87E" box="[246,303,1984,2006]" pageId="5" pageNumber="584" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, paratype male (18.2 × 10.8 mm) (NTOU), Guam; C, 
<taxonomicName id="09BB05D0FF9AFFCCB40443C6FAD0F87E" authority="Forest &amp; Guinot, 1961" authorityName=", Forest &amp; Guinot" authorityYear="1961" box="[856,1296,1984,2006]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="584" phylum="Arthropoda" rank="species" species="demani">
<emphasis id="FCCFA241FF9AFFCCB40443C6FBE1F87D" box="[856,1057,1984,2005]" italics="true" pageId="5" pageNumber="584">Neoliomera demani</emphasis>
<bibRefCitation id="AA2A03A2FF9AFFCCB37B43C6FAD0F87E" author="Forest, J. &amp; Guinot, D." box="[1063,1296,1984,2006]" pageId="5" pageNumber="584" pagination="1 - 195" refId="ref5843" refString="Forest, J. &amp; Guinot, D. (1961) Crustaces Decapodes Brachyoures de Tahiti et des Tuamotu. In: Expedition francaise sur les recifs coralliens de la Nouvelle-Caledonie. Volume preliminaire. Editions de la Fondation Singer Polignac, Paris, 9 - 11, 1 - 195, pls. 1 - 18." type="journal article" year="1961">Forest &amp; Guinot, 1961</bibRefCitation>
</taxonomicName>
, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, dorsal overall views; B, anterior thoracic sternum and abdomen.
</paragraph>
</caption>
<caption id="9AC42EDBFF99FFCFB7CB40F5FCC1FAEE" ID-DOI="http://doi.org/10.5281/zenodo.4920424" ID-Zenodo-Dep="4920424" httpUri="https://zenodo.org/record/4920424/files/figure.png" pageId="6" pageNumber="585" startId="6.[151,250,1267,1289]" targetBox="[164,1428,193,1248]" targetPageId="6">
<paragraph id="CE047E53FF99FFCFB7CB40F5FCC1FAEE" blockId="6.[151,1436,1267,1350]" pageId="6" pageNumber="585">
<emphasis id="FCCFA241FF99FFCFB7CB40F5FED1FAA1" bold="true" box="[151,273,1267,1289]" pageId="6" pageNumber="585">FIGURE 4</emphasis>
. A, B, E, 
<taxonomicName id="09BB05D0FF99FFCFB6D540F5FDA5FAA0" authority="P.-H &amp; Ng, 2014" authorityName="P.-H &amp; Ng" authorityYear="2014" box="[393,613,1267,1288]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="fragraea" status="sp. nov.">
<emphasis id="FCCFA241FF99FFCFB6D540F5FDA5FAA0" box="[393,613,1267,1288]" italics="true" pageId="6" pageNumber="585">Neoliomera fragraea</emphasis>
</taxonomicName>
<emphasis id="FCCFA241FF99FFCFB53340F5FD6DFAA1" bold="true" box="[623,685,1267,1289]" pageId="6" pageNumber="585">
<taxonomicNameLabel id="E7FC1F3AFF99FFCFB53340F5FD6DFAA1" box="[623,685,1267,1289]" pageId="6" pageNumber="585" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam; C, D, F, 
<taxonomicName id="09BB05D0FF99FFCFB7CB4114FD90FA80" authority="Forest &amp; Guinot, 1961" authorityName=", Forest &amp; Guinot" authorityYear="1961" box="[151,592,1298,1320]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="demani">
<emphasis id="FCCFA241FF99FFCFB7CB4114FEA0FA8F" box="[151,352,1298,1319]" italics="true" pageId="6" pageNumber="585">Neoliomera demani</emphasis>
<bibRefCitation id="AA2A03A2FF99FFCFB63B4114FD90FA80" author="Forest, J. &amp; Guinot, D." box="[359,592,1298,1320]" pageId="6" pageNumber="585" pagination="1 - 195" refId="ref5843" refString="Forest, J. &amp; Guinot, D. (1961) Crustaces Decapodes Brachyoures de Tahiti et des Tuamotu. In: Expedition francaise sur les recifs coralliens de la Nouvelle-Caledonie. Volume preliminaire. Editions de la Fondation Singer Polignac, Paris, 9 - 11, 1 - 195, pls. 1 - 18." type="journal article" year="1961">Forest &amp; Guinot, 1961</bibRefCitation>
</taxonomicName>
, male (35.7 × 18.7 mm) (NTOU), Taiwan. A, C, outer views of right chelae; B, D, inner views of right chelae; E, F, right fourth ambulatory leg.
</paragraph>
</caption>
<paragraph id="CE047E53FF99FFCFB79B4172FA8AF9CC" blockId="6.[151,1437,1395,2032]" pageId="6" pageNumber="585">
<bibRefCitation id="AA2A03A2FF99FFCFB79B4172FE10FA24" author="Forest, J. &amp; Guinot, D." box="[199,464,1395,1420]" pageId="6" pageNumber="585" pagination="1 - 195" refId="ref5843" refString="Forest, J. &amp; Guinot, D. (1961) Crustaces Decapodes Brachyoures de Tahiti et des Tuamotu. In: Expedition francaise sur les recifs coralliens de la Nouvelle-Caledonie. Volume preliminaire. Editions de la Fondation Singer Polignac, Paris, 9 - 11, 1 - 195, pls. 1 - 18." type="journal article" year="1961">Forest &amp; Guinot (1961)</bibRefCitation>
suggested that 
<taxonomicName id="09BB05D0FF99FFCFB5224172FC7AFA24" authority="Odhner, 1925" authorityName="Odhner" authorityYear="1925" box="[638,954,1395,1420]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="FCCFA241FF99FFCFB5224172FCD9FA23" box="[638,793,1395,1419]" italics="true" pageId="6" pageNumber="585">N. intermedia</emphasis>
<bibRefCitation id="AA2A03A2FF99FFCFB4434175FC7AFA24" author="Odhner, T." box="[799,954,1395,1420]" pageId="6" pageNumber="585" pagination="1 - 92" refId="ref6705" refString="Odhner, T. (1925) Monographierte Gattungen der Krabbenfamilie Xanthidae. I. Goteborgs Kungliga Vetenskaps-och Vitterhetssamhalles Handlingar, Series 4, 29 (1), 1 - 92, figs. 1 - 7, pls. 1 - 5." type="journal article" year="1925">Odhner, 1925</bibRefCitation>
</taxonomicName>
, described from the 
<collectingCountry id="B6AC3EC3FF99FFCFB3FC4175FAE1FA24" box="[1184,1313,1395,1420]" name="Philippines" pageId="6" pageNumber="585">Philippines</collectingCountry>
, is a junior synonym of 
<taxonomicName id="09BB05D0FF99FFCFB67B419EFE7DFA07" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" box="[295,445,1431,1455]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="FCCFA241FF99FFCFB67B419EFE7DFA07" box="[295,445,1431,1455]" italics="true" pageId="6" pageNumber="585">N. pubescens</emphasis>
</taxonomicName>
, and included records of this species from the 
<collectingCountry id="B6AC3EC3FF99FFCFB4804191FB9AFA18" box="[988,1114,1431,1456]" name="Philippines" pageId="6" pageNumber="585">Philippines</collectingCountry>
and 
<collectingCountry id="B6AC3EC3FF99FFCFB3C9419EFB15FA07" box="[1173,1237,1432,1455]" name="Japan" pageId="6" pageNumber="585">Japan</collectingCountry>
as 
<taxonomicName id="09BB05D0FF99FFCFB3A3419EFA55FA07" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" box="[1279,1429,1431,1455]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="FCCFA241FF99FFCFB3A3419EFA55FA07" box="[1279,1429,1431,1455]" italics="true" pageId="6" pageNumber="585">N. pubescens</emphasis>
</taxonomicName>
. 
<bibRefCitation id="AA2A03A2FF99FFCFB7CB41BDFEB1FA7C" author="Guinot, D." box="[151,369,1467,1492]" pageId="6" pageNumber="585" pagination="223 - 267" refId="ref6044" refString="Guinot, D. (1969) Sur divers Xanthidae, notamment sur Actaea de Haan et Paractaea gen. nov. (Crustacea Decapoda Brachyura). Cahiers du Pacifique, 13, 223 - 267, pls. 17 - 19." type="journal article" year="1969">Guinot (1969: 232</bibRefCitation>
, fig. 14), however, examined the 
<typeStatus id="1100C0F1FF99FFCFB5A341BAFCF2FA7C" box="[767,818,1468,1492]" pageId="6" pageNumber="585">type</typeStatus>
, figured the G1 and noted it was a separate species. 
<taxonomicName id="09BB05D0FF99FFCFB7CB41D9FE5BFA5F" authorityName="Odhner" authorityYear="1925" box="[151,411,1503,1527]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="FCCFA241FF99FFCFB7CB41D9FE5BFA5F" box="[151,411,1503,1527]" italics="true" pageId="6" pageNumber="585">Neoliomera intermedia</emphasis>
</taxonomicName>
is very distinct from 
<taxonomicName id="09BB05D0FF99FFCFB5D741E6FCDFFA5F" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" box="[651,799,1503,1527]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="FCCFA241FF99FFCFB5D741E6FCDFFA5F" box="[651,799,1503,1527]" italics="true" pageId="6" pageNumber="585">N. pubescens</emphasis>
</taxonomicName>
in the form of the carapace (relatively less wide) and the G1 is more elongated, with a differently structured distal part and setation pattern (
<bibRefCitation id="AA2A03A2FF99FFCFB3624205FB12F9B4" author="Guinot, D." box="[1086,1234,1539,1564]" pageId="6" pageNumber="585" pagination="223 - 267" refId="ref6044" refString="Guinot, D. (1969) Sur divers Xanthidae, notamment sur Actaea de Haan et Paractaea gen. nov. (Crustacea Decapoda Brachyura). Cahiers du Pacifique, 13, 223 - 267, pls. 17 - 19." type="journal article" year="1969">Guinot 1969</bibRefCitation>
: fig. 14). The live colours of 
<taxonomicName id="09BB05D0FF99FFCFB653422EFE69F997" authorityName="Odhner" authorityYear="1925" box="[271,425,1575,1599]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="FCCFA241FF99FFCFB653422EFE69F997" box="[271,425,1575,1599]" italics="true" pageId="6" pageNumber="585">N. intermedia</emphasis>
</taxonomicName>
, as figured by 
<bibRefCitation id="AA2A03A2FF99FFCFB50E4221FD1DF9E8" author="Sakai, T." box="[594,733,1575,1600]" pageId="6" pageNumber="585" pagination="1 - 251" refId="ref7044" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Vol. 1. - 3. Kodansha, Tokyo, English text: xxxix + 773 pp., Japanese text: 461 pp., plates: 16 pp., pls. 1 - 251. (plates)" type="book chapter" year="1976">Sakai (1976</bibRefCitation>
: pl. 142 fig. 3) are also quite different, being a uniform orange on the dorsal surfaces, without any white spots like in 
<taxonomicName id="09BB05D0FF99FFCFB5A4424AFC4BF9CB" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" box="[760,907,1611,1635]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="FCCFA241FF99FFCFB5A4424AFC4BF9CB" box="[760,907,1611,1635]" italics="true" pageId="6" pageNumber="585">N. pubescens</emphasis>
</taxonomicName>
(see also 
<bibRefCitation id="AA2A03A2FF99FFCFB4A6424DFB7BF9CC" author="Odhner, T." box="[1018,1211,1611,1636]" pageId="6" pageNumber="585" pagination="1 - 92" refId="ref6705" refString="Odhner, T. (1925) Monographierte Gattungen der Krabbenfamilie Xanthidae. I. Goteborgs Kungliga Vetenskaps-och Vitterhetssamhalles Handlingar, Series 4, 29 (1), 1 - 92, figs. 1 - 7, pls. 1 - 5." type="journal article" year="1925">Odhner 1925: 29</bibRefCitation>
, pl. 2 fig. 8).
</paragraph>
<paragraph id="CE047E53FF99FFCFB79B4276FC3AF858" blockId="6.[151,1437,1395,2032]" pageId="6" pageNumber="585">
Many early records of 
<taxonomicName id="09BB05D0FF99FFCFB6874276FDB4F92F" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" box="[475,628,1647,1671]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="FCCFA241FF99FFCFB6874276FDB4F92F" box="[475,628,1647,1671]" italics="true" pageId="6" pageNumber="585">N. pubescens</emphasis>
</taxonomicName>
have since been referred to 
<taxonomicName id="09BB05D0FF99FFCFB48C4276FAA2F920" authority="Forest &amp; Guinot, 1961" authorityName=", Forest &amp; Guinot" authorityYear="1961" box="[976,1378,1647,1672]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="demani">
<emphasis id="FCCFA241FF99FFCFB48C4276FB88F92F" box="[976,1096,1647,1671]" italics="true" pageId="6" pageNumber="585">N. demani</emphasis>
<bibRefCitation id="AA2A03A2FF99FFCFB3084276FAA2F920" author="Forest, J. &amp; Guinot, D." box="[1108,1378,1647,1672]" pageId="6" pageNumber="585" pagination="1 - 195" refId="ref5843" refString="Forest, J. &amp; Guinot, D. (1961) Crustaces Decapodes Brachyoures de Tahiti et des Tuamotu. In: Expedition francaise sur les recifs coralliens de la Nouvelle-Caledonie. Volume preliminaire. Editions de la Fondation Singer Polignac, Paris, 9 - 11, 1 - 195, pls. 1 - 18." type="journal article" year="1961">Forest &amp; Guinot, 1961</bibRefCitation>
</taxonomicName>
(see discussion below). We have on hand a series of specimens from 
<collectingCountry id="B6AC3EC3FF99FFCFB42B4295FC09F904" box="[887,969,1683,1708]" name="Taiwan" pageId="6" pageNumber="585">Taiwan</collectingCountry>
and 
<collectingCountry id="B6AC3EC3FF99FFCFB35D4295FB86F904" box="[1025,1094,1683,1708]" name="Guam" pageId="6" pageNumber="585">Guam</collectingCountry>
that superficially resemble 
<taxonomicName id="09BB05D0FF99FFCFB2DE4292FECAF967" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="FCCFA241FF99FFCFB2DE4292FECAF967" italics="true" pageId="6" pageNumber="585">N. pubescens</emphasis>
</taxonomicName>
but can be separated by the form of the frontal and anterolateral margins, chelea and fingers, merus of the ambulatory legs and structure of the G1s (
<tableCitation id="83394BE8FF99FFCFB5CD42DDFD25F95C" box="[657,741,1755,1780]" captionStart="TABLE 1" captionStartId="7.[151,239,152,175]" captionTargetPageId="7" captionText="TABLE 1. Differences between Neoliomera pubescens s. str., N. demani and N. fragaea n. sp." httpUri="http://table.plazi.org/id/9AC42EDBFF98FFCEB7CB449EFB46FF07" pageId="6" pageNumber="585" tableUuid="9AC42EDBFF98FFCEB7CB449EFB46FF07">Table 1</tableCitation>
). As such, they are here referred to a new species, 
<taxonomicName id="09BB05D0FF99FFCFB24642DAFA5CF95B" authority="P.-H &amp; Ng, 2014" authorityName="P.-H &amp; Ng" authorityYear="2014" box="[1306,1436,1755,1779]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="fragraea" status="sp. nov.">
<emphasis id="FCCFA241FF99FFCFB24642DAFA5CF95B" box="[1306,1436,1755,1779]" italics="true" pageId="6" pageNumber="585">N. fragraea</emphasis>
</taxonomicName>
<emphasis id="FCCFA241FF99FFCFB7CB42F9FF16F8B0" bold="true" box="[151,214,1791,1816]" pageId="6" pageNumber="585">
<taxonomicNameLabel id="E7FC1F3AFF99FFCFB7CB42F9FF16F8B0" box="[151,214,1791,1816]" pageId="6" pageNumber="585" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
The records of “ 
<taxonomicName id="09BB05D0FF99FFCFB6CA4306FDEBF8BF" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" box="[406,555,1791,1815]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="FCCFA241FF99FFCFB6CA4306FDEBF8BF" box="[406,555,1791,1815]" italics="true" pageId="6" pageNumber="585">N. pubescens</emphasis>
</taxonomicName>
” from the Hawaiian Islands by 
<bibRefCitation id="AA2A03A2FF99FFCFB4C842F9FB41F8B0" author="Rathbun, M. J." box="[916,1153,1791,1816]" pageId="6" pageNumber="585" pagination="827 - 930" refId="ref6950" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23 (3), 827 - 930, pls. 1 - 24. [1903]" type="journal article" year="1906">Rathbun (1906: 844)</bibRefCitation>
, 
<bibRefCitation id="AA2A03A2FF99FFCFB3D142F9FA58F8B0" author="Edmondson, C. H." box="[1165,1432,1791,1816]" pageId="6" pageNumber="585" pagination="1 - 381" refId="ref5765" refString="Edmondson, C. H. (1946) Reef and Shore Fauna of Hawaii. Special Publication of the Bernice P. Bishop Museum, 22, i-iii, 1 - 381, figs. 1 - 223." type="journal article" year="1946">Edmondson (1946: 292</bibRefCitation>
, fig. 178b), 
<bibRefCitation id="AA2A03A2FF99FFCFB6404325FDEFF894" author="Edmondson, C. H." box="[284,559,1827,1852]" pageId="6" pageNumber="585" pagination="215 - 309" refId="ref5805" refString="Edmondson, C. H. (1962) Xanthidae of Hawaii. Occasional Papers of the Bernice P. Bishop Museum, 22 (13), 215 - 309, figs. 1 - 34." type="journal article" year="1962">Edmondson (1962: 252</bibRefCitation>
, fig. 9d) and 
<bibRefCitation id="AA2A03A2FF99FFCFB5894325FC61F894" author="Sakai, T." box="[725,929,1827,1852]" pageId="6" pageNumber="585" pagination="1 - 251" refId="ref7044" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Vol. 1. - 3. Kodansha, Tokyo, English text: xxxix + 773 pp., Japanese text: 461 pp., plates: 16 pp., pls. 1 - 251. (plates)" type="book chapter" year="1976">Sakai (1976: 399</bibRefCitation>
, part) are probably 
<taxonomicName id="09BB05D0FF99FFCFB3D14322FAD4F893" authority="P.-H &amp; Ng, 2014" authorityName="P.-H &amp; Ng" authorityYear="2014" box="[1165,1300,1827,1851]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="fragraea" status="sp. nov.">
<emphasis id="FCCFA241FF99FFCFB3D14322FAD4F893" box="[1165,1300,1827,1851]" italics="true" pageId="6" pageNumber="585">N. fragraea</emphasis>
</taxonomicName>
<emphasis id="FCCFA241FF99FFCFB27C4325FAA3F894" bold="true" box="[1312,1379,1827,1852]" pageId="6" pageNumber="585">
<taxonomicNameLabel id="E7FC1F3AFF99FFCFB27C4325FAA3F894" box="[1312,1379,1827,1852]" pageId="6" pageNumber="585" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(see discussion for 
<taxonomicName id="09BB05D0FF99FFCFB662434EFE02F8F7" authority="P.-H &amp; Ng, 2014" authorityName="P.-H &amp; Ng" authorityYear="2014" box="[318,450,1863,1887]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="fragraea" status="sp. nov.">
<emphasis id="FCCFA241FF99FFCFB662434EFE02F8F7" box="[318,450,1863,1887]" italics="true" pageId="6" pageNumber="585">N. fragraea</emphasis>
</taxonomicName>
<emphasis id="FCCFA241FF99FFCFB6974341FDCCF8C8" bold="true" box="[459,524,1863,1888]" pageId="6" pageNumber="585">
<taxonomicNameLabel id="E7FC1F3AFF99FFCFB6974341FDCCF8C8" box="[459,524,1863,1888]" pageId="6" pageNumber="585" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
), although it is possible their material may be mixed because 
<taxonomicName id="09BB05D0FF99FFCFB389434EFA8AF8F7" authorityName=", Forest &amp; Guinot" authorityYear="1961" box="[1237,1354,1863,1887]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="demani">
<emphasis id="FCCFA241FF99FFCFB389434EFA8AF8F7" box="[1237,1354,1863,1887]" italics="true" pageId="6" pageNumber="585">N. demani</emphasis>
</taxonomicName>
is also known from the Hawaiian Islands. 
<bibRefCitation id="AA2A03A2FF99FFCFB571436DFD36F82C" author="Castro, P." box="[557,758,1899,1924]" pageId="6" pageNumber="585" pagination="1 - 154" refId="ref5729" refString="Castro, P. (2011) Catalog of the anomuran and brachyuran crabs (Crustacea: Decapoda: Anomura, Brachyura) of the Hawaiian Islands. Zootaxa, 2947, 1 - 154." type="journal article" year="2011">Castro (2011: 99)</bibRefCitation>
had already commented that these Hawaiian records of 
<taxonomicName id="09BB05D0FF99FFCFB2DE436AFECAF80F" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="FCCFA241FF99FFCFB2DE436AFECAF80F" italics="true" pageId="6" pageNumber="585">N. pubescens</emphasis>
</taxonomicName>
should be referred to a new species. The relatively large Hawaiian female specimen on hand (ZRC 2000.0483) (
<figureCitation id="568062D6FF99FFCFB67A43B5FEA8F864" box="[294,360,1971,1996]" captionStart="FIGURE 5" captionStartId="8.[151,250,1727,1749]" captionTargetBox="[151,1436,193,1706]" captionTargetId="figure-48@8.[151,1436,193,1706]" captionTargetPageId="8" captionText="FIGURE 5. Neoliomera fragraea n. sp., female (23.9 × 15.0 mm) (ZRC 2000.0483), Hawai‘i. A, dorsal view; B, thoracic sternum showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.4920426" httpUri="https://zenodo.org/record/4920426/files/figure.png" pageId="6" pageNumber="585">Fig. 5</figureCitation>
) agrees with the 
<collectingCountry id="B6AC3EC3FF99FFCFB57B43B5FDACF864" box="[551,620,1971,1996]" name="Guam" pageId="6" pageNumber="585">Guam</collectingCountry>
and 
<collectingCountry id="B6AC3EC3FF99FFCFB5FD43B5FD33F864" box="[673,755,1971,1996]" name="Taiwan" pageId="6" pageNumber="585">Taiwan</collectingCountry>
material, including the colour when alive, carapace structure and leg proportions, and we conclude it is conspecific with 
<taxonomicName id="09BB05D0FF99FFCFB46D43DEFC74F847" authority="P.-H &amp; Ng, 2014" authorityName="P.-H &amp; Ng" authorityYear="2014" box="[817,948,2007,2031]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="585" phylum="Arthropoda" rank="species" species="fragraea" status="sp. nov.">
<emphasis id="FCCFA241FF99FFCFB46D43DEFC74F847" box="[817,948,2007,2031]" italics="true" pageId="6" pageNumber="585">N. fragraea</emphasis>
</taxonomicName>
<emphasis id="FCCFA241FF99FFCFB4E743D1FC3AF858" bold="true" box="[955,1018,2007,2032]" pageId="6" pageNumber="585">
<taxonomicNameLabel id="E7FC1F3AFF99FFCFB4E743D1FC3AF858" box="[955,1018,2007,2032]" pageId="6" pageNumber="585" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
<caption id="9AC42EDBFF98FFCEB7CB449EFB46FF07" ID-Table-UUID="9AC42EDBFF98FFCEB7CB449EFB46FF07" box="[151,1158,151,175]" httpUri="http://table.plazi.org/id/9AC42EDBFF98FFCEB7CB449EFB46FF07" pageId="7" pageNumber="586" startId="7.[151,239,152,175]" targetBox="[159,1423,203,932]" targetIsTable="true" targetPageId="7">
<paragraph id="CE047E53FF98FFCEB7CB449EFB46FF07" blockId="7.[151,1158,151,175]" box="[151,1158,151,175]" pageId="7" pageNumber="586">
<emphasis id="FCCFA241FF98FFCEB7CB449EFECAFF07" bold="true" box="[151,266,152,175]" pageId="7" pageNumber="586">TABLE 1.</emphasis>
Differences between 
<taxonomicName id="09BB05D0FF98FFCEB6AE4491FD24FF07" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" box="[498,740,151,175]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" sensu="stricto" species="pubescens">
<emphasis id="FCCFA241FF98FFCEB6AE4491FD24FF07" box="[498,740,151,175]" italics="true" pageId="7" pageNumber="586">Neoliomera pubescens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E7FC1F3AFF98FFCEB5B7449EFCE5FF07" box="[747,805,152,175]" pageId="7" pageNumber="586" sensu="stricto">s. str.</taxonomicNameLabel>
, 
<taxonomicName id="09BB05D0FF98FFCEB46C449EFC5DFF07" authorityName=", Forest &amp; Guinot" authorityYear="1961" box="[816,925,151,175]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="demani">
<emphasis id="FCCFA241FF98FFCEB46C449EFC5DFF07" box="[816,925,151,175]" italics="true" pageId="7" pageNumber="586">N. demani</emphasis>
</taxonomicName>
and 
<taxonomicName id="09BB05D0FF98FFCEB48D449EFB84FF07" authority="P.-H &amp; Ng, 2014" authorityName="P.-H &amp; Ng" authorityYear="2014" box="[977,1092,151,175]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="fragaea" status="sp. nov.">
<emphasis id="FCCFA241FF98FFCEB48D449EFB84FF07" box="[977,1092,151,175]" italics="true" pageId="7" pageNumber="586">N. fragaea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E7FC1F3AFF98FFCEB316449EFB46FF07" box="[1098,1158,152,175]" pageId="7" pageNumber="586" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="CE047E53FF98FFCEB6FF44CDFA85FC0C" pageId="7" pageNumber="586">
<table id="BCBB8CF3FF980036B7C344CDFA4FFC0C" box="[159,1423,203,932]" gridcols="4" gridrows="7" pageId="7" pageNumber="586">
<tr id="708B7C11FF980036B7C344CDFA4FFF48" box="[159,1423,203,224]" gridrow="0" pageId="7" pageNumber="586" rowspan-0="1">
<th id="335A156DFF980036B6FF44CDFD30FF48" box="[419,752,203,224]" gridcol="1" gridrow="0" pageId="7" pageNumber="586">
<taxonomicName id="09BB05D0FF98FFCEB6FF44CDFD49FF48" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" box="[419,649,203,224]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="FCCFA241FF98FFCEB6FF44CDFD49FF48" box="[419,649,203,224]" italics="true" pageId="7" pageNumber="586">Neoliomera pubescens</emphasis>
</taxonomicName>
</th>
<th id="335A156DFF980036B45544CDFB84FF48" box="[777,1092,203,224]" gridcol="2" gridrow="0" pageId="7" pageNumber="586">
<taxonomicName id="09BB05D0FF98FFCEB45544CDFC11FF48" authorityName=", Forest &amp; Guinot" authorityYear="1961" box="[777,977,203,224]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="demani">
<emphasis id="FCCFA241FF98FFCEB45544CDFC11FF48" box="[777,977,203,224]" italics="true" pageId="7" pageNumber="586">Neoliomera demani</emphasis>
</taxonomicName>
</th>
<th id="335A156DFF980036B30544CDFA4FFF48" box="[1113,1423,203,224]" gridcol="3" gridrow="0" pageId="7" pageNumber="586">
<taxonomicName id="09BB05D0FF98FFCEB30544CDFAE6FF48" authorityName="P.-H &amp; Ng" authorityYear="2014" box="[1113,1318,203,224]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="fragaea">
<emphasis id="FCCFA241FF98FFCEB30544CDFAE6FF48" box="[1113,1318,203,224]" italics="true" pageId="7" pageNumber="586">Neoliomera fragaea</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="708B7C11FF980036B7C344F2FA4FFEEF" box="[159,1423,244,327]" gridrow="1" pageId="7" pageNumber="586">
<th id="335A156DFF980036B7C344F2FEB2FEEF" box="[159,370,244,327]" gridcol="0" gridrow="1" pageId="7" pageNumber="586">Anterolateral margin</th>
<td id="335A156DFF980036B6FF44F2FD30FEEF" box="[419,752,244,327]" gridcol="1" gridrow="1" pageId="7" pageNumber="586">Almost entire, lateral lobes not discernible, separated only by shallow fissures (Fig. 6A, C)</td>
<td id="335A156DFF980036B45544F2FB84FEEF" box="[777,1092,244,327]" gridcol="2" gridrow="1" pageId="7" pageNumber="586">Anterolateral margin with 3 well defined lobes (Figs. 3C, 6E)</td>
<td id="335A156DFF980036B30544F2FA4FFEEF" box="[1113,1423,244,327]" gridcol="3" gridrow="1" pageId="7" pageNumber="586">Anterolateral margin with 3 well defined lobes (Fig. 1A, D, E, 2A, B, 3A, 5A)</td>
</tr>
<tr id="708B7C11FF980036B7C3455CFA4FFE64" box="[159,1423,346,460]" gridrow="2" pageId="7" pageNumber="586">
<th id="335A156DFF980036B7C3455CFEB2FE64" box="[159,370,346,460]" gridcol="0" gridrow="2" pageId="7" pageNumber="586">Frontal margin</th>
<td id="335A156DFF980036B6FF455CFD30FE64" box="[419,752,346,460]" gridcol="1" gridrow="2" pageId="7" pageNumber="586">Median lobes short, truncate, not projecting anteriorly (Figs. 6A, C)</td>
<td id="335A156DFF980036B455455CFB84FE64" box="[777,1092,346,460]" gridcol="2" gridrow="2" pageId="7" pageNumber="586">Median lobes relatively more salient, triangular, projecting anteriorly (Figs. 3C, 6E)</td>
<td id="335A156DFF980036B305455CFA4FFE64" box="[1113,1423,346,460]" gridcol="3" gridrow="2" pageId="7" pageNumber="586">Median lobes relatively salient, triangular, projecting anteriorly (Figs. 1A, D, E, 2, 3A, 5A)</td>
</tr>
<tr id="708B7C11FF980036B7C345E6FA4FFD27" box="[159,1423,480,655]" gridrow="3" pageId="7" pageNumber="586">
<th id="335A156DFF980036B7C345E6FEB2FD27" box="[159,370,480,655]" gridcol="0" gridrow="3" pageId="7" pageNumber="586">Palm of chela</th>
<td id="335A156DFF980036B6FF45E6FD30FD27" box="[419,752,480,655]" gridcol="1" gridrow="3" pageId="7" pageNumber="586">Length of dactylus greater than width of palm; black pigmentation of finger extends only to distal margins of palm (Fig. 6D)</td>
<td id="335A156DFF980036B45545E6FB84FD27" box="[777,1092,480,655]" gridcol="2" gridrow="3" pageId="7" pageNumber="586">Length of dactylus subequal to width of palm; black pigmentation of finger extensive, forming band around palm, covering much of distal half of palm (Figs. 4C, D, 6G)</td>
<td id="335A156DFF980036B30545E6FA4FFD27" box="[1113,1423,480,655]" gridcol="3" gridrow="3" pageId="7" pageNumber="586">Length of dactylus greater than width of palm; black pigmentation of finger extends only to distal margins of palm (Fig. 4A, B)</td>
</tr>
<tr id="708B7C11FF980036B7C346A4FA4FFD5E" box="[159,1423,674,758]" gridrow="4" pageId="7" pageNumber="586">
<th id="335A156DFF980036B7C346A4FEB2FD5E" box="[159,370,674,758]" gridcol="0" gridrow="4" pageId="7" pageNumber="586">Fingers of chela</th>
<td id="335A156DFF980036B6FF46A4FD30FD5E" box="[419,752,674,758]" gridcol="1" gridrow="4" pageId="7" pageNumber="586">Gently curved throughout length (Fig. 6D)</td>
<td id="335A156DFF980036B45546A4FB84FD5E" box="[777,1092,674,758]" gridcol="2" gridrow="4" pageId="7" pageNumber="586">Straight along proximal part before strongly curving distally (Figs. 4C, D, 6F, G)</td>
<td id="335A156DFF980036B30546A4FA4FFD5E" box="[1113,1423,674,758]" gridcol="3" gridrow="4" pageId="7" pageNumber="586">Straight along proximal part before curving distally (Fig. 4A, B)</td>
</tr>
<tr id="708B7C11FF980036B7C3470FFA4FFC96" box="[159,1423,777,830]" gridrow="5" pageId="7" pageNumber="586">
<th id="335A156DFF980036B7C3470FFEB2FC96" box="[159,370,777,830]" gridcol="0" gridrow="5" pageId="7" pageNumber="586">Merus of last ambulatory leg</th>
<td id="335A156DFF980036B6FF470FFD30FC96" box="[419,752,777,830]" gridcol="1" gridrow="5" pageId="7" pageNumber="586">Short, broad (Fig. 6A)</td>
<td id="335A156DFF980036B455470FFB84FC96" box="[777,1092,777,830]" gridcol="2" gridrow="5" pageId="7" pageNumber="586">Long, broad (Figs. 1F, 3C)</td>
<td id="335A156DFF980036B305470FFA4FFC96" box="[1113,1423,777,830]" gridcol="3" gridrow="5" pageId="7" pageNumber="586">Short, broad (Figs. 1A, D, E, 3A)</td>
</tr>
<tr id="708B7C11FF980036B7C34757FA4FFC0C" box="[159,1423,849,932]" gridrow="6" pageId="7" pageNumber="586">
<th id="335A156DFF980036B7C34757FEB2FC0C" box="[159,370,849,932]" gridcol="0" gridrow="6" pageId="7" pageNumber="586">G1</th>
<td id="335A156DFF980036B6FF4757FD30FC0C" box="[419,752,849,932]" gridcol="1" gridrow="6" pageId="7" pageNumber="586">Subdistal lobe not extending beyond gently curving tip (Fig. 7A, C)</td>
<td id="335A156DFF980036B4554757FB84FC0C" box="[777,1092,849,932]" gridcol="2" gridrow="6" pageId="7" pageNumber="586">Subdistal lobe subtruncate, not extending beyond almost straight tip (Fig. 7D)</td>
<td id="335A156DFF980036B3054757FA4FFC0C" box="[1113,1423,849,932]" gridcol="3" gridrow="6" pageId="7" pageNumber="586">Subdistal lobe subovate, long, extending beyond almost straight tip (Fig. 7B, E)</td>
</tr>
</table>
</paragraph>
<paragraph id="CE047E53FF98FFCEB79B47EFFEAEFB3A" blockId="7.[151,1437,1001,1278]" pageId="7" pageNumber="586">
The specimen of 
<taxonomicName id="09BB05D0FF98FFCEB6CD47ECFDD6FBA9" authority="P.-H &amp; Ng, 2014" authorityName="P.-H &amp; Ng" authorityYear="2014" box="[401,534,1001,1025]" class="Malacostraca" family="Xanthidae" genus="Neoliomera" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="586" phylum="Arthropoda" rank="species" species="fragraea" status="sp. nov.">
<emphasis id="FCCFA241FF98FFCEB6CD47ECFDD6FBA9" box="[401,534,1001,1025]" italics="true" pageId="7" pageNumber="586">N. fragraea</emphasis>
</taxonomicName>
<emphasis id="FCCFA241FF98FFCEB57C47EFFDA2FBAA" bold="true" box="[544,610,1001,1026]" pageId="7" pageNumber="586">
<taxonomicNameLabel id="E7FC1F3AFF98FFCEB57C47EFFDA2FBAA" box="[544,610,1001,1026]" pageId="7" pageNumber="586" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from O‘ahu, Hawaiian Islands (ZRC 2000.0483) was obtained at night from a reef with strong currents (D. Takaoka, pers. comm.). The second author has also observed several specimens from shallow water in the intertidal in Maui at night, hiding among the eroded limestone formations of dead reefs that are exposed to very strong waves and tidal surges (see habitat notes in 
<bibRefCitation id="AA2A03A2FF98FFCEB4BA4050FB8CFBC6" author="Ng, P. K. L." box="[998,1100,1109,1134]" pageId="7" pageNumber="586" pagination="34 - 48" refId="ref6605" refString="Ng, P. K. L. (2011) Pele ramseyi, a new genus and new species of anchialine swimming crab (Crustacea: Brachyura: Portunidae) from the Hawaiian Islands. Zootaxa, 2737, 34 - 48." type="journal article" year="2011">Ng 2011</bibRefCitation>
). These conditions make their collection difficult.
</paragraph>
<paragraph id="CE047E53FF98FFCEB79B4098FDCAFB56" blockId="7.[151,1437,1001,1278]" pageId="7" pageNumber="586">
<materialsCitation id="7ED3740EFF98FFCEB79B4098FDC6FB56" ID-GBIF-Occurrence="3326196316" collectionCode="NTOU" pageId="7" pageNumber="586" specimenCode="ZRC 2014.0206" specimenCount="2" typeStatus="holotype">
Two of the specimens, the 
<typeStatus id="1100C0F1FF98FFCEB6A8409BFD96FB1E" box="[500,598,1181,1206]" pageId="7" pageNumber="586" type="holotype">holotype</typeStatus>
male (18.3 × 
<quantity id="0943D3B6FF98FFCEB5AF409BFC99FB1D" box="[755,857,1181,1206]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="7" pageNumber="586" unit="mm" value="10.1">10.1 mm</quantity>
, 
<specimenCode id="9E1DD628FF98FFCEB439409BFBDBFB1E" box="[869,1051,1181,1206]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="586">ZRC 2014.0206</specimenCode>
) and a 
<typeStatus id="1100C0F1FF98FFCEB3314098FB0DFB1E" box="[1133,1229,1182,1206]" pageId="7" pageNumber="586" type="paratype">paratype</typeStatus>
male (18.2 × 
<quantity id="0943D3B6FF98FFCEB237409BFF05FB71" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="7" pageNumber="586" unit="mm" value="10.8">10.8 mm</quantity>
, 
<collectionCode id="A8AAE696FF98FFCEB78940C7FEE7FB72" box="[213,295,1217,1242]" country="Taiwan" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="7" pageNumber="586">NTOU</collectionCode>
), are most probably infected with a bopyrid in their gill chambers; their branchial regions being distinctly swollen (
<figureCitation id="568062D6FF98FFCEB63140E3FE0CFB56" box="[365,460,1253,1278]" captionStart="FIGURE 1" captionStartId="3.[151,250,1905,1927]" captionTargetBox="[151,1436,193,1884]" captionTargetId="figure-95@3.[151,1436,193,1884]" captionTargetPageId="3" captionText="FIGURE 1. Colours in life. A–C, Neoliomera fragraea n. sp., female (25.3 × 14.3 mm) (NTOU), Taiwan; D, Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam [photograph by G. Paulay]; E, Neoliomera fragraea n. sp., paratype male (18.2 × 10.8 mm) (NTOU) [photograph by G. Paulay]; F, Neoliomera demani Forest &amp; Guinot, 1961, male (35.7 × 18.7 mm) (NTOU), Taiwan [photograph by Wang Kuei-Ching]." figureDoi="http://doi.org/10.5281/zenodo.4920418" httpUri="https://zenodo.org/record/4920418/files/figure.png" pageId="7" pageNumber="586">Figs. 1E</figureCitation>
, 
<figureCitation id="568062D6FF98FFCEB68540E3FE3DFB56" box="[473,509,1253,1278]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[241,1345,193,1932]" captionTargetId="figure-54@4.[241,1345,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Neoliomera fragraea n. sp., holotype male (18.3 × 10.1 mm) (ZRC 2014.0206), Guam. A, dorsal view; B, dorsal view of carapace; C, frontal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4920420" httpUri="https://zenodo.org/record/4920420/files/figure.png" pageId="7" pageNumber="586">2C</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>