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<mods:title id="E491886A7F52F19E272F0FC0529F247A">Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
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<mods:namePart id="7845BF476018FA5116F6E8655C97EA23">Bistroem, Olof</mods:namePart>
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<mods:namePart id="8B71C0DBA4116AD28D4C923F3132CFBC">Bergsten, Johannes</mods:namePart>
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<treatment id="45FBBD15AB40ACE1D92D721E91133658" ID-GBIF-Taxon="127862781" LSID="urn:lsid:plazi:treatment:45FBBD15AB40ACE1D92D721E91133658" httpUri="http://treatment.plazi.org/id/45FBBD15AB40ACE1D92D721E91133658" lastPageId="183" lastPageNumber="184" pageId="176" pageNumber="177">
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<paragraph id="F084708B69993ADB570E4E315FF08E07" pageId="176" pageNumber="177">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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<subSubSection id="4AD55B1E0B24F884EED70182FB0BB8DD" pageId="176" pageNumber="177" type="nomenclature">
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<taxonomicName id="AAFEB80624516D33C45D48DF5516F8EF" ID-CoL="3RM6H" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="176" pageNumber="177" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis Sharp, 1882</taxonomicName>
Figs 147-149, 332, 478-480, 564
</paragraph>
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<subSubSection id="D4DA35A5BE912AF869D9DA1B5BA93655" pageId="177" pageNumber="178" type="reference_group">
<paragraph id="F8FF17939086EF24DA9ECBC137D57127" pageId="177" pageNumber="178">
<taxonomicName id="9C701BDF91D0EDF42953BC77618802F9" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="species" species="cyclopis">
<pageBreakToken id="8A51F1199986D0616D257DB4B094683F" pageId="177" pageNumber="178" start="start">Laccophilus</pageBreakToken>
cyclopis
</taxonomicName>
<bibRefCitation id="950331B153D54E7F71BD206D172C2465" author="Sharp, D" journalOrPublisher="The Scientific Transactions of the Royal Dublin Society" pageId="263" pageNumber="264" pagination="179 - 1003" title="On aquatic carnivorous Coleoptera or Dytiscidae." volume="2" year="1882">Sharp 1882</bibRefCitation>
: 308 (original description, faunistics); v. d.
<bibRefCitation id="50333A3CCEA636E5093F6830629560E5" author="Branden, C van den" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="255" pageNumber="256" pagination="1 - 118" title="Catalogue des coleopteres carnassiers aquatiques (Haliplidae, Amphizoidae, Pelobiidae et Dytiscidae)." volume="28" year="1885">Branden 1885</bibRefCitation>
: 20 (catalogue, faunistics);
<bibRefCitation id="4846222A6193C337F296B95DA725B375" author="Regimbart, M" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="261" pageNumber="262" pagination="227 - 240" title="Haliplidae, Dytiscidae &amp; Gyrinidae. Voyage de M. E. Simon dans l'Afrique australe (Decembre-Mars 1893)." volume="63" year="1894">
<normalizedToken id="77DB0FA038F8A0E6022ED7C52805B1F2" originalValue="Régimbart">Regimbart</normalizedToken>
1894
</bibRefCitation>
: 237: (description, faunistics);
<bibRefCitation id="73A9EC8823FBECC45E1CDD1C3930046F" author="Regimbart, M" journalOrPublisher="Memoires de la Societe entomologique de Belgique" pageId="262" pageNumber="263" pagination="1 - 244" title="Revision des Dytiscidae et Gyrinidae d'Afrique, Madagascar et iles voisines. En contribution a la faune entomologique du Congo." volume="4" year="1895">
<normalizedToken id="0553866D05284A9C2DFB41F99090B6BD" originalValue="Régimbart">Regimbart</normalizedToken>
1895
</bibRefCitation>
: 140 (description, faunistics);
<bibRefCitation id="4E475BF92929B788E0B00B6C19E561F3" author="Zimmermann, A" journalOrPublisher="Junk, Berlin" pageId="263" pageNumber="264" title="Dytiscidae, Haliplidae, Hygrobiidae, Amphizoidae. In: Schenkling S (Ed.) Coleopterorum Catalogus. Vol. 4 pars 71. W." year="1920 a">Zimmermann 1920a</bibRefCitation>
:18 (catalogue, faunistics);
<bibRefCitation id="DBDD038C05D3D28262317D3CC19EBB17" author="Zimmermann, A" journalOrPublisher="Entomologische Blaetter" pageId="263" pageNumber="264" pagination="224 - 234" title="Bemerkenswerte Neuerwerbungen des Zoologischen Museums in Hamburg. Haliplidae, Dytiscidae et Gyrinidae." volume="16" year="1920 b">Zimmermann 1920b</bibRefCitation>
: 225 (faunistics);
<bibRefCitation id="1DDC8750EE6A2EF0DE0CF0B64977FC82" author="Gschwendtner, L" journalOrPublisher="Annals of the Transvaal Museum" pageId="256" pageNumber="257" pagination="15 - 25" title="Wissenschaftliche Ergebnisse der Vernay-Lang Kalahari-Expedition, Maerz bis September, 1930. Haliplidae et Dytiscidae." volume="17" year="1935 a">Gschwendtner 1935a</bibRefCitation>
: 18 (faunistics);
<bibRefCitation id="09E161944B3A3393A2DC14BB713CD381" author="Guignot, F" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle Paris" pageId="256" pageNumber="257" pagination="215 - 322" title="Mission scientifique de l'Omo 6 (58). ColeopteraDytiscidae (2 e partie)." volume="19" year="1946 c">Guignot 1946c</bibRefCitation>
: 269 (description, faunistics);
<bibRefCitation id="4DBFC1E5DF6C70D4FE2210993EAD9C90" author="Omer-Cooper, Joyce" journalOrPublisher="Memoirs of the Entomological Society of Southern Africa" pageId="260" pageNumber="261" pagination="1 - 90" title="Dytiscidae (Coleoptera) collected in the Transvaal (Rhodes University Expedition 1948)." volume="5" year="1957">Omer-Cooper 1957</bibRefCitation>
: 15, 90 (description, discussion, faunistics);
<bibRefCitation id="6E255AB5DFCAFAEA81109CEB33164251" author="Guignot, F" journalOrPublisher="Annales du Musee Royal du Congo Belge (Serie 8 vo) Sciences Zoologiques" pageId="258" pageNumber="259" pagination="323 - 648" title="Revision des hydrocanthares d'Afrique (ColeopteraDytiscoidea). 2." volume="78" year="1959 a">Guignot 1959a</bibRefCitation>
: 558, 562, 564 (description, faunistics);
<bibRefCitation id="0CC932BE9666CEED4B6262BC54351201" author="Omer-Cooper, Joyce" journalOrPublisher="Annals of the Cape Provincial Museums" pageId="261" pageNumber="262" pagination="292 - 296" title="The distribution of the Cape Province Dytiscidae and speculations as to their possible origins." volume="2" year="1962">Omer-Cooper 1962</bibRefCitation>
: 295 (faunistics);
<bibRefCitation id="6336CF16700AE7E0095495CC8580CA55" author="Omer-Cooper, Joyce" journalOrPublisher="South African Animal Life" pageId="261" pageNumber="262" pagination="59 - 214" title="Chapter 2. Coleoptera: Dytiscidae." volume="11" year="1965">Omer-Cooper 1965</bibRefCitation>
: 76, 83, 85 (description, biol.);
<bibRefCitation id="3F274AF99086E8B091A6FF1AEEB269D8" author="Betrand, H" journalOrPublisher="Bulletin du Museum National d'Histoire Naturelle (ser. 2)" pageId="253" pageNumber="254" pagination="861 - 870" title="Coleopteres hydrocanthares recoltes en Afrique du Sud et Rhodesies (Bertrand, 1959)." volume="39" year="1967">Bertrand and Legros 1967</bibRefCitation>
: 862 (faunistics);
<bibRefCitation id="0D7432414AC7802150B7EC966CB681B1" author="Curtis, BA" journalOrPublisher="Madouqua" pageId="255" pageNumber="256" pagination="163 - 187" title="Freshwater macro-invertebrates of Namibia." volume="17" year="1991">Curtis 1991</bibRefCitation>
: 186 (faunistics);
<bibRefCitation id="C099339A19CDEBB9EDB2BB0C49DC11E3" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
: 242 (catalogue, faunistics);
<bibRefCitation id="9FCDFF65F76C3E94D6CFEF45BA6E3403" author="Pederzani, F" journalOrPublisher="Quaderno di Studi e Notizie di Storia Naturale della Romana" pageId="261" pageNumber="262" pagination="35 - 41" title="Laccophiluscomoensis sp. n. from Ivory Coast (InsectaColeopteraDytiscidae)." volume="16" year="2002">Pederzani and Reintjes 2002</bibRefCitation>
: 38 (faunistics);
<bibRefCitation id="4BACE8512DE95135A1AB1354DA91195F" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
: 210 (catalogue, faunistics).
</paragraph>
<paragraph id="85781AA601607F466CB0CD222A7A2196" pageId="177" pageNumber="178">
<taxonomicName id="85EE7F45CE0D984DAE92E325C5F90EAB" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus shephardi" order="Coleoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="species" species="shephardi">Laccophilus shephardi</taxonomicName>
<bibRefCitation id="4E649B7DA5C50FD3B0711C4771E0FCE6" author="Omer-Cooper, Joyce" journalOrPublisher="South African Animal Life" pageId="261" pageNumber="262" pagination="59 - 214" title="Chapter 2. Coleoptera: Dytiscidae." volume="11" year="1965">Omer-Cooper 1965</bibRefCitation>
: 76, 84 (original description, faunistics);
<bibRefCitation id="81C01A9508FF2F4A8D5813A1106DF4AF" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
: 251 (catalogue, faunistics)
<bibRefCitation id="1F0737AF770B1EBEFCBD8D7BFABC80F7" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
: 217 (catalogue, faunistics). New synonym.
</paragraph>
</subSubSection>
<subSubSection id="9994CBB02C7FFF1F46CDA8468108DEAE" pageId="177" pageNumber="178" type="type localities">
<paragraph id="B432CCC91CFE30DD069EB5C915257646" pageId="177" pageNumber="178">Type localities.</paragraph>
<paragraph id="28988318CC22103FBE52257CF0A8AF5B" pageId="177" pageNumber="178">
<taxonomicName id="536C65FE602AB1AEEE47FA12DB694EDA" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
: South Africa: Grahamstown.
</paragraph>
<paragraph id="F5DA5853AFE29CE60A112350C0EC8E69" pageId="177" pageNumber="178">
<taxonomicName id="1D155642C41D547A978411480DD98F53" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus shephardi" order="Coleoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="species" species="shephardi">Laccophilus shephardi</taxonomicName>
: South Africa: Mt Currie district, nr. Zwartberg.
</paragraph>
</subSubSection>
<subSubSection id="62FC45020ED276616CA01A97C18CEB2C" pageId="177" pageNumber="178" type="type material">
<paragraph id="B9207BF7D244EB32A7C1F380B6666169" pageId="177" pageNumber="178">Type material studied</paragraph>
<paragraph id="81F84EA814B5BE6470AE8CF45FAE9793" pageId="177" pageNumber="178">
(18 exs.).
<taxonomicName id="2B697BC9CCF7F61285FC535D20366401" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
: Lectotype (by present designation): male: &quot;Type / S. Africa / Grahamstown / Type 573
<taxonomicName id="F52D5372B9AB20FA54D49B2809D95E38" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
sp. Afr. mer. / Sharp Coll. 1905-313&quot; (BMNH). - Paralectotypes: Same data, but
<normalizedToken id="3476CFB38AD4C53295CCF612BB718D03" originalValue="“Cotype”">&quot;Cotype&quot;</normalizedToken>
(1 ex. BMNH); same data but &quot;Cotype Caffrarie&quot; (1 ex. BMNH); &quot;Cotype / S. Africa /
<taxonomicName id="2DCDB2CB690E47D558717FF04F239E76" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
Sharp Co-type&quot; (3 exs. BMNH).
</paragraph>
<paragraph id="53BD0DA0852CD8C90A36B536B8168EAF" pageId="177" pageNumber="178">
<taxonomicName id="E76613C6408F80CB833FF41B85ADCF2C" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus shephardi" order="Coleoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="species" species="shephardi">Laccophilus shephardi</taxonomicName>
: Lectotype (by present designation): male: &quot;Syntype /
<taxonomicName id="A666C78C6F9685276C02EE3C587EB6A6" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus shephardi" order="Coleoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="species" species="shephardi">Laccophilus shephardi</taxonomicName>
male type / E Cape province Mt Currie Dist. nr Zwartberg 14-XI-1957 J. O-C.&quot; (BMNH). - Paralectotypes: Same as lectotype but labelled as &quot;female type&quot; (1 ex. BMNH); &quot;Cotype / E. Cape Prov. Maclear 9.V.1926 No 275 J. Omer-Cooper /det. J. Omer-Cooper
<taxonomicName id="2060597A195ABF8CDC11D16268FDA832" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus shephardi" order="Coleoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="species" species="shephardi">Laccophilus shephardi</taxonomicName>
O-C.&quot; (5 exs. AMGS); &quot;E. Cape Prov. Mount Currie 13.II. 1957 No. 400 J. Omer-Cooper&quot; (5 exs. AMGS); &quot;S. Africa E. Cape Prov. Barkley East 14.II. 1948 JOC. / det. J. Omer-Cooper
<taxonomicName id="F97B7C13CC1D00508073657D48379A11" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus shephardi" order="Coleoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="species" species="shephardi">Laccophilus shephardi</taxonomicName>
O-C.&quot; (1 ex. AMGS).
</paragraph>
</subSubSection>
<subSubSection id="3BC7133F8C2260DC6406DDFFD7833C28" lastPageId="182" lastPageNumber="183" pageId="177" pageNumber="178" type="materials_examined">
<paragraph id="E09903B3D892071D9F5A0AAAC4D28AD3" pageId="177" pageNumber="178">Additional material studied</paragraph>
<paragraph id="607A6EA7BCC17AAB8A139C50FB7FF085" lastPageId="182" lastPageNumber="183" pageId="177" pageNumber="178">
(655 exs.). Namibia: &quot;Windhoek Town Dam 7.7. 1939 JOC.&quot; (1 ex. AMGS); &quot;Windhoek New Dam 7.7. 1939 JOC.&quot; (1 ex. AMGS); &quot;Windhoek River Bed 9 July 1937&quot; (2 exs. AMGS); &quot;Windhoek 5.7. 1939&quot; (1 ex. AMGS); &quot;Windhoek Eros Mt. 1600 m
<geoCoordinate id="E489EA460D6F9CAAF182D6FA391A6E05" direction="south" orientation="latitude" precision="555" value="-22.34">22.34S-</geoCoordinate>
<geoCoordinate id="31D9464412A811AFFC39742F384D0637" direction="east" orientation="longitude" precision="555" value="17.06">17.06E</geoCoordinate>
/ 10.9. 1974 shorewashing
<normalizedToken id="C127BA29F038250BEBA3099D8644F3FA" originalValue="Endrödy-Younga">Endroedy-Younga</normalizedToken>
: 365&quot; (1 ex. TMSA); &quot;Windhoek Distr., Valencia Ranch / 14-24.4. 1972 Strydom&quot; (1 ex. TMSA); &quot;Osona b. Okahandja 19-20.10. 1991
<normalizedToken id="C746E9CC10DE7EA4DA54E50A365D3FD1" originalValue="Göllner">Goellner</normalizedToken>
leg.&quot; (1 ex. ZMHB); &quot;Osona b. Okahandja p. III. -m IV 1988 leg. J. Irish&quot; (1 ex. ZMHB); &quot;Pools in overflow stream from dam, much weed &amp; algae / Okarupa, ca. 17 mi E of Okahandja 4900 ft, 22.5. 1954 / J. Balfour-Browne&quot; (21 exs. BMNH, 2 exs. MZH); &quot;Okahandja 4700 ft. 22.5. 1954 / Water-hole and seepage through sand J. Balfour-Browne&quot; (8 exs. BMNH, 1 ex. MZH); &quot;Okahandja 4400 ft. 21.5. 1954 Small, deep water-hole, thick
<taxonomicName id="497C3F84347CF0E091866228590DF75D" class="Liliopsida" family="Araceae" genus="Lemna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lemna" order="Alismatales" pageId="177" pageNumber="178" phylum="Tracheophyta" rank="genus">Lemna</taxonomicName>
cover J. Balfour-Browne&quot; (3 exs. BMNH); &quot;Khomashochl, Farm Wissenfels
<geoCoordinate id="F4790EC9924626E7BB4B7ED1DE049D4B" direction="south" orientation="latitude" precision="925" value="-23.333334">23°20'S-</geoCoordinate>
<geoCoordinate id="0C5B093E78B4F203E723215C4F130754" direction="east" orientation="longitude" precision="925" value="16.416666">16°25'E</geoCoordinate>
/ 14.9. 1974 shore washing E-Y: 371&quot; (5 exs. TMSA, 1 ex. MZH); &quot;Hardap Dam, Mariental Dist. 10-14. 4. 1972 Strydom &amp; Jones&quot; (6 exs. TMSA); &quot;Hardap GR: Water Institute, pool shore waterplant treating, water catcher 1.12. 1997, 24°
<geoCoordinate id="EBE3B296F99EEB1F0B7B40391576B92A" direction="south" orientation="latitude" precision="555" value="-29.41">29.41S</geoCoordinate>
/17°
<geoCoordinate id="062578A9DB9839DBFA3D40E2D14084E0" direction="east" orientation="longitude" precision="555" value="51.52">51.52E</geoCoordinate>
, leg. M. Uhlig&quot; (9 exs. ZMHB, 4 exs. MZH, 2 exs. NMNW); &quot;Otjozondjupa Dist., Toggekry 250 (Omatako)
<geoCoordinate id="71A54C4C2D90B260F16D0C2639D990A3" direction="south" orientation="latitude" precision="1" value="-21.511917">21°30'42.9&quot;S</geoCoordinate>
,
<geoCoordinate id="0CBA211783176760613780B12C8D0D13" direction="east" orientation="longitude" precision="1" value="16.73239">16°43'56.6&quot;E</geoCoordinate>
, 1520 m NN, 8.3. 2003, hand light trap Frisch &amp; Vohland leg.&quot; (1 ex. ZMHB); same data, but &quot;1100 m, 5.3. 2003&quot; (1 ex. ZMHB); &quot;Otjozondjupa Dist., Otjiamongombe West 44,
<geoCoordinate id="B0752FE5147EF6087F03CAC3C57A700D" direction="south" orientation="latitude" precision="1" value="-21.59575">21°35'44.7&quot;S</geoCoordinate>
,
<geoCoordinate id="3F2E11005FC7F0E312A85616BFC438ED" direction="east" orientation="longitude" precision="1" value="16.938166">16°56'17.4&quot;E</geoCoordinate>
, 1498 m, NN 28.2.2003 hand light trap Frisch &amp; Vohland leg.&quot; (1 ex. ZMHB); &quot;Keetmanshoop Dist., Gellap Ost 3, 23 km NW Keetsmanshoop,
<pageBreakToken id="DFFD90DA422181D0192F9478E9D08480" pageId="178" pageNumber="179" start="start">dwarf</pageBreakToken>
shrub savannah (Nama-Karoo) / 4.4. 2001, 34-28˚C,
<geoCoordinate id="9B94030627680EB42AA84F019A27833D" direction="south" orientation="latitude" precision="1" value="-26.404917">26°24'17.7&quot;S</geoCoordinate>
, 18°00'41.9&quot; lux 18.00-21.00 leg. Uhlig et al.&quot; (1 ex. ZMHB); &quot;Damaral. Groot Barmen
<geoCoordinate id="188E1E463D472024B642D71D1DC394AF" direction="south" orientation="latitude" precision="555" value="-22.05">22.05S-</geoCoordinate>
<geoCoordinate id="2C3558478142375FE5B1C4E920073680" direction="east" orientation="longitude" precision="555" value="16.4">16.40E</geoCoordinate>
/ 12.9. 1975, shorewashing,
<normalizedToken id="0D3B3333DDEF467C20EF722F8052DA4B" originalValue="Endrödy-Younga">Endroedy-Younga</normalizedToken>
: 370&quot; (1 ex. TMSA); Namib Tinkas Dam
<geoCoordinate id="8DEF5B780B6842281B349A2AABE4A9B3" direction="south" orientation="latitude" precision="555" value="-22.5">22.50S-</geoCoordinate>
<geoCoordinate id="76D2A81FA400D374BC4AE920BBE54722" direction="east" orientation="longitude" precision="555" value="15.3">15.30E</geoCoordinate>
/ 1.11. 1974 shore washing
<normalizedToken id="F9EDA2FECBCEDE4740E66521DDEC0411" originalValue="Endrödy-Younga">Endroedy-Younga</normalizedToken>
440&quot; (1 ex. TMSA); &quot;Grootfontein v. Erffa S.G.&quot; (1 ex. NHMB); &quot;DSWA,?-erseba 8. 1905 Schultze&quot; (1 ex. NHMB); &quot;DSWA Gr. Etemba Casper S.G. /
<taxonomicName id="2AD7CD4AFD19781F6EF20874E1950D6B" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="178" pageNumber="179" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
Sharp det. Brancucci 1982&quot; (2 exs. ZMHB). - Botswana: &quot;Metsimaklaba 7-12.3. 1930 /
<taxonomicName id="CE5DDCC6D2DD45C163F93D44B1C2F0F4" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="178" pageNumber="179" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
Sharp det. J. Omer-Cooper&quot; (1 ex. TMSA); &quot;Tiokweng 15-21.3. 1988 Ward /
<taxonomicName id="B355A84DA0DFCD4E4720951CE05D2EF8" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="178" pageNumber="179" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
Sharp det.
<bibRefCitation id="6678E8010E376BD01E0D8079293E3173" author="Rocchi, S" journalOrPublisher="Frustula Entomologica (n. s.)" pageId="262" pageNumber="263" pagination="71 - 89" title="Contributo alla conoscenza degli Haliplidae e dei Dytiscidae del Madagascar con descrizione di due nuove specie (Coleoptera)." volume="14" year="1991">Rocchi 1991</bibRefCitation>
&quot; (1 ex. CSR). - South Africa: &quot;Pretoria, van Son /
<taxonomicName id="F545F5B64F5340DD2EE5F32FEA0D29EE" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="178" pageNumber="179" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
Shp det. J. Omer-Cooper&quot; (1 ex. TMSA); &quot;Pretoria 8.2., A.J.T. Janse /
<taxonomicName id="81D9E8BC734F9EB1EFECAAAA3FBF879C" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="178" pageNumber="179" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
Shp det. J. Omer-Cooper&quot; (1 ex. TMSA); &quot;Pretoria 27.9. 1958 L.
<normalizedToken id="98D5F3D3469C3652DCA6731D7056C926" originalValue="Vári”">Vari&quot;</normalizedToken>
(1 ex. TMSA); &quot;Pretoria 1948&quot; (1 ex. TMSA); &quot;Moorddrift 9. 1924 G. v. Dam /
<taxonomicName id="FBF3B537125335927A4EA7DD30E3F5B5" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="178" pageNumber="179" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
Shp det. J. Omer-Cooper&quot; (13 exs. TMSA); same data but &quot;
<taxonomicName id="6FA5643D99E8CD5263A57C22A246E048" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="178" pageNumber="179" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
Shp det. J. Omer-Cooper&quot; (1 ex. TMSA); &quot;Transvaal gravel pits Ermelo 8.12. 1948 JOC. /
<taxonomicName id="BA2CE431BFF139EBB61335E19541C04C" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="178" pageNumber="179" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
Sharp J. Balfour-Browne det.&quot; (1 ex. AMGS); &quot;Transvaal gravel pits Ermelo 8.12. 1948&quot; (1 ex. AMGS) &quot;S. Afr. Johannesburg Rivonia 19.XI. 1952 Carpenter&quot; (3 exs. AMGS); &quot;Gauteng, Natalspruit Vlei, Johannesburg,
<geoCoordinate id="5701D0F5DAC7668CFB4FE29EDC6CFB93" direction="north" orientation="latitude" precision="55" value="26.28">N-26.280</geoCoordinate>
,
<geoCoordinate id="94D346F7C283F592C94D8D965A870B91" direction="east" orientation="longitude" precision="55" value="28.15">E28.150</geoCoordinate>
, 5.6. 1971 Reavell&quot; (4 exs. AMGS); &quot;Transvaal nr Standerton 8.IV. 1954/Pool by muddy stream: algae et grass&quot; (1 ex. BMNH; habitus in Fig. 480); &quot;Transvaal Middleburg 29.11. 1948 JOC.&quot; (2 exs. AMGS); &quot;Middelburg Dam 22.2. 47 JOC&quot; (3 exs. AMGS); &quot;Freddy van Zyl Bridge, Oorsloog Spruit 25.2. 1947 JOC.&quot; (1 ex. AMGS); &quot;Trsvl, Jukskei R at Sandspruit
<geoCoordinate id="9C84F9E7F2657BFC16D635D71F24CDC2" direction="north" orientation="latitude" precision="55" value="26.01">N-26.010</geoCoordinate>
,
<geoCoordinate id="8D88628BEA444A0051004702F392E0B0" direction="east" orientation="longitude" precision="55" value="28.05">E28.050</geoCoordinate>
, 9.2. 1956 Allansaon&quot; (2 exs. AMGS); &quot;Trsvl, Jukskei R, at Alexandra,
<geoCoordinate id="1267CB9336A74DF5476E08AB2C55A101" direction="north" orientation="latitude" precision="55" value="26.1">N-26.100</geoCoordinate>
,
<geoCoordinate id="3031EE3BF1A9B8A9E86CF0B8F757E529" direction="east" orientation="longitude" precision="55" value="28.11">E28.110</geoCoordinate>
, 7.2. 1956 Allanson&quot; (2 exs. AMGS); &quot;Trsvl Sandfonteinspruit at Bramley Bridge
<geoCoordinate id="1A8637A17DACA1AA802180EDCA60438B" direction="north" orientation="latitude" precision="55" value="26.11">N-26.110</geoCoordinate>
,
<geoCoordinate id="59324BCD96E1908467D6E4AEAEEF6814" direction="east" orientation="longitude" precision="55" value="28.07">E28.070</geoCoordinate>
, 14.3. 1956 Allanson&quot; (2 exs. AMGS); &quot;Trsvl., Germiston Lake,
<geoCoordinate id="CC5E9B6A18A90FF05250C3F9685D3E42" direction="north" orientation="latitude" precision="55" value="26.23">N-26.230</geoCoordinate>
,
<geoCoordinate id="5C52213A13BFAFFEA23D008AB156F4A5" direction="east" orientation="longitude" precision="55" value="28.17">E28.170</geoCoordinate>
, 12.8. 1971 Reavell&quot; (1 ex. AMGS); &quot;Trsvl, stream,
<normalizedToken id="8AA8FCC4DEFDBF4DCBC3B1582B47C7FC" originalValue="Cowle’s">Cowle's</normalizedToken>
Dam, Brakpan
<geoCoordinate id="61CD9C3C12431C2877B4523F0DAF1ED3" direction="north" orientation="latitude" precision="55" value="26.23">N-26.230</geoCoordinate>
,
<geoCoordinate id="13F3DB7CA68F2A6661D0F7A8ED817ACF" direction="east" orientation="longitude" precision="55" value="28.31">E28.310</geoCoordinate>
, 11.8. 1971 Reavell&quot; (1 ex. AMGS); &quot;Transvaal Belfast N. 1948 O-C.&quot; (1 ex. AMGS); &quot;Belfast N. 30. 1948 JOC.&quot; (4 exs. AMGS); &quot;Belfast pond 23. N. 1948 OC.&quot; (4 exs. AMGS); &quot;Belfast 29.11. 1948 JOC.&quot; (5 exs. AMGS); &quot;Transvaal Standerton 8.12. 1948 JOC.&quot; (1 ex. AMGS); &quot;Transvaal Sand R. Pietersburg distr. 27.7. 1948&quot; (3 exs. AMGS); &quot;Transvaal Dynamite Factory Dams Wormsley April 1946&quot; (4 exs. AMGS); &quot;Koster 10. 1924 G. v. Dam / J. Omer-Cooper&quot; (1 ex. TMSA); &quot;pond Breyton rd nr. Chrissie Dec. 1948&quot; (1 ex. AMGS); &quot;Trib. R. Koop nr Barberton 1.12. 1948&quot; (1 ex. AMGS); &quot;Bandolierkop 23. N. 1948 OC.&quot; (1 ex. AMGS); &quot;Transvaal Springsl. (?) 20.6. 1950 G.B.H.&quot; (1 ex. AMGS); &quot;Trsvl nr Standerton / pool by muddy stream, algae &amp; grass J. Balfour-Browne leg.&quot; (1 ex. BMNH); &quot;NW Distr. Klerksdorph 20 km W of Botha Ville, Vaal Riv. 31.1. 2001 Snizek leg.&quot; (2 exs. NMW); &quot;Fount. Grove 17.08 05 C. Swiestrai /
<taxonomicName id="D2CFF4DAE1A89B1A1DEA606B3E4FED23" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus lineatus" order="Coleoptera" pageId="178" pageNumber="179" phylum="Arthropoda" rank="species" species="lineatus">Laccophilus lineatus</taxonomicName>
<normalizedToken id="6BC4928EBEDF4232693ACB90545231D7" originalValue="Aubé">Aube</normalizedToken>
C. Swiestra&quot; (1 ex. TMSA); &quot;NW, 70 km SW Mmabatho Setlagole 21.1. 2000 Halada leg.&quot; (1 ex. NMW); &quot;NW, 50 km S Kimberley, Ritchie 12.1. 2000 Halada leg.&quot; (1 ex. NMW); &quot;TRSVL, trib. of Sand river, Peekes /
<taxonomicName id="E84640D43B59D729F035F02AC38A588B" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="178" pageNumber="179" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
Shp det. J. Omer-Cooper&quot; (1 ex. TMSA); &quot;Fountains Pta 5.10. 1932 G. van Son /
<taxonomicName id="4A8675AEF216BEE76F39C9395FB36836" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="178" pageNumber="179" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
Sharp det. Gschwendtner / Omer-Cooper&quot; (2 exs. TMSA); &quot;Trsvl, Pond So Ermelo, Hwy N11, 1.12. 1995 Challet&quot; (1 ex. CGC); &quot;Mpumalanga Ezemvelo, Nat. R. 1374 m,
<geoCoordinate id="88C9F0A9A1E80828E153EFCC1EA41EE4" direction="south" orientation="latitude" precision="555" value="-25.42">25.42S-</geoCoordinate>
<geoCoordinate id="154C5E508FF8EA54296B7226C97A14F5" direction="east" orientation="longitude" precision="555" value="29.01">29.01E</geoCoordinate>
/ 27.1.2004 light trap, High
<pageBreakToken id="A6EBD30D409DEAD987CB7F1886376D76" pageId="179" pageNumber="180" start="start">veld</pageBreakToken>
E-Y: 3609 TMSA Staff&quot; (1 ex. TMSA); &quot;OFS Vredefort nr Honingsspruit 29.VIII. 1947 JOC.&quot; (1 ex. AMGS); &quot;O.F.St. Sasolburg 11. 1982 D.M. Kroon&quot; (1 ex. TMSA, 1 ex. MZH); &quot;Van
<normalizedToken id="8260734A1C14CBA51AC2DB63F2ED15A1" originalValue="Rhyn’s">Rhyn's</normalizedToken>
Pass N. 1933 G. van Son&quot; (1 ex, TMS); &quot;OFS, Parys 4 km E,
<geoCoordinate id="1A1996D2542AE527CB7D99D2760BA83F" direction="south" orientation="latitude" precision="555" value="-26.54">26.54S-</geoCoordinate>
<geoCoordinate id="89A4610905CED1183B0F78F0D67AEA91" direction="east" orientation="longitude" precision="555" value="27.35">27.35E</geoCoordinate>
/Farm Abel 52, 4-7.12. 1992 M.
<normalizedToken id="5CF116A0902E3F36AB294EEF3B7A831F" originalValue="Krüger">Krueger</normalizedToken>
leg.&quot; (3 exs. TMSA); &quot;OFS, Entembeni Mission nr Kraansfontein, 30.3. 1991 Reavell&quot; (1 ex. AMGS); &quot;Colesburg Dam Outsloe Town 12.2. 1947&quot; (1 ex. AMGS); &quot;Norvals Point Colesburg 23.2. 1947 JOC.&quot; (6 exs. AMGS); same but &quot;24.2. 1947&quot; (1 ex. AMGS); &quot;van
<normalizedToken id="BE8F3164266B7CEA6DDD328BEE96AB6A" originalValue="Wyk´s">Wyk's</normalizedToken>
Fontein, Colesburg 23.2. 1947 JOC.&quot; (1 ex. AMGS); &quot;Colesburg, Outside Town 22.2. 1947 JOC.&quot; (1 ex. AMGS); &quot;Kw. Natal, Karkloof R., Mgeni Confluence,
<geoCoordinate id="B510F7AA681861A556C3093753CD35F9" direction="north" orientation="latitude" precision="5" value="29.4438">N-29.4438</geoCoordinate>
,
<geoCoordinate id="2C8BF8519AF1D31D635FAEC59A05AFDE" direction="east" orientation="longitude" precision="1" value="30.301832">E30.301833</geoCoordinate>
, 11.11. 2003 Graham &amp; Dickens&quot; (1 ex. AMGS); &quot;Kw. Natal, Mooi R at Mearns Weir,
<geoCoordinate id="C04A7747E19AD45525F54219A08D129B" direction="north" orientation="latitude" precision="55" value="29.25">N-29.250</geoCoordinate>
,
<geoCoordinate id="EF15CCFB9C2791580A15D646EC004281" direction="east" orientation="longitude" precision="55" value="29.97">E29.970</geoCoordinate>
, light trap, 5.1. 1996 De Moor &amp; Dickens&quot; (2 exs. AMGS); &quot;Kw. Natal, Mooi R, at Retreat Farm,
<geoCoordinate id="4F62F22AD0C804088118B05C063C7353" direction="north" orientation="latitude" precision="55" value="29.27">N-29.270</geoCoordinate>
,
<geoCoordinate id="C0107FC7E1F1EFB1078024FE69CEE336" direction="east" orientation="longitude" precision="55" value="29.97">E29.970</geoCoordinate>
, light trap, 4.1. 1996 De Moor &amp; Dickens&quot; (2 exs. AMGS); &quot;Kw. Natal, Mooi R, at Dalcrue Farm,
<geoCoordinate id="6DABA7874E462C977D6451345DB37014" direction="north" orientation="latitude" precision="55" value="29.36">N-29.360</geoCoordinate>
,
<geoCoordinate id="F947BA30976C286EBEFDE36C7E698C76" direction="east" orientation="longitude" precision="55" value="29.89">E29.890</geoCoordinate>
, light trap, 4.1. 1996 De Moor &amp; Dickens&quot; (5 exs. AMGS); &quot;Kw. Natal, Klein Mooi R, at Dartington Vlei,
<geoCoordinate id="4DF11091AC9EC0CEAA3A69AEF15458CA" direction="north" orientation="latitude" precision="55" value="29.26">N-29.260</geoCoordinate>
,
<geoCoordinate id="F1B26C347625DC99FE71DA0A7F7CE626" direction="east" orientation="longitude" precision="55" value="29.87">E29.870</geoCoordinate>
, light trap, 5.1. 1996 De Moor &amp; Dickens&quot; (2 exs. AMGS); &quot;Kw. Natal,
<normalizedToken id="AA8EBEB349554B8F3DA6EF36A471691A" originalValue="Mcleod’s">Mcleod's</normalizedToken>
Farm nr Dargle, 4.2. 1989 Reavell&quot; (2 exs. AMGS); &quot;Kw. Natal, Ngagane R., Steildrift rd,
<geoCoordinate id="DF2924C35E996953C31F4A37149AE77B" direction="north" orientation="latitude" precision="55" value="27.77">N-27.770</geoCoordinate>
,
<geoCoordinate id="34B5B71EF6F9A16E22C10B5F5F7C563C" direction="east" orientation="longitude" precision="55" value="30.02">E30.020</geoCoordinate>
, 27.8. 1994 Metz&quot; (1 ex. AMGS); &quot;Kw. Natal, Lions R.,
<geoCoordinate id="46358B2B129AC091E6BA9F2C90BBCF79" direction="north" orientation="latitude" precision="55" value="29.47">N-29.470</geoCoordinate>
,
<geoCoordinate id="DE5B0552F7E88AC501523B0074C6AE6D" direction="east" orientation="longitude" precision="55" value="30.15">E30.150</geoCoordinate>
, 4.7. 1995 Dickens&quot; (3 exs. AMGS); &quot;Kw. Natal, Yorkshire Wolds Farm,
<geoCoordinate id="BC9638B482F84BA5D2C8CE61663F2985" direction="north" orientation="latitude" precision="5" value="29.3044">N-29.3044</geoCoordinate>
,
<geoCoordinate id="9C8CCF1356F43278167AC19FD5B2F6D3" direction="east" orientation="longitude" precision="1" value="29.773054">E29.773055</geoCoordinate>
, 10.6. 1998 De Moor &amp; al.&quot; (5 exs. AMGS); &quot;Natal Engelbrechts Drift Pond, Zaaihoek Dam, 272750S, 300510E, alt. 1740 m, Coke, Ngwenya&quot; (1 ex. CCT); &quot;Natal: Bergville Mont-Aux-Sources 4000 ft. 6.IV. 1954 / farm dam, thick weed at edge /
<taxonomicName id="A6C3C6204CAD3AF7E464149521E19F03" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
Sharp J. Balfour-Browne det. VIII, 1958&quot; (1 ex. BMNH); &quot;Natal Kokstad 14.4. 1947 JOC.&quot; (2 exs. AMGS); &quot;Nqutu Zululand 5.8. 1949 Newton&quot; (3 exs. SAMC); &quot;Nqutu Zululand 21.9. 1949 Newton&quot; (1 ex. SAMC); &quot;Durban Natal 20.1. 1903&quot; (1 ex. SAMC); &quot;Natal Bergville Mont-aux-Sources, 4000 ft.6.4. 1954 / farm dam, thick weed at edges J. Balfour-Browne /
<taxonomicName id="E6EA2C196652D710A09BCC601ABAF805" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
Shp det. J. Balfour-Browne&quot; (1 ex. BMNH); &quot;Natal Crocodile R. 13.VI. 1956&quot; (1 ex. AMGS); &quot;Natal, reservoir next of Pickle Pot on R617, 1000 m 2.2. 1997 Turner&quot; (1 ex. CCT); &quot;NC., drying ponds in stream, 12 km SE Garies 26.2. 1997 Challet leg.&quot;(2 exs. CNG, 1 ex. MZH); &quot;CP, Creek 9 km W Ft Beaufort 6.3. 1997 Challet&quot; (1 ex. CNC); &quot;Hutchinson, Cape, Marshall&quot; (1 ex. SAMC); &quot;Kimberley Bro. Power 1.6. 1912&quot; (1 ex. SAMC); &quot;MIL 9 A&quot; (= Scott &amp; Millard 1947 Estuary at Milnerton near Cape Town) (3 exs. AMGS); &quot;U.C.T. FR. W. 72&quot; (1 ex. AMGS); &quot;WC. Pr. Arniston July 1946&quot; (4 exs. AMGS); &quot;ERS 26B&quot; (= WCPr. Krom River 33,56,29S, 18,50,56 E) &quot;8.1. 1953 A.D. Harrison&quot; (2 exs. AMGS); &quot;WC, Franschhoek, rd R45, 25.3. 2001, river 3 km SE Franschhoek Ribera &amp; Cieslak leg.&quot; (1 ex. CIR); &quot;WC Du Toits Kloof rd N1, pond and River Wit in resort 24.3. 2001 Ribera &amp; Cieslak leg.&quot; (2 exs. CIR); &quot;WC Rd 310 before Khayelitsha artificial lagoon 21.3. 2001 Ribera &amp; Cieslak leg.&quot; (1 ex. CIR); &quot;Ceres Pond (964 m),
<geoCoordinate id="542BFB6E7B563043A51AE1A863993CFD" direction="south" orientation="latitude" precision="1" value="-32.9666">S32°57'59.75&quot;</geoCoordinate>
<geoCoordinate id="D54C8C5101FABCD179022E14C9B6D59B" direction="east" orientation="longitude" precision="1" value="19.383297">E19°22'59.87&quot;</geoCoordinate>
Hidalgo-Galiana &amp; Kleynhans leg.&quot; (1 ex. CIR); &quot;W. Cape, Meerlust dam, farm and fish dam surrounded by indigenous and alien plants,
<geoCoordinate id="1FEA72DA930027A7A18E7E6110C32607" direction="east" orientation="longitude" precision="1" value="18.74692">18.74692E</geoCoordinate>
, -
<geoCoordinate id="71F05371FB8A59F4129B3368CF7613AC" direction="north" orientation="latitude" precision="1" value="34.01554">34.01554N</geoCoordinate>
, alt. 20 m, 2006-2007&quot; (1 ex. CTT); &quot;Victoria West 23.6. 1911 Morris&quot; (2 exs. AMGS); &quot;nr Wellington 300ft. 10.VIII. 1954 J. Balfour-Browne / Deep dam pool with grass and
<taxonomicName id="3C75522C91A981FEABF9A906631CE61F" class="Liliopsida" family="Juncaceae" genus="Juncus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Juncus" order="Poales" pageId="179" pageNumber="180" phylum="Tracheophyta" rank="genus">Juncus</taxonomicName>
edging /
<taxonomicName id="EA12D508F444C5B8A5FB41154B17AAAC" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="species" species="cyclopis">
<pageBreakToken id="9A7710A96A41C7482531598FDA8BF95D" pageId="180" pageNumber="181" start="start">Laccophilus</pageBreakToken>
cyclopis
</taxonomicName>
Sharp J. Balfour-Browne det. IV. 1962&quot; (1 ex. BMNH);&quot;Paarl 16.10. 1949 Malkin / river&quot; (1 ex. BMNH); &quot;ECpr. Gt Fish R. Res. 33;6;59S, 26;39;3E (ECR 732) sm. river dam, clay, vegetation 7. XI. 2008 O.
<normalizedToken id="F0A0392CE7CF3C663DC8B8DE43C4303B" originalValue="Biström">Bistroem</normalizedToken>
leg.&quot; (1 ex. MZH); &quot;ECPr. Glenstone Farm nr Grahamstown ca. 1400 ft a.s.l. (ECR 731) in Suurberg Quartzite Fynbos, pool below spring 16.XI. 2008 O.
<normalizedToken id="B93CD96ACCBCBE352CF21A3C2D80B705" originalValue="Biström">Bistroem</normalizedToken>
leg.&quot; (2 exs. MZH); &quot;Willowmore 21.2. 1947 JOC.&quot; (2 exs. AMGS); &quot;Elliot 11.V. 1956 JOC.&quot; (1 ex. AMGS); &quot;Kowie Riv. nr. Grahamstown Uys leg.&quot; (1 ex. MZH); &quot;C. Prov. Albany District Grahamstown April 1951&quot; (1 ex. AMGS); &quot;Cape Prov. Albany Distr. 5 m. Hedam, Grahamstown April 1951&quot; (1 ex. AMGS); &quot;EC. Ft Fordyce NR, pond,
<geoCoordinate id="CF5CA519D4C6C4469D5B63DDB7E6BE53" direction="south" orientation="latitude" precision="925" value="-32.666668">32°40'S</geoCoordinate>
,
<geoCoordinate id="5C83E1BEC8CA71AEAD6E1B9812A6013E" direction="east" orientation="longitude" precision="925" value="26.483334">26°29'E</geoCoordinate>
, 1.12. 2009 P. Bulirsch leg.&quot; (2 exs. NMPC); &quot;WC. Witsenberg Valley in ditch 25.5. 2005 Challet&quot; (1 ex. CGC); &quot;E. C. Pr. Cathcart 3.IV. 1955&quot; (1 ex. AMGS); &quot;E. C. Pr. Cradock July 1946&quot; (2 exs. AMGS); &quot;E. C. Pr. Plutos Vlei 20.VII. 1946 JOC.&quot; (1 ex. AMGS); &quot;E. Cape Dam at head of stream, Lang? Barn? Dist. 14.2. 1948 JOC.&quot; (1 ex. AMGS); &quot;E. C. Pr. Glengrey nr Ladyfrere 6.4. 1955&quot; (1 ex. AMGS); &quot;E. C. Pr. Indwe small pond 6.IV. 1955&quot; (1 ex. AMGS); &quot;E C.Pr. Graaf Reinet Nieu Bethseda 28.1.2001 Snizek leg.&quot; (2 exs. NMW, 1 ex. MZH); &quot;E CPr. Bethesda Rd, Graaf Reinet distr. 29.2. 1941 JOC.&quot; (3 exs. AMGS); &quot;ECPr., 34 km SE Idutywa N of Willowvale, shallow pool, S32°12.577, E28°36.192 alt. 447 m, 23.1. 2005 Bergsten&quot; (1 ex. NHRS); &quot;Barkley East dam at head of Lang Kloof 14.2. 1948&quot; (1 ex. AMGS); &quot;Aliwal North 18-25.8. 1954 H. Andreae&quot; (3 exs. SAMC); &quot;C.Pr.Karoo NP, Mountain View River 32°
<geoCoordinate id="533AE4F61C343BDAE63638F8566265EC" direction="south" orientation="latitude" precision="5555" value="-13.6">13.6S</geoCoordinate>
,
<geoCoordinate id="ED41E00482F0F8913E9FDA64055B410D" direction="east" orientation="longitude" precision="92" value="22.526667">22°31.6'E</geoCoordinate>
, 17.11. 1997,
<taxonomicName id="0E8BAE3ADDB91E8125345D1964478000" class="Liliopsida" family="Poaceae" genus="Phragmites" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Phragmites" order="Poales" pageId="180" pageNumber="181" phylum="Tracheophyta" rank="genus">Phragmites</taxonomicName>
grass + litter sievings, 900 m Uhlig, Ndamane et Ari leg.&quot; (2 exs. ZMHB); &quot;C.Pr., Karoo NP, pond + shore 13.11. 1993,
<geoCoordinate id="2636FF8941A17ED7D477E88F45C4F98D" direction="south" orientation="latitude" precision="925" value="-32.316666">32°19'S</geoCoordinate>
,
<geoCoordinate id="06648C2DFC258B8AAC71E9540E0B4907" direction="east" orientation="longitude" precision="925" value="22.5">22°30'E</geoCoordinate>
, Uhlig leg.&quot; (1 ex. ZMHB; habitus in Fig. 479); &quot;C.Pr., Karoo NP,
<geoCoordinate id="9337BCF36EF23EE9B63A86CC4283BFAD" direction="south" orientation="latitude" precision="925" value="-32.316666">32°19'S</geoCoordinate>
,
<geoCoordinate id="F23610457040581B41A73F91C15450A5" direction="east" orientation="longitude" precision="925" value="22.5">22°30'E</geoCoordinate>
, 12-14.11. 1993 Deckert leg.&quot; (1 ex. ZMHB; habitus in Fig. 478); &quot;C.Pr., 20 mi SE Swellendam 3.1. 1951 Brinck-Rudebeck exp.&quot; (6 exs. MZLU); &quot;C.Pr., Albertinia 10.1. 1951 Brinck-Rudebeck Exp.&quot; (4 exs. MZLU); &quot;C.Pr., Zuurberg Pass 15 mi N Addo 16.1. 1951 Brinck-Rudebeck Exp.&quot; (1 ex. MZLU); &quot;C.Pr. Brandvlei 6 mi SW Worchester 11.2. 1951 Brinck-Rudebeck Exp.&quot; (1 ex. MZLU); &quot;C. Pr. nr. Wellington 300 ft. 10.8. 1954 / Deep, dam pool with grass and
<taxonomicName id="A1896134E03A321641484520D8020F13" class="Liliopsida" family="Juncaceae" genus="Juncus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Juncus" order="Poales" pageId="180" pageNumber="181" phylum="Tracheophyta" rank="genus">Juncus</taxonomicName>
edging J. Balfour-Browne leg. /
<taxonomicName id="C778E716E5AFF5853A7466754896812E" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
Shp det. J. Balfour-Browne&quot; (1 ex. BMNH); &quot;C.Pr. Heidelberg 4.1. 1994 Wewalka leg.&quot; (1 ex. NMW); &quot;ECPr., Sundays R. below Graaf Reinet,
<geoCoordinate id="8B9A40B8D60893A351EAF53D99CAEBD3" direction="north" orientation="latitude" precision="55" value="32.27">N-32.270</geoCoordinate>
,
<geoCoordinate id="39742A7F1DDCD1C67BEE4E11EBF56A23" direction="east" orientation="longitude" precision="55" value="24.55">E24.550</geoCoordinate>
, 20.2. 1967 Allanson &amp; Forbes&quot; (1 ex. AMGS); &quot;ECPr., Sundays R. at Letskraal,
<geoCoordinate id="D06A3692F38FE9EE3CB41EDD2D96513B" direction="north" orientation="latitude" precision="55" value="32.06">N-32.060</geoCoordinate>
,
<geoCoordinate id="8F07208AF3B14FF6F4D55215D16B33A9" direction="east" orientation="longitude" precision="55" value="24.83">E24.830</geoCoordinate>
, 14.2. 1968 Allanson &amp; Forbes&quot; (2 exs. AMGS); &quot;ECPr., Ronan Vlei, Kuntwanazana R., N-31.20.16, E28.03.33, 26.3. 1993 De Moor &amp; Barber-James&quot; (2 exs. AMGS); &quot;ECPr., Mncotsho R, N-32.64.48, E27.36.52, 11.8. 2003 De Moor &amp; Barber-James&quot; (1 ex. AMGS); &quot;ECPr., Mncotsho R, Trib. Buffalo R., N-32.54.43, E27.36.48, 18.5. 2004 De Moor &amp; Barber-James&quot; (1 ex. AMGS); &quot;ECPr., Xolo R., small dam, N-32.52.43, E27.37.05, 10.12. 2002, De Moor&quot; (1 ex. AMGS); &quot;Karree Douw Pass, Humansdorp, 17.2. 1947&quot; (1 ex. AMGS); &quot;Ca. 12 mi from Queenstown on road to Lady Frere, ca. 3800 ft., 26.3. 1954 / in small grassy dam / J. Balfour-Browne&quot; (5 exs. BMNH, 1 ex. MZH); &quot;ECPr. ca 5 km N Queenstown, Alan Marsh Prop, small pond, muddish water, S31°48.859, E26°49.715 alt.1208 m 19.1. 2005 Bergsten&quot; (11 exs. NHRS); &quot;ECPr., ca. 5 km N Queenstown, Alan Marsh Prop., muddy pond with grass beds, S31°48.861, E26°50.067, alt. 1232 m, 19.1. 2005
<pageBreakToken id="B42D212CA52CF9BA108586E28AF8FA89" pageId="181" pageNumber="182" start="start">Bergsten</pageBreakToken>
&quot; (17 exs. NHRS); same data but &quot;muddy pond with sand and grass bed, S31°48.887, E26°49.529, alt. 1191 m&quot; (5 exs. NHRS); &quot;ECPr., Ca. 2 km N Queenstown, Longhill Game Reserve, pond with muddy water, S31°51.317, E26°51.322, alt. 1175 m, 19.1. 2005 Bergsten&quot; (3 exs. NHRS); &quot;EC., stream NW Burgersdorp 18.5. 2005 Challet&quot; (1 ex. CGC); &quot;EC., road to Venterstad from Burgersdorp 18.5. 2005 Challet&quot; (2 exs. CGC); &quot;EC., muddy pond So of Burgersdorp 18.5. 2005 Challet&quot; (2 exs. CGC); &quot;Nyebo Stream Transkei 5.4. 1947 JOC.&quot; (1 ex. AMGS). &quot;ECPr. Hogsback
<geoCoordinate id="409E2C3346E1F072F608011B8DC9B283" direction="south" orientation="latitude" precision="555" value="-32.35">32.35S</geoCoordinate>
<geoCoordinate id="20C01FD1F2F1E8E94FC9B4BDBF896576" direction="east" orientation="longitude" precision="555" value="26.56">26.56E</geoCoordinate>
, 6.9. van Noort&quot; (1 ex. SAMC); &quot;ECape prov, King Williams Town 4.4. 1947&quot; (1 ex AMGS); &quot;ECPr., close to Dwesa nature Reserve, muddy pond with vegetation edges S32°18.582, E28°49.002, alt. 76 m, 24-25.1. 2005 Bergsten leg.&quot; (1 ex. NHRS); &quot;EC., 78.7 km So Tarkastadr, rain pond, 21.5. 2005 Challet leg.&quot; (1 ex. CGC); &quot;EC., Gt Brak R., Hwy 56 at Schoombe, 17.5. 2005 Challet leg.&quot; (2 exs. CGC); &quot;EC., Pond 20 km No. Adelaide Hwy 344, 29.8. 2004 Challet&quot; (3 exs. CGC); &quot;EC., stream So of Adelaide 17.5. 2005 Challet&quot; (1 ex. CGC); &quot;ECPr. Farm at Bedford,
<geoCoordinate id="CC4C9815FEDD0100480E01E7A64B2E54" direction="north" orientation="latitude" precision="55" value="32.67">N-32.670</geoCoordinate>
,
<geoCoordinate id="CD0D8ACCC69C099CAD0DF78B7710BD09" direction="east" orientation="longitude" precision="55" value="26.05">E26.050</geoCoordinate>
, 5.12. 1972 Stuart &amp; Greig&quot; (15 exs. AMGS); &quot;ECPr., 3 km N Sterkstroom, Ivan Hansen Prop., small shallow pond, S31°30.337, E26°32.523, alt. 1380 m, 20.1. 2005 Bergsten&quot; (5 exs. NHRS); &quot;ECPr, 2 km N Sterkstroom, Ivan Hansen Prop., small pond with vegetation S31°30.233, E26°32.160, alt 1414 m 20, 28.1. 2005 Bergsten leg.&quot; (5 exs. NHRS); &quot;ECPr. 3 km N Sterkstroom, Ivan Hansen Prop., small, shallow pond, S31°30.337, E26°32.523, alt. 1380 m, 20.1. 2005 Bergsten&quot; (3 exs. NHRS); &quot;ECPr., 2 km N Sterkstroom, Stretton Prop., vegetation rich pond (at sides and in water), S31°29.221, E26°43.984, alt 1533 m, 21.1. 2005 Bergsten&quot; (7 exs. NHRS); &quot;ECPr., N of Sterkstroom, small pond, S31°31.297, E26°32.259, alt.1388 m, 28.1. 2005 Bergsten&quot; (4 exs. NHRS); &quot;ECPr., N of Sterkstroom, Large dam with vegetation at edges, S31°28.571, E26°29.602, alt. 1975 m, 29.1. 2005 Bergsten&quot; (1 ex. NHRS); &quot;EC., pond @HwyR350, No Hellsport Pass 8.12. 1995 Challet&quot; (6 exs. CGC, 1 ex. MZH); &quot;SA NW, CPr., 1 km No Palersheuwel, 28.2. 1997 Challet&quot; (5 exs. CGC, 1 ex. MZH); &quot;EC., rain pond 20 km SO Indutwe 29.8. 2004 Challet&quot; (3 exs. CGC); &quot;ECPr., 7 km E Idutywa, small dirt pool, S32°07.169, E28°22.563, alt. 772 m, 23.1. 2005 Bergsten&quot; (4 exs. NHRS); &quot;ECPr., S31°24.592, E26°54.700, alt. 1745 m, Bradgate Wilson prop., Large Dam, Ca. 15 km W Dordrecht 22.1. 2005 Bergsten&quot; (78 exs. NHRS, 5 exs. MZH); &quot;Cape Col. Victoria West / P.D. Morris coll.&quot; (2 exs. NMW); &quot;Somerset West 28-29.12.1991 Mazzoldi / Marshy area nr Firgrove /
<taxonomicName id="A6266E2C70F46591D3834CAE8248A54C" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
Shap. det. Rocchi&quot; (1 ex. CSR); &quot;WCPr., Somerset West Mall, Azolla pond, 28.3.2005 Reavell&quot; (1 ex. AMGS); &quot;Cape Town&quot; (4 exs. SAMC); &quot;WCape, pond, Cape Good Hope Res. 26.2. 1997 Turner&quot; (2 exs. CCT); &quot;WC, Townsriver rd, N Guydo Pass, Ceres, N-33.10.27, E19.23.28, sandy stream, 3.9. 2003 Turner, Mann &amp; Reavell&quot; (12 exs. CCT); &quot;CPr., pools in riverbed, Williston to Calvinia on R65, Gt Karoo 11.2. 1997 Turner&quot; (5 exs. CCT); &quot;WCp., reservoir on roadside, Bordjilesri, C. G.H. Res., Cape Town 15..2. 1997 Turner&quot; (1 ex. CCT); &quot;WCPr., Zekoevlei Fm, 14 ks S Bremersdorp, Cape
<normalizedToken id="B9AFEEFB7EE886839A4CA7B7D8E20E1D" originalValue="L’Algulhas">L'Algulhas</normalizedToken>
, river, 25.2. 1997 Turner&quot; (1 ex. CCT); &quot;WC, E barrydale on R62, River, slow flowing, silty with sandy rock bottom and emergent veg. N-33.49.05, E20.53.37, 386 m, 4.9. 2003 Turner. Mann &amp; Reavell&quot; (2 exs. CCT); WC. Reservoir, Road from Ouplas to Bredasdorp,
<pageBreakToken id="395C5E1A4D50F4EB4A1FDE164E90371A" pageId="182" pageNumber="183" start="start">N-</pageBreakToken>
34.27.02, E20.10.51, 62 m, 10.9. 2003 Turner. Mann &amp; Reavell&quot; (1 ex, CCT); &quot;WC, sandy bottomed stream N Guydo Pass on Toawsrivier rd., N-33.10.27, E19.23.28, 305 m, 3.9. 2003 Turner. Mann &amp; Reavell&quot; (5 exs. CCT); &quot;WC. De Hoop Res., N-34.27.23, E20.26.19, 13 m, 9.9. 2003, Turner. Mann &amp; Reavell&quot; (63 exs. CCT, 10 exs. MZH); &quot;CPr., Stellenbsch, Fauna Aponogeton Vlei 18.9. 1994 Reavell&quot; (1 ex. CCT); &quot;WCPr., Reservoir on R316 ca 20 km S Caledon N-34.15.86, E19.35.54, 122 m, 10.9. 2003 Turner, Mann, Reavell&quot; (17 exs. CCT); &quot;WCPr. Reservoir, Rd N15 E Worcester, N-33.37.05, E19.21.45, 4.9. 2003 Turner, Mann, Reavell&quot; (1 ex. CCT); &quot;WC., River to E Ashton on Mnt. Rd, N-33.49.35, E20.05.31, 263 m, Turner, Mann, Reavell&quot; (3 exs. CCT); &quot;CP., Cape Francis in ditch, 7.12. 1995 Challet&quot; (1 ex. CGC); &quot;C.Pr. Simons Town 12-20.4. 1915 Cameron leg.&quot; (1 ex. BMNH); &quot;C.P. Grootvaderbos 1-6.11. 1940 G. van Son /
<taxonomicName id="DC80258049757FE84221B6D111927D96" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
Shp det. J. Omer-Cooper&quot; (9 exs. TSMA); &quot;Cape, swampy dam nr Pot R.,
<geoCoordinate id="8CED0D3F1BBE86FA6D2112BB8220490D" direction="north" orientation="latitude" precision="55" value="31.99">N-31.990</geoCoordinate>
,
<geoCoordinate id="BDBAB3470FE04AB421B5ADF3E94CC549" direction="east" orientation="longitude" precision="55" value="28.26">E28.260</geoCoordinate>
, 27.3. 1993 De Moor &amp; al&quot; (2 exs. AMGS); &quot;WCPr., Klein Berg R., N-13.30.03, E18.58.35, 26.4. 1951 Harrison&quot; (2 exs. AMGS); &quot;WCPr., Muizenberg, Pond, N,-33.9505, E, 18.510, 3.1. 1962 M Harrison&quot; (4 exs. AMGS); &quot;Cape, Kroomie R. Ft. Baeufort-Adelaide Rd.,
<geoCoordinate id="828B1C1485F9E2C17A4C20A4A382136B" direction="north" orientation="latitude" precision="555" value="32.77">N-32.77</geoCoordinate>
,
<geoCoordinate id="E7A581ABE7710266B49F03E0CAAFFBB4" direction="east" orientation="longitude" precision="555" value="26.43">E26.43</geoCoordinate>
, 5.12.1992 Stuart &amp; Greig&quot; (5 exs. MZH); &quot;Cape, Gorah Farm Dam,
<geoCoordinate id="760760189A164A1B8EC2514E70FEEE36" direction="north" orientation="latitude" precision="555" value="33.63">N-33,63</geoCoordinate>
,
<geoCoordinate id="EF7B894D13E01364606F51407A66D286" direction="east" orientation="longitude" precision="55" value="26.61">E26.610</geoCoordinate>
, 30.10. 1972 Stuart &amp; Greig&quot; (1 ex. AMGS); &quot;Cape, Wit R at Slagbloom,
<geoCoordinate id="80493ABEA98925A048558D0D378C6AE6" direction="north" orientation="latitude" precision="55" value="33.37">N-33.370</geoCoordinate>
<geoCoordinate id="F37109B074FA78C69A873A8FBEF90FE0" direction="east" orientation="longitude" precision="55" value="25.67">E25.670</geoCoordinate>
, 20.10. 1972 Stobbs&quot; (1 ex. AMGS); &quot;Cape, Stream, 19 km of Brand R Rd.,
<geoCoordinate id="B682BB31F08D6B21FCF6D5E541A8C954" direction="north" orientation="latitude" precision="55" value="33.89">N-33.890</geoCoordinate>
,
<geoCoordinate id="B9FF5CD6241F0EF2DF04DE5ED7E009FA" direction="east" orientation="longitude" precision="55" value="21.06">E21.060</geoCoordinate>
, 30.9.1972 Stobbs&quot; (6 exs. AMGS); &quot;Still Bay C.P. 9-12.11. 1940 G. van Son /
<taxonomicName id="E59DC33CF2354B805F5BACB14FB0A780" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
Sharp det. J. Omer-Cooper&quot; (1 ex. TMSA); &quot;Capland Willowmore Dr. Brauns&quot; (5 exs. TMSA); &quot;Golden Gate G.G.H.N.P. 16-25.1. 1968 Potgieter &amp; Jones&quot; (1 ex. TMSA). - Lesotho: &quot;Teyateyaneng 2.12. 1955 JOC.&quot; (9 exs. AMGS); &quot;Basutoland Maputomy?, small dam 7.6. 1956 G. Ewer&quot; (1 ex. AMGS); &quot;Sani Pass, stream, riffles and pools, 2864 m 1.2. 1997 Turner&quot; (6 exs. CCT).
</paragraph>
</subSubSection>
<subSubSection id="121A2331D8DF2DADE95BC35BF878F08F" pageId="182" pageNumber="183" type="specimens with unknown country association">
<paragraph id="5C77EF0DA5E867A93738504E03BF0B8F" pageId="182" pageNumber="183">Specimens with unknown country association.</paragraph>
<paragraph id="9F5BABA1D1022D3473184DFC2CC5AA89" pageId="182" pageNumber="183">&quot;Africa / J. Hope Coll. B.M. 1948-217&quot; (4 exs. BMNH).</paragraph>
</subSubSection>
<subSubSection id="6535DE0F3E0AF8A4BA683C2FE53F5964" pageId="182" pageNumber="183" type="comments on synonymy">
<paragraph id="88FC7B592DFA25B3C0F3052DB0C250D5" pageId="182" pageNumber="183">Comments on synonymy.</paragraph>
<paragraph id="1023A9DD760AAC70EB00B46534AF002D" pageId="182" pageNumber="183">
Type material of
<taxonomicName id="A01B1A87F79EBB28617E87634F6CE094" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
and
<taxonomicName id="174B1D7C9BEE8A1044B922B9B8B94858" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus shephardi" order="Coleoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="shephardi">Laccophilus shephardi</taxonomicName>
have been examined and compared.
<taxonomicName id="14325AC48C851E8851D2AF4694BF3CBB" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
, being a quite widely distributed species exhibits distinct variation in appearance of dorsal colour pattern of body. Being so, it is evident that
<taxonomicName id="8E37EC6EDBB2302E00882C4EEFBA5C29" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus shephardi" order="Coleoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="shephardi">Laccophilus shephardi</taxonomicName>
represents a colour morph of the species. We have not observed the differences in male genitalia and last abdominal ventrite of male, stressed by Omer-Cooper in the original description of
<taxonomicName id="4A4097A30BBD220196035E2EF1819949" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus shephardi" order="Coleoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="shephardi">Laccophilus shephardi</taxonomicName>
. Accordingly, we consider
<taxonomicName id="AFA6C61AE6D4D926D825A8605BAFAD2F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
and
<taxonomicName id="E1A31B24EEB0D47B9FD7E190620890E0" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus shephardi" order="Coleoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="shephardi">Laccophilus shephardi</taxonomicName>
as synonyms.
<taxonomicName id="EC2B8B4022E9DC03F942E8C87C05352D" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
, which is the older name, is the valid name of this species.
</paragraph>
</subSubSection>
<subSubSection id="13451D531CFCDF2AC10002AB0EA4EFDC" pageId="182" pageNumber="183" type="diagnosis">
<paragraph id="C2E2F0402097CC0F752DFDA93FE373D5" pageId="182" pageNumber="183">Diagnosis.</paragraph>
<paragraph id="FBEC1980E7BA03A9B256C2EEB48FA12E" pageId="182" pageNumber="183">
<taxonomicName id="93CBBC65C10FD8CEED4245C90D59552E" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
is a distinct, although, externally a variable species. It resembles quite much of species exhibiting distinct, dark, longitudinal, elytral markings, e.g. as
<taxonomicName id="64A9C0FF025CD030F1E69D06F14974AF" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus incrassatus" order="Coleoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="incrassatus">Laccophilus incrassatus</taxonomicName>
and
<taxonomicName id="C78B19137A5B4DAAEE87270FA5A3B064" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus quindecimvittatus" order="Coleoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="quindecimvittatus">Laccophilus quindecimvittatus</taxonomicName>
. Longitudinal markings, however, when present, are strongly undulate in
<taxonomicName id="B7FC357CD199DA3D9579F9EF483EA542" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
. In some specimens of
<taxonomicName id="87FF2416394460F5FAC191AADD65DC05" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
elytral dark markings strongly reduced and are lacking. Double elytral reticulation exhibited by
<taxonomicName id="391C7D6074748924B9FC70F855CE3951" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="182" pageNumber="183" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
is also a useful feature at identification because large meshes often strongly reduced in a number of close species, and microsculpture appears in such cases, simple or almost simple.
</paragraph>
</subSubSection>
<subSubSection id="60D90CBC30156066223BBB0C85DA642A" pageId="183" pageNumber="184" type="description">
<paragraph id="8D94A7CC296E157F4372EE562D6C1C40" pageId="183" pageNumber="184">
<pageBreakToken id="41DF2B77CF2EC2874EED148EB4E41021" pageId="183" pageNumber="184" start="start">Description</pageBreakToken>
.
</paragraph>
<paragraph id="A7AF5A3FA89AEB8456D5CD75ECF199C7" pageId="183" pageNumber="184">Body length 3.8-4.8 mm, width 2.2-2.6 mm. Body dorsally with distinct, somewhat variable colour pattern. Dark longitudinal irrorations of elytra rarely strongly reduced (Figs 478-480).</paragraph>
<paragraph id="036A0CC849B0FA2DDD72F792D7810539" pageId="183" pageNumber="184">Head: Pale ferrugineous. At pronotum, generally with vague darker area. Rather shiny, although distinctly microsculptured. Reticulation double; finer meshes often reduced, and hardly visible. At eyes, with minute irregular punctures.</paragraph>
<paragraph id="8B4EC32F637F6A7AA043D645FC5EA36E" pageId="183" pageNumber="184">Pronotum: Pale ferrugineous. Frontally and posteriorly often extensively darker; dark areas sometimes absent. Rather shiny, although distinctly microsculptured. Reticulation double; fine meshes in part indistinct or totally absent. At margins, with fine, scattered punctures.</paragraph>
<paragraph id="31727D12DC5575279428210B6C24A5AC" pageId="183" pageNumber="184">Elytra: Pale ferrugineous with dark ferrugineous, almost uniform, longitudinal irrorations (Fig. 475). Irrorations sometimes reduced (Figs 478-480). Submat, with distinct reticulation which is double. In general, large meshes contain 3-8 fine meshes. Sporadic fine punctures form vague, longitudinal, punctured areas (not distinct rows). Lateral punctate area continues towards apex as a densely punctate and pubescent row.</paragraph>
<paragraph id="2D5AED588CBF9D267FA1B0FBAE92C658" pageId="183" pageNumber="184">Ventral aspect: Pale ferrugineous, metathorax and base of metacoxal plates slightly darker; colour variation vague. Submat, finely microsculptured. Metacoxal plates with some transversely located shallow depressions. Base of metacoxal plates somewhat elevated in comparison with base of abdomen. Abdomen densely striated. Apical ventrite (Figs 147-148); apex bifid and on one side provided with a low knob. Prosternal process long, slender and pointed.</paragraph>
<paragraph id="521F5EA17E5EC508E01AC17319200436" pageId="183" pageNumber="184">Legs: Pro- and mesotarsus long, at base slightly enlarged, narrows gradually towards apex. Segments provided with distinct suckers.</paragraph>
<paragraph id="7E2ADD6CC98DB15F0758F8FA4AFCF523" pageId="183" pageNumber="184">Male genitalia: Penis in lateral aspect broad, from base to apex slightly curved; extreme apex formed as a short, medium broad and apically, sharp extension (Fig. 332).</paragraph>
<paragraph id="B1857CF96EE51D0A9DF0AF56CB45753E" pageId="183" pageNumber="184">Female: Apical ventrite symmetric, lacks knob and apex not bifid (Fig. 149). Pro- and mesotarsus slender.</paragraph>
</subSubSection>
<subSubSection id="97BFCBD5C985C47DF7D425D806B3B78B" pageId="183" pageNumber="184" type="distribution">
<paragraph id="9260CD20773E0CFAE7F4A6D92979FD03" pageId="183" pageNumber="184">Distribution.</paragraph>
<paragraph id="90FC0B83D3882CBFFFF7C16D9291771A" pageId="183" pageNumber="184">
Namibia, Botswana, South Africa, Lesotho (Fig. 564).
<bibRefCitation id="43CB9AE2624FF92669031648DD501AC1" author="Omer-Cooper, Joyce" journalOrPublisher="South African Animal Life" pageId="261" pageNumber="262" pagination="59 - 214" title="Chapter 2. Coleoptera: Dytiscidae." volume="11" year="1965">Omer-Cooper (1965)</bibRefCitation>
adds Swaziland. Records outside southern Africa are considered doubtful.
</paragraph>
</subSubSection>
<subSubSection id="75E31DB6C796BFB9C29A6ADABE4F68CE" pageId="183" pageNumber="184" type="collecting circumstances">
<paragraph id="9BFC232528E8CF41B48DF79C30AC70E2" pageId="183" pageNumber="184">Collecting circumstances.</paragraph>
<paragraph id="E4CD72C815A795BA21022694B09B93C2" pageId="183" pageNumber="184">
According to label data, the species has been sampled in dams, pools, water-holes etc., with various, often thick vegetation of weed, algae, grass,
<taxonomicName id="57FD1B2953217850238D37088050CA22" class="Liliopsida" family="Juncaceae" genus="Juncus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Juncus" order="Poales" pageId="183" pageNumber="184" phylum="Tracheophyta" rank="genus">Juncus</taxonomicName>
and
<taxonomicName id="E066A4D80EAF8794B6812682037EFBBB" class="Liliopsida" family="Araceae" genus="Lemna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lemna" order="Alismatales" pageId="183" pageNumber="184" phylum="Tracheophyta" rank="genus">Lemna</taxonomicName>
. Also collected at light.
<bibRefCitation id="F31830107A42A6767C677E4E688F2490" author="Omer-Cooper, Joyce" journalOrPublisher="South African Animal Life" pageId="261" pageNumber="262" pagination="59 - 214" title="Chapter 2. Coleoptera: Dytiscidae." volume="11" year="1965">Omer-Cooper (1965)</bibRefCitation>
reports the species as occurring &quot;in every type of fresh water habitat&quot;.
</paragraph>
</subSubSection>
</treatment>
</document>
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