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<document id="65816C0BD83433D7EBB619B6B6ACE3C1" ID-CLB-Dataset="54304" ID-DOI="10.5852/ejt.2023.888.2207" ID-GBIF-Dataset="f7b20254-7a43-4153-b471-721bb1a77e53" ID-ISSN="2118-9773" ID-Zenodo-Dep="8246747" ID-ZooBank="76AE9DED-146D-49D0-96E8-4B2A74412F20" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1692021898274" checkinUser="felipe" docAuthor="Gnezdilov, Vladimir M. &amp; Malenovský, Igor" docDate="2023" docId="455A87DB446AFFA22C46236F8AECFA0E" docLanguage="en" docName="ejt-888_46-63.pdf" docOrigin="European Journal of Taxonomy 888 (1)" docSource="http://dx.doi.org/10.5852/ejt.2023.888.2207" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Oceatra litoralis Gnezdilov &amp; Malenovský 2023, gen. et sp. nov." docType="treatment" docUuid="C3F31EA9-7579-425A-BA43-C2A916A36F78" docUuidSource="ZooBank" docVersion="6" lastPageNumber="52" masterDocId="B963FFA34469FFA42E1A27558B25FFE1" masterDocTitle="First record of the planthopper tribe Adenissini (Hemiptera: Caliscelidae) from Socotra Island, with description of a new genus and two new species" masterLastPageNumber="63" masterPageNumber="46" pageNumber="49" updateTime="1699557549247" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0E2704E42603C7712A23D4EB5E1FE52E">First record of the planthopper tribe Adenissini (Hemiptera: Caliscelidae) from Socotra Island, with description of a new genus and two new species</mods:title>
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<mods:namePart id="E4DC8A83CBF1E093DCF63D3247B41710">Gnezdilov, Vladimir M.</mods:namePart>
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<mods:affiliation id="3FBF00C9FA7733853934A7BF74A8315D">Zoological Institute, Russian Academy of Sciences, 1 Universitetskaya Emb., Saint Petersburg 199034, Russia.</mods:affiliation>
<mods:nameIdentifier id="4F8504984E0E1B8B620B68F6E1B7CFC1" type="email">vmgnezdilov@mail.ru,vgnezdilov@zin.ru</mods:nameIdentifier>
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<mods:namePart id="39C96A633755D1E971FFA3BF3B32BAEB">Malenovský, Igor</mods:namePart>
<mods:nameIdentifier id="A29F3EE7A65A715A0C8A8F1FC8318D80" type="LSID">8AF430DB-BBBD-4F48-92CA-259290924F71</mods:nameIdentifier>
<mods:affiliation id="3168712C07458AB58AD20F0390207649">Department of Botany and Zoology, Faculty of Science, Masaryk University, Kotlářská 2, CZ- 611 37 Brno, Czech Republic. &amp; Department of Entomology, Moravian Museum, Zelný trh 6, CZ- 659 37 Brno, Czech Republic.</mods:affiliation>
<mods:nameIdentifier id="73ABF1B4DF6629B2E5C7201158298D3A" type="email">malenovsky@sci.muni.cz,imalenovsky@mzm.cz</mods:nameIdentifier>
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<mods:title id="66846FBF9C7994BCD67165D7E65E3CAC">European Journal of Taxonomy</mods:title>
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<mods:date id="0C276DEFA95767C3B15EFBD441D1C1B8">2023</mods:date>
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<mods:number id="49A6C8157C482DB4A092162EAECAF939">2023-08-08</mods:number>
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<treatment id="455A87DB446AFFA22C46236F8AECFA0E" ID-DOI="http://doi.org/10.5281/zenodo.8246743" ID-GBIF-Taxon="212429543" ID-Zenodo-Dep="8246743" LSID="urn:lsid:zoobank.org:act:C3F31EA9-7579-425A-BA43-C2A916A36F78" httpUri="http://treatment.plazi.org/id/455A87DB446AFFA22C46236F8AECFA0E" lastPageId="6" lastPageNumber="52" pageId="3" pageNumber="49">
<subSubSection id="85E96546446AFFA72C46236F88FDFBB4" box="[604,984,1082,1109]" pageId="3" pageNumber="49" type="nomenclature">
<paragraph id="CD4C36CD446AFFA72C46236F88FDFBB4" blockId="3.[366,1222,1082,1180]" box="[604,984,1082,1109]" pageId="3" pageNumber="49">
<heading id="960481A1446AFFA72C46236F88FDFBB4" box="[604,984,1082,1109]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="49" reason="2">
<taxonomicName id="0AF34D4E446AFFA72C46236F8805FBB4" authority="Gnezdilov &amp; Malenovský, 2023" authorityName="Gnezdilov &amp; Malenovský" authorityYear="2023" box="[604,800,1082,1109]" class="Insecta" family="Caliscelidae" genus="Oceatra" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="litoralis" status="gen. et sp. nov.">
<emphasis id="FF87EADF446AFFA72C46236F8805FBB4" bold="true" box="[604,800,1082,1109]" italics="true" pageId="3" pageNumber="49">Oceatra litoralis</emphasis>
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<taxonomicNameLabel id="E4B457A4446AFFA72D32236E88FDFBB4" box="[808,984,1083,1109]" pageId="3" pageNumber="49" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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<uri id="B9623ACF446AFFA72F7423088FE3FB99" box="[366,1222,1117,1144]" pageId="3" pageNumber="49">
urn:lsid:zoobank.org:act:
<uuid id="B9550C18446AFFA72C9723088FE3FB99" box="[653,1222,1117,1144]" pageId="3" pageNumber="49">C3F31EA9-7579-425A-BA43-C2A916A36F78</uuid>
</uri>
</paragraph>
<paragraph id="CD4C36CD446AFFA72C6F23D4889BFB7A" blockId="3.[366,1222,1082,1180]" box="[629,958,1153,1180]" pageId="3" pageNumber="49">
<figureCitation id="55C82A48446AFFA72C6F23D489FBFB7A" box="[629,734,1153,1179]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="4.[189,232,1963,1989]" captionStartId-1="5.[189,232,1963,1989]" captionTargetBox-0="[204,1384,718,1909]" captionTargetBox-1="[189,1398,747,1903]" captionTargetId-0="figure-173@4.[189,1398,703,1924]" captionTargetId-1="figure-170@5.[189,1398,747,1923]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Fig. 1. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC). A. Dorsal view. B. Lateral view. C. Frontal view. D. Genital block, caudal view." captionText-1="Fig. 2. Oceatra litoralis gen. et sp. nov., paratype, ♀ (MMBC). A. Dorsal view. B. Frontal view. C. Terminalia, ventral view. D. Terminalia, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.8246749" figureDoi-1="http://doi.org/10.5281/zenodo.8246751" httpUri-0="https://zenodo.org/record/8246749/files/figure.png" httpUri-1="https://zenodo.org/record/8246751/files/figure.png" pageId="3" pageNumber="49">Figs 1–2</figureCitation>
,
<figureCitation id="55C82A48446AFFA72CF023D48814FB7A" box="[746,817,1153,1179]" captionStart="Fig" captionStartId="7.[189,232,1960,1986]" captionTargetBox="[304,1287,1319,1898]" captionTargetId="figure-39@7.[280,1308,1316,1918]" captionTargetPageId="7" captionText="Fig. 4. Adenissina Dlabola, 1980, forewings. A. Perissana dlabolai Gnezdilov &amp; Wilson, 2006, ♂. B–C. Oceatra litoralis gen. et sp. nov., paratype, ♂ (ZIN). B. Left wing. C. Right wing. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246755" httpUri="https://zenodo.org/record/8246755/files/figure.png" pageId="3" pageNumber="49">4B–C</figureCitation>
,
<figureCitation id="55C82A48446AFFA72D2723D488A3FB7D" box="[829,902,1153,1180]" captionStart="Fig" captionStartId="8.[191,234,1928,1954]" captionTargetBox="[216,1375,950,1853]" captionTargetId="figure-197@8.[189,1398,948,1886]" captionTargetPageId="8" captionText="Fig. 5. A–C. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC). D–E. Oceatra scandens gen. et sp. nov., holotype, ♂ (MMBC). A, D. Head and pronotum, dorsal view. B, E. Head, frontal view. C. Rostrum, lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246759" httpUri="https://zenodo.org/record/8246759/files/figure.png" pageId="3" pageNumber="49">5A–C</figureCitation>
,
<figureCitation id="55C82A48446AFFA72D8823D4889BFB7A" box="[914,958,1153,1179]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="9.[189,232,1858,1884]" captionStartId-1="10.[189,232,1963,1989]" captionTargetBox-0="[220,1339,277,1790]" captionTargetBox-1="[208,1395,1401,1904]" captionTargetId-0="figure-14@9.[189,1398,266,1810]" captionTargetId-1="figure-439@10.[189,1398,1368,1923]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Fig. 6. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC), terminalia. A. Genital block, lateral view. B. Pygofer, lateral view. C. Anal tube, dorsal view. D. Style, lateral view. E. Style, dorsal view. F. Penis, connective, and anal tube, lateral view. G. Penis, ventral view. H. Penis and connective, dorsal view. I. Apex of penis, dorsal view. Abbreviations: aep = apical aedeagal process; mot = mushroomshaped outgrowth of phallobase; vh = ventral aedeagal hooks; vl = ventral phallobase lobe. Not to scale." captionText-1="Fig. 7. Oceatra litoralis gen. et sp. nov., paratype, ♀ (MMBC), terminalia. A. Pygofer and anal tube, dorsal view. B. Genital block, lateral view. C. Sternite VII, ventral view. Not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.8246761" figureDoi-1="http://doi.org/10.5281/zenodo.8246763" httpUri-0="https://zenodo.org/record/8246761/files/figure.png" httpUri-1="https://zenodo.org/record/8246763/files/figure.png" pageId="3" pageNumber="49">6–7</figureCitation>
</paragraph>
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<subSubSection id="85E96546446AFFA72EA72397888AFAEA" box="[189,943,1218,1291]" pageId="3" pageNumber="49" type="diagnosis">
<paragraph id="CD4C36CD446AFFA72EA723978A13FB3D" blockId="3.[189,310,1218,1244]" box="[189,310,1218,1244]" pageId="3" pageNumber="49">
<heading id="960481A1446AFFA72EA723978A13FB3D" bold="true" box="[189,310,1218,1244]" fontSize="11" level="3" pageId="3" pageNumber="49" reason="3">
<emphasis id="FF87EADF446AFFA72EA723978A13FB3D" bold="true" box="[189,310,1218,1244]" pageId="3" pageNumber="49">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="CD4C36CD446AFFA72EA723A4888AFAEA" blockId="3.[189,943,1265,1291]" box="[189,943,1265,1291]" pageId="3" pageNumber="49">
See the identification key to species of
<taxonomicName id="0AF34D4E446AFFA72C9223A489CDFAEA" authority="Gnezdilov &amp; Malenovský, 2023" authorityName="Gnezdilov &amp; Malenovský" authorityYear="2023" box="[648,744,1265,1291]" class="Insecta" family="Caliscelidae" genus="Oceatra" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="49" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="FF87EADF446AFFA72C9223A489CDFAEA" box="[648,744,1265,1291]" italics="true" pageId="3" pageNumber="49">Oceatra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4B457A4446AFFA72CF523A4887FFAEA" box="[751,858,1265,1291]" pageId="3" pageNumber="49" rank="genus">gen. nov.</taxonomicNameLabel>
above.
</paragraph>
</subSubSection>
<subSubSection id="85E96546446AFFA72EA722678970FA7F" pageId="3" pageNumber="49" type="etymology">
<paragraph id="CD4C36CD446AFFA72EA722678A61FAAD" blockId="3.[189,324,1330,1356]" box="[189,324,1330,1356]" pageId="3" pageNumber="49">
<heading id="960481A1446AFFA72EA722678A61FAAD" bold="true" box="[189,324,1330,1356]" fontSize="11" level="3" pageId="3" pageNumber="49" reason="3">
<emphasis id="FF87EADF446AFFA72EA722678A61FAAD" bold="true" box="[189,324,1330,1356]" pageId="3" pageNumber="49">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="CD4C36CD446AFFA72EA722348970FA7F" blockId="3.[189,1399,1377,1438]" pageId="3" pageNumber="49">
From the Latin adjective ‘
<taxonomicName id="0AF34D4E446AFFA72FE222348970FA9A" authorityName="Gnezdilov &amp; Malenovský" authorityYear="2023" box="[504,597,1377,1403]" class="Insecta" family="Caliscelidae" genus="Oceatra" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="litoralis">
<emphasis id="FF87EADF446AFFA72FE222348970FA9A" box="[504,597,1377,1403]" italics="true" pageId="3" pageNumber="49">litoralis</emphasis>
</taxonomicName>
’ (= ‘of the seashore’, ‘littoral’), referring to the habitat of the new species in the coastal salt marshes.
</paragraph>
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<subSubSection id="85E96546446AFFA72EA72290894DF8E1" pageId="3" pageNumber="49" type="materials_examined">
<paragraph id="CD4C36CD446AFFA72EA722908A49FA3E" blockId="3.[189,364,1477,1503]" box="[189,364,1477,1503]" pageId="3" pageNumber="49">
<heading id="960481A1446AFFA72EA722908A49FA3E" bold="true" box="[189,364,1477,1503]" fontSize="11" level="3" pageId="3" pageNumber="49" reason="3">
<emphasis id="FF87EADF446AFFA72EA722908A49FA3E" bold="true" box="[189,364,1477,1503]" pageId="3" pageNumber="49">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="CD4C36CD446AFFA72EF022A18F8FF999" blockId="3.[189,1398,1524,1656]" pageId="3" pageNumber="49">
<materialsCitation id="7D9B3C90446AFFA72EF022A18F8FF999" ID-GBIF-Occurrence="4173763301" collectedFrom="Avicennia marina mangrove, salt marsh and sand dunes" collectingDate="2012-06-20" collectingDateMax="2012-06-21" collectingDateMin="2012-06-20" collectingMethod="sweeping and suction" collectionCode="MMBC" collectorName="I. Malenovsky &amp; P. Kment &amp; J. Bezdek &amp; J. Hajek &amp; V. Hula &amp; J. Niedobova &amp; L. Purchart" country="Yemen" elevation="3" latitude="12.568334" location="Socotra Island" longLatPrecision="129" longitude="53.398335" municipality="Socotra Island" pageId="3" pageNumber="49" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="FF87EADF446AFFA72EF022A18A79F9EF" bold="true" box="[234,348,1524,1550]" pageId="3" pageNumber="49">
<typeStatus id="1248886F446AFFA72EF022A18A79F9EF" box="[234,348,1524,1550]" pageId="3" pageNumber="49" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="B5E4765D446AFFA72EA7214D8A00F9D3" box="[189,293,1560,1586]" name="Yemen" pageId="3" pageNumber="49">YEMEN</collectingCountry>
•
<specimenCount id="DBF5FD44446AFFA72F27214D8A73F9D3" box="[317,342,1560,1586]" pageId="3" pageNumber="49" type="male">♂</specimenCount>
;
<collectingMunicipality id="2D28ACB7446AFFA72F7821428928F9D0" box="[354,525,1559,1585]" country="Yemen" latitude="12.568334" longLatPrecision="129" longitude="53.398335" name="Socotra Island" pageId="3" pageNumber="49">Socotra Island</collectingMunicipality>
, ca
<locationDeviation id="DCFE2FEF446AFFA72C2221428836F9D0" box="[568,787,1559,1586]" location="Shuab" pageId="3" pageNumber="49">
<quantity id="0A0B9B28446AFFA72C2221428957F9D0" box="[568,626,1559,1585]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="49" unit="km" value="3.0">3 km</quantity>
NE of Shuab
</locationDeviation>
;
<geoCoordinate id="A8C7500A446AFFA72D0421408885F9D3" box="[798,928,1557,1586]" degrees="12" direction="north" minutes="34.1" orientation="latitude" pageId="3" pageNumber="49" precision="92" value="12.568334">12°34.1′ N</geoCoordinate>
,
<geoCoordinate id="A8C7500A446AFFA72DB121408F0CF9D3" box="[939,1065,1557,1586]" degrees="53" direction="east" minutes="23.9" orientation="longitude" pageId="3" pageNumber="49" precision="92" value="53.398335">53°23.9′ E</geoCoordinate>
;
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a.s.l.
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;
<date id="B94D100D446AFFA72AB721428E57F9D0" box="[1197,1394,1559,1586]" pageId="3" pageNumber="49" value="2012-06-20" valueMax="2012-06-21" valueMin="2012-06-20">
<collectingDate id="A909E9E5446AFFA72AB721428E57F9D0" box="[1197,1394,1559,1586]" pageId="3" pageNumber="49" value="2012-06-20" valueMax="2012-06-21" valueMin="2012-06-20">20–21 Jun. 2012</collectingDate>
</date>
;
<collectorName id="6006531B446AFFA72EA7216E8A4CF9B5" box="[189,361,1594,1621]" pageId="3" pageNumber="49">I. Malenovský</collectorName>
,
<collectorName id="6006531B446AFFA72F6E216E8AC7F9B4" box="[372,482,1595,1621]" pageId="3" pageNumber="49">P. Kment</collectorName>
,
<collectorName id="6006531B446AFFA72FF7216E8947F9B5" box="[493,610,1594,1621]" pageId="3" pageNumber="49">J. Bezděk</collectorName>
,
<collectorName id="6006531B446AFFA72C77216E89EAF9B5" box="[621,719,1594,1621]" pageId="3" pageNumber="49">J. Hájek</collectorName>
,
<collectorName id="6006531B446AFFA72CC0216E8810F9B5" box="[730,821,1594,1621]" pageId="3" pageNumber="49">V. Hula</collectorName>
,
<collectorName id="6006531B446AFFA72D5A216E88FEF9B5" box="[832,987,1594,1621]" pageId="3" pageNumber="49">J. Niedobová</collectorName>
and
<collectorName id="6006531B446AFFA72A08216E8FB0F9B5" box="[1042,1173,1594,1621]" pageId="3" pageNumber="49">L. Purchart</collectorName>
leg.;
<collectingMethod id="14B24EDA446AFFA72ACD216F8A37F996" pageId="3" pageNumber="49">sweeping and suction</collectingMethod>
sampling in
<collectedFrom id="7EEF7530446AFFA72FB0210B8F1EF996" box="[426,1083,1629,1656]" pageId="3" pageNumber="49">
<taxonomicName id="0AF34D4E446AFFA72FB0210B895FF996" box="[426,634,1630,1656]" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="49" phylum="Tracheophyta" rank="species" species="marina">
<emphasis id="FF87EADF446AFFA72FB0210B895FF996" box="[426,634,1630,1656]" italics="true" pageId="3" pageNumber="49">Avicennia marina</emphasis>
</taxonomicName>
mangrove, salt marsh and sand dunes
</collectedFrom>
;
<collectionCode id="ABE2AE08446AFFA72A5D210B8F83F999" box="[1095,1190,1630,1656]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="3" pageNumber="49">MMBC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="CD4C36CD446AFFA72EF021CA894DF8E1" blockId="3.[189,1398,1695,1792]" pageId="3" pageNumber="49">
<materialsCitation id="7D9B3C90446AFFA72EF021CA8FB6F93D" ID-GBIF-Occurrence="4173763302" collectedFrom="Avicennia marina mangrove, salt marsh and sand dunes" collectingDate="2012-06-20" collectingDateMax="2012-06-21" collectingDateMin="2012-06-20" collectingMethod="sweeping and suction" collectionCode="MMBC" collectorName="I. Malenovsky &amp; P. Kment &amp; J. Bezdek &amp; J. Hajek &amp; V. Hula &amp; J. Niedobova &amp; L. Purchart" country="Yemen" elevation="3" latitude="12.568334" location="Socotra Island" longLatPrecision="129" longitude="53.398335" municipality="Socotra Island" pageId="3" pageNumber="49" specimenCount="1" specimenCount-female="1" stateProvince="Socotra Island" typeStatus="paratype">
<emphasis id="FF87EADF446AFFA72EF021CA8A42F958" bold="true" box="[234,359,1695,1721]" pageId="3" pageNumber="49">
<typeStatus id="1248886F446AFFA72EF021CA8A42F958" box="[234,359,1695,1721]" pageId="3" pageNumber="49" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="B5E4765D446AFFA72EA721968A00F93C" box="[189,293,1731,1757]" name="Yemen" pageId="3" pageNumber="49">YEMEN</collectingCountry>
–
<emphasis id="FF87EADF446AFFA72F5121978921F93D" bold="true" box="[331,516,1730,1756]" pageId="3" pageNumber="49">
<collectingRegion id="0F37F82F446AFFA72F5121978921F93D" box="[331,516,1730,1756]" pageId="3" pageNumber="49">Socotra Island</collectingRegion>
</emphasis>
•
<specimenCount id="DBF5FD44446AFFA72C3F21978972F93C" box="[549,599,1730,1757]" pageId="3" pageNumber="49" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="ABE2AE08446AFFA72A2221978FB6F93D" box="[1080,1171,1730,1756]" collectionName="Czech Republic, Brno, Moravske Muzeum [Moravian Museum]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="3" pageNumber="49">MMBC</collectionCode>
</materialsCitation>
•
<materialsCitation id="7D9B3C90446AFFA72AAE21978940F8E1" ID-GBIF-Occurrence="4173763303" collectedFrom="Avicennia marina mangrove, salt marsh and sand dunes" collectingDate="2012-06-20" collectingDateMax="2012-06-21" collectingDateMin="2012-06-20" collectingMethod="sweeping and suction" collectionCode="ZIN" collectorName="I. Malenovsky &amp; P. Kment &amp; J. Bezdek &amp; J. Hajek &amp; V. Hula &amp; J. Niedobova &amp; L. Purchart" country="Yemen" elevation="3" latitude="12.568334" location="Socotra Island" longLatPrecision="129" longitude="53.398335" municipality="Socotra Island" pageId="3" pageNumber="49" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="DBF5FD44446AFFA72AAE21978FC3F93C" box="[1204,1254,1730,1757]" pageId="3" pageNumber="49" type="male">1 ♂</specimenCount>
,
<specimenCount id="DBF5FD44446AFFA72AED21978E0DF93C" box="[1271,1320,1730,1757]" pageId="3" pageNumber="49" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="ABE2AE08446AFFA72C2A21B38940F8E1" box="[560,613,1766,1792]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3" pageNumber="49" type="Museum">ZIN</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="85E96546446AFFA22EA720738888FB68" lastPageId="6" lastPageNumber="52" pageId="3" pageNumber="49" type="description">
<paragraph id="CD4C36CD446AFFA72EA720738A6BF8A1" blockId="3.[189,334,1830,1856]" box="[189,334,1830,1856]" pageId="3" pageNumber="49">
<heading id="960481A1446AFFA72EA720738A6BF8A1" bold="true" box="[189,334,1830,1856]" fontSize="11" level="3" pageId="3" pageNumber="49" reason="3">
<emphasis id="FF87EADF446AFFA72EA720738A6BF8A1" bold="true" box="[189,334,1830,1856]" pageId="3" pageNumber="49">Description</emphasis>
</heading>
</paragraph>
<paragraph id="CD4C36CD446AFFA72EA7200388DAF891" blockId="3.[189,1023,1877,1904]" box="[189,1023,1877,1904]" pageId="3" pageNumber="49">
<smallCapsWord id="CBAAA011446AFFA72EA720038A4BF88F" baselines="1897,1897" box="[189,366,1878,1904]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="3" pageNumber="49">MEASUREMENTS</smallCapsWord>
. Total body length: males – 3.0 mm, females – 4.0 mm.
</paragraph>
<paragraph id="CD4C36CD446AFFA12EA720C28A70FEA7" blockId="3.[187,1398,1943,2004]" lastBlockId="5.[189,1399,265,326]" lastPageId="5" lastPageNumber="51" pageId="3" pageNumber="49">
<smallCapsWord id="CBAAA011446AFFA72EA720C28A6DF84E" baselines="1962,1962" box="[189,328,1943,1969]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coloration" pageId="3" pageNumber="49">COLORATION</smallCapsWord>
. Male (
<figureCitation id="55C82A48446AFFA72F8520C28AC6F850" box="[415,483,1943,1969]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[204,1384,718,1909]" captionTargetId="figure-173@4.[189,1398,703,1924]" captionTargetPageId="4" captionText="Fig. 1. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC). A. Dorsal view. B. Lateral view. C. Frontal view. D. Genital block, caudal view." figureDoi="http://doi.org/10.5281/zenodo.8246749" httpUri="https://zenodo.org/record/8246749/files/figure.png" pageId="3" pageNumber="49">Fig. 1</figureCitation>
). General coloration light ochreous. Upper half of metope dark brown to black, carinae light ochreous (
<figureCitation id="55C82A48446AFFA72FD120EF8903F835" box="[459,550,1978,2004]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[204,1384,718,1909]" captionTargetId="figure-173@4.[189,1398,703,1924]" captionTargetPageId="4" captionText="Fig. 1. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC). A. Dorsal view. B. Lateral view. C. Frontal view. D. Genital block, caudal view." figureDoi="http://doi.org/10.5281/zenodo.8246749" httpUri="https://zenodo.org/record/8246749/files/figure.png" pageId="3" pageNumber="49">Fig. 1C</figureCitation>
). Postclypeus with dark brown dots and spots laterally. Anteclypeus dark brown. Rostrum dark brown ventrally, black apically. Pedicel brown to dark brown. Coryphe with dark brown dots (
<figureCitation id="55C82A48446DFFA02F4826798A8AFEA7" box="[338,431,300,326]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[204,1384,718,1909]" captionTargetId="figure-173@4.[189,1398,703,1924]" captionTargetPageId="4" captionText="Fig. 1. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC). A. Dorsal view. B. Lateral view. C. Frontal view. D. Genital block, caudal view." figureDoi="http://doi.org/10.5281/zenodo.8246749" httpUri="https://zenodo.org/record/8246749/files/figure.png" pageId="4" pageNumber="50">Fig. 1A</figureCitation>
). Preocular fields dark brown. Paranotal lobes of pronotum with dark brown dots. Mesonotum with dark brown spot in each anterior angle. Forewing with dark brown pattern concentrated along outer clavus margin between CuP and Pcu and in cells of corium in its apical half (
<figureCitation id="55C82A48446DFFA02AD326278E01FE6C" box="[1225,1316,370,397]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[204,1384,718,1909]" captionTargetId="figure-173@4.[189,1398,703,1924]" captionTargetPageId="4" captionText="Fig. 1. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC). A. Dorsal view. B. Lateral view. C. Frontal view. D. Genital block, caudal view." figureDoi="http://doi.org/10.5281/zenodo.8246749" httpUri="https://zenodo.org/record/8246749/files/figure.png" pageId="4" pageNumber="50">Fig. 1B</figureCitation>
). Hind coxae brown posteriorly. Femora and tibiae with dark brown bands. Third tarsomeres with dark brown apical halves. Claws and dorso-lateral plates of pretarsus dark brown. Apices of spines on legs black. Each laterotergite with large dark brown to black spot. Style light yellow, with dark brown capitulum. Penis dark brown (
<figureCitation id="55C82A48446DFFA02FBE26AA8921FDF8" box="[420,516,511,537]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[204,1384,718,1909]" captionTargetId="figure-173@4.[189,1398,703,1924]" captionTargetPageId="4" captionText="Fig. 1. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC). A. Dorsal view. B. Lateral view. C. Frontal view. D. Genital block, caudal view." figureDoi="http://doi.org/10.5281/zenodo.8246749" httpUri="https://zenodo.org/record/8246749/files/figure.png" pageId="4" pageNumber="50">Fig. 1D</figureCitation>
). Female (
<figureCitation id="55C82A48446DFFA02C9F26AA89EAFDF8" box="[645,719,511,537]" captionStart="Fig" captionStartId="5.[189,232,1963,1989]" captionTargetBox="[189,1398,747,1903]" captionTargetId="figure-170@5.[189,1398,747,1923]" captionTargetPageId="5" captionText="Fig. 2. Oceatra litoralis gen. et sp. nov., paratype, ♀ (MMBC). A. Dorsal view. B. Frontal view. C. Terminalia, ventral view. D. Terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8246751" httpUri="https://zenodo.org/record/8246751/files/figure.png" pageId="4" pageNumber="50">Fig. 2</figureCitation>
) similar to male. Forewing with dark brown cells along outer clavus margin and in posterior half of corium (
<figureCitation id="55C82A48446DFFA02D5A25778885FDDC" box="[832,928,546,573]" captionStart="Fig" captionStartId="5.[189,232,1963,1989]" captionTargetBox="[189,1398,747,1903]" captionTargetId="figure-170@5.[189,1398,747,1923]" captionTargetPageId="5" captionText="Fig. 2. Oceatra litoralis gen. et sp. nov., paratype, ♀ (MMBC). A. Dorsal view. B. Frontal view. C. Terminalia, ventral view. D. Terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8246751" httpUri="https://zenodo.org/record/8246751/files/figure.png" pageId="4" pageNumber="50">Fig. 2A</figureCitation>
) or only with sparse dark brown spots scattered across whole wing. Visible abdominal tergites VI–VIII with dark brown spots. Gonoplacs brown (
<figureCitation id="55C82A48446CFFA12F00265C8ABAFEC2" box="[282,415,265,291]" captionStart="Fig" captionStartId="5.[189,232,1963,1989]" captionTargetBox="[189,1398,747,1903]" captionTargetId="figure-170@5.[189,1398,747,1923]" captionTargetPageId="5" captionText="Fig. 2. Oceatra litoralis gen. et sp. nov., paratype, ♀ (MMBC). A. Dorsal view. B. Frontal view. C. Terminalia, ventral view. D. Terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8246751" httpUri="https://zenodo.org/record/8246751/files/figure.png" pageId="5" pageNumber="51">Fig. 2C–D</figureCitation>
). Anal tube with black median stripe running and expanding from paraproct to its hind margin.
</paragraph>
<caption id="998C6645446DFFA02EA720FE89C2F809" ID-DOI="http://doi.org/10.5281/zenodo.8246749" ID-Zenodo-Dep="8246749" httpUri="https://zenodo.org/record/8246749/files/figure.png" pageId="4" pageNumber="50" startId="4.[189,232,1963,1989]" targetBox="[204,1384,718,1909]" targetPageId="4">
<paragraph id="CD4C36CD446DFFA02EA720FE89C2F809" blockId="4.[189,1399,1963,2025]" pageId="4" pageNumber="50">
<emphasis id="FF87EADF446DFFA02EA720FE8A35F824" bold="true" box="[189,272,1963,1989]" pageId="4" pageNumber="50">Fig. 1.</emphasis>
<taxonomicName id="0AF34D4E446DFFA02F0720F98AC3F824" authority="Gnezdilov &amp; Malenovský, 2023" authorityName="Gnezdilov &amp; Malenovský" authorityYear="2023" box="[285,486,1963,1990]" class="Insecta" family="Caliscelidae" genus="Oceatra" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="litoralis" status="gen. et sp. nov.">
<emphasis id="FF87EADF446DFFA02F0720F98AC3F824" box="[285,486,1963,1990]" italics="true" pageId="4" pageNumber="50">Oceatra litoralis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4B457A4446DFFA02FE820F98991F827" box="[498,692,1964,1990]" pageId="4" pageNumber="50" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype, ♂ (MMBC).
<emphasis id="FF87EADF446DFFA02DF920FE88DDF824" bold="true" box="[995,1016,1963,1989]" pageId="4" pageNumber="50">A</emphasis>
. Dorsal view.
<emphasis id="FF87EADF446DFFA02AAB20FE8FE0F824" bold="true" box="[1201,1221,1963,1989]" pageId="4" pageNumber="50">B</emphasis>
. Lateral view.
<emphasis id="FF87EADF446DFFA02EA7209B8BF7F809" bold="true" box="[189,210,1998,2024]" pageId="4" pageNumber="50">C</emphasis>
. Frontal view.
<emphasis id="FF87EADF446DFFA02F99209B8ABDF809" bold="true" box="[387,408,1998,2024]" pageId="4" pageNumber="50">D</emphasis>
. Genital block, caudal view.
</paragraph>
</caption>
<paragraph id="CD4C36CD446CFFA22EA7263B8AD2FDFB" blockId="5.[187,1400,365,674]" lastBlockId="6.[188,1400,265,538]" lastPageId="6" lastPageNumber="52" pageId="5" pageNumber="51">
<smallCapsWord id="CBAAA011446CFFA12EA7263B8A15FE67" baselines="385,385" box="[189,304,366,392]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="External" pageId="5" pageNumber="51">EXTERNAL</smallCapsWord>
<smallCapsWord id="CBAAA011446CFFA12F2326268A97FE67" baselines="385" box="[313,434,371,390]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="structure" pageId="5" pageNumber="51">STRUCTURE</smallCapsWord>
. Metope (
<figureCitation id="55C82A48446CFFA12C37263B89AAFE69" box="[557,655,366,392]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[204,1384,718,1909]" captionTargetId="figure-173@4.[189,1398,703,1924]" captionTargetPageId="4" captionText="Fig. 1. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC). A. Dorsal view. B. Lateral view. C. Frontal view. D. Genital block, caudal view." figureDoi="http://doi.org/10.5281/zenodo.8246749" httpUri="https://zenodo.org/record/8246749/files/figure.png" pageId="5" pageNumber="51">Figs 1C</figureCitation>
,
<figureCitation id="55C82A48446CFFA12C87263B89E7FE69" box="[669,706,366,392]" captionStart="Fig" captionStartId="5.[189,232,1963,1989]" captionTargetBox="[189,1398,747,1903]" captionTargetId="figure-170@5.[189,1398,747,1923]" captionTargetPageId="5" captionText="Fig. 2. Oceatra litoralis gen. et sp. nov., paratype, ♀ (MMBC). A. Dorsal view. B. Frontal view. C. Terminalia, ventral view. D. Terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8246751" httpUri="https://zenodo.org/record/8246751/files/figure.png" pageId="5" pageNumber="51">2B</figureCitation>
,
<figureCitation id="55C82A48446CFFA12CCA263889D1FE69" box="[720,756,365,392]" captionStart="Fig" captionStartId="8.[191,234,1928,1954]" captionTargetBox="[216,1375,950,1853]" captionTargetId="figure-197@8.[189,1398,948,1886]" captionTargetPageId="8" captionText="Fig. 5. A–C. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC). D–E. Oceatra scandens gen. et sp. nov., holotype, ♂ (MMBC). A, D. Head and pronotum, dorsal view. B, E. Head, frontal view. C. Rostrum, lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246759" httpUri="https://zenodo.org/record/8246759/files/figure.png" pageId="5" pageNumber="51">5B</figureCitation>
) about as wide as long, with lateral margins obtusely angular below eyes; upper margin sharply trapezoidal, concave. Median carina distinct from the level of eye middle through whole postclypeus, interrupted by metopoclypeal suture; sublateral carinae extending from upper margin of metope almost to metopoclypeal suture. Metopoclypeal suture distinct. Coryphe transverse, nearly twice as wide as long medially (
<figureCitation id="55C82A48446CFFA12D9326AF88CCFDF4" box="[905,1001,506,533]" captionStart="Fig" captionStartId="8.[191,234,1928,1954]" captionTargetBox="[216,1375,950,1853]" captionTargetId="figure-197@8.[189,1398,948,1886]" captionTargetPageId="8" captionText="Fig. 5. A–C. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC). D–E. Oceatra scandens gen. et sp. nov., holotype, ♂ (MMBC). A, D. Head and pronotum, dorsal view. B, E. Head, frontal view. C. Rostrum, lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246759" httpUri="https://zenodo.org/record/8246759/files/figure.png" pageId="5" pageNumber="51">Fig. 5A</figureCitation>
). Ocelli absent. Pedicel globular. Rostrum long, reaching sternite VI, not narrowing apically; second segment 0.25 times as long as third one (
<figureCitation id="55C82A48446CFFA12EE025148A7EFDBD" box="[250,347,577,604]" captionStart="Fig" captionStartId="8.[191,234,1928,1954]" captionTargetBox="[216,1375,950,1853]" captionTargetId="figure-197@8.[189,1398,948,1886]" captionTargetPageId="8" captionText="Fig. 5. A–C. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC). D–E. Oceatra scandens gen. et sp. nov., holotype, ♂ (MMBC). A, D. Head and pronotum, dorsal view. B, E. Head, frontal view. C. Rostrum, lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246759" httpUri="https://zenodo.org/record/8246759/files/figure.png" pageId="5" pageNumber="51">Fig. 5C</figureCitation>
). Pronotum wide, depressed in the middle, without carinae or with very short median carina at its hind margin (
<figureCitation id="55C82A48446CFFA12FF62531896EFD9E" box="[492,587,612,639]" captionStart="Fig" captionStartId="8.[191,234,1928,1954]" captionTargetBox="[216,1375,950,1853]" captionTargetId="figure-197@8.[189,1398,948,1886]" captionTargetPageId="8" captionText="Fig. 5. A–C. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC). D–E. Oceatra scandens gen. et sp. nov., holotype, ♂ (MMBC). A, D. Head and pronotum, dorsal view. B, E. Head, frontal view. C. Rostrum, lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246759" httpUri="https://zenodo.org/record/8246759/files/figure.png" pageId="5" pageNumber="51">Fig. 5A</figureCitation>
). Paradiscal fields of pronotum very narrow, not visible behind eyes. Paranotal lobes nearly rectangular. Mesonotum as long as pronotum medially, with median and lateral carinae. Forewing (
<figureCitation id="55C82A48446FFFA22FBF265C8906FEC2" box="[421,547,265,291]" captionStart="Fig" captionStartId="7.[189,232,1960,1986]" captionTargetBox="[304,1287,1319,1898]" captionTargetId="figure-39@7.[280,1308,1316,1918]" captionTargetPageId="7" captionText="Fig. 4. Adenissina Dlabola, 1980, forewings. A. Perissana dlabolai Gnezdilov &amp; Wilson, 2006, ♂. B–C. Oceatra litoralis gen. et sp. nov., paratype, ♂ (ZIN). B. Left wing. C. Right wing. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246755" httpUri="https://zenodo.org/record/8246755/files/figure.png" pageId="6" pageNumber="52">Fig. 4B–C</figureCitation>
) widely truncate apically, reaching hind margin of pygofer in male and surpassing hind margin of tergite VII in female. Hypocostal plate relatively wide and long, extending along three quarters of wing length. Radius and median veins running from basal cell in one short stem. R, M, CuA, and Pcu furcating apically. Hind tibia with two lateral spines in apical half and seven apical spines. First metatarsomere 1.5 times as long as second one, with two latero-apical and five intermediate spines arranged in arc. Second metatarsomere with only two latero-apical spines. Claws, in dorsal view, nearly twice as long as arolium of pretarsus, each with a single long seta. Dorso-lateral plates of pretarsus narrow.
</paragraph>
<caption id="998C6645446CFFA12EA720FE8869F809" ID-DOI="http://doi.org/10.5281/zenodo.8246751" ID-Zenodo-Dep="8246751" httpUri="https://zenodo.org/record/8246751/files/figure.png" pageId="5" pageNumber="51" startId="5.[189,232,1963,1989]" targetBox="[189,1398,747,1903]" targetPageId="5">
<paragraph id="CD4C36CD446CFFA12EA720FE8869F809" blockId="5.[189,1399,1963,2025]" pageId="5" pageNumber="51">
<emphasis id="FF87EADF446CFFA12EA720FE8A35F824" bold="true" box="[189,272,1963,1989]" pageId="5" pageNumber="51">Fig. 2.</emphasis>
<taxonomicName id="0AF34D4E446CFFA12F0720F98AC3F824" authority="Gnezdilov &amp; Malenovský, 2023" authorityName="Gnezdilov &amp; Malenovský" authorityYear="2023" box="[285,486,1963,1990]" class="Insecta" family="Caliscelidae" genus="Oceatra" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="litoralis" status="gen. et sp. nov.">
<emphasis id="FF87EADF446CFFA12F0720F98AC3F824" box="[285,486,1963,1990]" italics="true" pageId="5" pageNumber="51">Oceatra litoralis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4B457A4446CFFA12FE820F98991F827" box="[498,692,1964,1990]" pageId="5" pageNumber="51" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, paratype, ♀ (MMBC).
<emphasis id="FF87EADF446CFFA12DF820FE88D2F824" bold="true" box="[994,1015,1963,1989]" pageId="5" pageNumber="51">A</emphasis>
. Dorsal view.
<emphasis id="FF87EADF446CFFA12AAA20FE8FE1F824" bold="true" box="[1200,1220,1963,1989]" pageId="5" pageNumber="51">B</emphasis>
. Frontal view.
<emphasis id="FF87EADF446CFFA12EA7209B8BF7F809" bold="true" box="[189,210,1998,2024]" pageId="5" pageNumber="51">C</emphasis>
. Terminalia, ventral view.
<emphasis id="FF87EADF446CFFA12C16209B8904F809" bold="true" box="[524,545,1998,2024]" pageId="5" pageNumber="51">D</emphasis>
. Terminalia, lateral view.
</paragraph>
</caption>
<paragraph id="CD4C36CD446FFFA22EA72514892DFBE0" blockId="6.[188,1401,577,1025]" pageId="6" pageNumber="52">
<smallCapsWord id="CBAAA011446FFFA22EA725148BDAFDB8" baselines="596,596" box="[189,255,577,603]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="6" pageNumber="52">MALE</smallCapsWord>
<smallCapsWord id="CBAAA011446FFFA22F1325138AAFFDB8" baselines="596" box="[265,394,582,601]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="terminalia" pageId="6" pageNumber="52">TERMINALIA</smallCapsWord>
(
<figureCitation id="55C82A48446FFFA22F8425148925FDBA" box="[414,512,577,603]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[204,1384,718,1909]" captionTargetId="figure-173@4.[189,1398,703,1924]" captionTargetPageId="4" captionText="Fig. 1. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC). A. Dorsal view. B. Lateral view. C. Frontal view. D. Genital block, caudal view." figureDoi="http://doi.org/10.5281/zenodo.8246749" httpUri="https://zenodo.org/record/8246749/files/figure.png" pageId="6" pageNumber="52">Figs 1D</figureCitation>
,
<figureCitation id="55C82A48446FFFA22C1525148905FDBA" box="[527,544,577,603]" captionStart="Fig" captionStartId="9.[189,232,1858,1884]" captionTargetBox="[220,1339,277,1790]" captionTargetId="figure-14@9.[189,1398,266,1810]" captionTargetPageId="9" captionText="Fig. 6. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC), terminalia. A. Genital block, lateral view. B. Pygofer, lateral view. C. Anal tube, dorsal view. D. Style, lateral view. E. Style, dorsal view. F. Penis, connective, and anal tube, lateral view. G. Penis, ventral view. H. Penis and connective, dorsal view. I. Apex of penis, dorsal view. Abbreviations: aep = apical aedeagal process; mot = mushroomshaped outgrowth of phallobase; vh = ventral aedeagal hooks; vl = ventral phallobase lobe. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246761" httpUri="https://zenodo.org/record/8246761/files/figure.png" pageId="6" pageNumber="52">6</figureCitation>
). Anal tube, in dorsal view, wide, 0.7 times as wide as long at midline, slightly widening apically, with slightly obtusely angulate anterior margin (
<figureCitation id="55C82A48446FFFA22A3525318FA9FD9F" box="[1071,1164,612,638]" captionStart="Fig" captionStartId="9.[189,232,1858,1884]" captionTargetBox="[220,1339,277,1790]" captionTargetId="figure-14@9.[189,1398,266,1810]" captionTargetPageId="9" captionText="Fig. 6. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC), terminalia. A. Genital block, lateral view. B. Pygofer, lateral view. C. Anal tube, dorsal view. D. Style, lateral view. E. Style, dorsal view. F. Penis, connective, and anal tube, lateral view. G. Penis, ventral view. H. Penis and connective, dorsal view. I. Apex of penis, dorsal view. Abbreviations: aep = apical aedeagal process; mot = mushroomshaped outgrowth of phallobase; vh = ventral aedeagal hooks; vl = ventral phallobase lobe. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246761" httpUri="https://zenodo.org/record/8246761/files/figure.png" pageId="6" pageNumber="52">Fig. 6C</figureCitation>
). Anal column short and wide. Pygofer, in lateral view, narrow, with hind margins strongly convex in upper half (
<figureCitation id="55C82A48446FFFA22AF025D28E4FFD40" box="[1258,1386,647,673]" captionStart="Fig" captionStartId="9.[189,232,1858,1884]" captionTargetBox="[220,1339,277,1790]" captionTargetId="figure-14@9.[189,1398,266,1810]" captionTargetPageId="9" captionText="Fig. 6. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC), terminalia. A. Genital block, lateral view. B. Pygofer, lateral view. C. Anal tube, dorsal view. D. Style, lateral view. E. Style, dorsal view. F. Penis, connective, and anal tube, lateral view. G. Penis, ventral view. H. Penis and connective, dorsal view. I. Apex of penis, dorsal view. Abbreviations: aep = apical aedeagal process; mot = mushroomshaped outgrowth of phallobase; vh = ventral aedeagal hooks; vl = ventral phallobase lobe. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246761" httpUri="https://zenodo.org/record/8246761/files/figure.png" pageId="6" pageNumber="52">Fig. 6A–B</figureCitation>
). Anal tube and phallobase with well-sclerotised connection (
<figureCitation id="55C82A48446FFFA22D9A25FF88FFFD25" box="[896,986,682,708]" captionStart="Fig" captionStartId="9.[189,232,1858,1884]" captionTargetBox="[220,1339,277,1790]" captionTargetId="figure-14@9.[189,1398,266,1810]" captionTargetPageId="9" captionText="Fig. 6. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC), terminalia. A. Genital block, lateral view. B. Pygofer, lateral view. C. Anal tube, dorsal view. D. Style, lateral view. E. Style, dorsal view. F. Penis, connective, and anal tube, lateral view. G. Penis, ventral view. H. Penis and connective, dorsal view. I. Apex of penis, dorsal view. Abbreviations: aep = apical aedeagal process; mot = mushroomshaped outgrowth of phallobase; vh = ventral aedeagal hooks; vl = ventral phallobase lobe. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246761" httpUri="https://zenodo.org/record/8246761/files/figure.png" pageId="6" pageNumber="52">Fig. 6F</figureCitation>
). Phallobase, in lateral view, wide, slightly enlarged apically (
<figureCitation id="55C82A48446FFFA22C1B259B89A3FD09" box="[513,646,717,744]" captionStart="Fig" captionStartId="9.[189,232,1858,1884]" captionTargetBox="[220,1339,277,1790]" captionTargetId="figure-14@9.[189,1398,266,1810]" captionTargetPageId="9" captionText="Fig. 6. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC), terminalia. A. Genital block, lateral view. B. Pygofer, lateral view. C. Anal tube, dorsal view. D. Style, lateral view. E. Style, dorsal view. F. Penis, connective, and anal tube, lateral view. G. Penis, ventral view. H. Penis and connective, dorsal view. I. Apex of penis, dorsal view. Abbreviations: aep = apical aedeagal process; mot = mushroomshaped outgrowth of phallobase; vh = ventral aedeagal hooks; vl = ventral phallobase lobe. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246761" httpUri="https://zenodo.org/record/8246761/files/figure.png" pageId="6" pageNumber="52">Fig. 6A, F</figureCitation>
). Dorso-lateral lobes of phallobase fused dorsally in one lobe with strongly convex margin (
<figureCitation id="55C82A48446FFFA22C0725A4894AFCEA" box="[541,623,753,779]" captionStart="Fig" captionStartId="9.[189,232,1858,1884]" captionTargetBox="[220,1339,277,1790]" captionTargetId="figure-14@9.[189,1398,266,1810]" captionTargetPageId="9" captionText="Fig. 6. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC), terminalia. A. Genital block, lateral view. B. Pygofer, lateral view. C. Anal tube, dorsal view. D. Style, lateral view. E. Style, dorsal view. F. Penis, connective, and anal tube, lateral view. G. Penis, ventral view. H. Penis and connective, dorsal view. I. Apex of penis, dorsal view. Abbreviations: aep = apical aedeagal process; mot = mushroomshaped outgrowth of phallobase; vh = ventral aedeagal hooks; vl = ventral phallobase lobe. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246761" httpUri="https://zenodo.org/record/8246761/files/figure.png" pageId="6" pageNumber="52">Fig. 6I</figureCitation>
); each dorso-lateral lobe with mushroom-shaped outgrowth above ventral phallobase lobe (
<figureCitation id="55C82A48446FFFA22FFC2441891AFCCF" box="[486,575,788,814]" captionStart="Fig" captionStartId="9.[189,232,1858,1884]" captionTargetBox="[220,1339,277,1790]" captionTargetId="figure-14@9.[189,1398,266,1810]" captionTargetPageId="9" captionText="Fig. 6. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC), terminalia. A. Genital block, lateral view. B. Pygofer, lateral view. C. Anal tube, dorsal view. D. Style, lateral view. E. Style, dorsal view. F. Penis, connective, and anal tube, lateral view. G. Penis, ventral view. H. Penis and connective, dorsal view. I. Apex of penis, dorsal view. Abbreviations: aep = apical aedeagal process; mot = mushroomshaped outgrowth of phallobase; vh = ventral aedeagal hooks; vl = ventral phallobase lobe. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246761" httpUri="https://zenodo.org/record/8246761/files/figure.png" pageId="6" pageNumber="52">Fig. 6F</figureCitation>
, mot). Ventral phallobase lobe short and narrow, narrowing apically (
<figureCitation id="55C82A48446FFFA22EDC24628A01FCB0" box="[198,292,823,849]" captionStart="Fig" captionStartId="9.[189,232,1858,1884]" captionTargetBox="[220,1339,277,1790]" captionTargetId="figure-14@9.[189,1398,266,1810]" captionTargetPageId="9" captionText="Fig. 6. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC), terminalia. A. Genital block, lateral view. B. Pygofer, lateral view. C. Anal tube, dorsal view. D. Style, lateral view. E. Style, dorsal view. F. Penis, connective, and anal tube, lateral view. G. Penis, ventral view. H. Penis and connective, dorsal view. I. Apex of penis, dorsal view. Abbreviations: aep = apical aedeagal process; mot = mushroomshaped outgrowth of phallobase; vh = ventral aedeagal hooks; vl = ventral phallobase lobe. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246761" httpUri="https://zenodo.org/record/8246761/files/figure.png" pageId="6" pageNumber="52">Fig. 6G</figureCitation>
, vl). Apical processes of aedeagus extending beyond phallobase margin, long and wide, each with one long spiny and several small teeth apically (
<figureCitation id="55C82A48446FFFA22D58240F88BFFC95" box="[834,922,858,884]" captionStart="Fig" captionStartId="9.[189,232,1858,1884]" captionTargetBox="[220,1339,277,1790]" captionTargetId="figure-14@9.[189,1398,266,1810]" captionTargetPageId="9" captionText="Fig. 6. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC), terminalia. A. Genital block, lateral view. B. Pygofer, lateral view. C. Anal tube, dorsal view. D. Style, lateral view. E. Style, dorsal view. F. Penis, connective, and anal tube, lateral view. G. Penis, ventral view. H. Penis and connective, dorsal view. I. Apex of penis, dorsal view. Abbreviations: aep = apical aedeagal process; mot = mushroomshaped outgrowth of phallobase; vh = ventral aedeagal hooks; vl = ventral phallobase lobe. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246761" httpUri="https://zenodo.org/record/8246761/files/figure.png" pageId="6" pageNumber="52">Fig. 6F</figureCitation>
, aep). Ventral aedeagal hooks arising at aedeagal apex, almost as long as aedeagus, directed basally, acute apically (
<figureCitation id="55C82A48446FFFA22A2F242B8F97FC76" box="[1077,1202,893,920]" captionStart="Fig" captionStartId="9.[189,232,1858,1884]" captionTargetBox="[220,1339,277,1790]" captionTargetId="figure-14@9.[189,1398,266,1810]" captionTargetPageId="9" captionText="Fig. 6. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC), terminalia. A. Genital block, lateral view. B. Pygofer, lateral view. C. Anal tube, dorsal view. D. Style, lateral view. E. Style, dorsal view. F. Penis, connective, and anal tube, lateral view. G. Penis, ventral view. H. Penis and connective, dorsal view. I. Apex of penis, dorsal view. Abbreviations: aep = apical aedeagal process; mot = mushroomshaped outgrowth of phallobase; vh = ventral aedeagal hooks; vl = ventral phallobase lobe. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246761" httpUri="https://zenodo.org/record/8246761/files/figure.png" pageId="6" pageNumber="52">Fig. 6F–G</figureCitation>
, vh). Connective with large, widely opened cup. Style massive, with large curved capitulum without neck, hind margin of style straight, caudal angle rounded (
<figureCitation id="55C82A48446FFFA22C97249189CFFC3F" box="[653,746,964,990]" captionStart="Fig" captionStartId="9.[189,232,1858,1884]" captionTargetBox="[220,1339,277,1790]" captionTargetId="figure-14@9.[189,1398,266,1810]" captionTargetPageId="9" captionText="Fig. 6. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC), terminalia. A. Genital block, lateral view. B. Pygofer, lateral view. C. Anal tube, dorsal view. D. Style, lateral view. E. Style, dorsal view. F. Penis, connective, and anal tube, lateral view. G. Penis, ventral view. H. Penis and connective, dorsal view. I. Apex of penis, dorsal view. Abbreviations: aep = apical aedeagal process; mot = mushroomshaped outgrowth of phallobase; vh = ventral aedeagal hooks; vl = ventral phallobase lobe. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246761" httpUri="https://zenodo.org/record/8246761/files/figure.png" pageId="6" pageNumber="52">Fig. 6D</figureCitation>
). Capitulum, in dorsal view, with small apical tooth and weak lateral tooth (
<figureCitation id="55C82A48446FFFA22F8524B28ADEFBE0" box="[415,507,999,1025]" captionStart="Fig" captionStartId="9.[189,232,1858,1884]" captionTargetBox="[220,1339,277,1790]" captionTargetId="figure-14@9.[189,1398,266,1810]" captionTargetPageId="9" captionText="Fig. 6. Oceatra litoralis gen. et sp. nov., holotype, ♂ (MMBC), terminalia. A. Genital block, lateral view. B. Pygofer, lateral view. C. Anal tube, dorsal view. D. Style, lateral view. E. Style, dorsal view. F. Penis, connective, and anal tube, lateral view. G. Penis, ventral view. H. Penis and connective, dorsal view. I. Apex of penis, dorsal view. Abbreviations: aep = apical aedeagal process; mot = mushroomshaped outgrowth of phallobase; vh = ventral aedeagal hooks; vl = ventral phallobase lobe. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246761" httpUri="https://zenodo.org/record/8246761/files/figure.png" pageId="6" pageNumber="52">Fig. 6E</figureCitation>
).
</paragraph>
<paragraph id="CD4C36CD446FFFA22EA7237C8888FB68" blockId="6.[188,1399,1065,1162]" pageId="6" pageNumber="52">
<smallCapsWord id="CBAAA011446FFFA22EA7237C8A30FBA0" baselines="1084,1084" box="[189,277,1065,1091]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Female" pageId="6" pageNumber="52">FEMALE</smallCapsWord>
<smallCapsWord id="CBAAA011446FFFA22F06237B8AB8FBA0" baselines="1084" box="[284,413,1070,1089]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="terminalia" pageId="6" pageNumber="52">TERMINALIA</smallCapsWord>
(
<figureCitation id="55C82A48446FFFA22FB4237C8914FBA2" box="[430,561,1065,1091]" captionStart="Fig" captionStartId="5.[189,232,1963,1989]" captionTargetBox="[189,1398,747,1903]" captionTargetId="figure-170@5.[189,1398,747,1923]" captionTargetPageId="5" captionText="Fig. 2. Oceatra litoralis gen. et sp. nov., paratype, ♀ (MMBC). A. Dorsal view. B. Frontal view. C. Terminalia, ventral view. D. Terminalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8246751" httpUri="https://zenodo.org/record/8246751/files/figure.png" pageId="6" pageNumber="52">Figs 2C–D</figureCitation>
,
<figureCitation id="55C82A48446FFFA22C27237C896BFBA2" box="[573,590,1065,1091]" captionStart="Fig" captionStartId="10.[189,232,1963,1989]" captionTargetBox="[208,1395,1401,1904]" captionTargetId="figure-439@10.[189,1398,1368,1923]" captionTargetPageId="10" captionText="Fig. 7. Oceatra litoralis gen. et sp. nov., paratype, ♀ (MMBC), terminalia. A. Pygofer and anal tube, dorsal view. B. Genital block, lateral view. C. Sternite VII, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246763" httpUri="https://zenodo.org/record/8246763/files/figure.png" pageId="6" pageNumber="52">7</figureCitation>
). Hind margin of sternite VII with two short teeth medially (
<figureCitation id="55C82A48446FFFA22B17237C8E4CFBA2" box="[1293,1385,1065,1091]" captionStart="Fig" captionStartId="10.[189,232,1963,1989]" captionTargetBox="[208,1395,1401,1904]" captionTargetId="figure-439@10.[189,1398,1368,1923]" captionTargetPageId="10" captionText="Fig. 7. Oceatra litoralis gen. et sp. nov., paratype, ♀ (MMBC), terminalia. A. Pygofer and anal tube, dorsal view. B. Genital block, lateral view. C. Sternite VII, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246763" httpUri="https://zenodo.org/record/8246763/files/figure.png" pageId="6" pageNumber="52">Fig. 7C</figureCitation>
). Anal tube, in dorsal view, as wide as long, with hind margin concave (
<figureCitation id="55C82A48446FFFA22DE023198F7CFB87" box="[1018,1113,1100,1126]" captionStart="Fig" captionStartId="10.[189,232,1963,1989]" captionTargetBox="[208,1395,1401,1904]" captionTargetId="figure-439@10.[189,1398,1368,1923]" captionTargetPageId="10" captionText="Fig. 7. Oceatra litoralis gen. et sp. nov., paratype, ♀ (MMBC), terminalia. A. Pygofer and anal tube, dorsal view. B. Genital block, lateral view. C. Sternite VII, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246763" httpUri="https://zenodo.org/record/8246763/files/figure.png" pageId="6" pageNumber="52">Fig. 7A</figureCitation>
); in lateral view, ventral part of anal tube convex (
<figureCitation id="55C82A48446FFFA22FF3233A8963FB68" box="[489,582,1135,1161]" captionStart="Fig" captionStartId="10.[189,232,1963,1989]" captionTargetBox="[208,1395,1401,1904]" captionTargetId="figure-439@10.[189,1398,1368,1923]" captionTargetPageId="10" captionText="Fig. 7. Oceatra litoralis gen. et sp. nov., paratype, ♀ (MMBC), terminalia. A. Pygofer and anal tube, dorsal view. B. Genital block, lateral view. C. Sternite VII, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8246763" httpUri="https://zenodo.org/record/8246763/files/figure.png" pageId="6" pageNumber="52">Fig. 7B</figureCitation>
). Anal column short and wide.
</paragraph>
</subSubSection>
<subSubSection id="85E96546446FFFA22EA723E58AECFA0E" pageId="6" pageNumber="52" type="distribution">
<paragraph id="CD4C36CD446FFFA22EA723E58AD4FB2A" blockId="6.[189,497,1200,1227]" box="[189,497,1200,1227]" pageId="6" pageNumber="52">
<heading id="960481A1446FFFA22EA723E58AD4FB2A" bold="true" box="[189,497,1200,1227]" fontSize="11" level="3" pageId="6" pageNumber="52" reason="3">
<emphasis id="FF87EADF446FFFA22EA723E58AD4FB2A" bold="true" box="[189,497,1200,1227]" pageId="6" pageNumber="52">Distribution and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="CD4C36CD446FFFA22EA7238A8AECFA0E" blockId="6.[189,1399,1247,1520]" pageId="6" pageNumber="52">
So far only known from the western coast of
<collectingCountry id="B5E4765D446FFFA22CF3238A88BDFB18" box="[745,920,1247,1273]" name="Socotra Island" pageId="6" pageNumber="52">Socotra Island</collectingCountry>
, close to the cape and village of Shuab (
<figureCitation id="55C82A48446FFFA22EDC22578A2AFAFD" box="[198,271,1282,1308]" captionStart="Fig" captionStartId="12.[189,232,1998,2024]" captionTargetBox="[196,1397,1346,1950]" captionTargetId="figure-420@12.[191,1400,1344,1955]" captionTargetPageId="12" captionText="Fig. 9. Distribution of Oceatra spp. in Socotra Island." figureDoi="http://doi.org/10.5281/zenodo.8246767" httpUri="https://zenodo.org/record/8246767/files/figure.png" pageId="6" pageNumber="52">Fig. 9</figureCitation>
). The
<typeStatus id="1248886F446FFFA22F4122568AABFAFC" box="[347,398,1283,1309]" pageId="6" pageNumber="52">type</typeStatus>
series was collected in coastal sand dunes and salt marshes in vegetation dominated by the semi-succulent plant species
<taxonomicName id="0AF34D4E446FFFA22C9922738F7CFADE" authority="(Vierh.) Radcl." authorityName="Radcl." baseAuthorityName="Vierh." box="[643,1113,1317,1343]" class="Magnoliopsida" family="Plumbaginaceae" genus="Limonium" kingdom="Plantae" order="Caryophyllales" pageId="6" pageNumber="52" phylum="Tracheophyta" rank="species" species="sokotranum">
<emphasis id="FF87EADF446FFFA22C99227388B6FADE" box="[643,915,1317,1343]" italics="true" pageId="6" pageNumber="52">Limonium sokotranum</emphasis>
(Vierh.) Radcl.
</taxonomicName>
-Sm. (
<taxonomicName id="0AF34D4E446FFFA22AB922708E4FFADE" box="[1187,1386,1317,1343]" class="Magnoliopsida" family="Plumbaginaceae" kingdom="Plantae" order="Caryophyllales" pageId="6" pageNumber="52" phylum="Tracheophyta" rank="family">Plumbaginaceae</taxonomicName>
),
<taxonomicName id="0AF34D4E446FFFA22EA7221C88F6FA82" authority="(Moric.) K. Koch" authorityName="K. Koch" baseAuthorityName="Moric." box="[189,979,1353,1379]" class="Magnoliopsida" family="Amaranthaceae" genus="Arthrocnemum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="6" pageNumber="52" phylum="Tracheophyta" rank="species" species="macrostachyum">
<emphasis id="FF87EADF446FFFA22EA7221C8908FA82" box="[189,557,1353,1379]" italics="true" pageId="6" pageNumber="52">Arthrocnemum macrostachyum</emphasis>
(Moric.) K. Koch (Amaranthaceae)
</taxonomicName>
and
<taxonomicName id="0AF34D4E446FFFA22A16221C8AA9FA67" authority="Hadidi" authorityName="Hadidi" class="Magnoliopsida" family="Zygophyllaceae" genus="Zygophyllum" kingdom="Plantae" order="Zygophyllales" pageId="6" pageNumber="52" phylum="Tracheophyta" rank="species" species="qatarense">
<emphasis id="FF87EADF446FFFA22A16221C8E05FA83" box="[1036,1312,1353,1379]" italics="true" pageId="6" pageNumber="52">Zygophyllum qatarense</emphasis>
Hadidi (Zygophyllaceae)
</taxonomicName>
(
<figureCitation id="55C82A48446FFFA22F8722398929FA67" box="[413,524,1388,1414]" captionStart="Fig" captionStartId="13.[189,232,1892,1918]" captionTargetBox="[200,1388,484,1848]" captionTargetId="figure-60@13.[200,1388,484,1852]" captionTargetPageId="13" captionText="Fig. 10. Habitats of Oceatra spp. in Socotra Island. A. Type locality of O. litoralis gen. et sp. nov., coastal sand dunes and salt marshes near Shuab (photograph by I. Malenovský, 20 Jun. 2012). B. Type locality of O. scandens gen. et sp. nov., open montane woodland and low shrub communities on slopes of Mt Scand in the Hagher Mountains (photograph by L. Purchart, 1 Feb. 2010)." figureDoi="http://doi.org/10.5281/zenodo.8246769" httpUri="https://zenodo.org/record/8246769/files/figure.png" pageId="6" pageNumber="52">Fig. 10A</figureCitation>
). Most specimens were swept from
<taxonomicName id="0AF34D4E446FFFA22DA622388F42FA67" box="[956,1127,1388,1414]" class="Magnoliopsida" family="Plumbaginaceae" genus="Limonium" kingdom="Plantae" order="Caryophyllales" pageId="6" pageNumber="52" phylum="Tracheophyta" rank="species" species="sokotranum">
<emphasis id="FF87EADF446FFFA22DA622388F42FA67" box="[956,1127,1388,1414]" italics="true" pageId="6" pageNumber="52">L. sokotranum</emphasis>
</taxonomicName>
which might be a host plant for
<taxonomicName id="0AF34D4E446FFFA22F3D22DA8A8DFA48" authority="Gnezdilov &amp; Malenovský, 2023" authorityName="Gnezdilov &amp; Malenovský" authorityYear="2023" box="[295,424,1423,1449]" class="Insecta" family="Caliscelidae" genus="Oceatra" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="litoralis" status="gen. et sp. nov.">
<emphasis id="FF87EADF446FFFA22F3D22DA8A8DFA48" box="[295,424,1423,1449]" italics="true" pageId="6" pageNumber="52">O. litoralis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4B457A4446FFFA22FB522C5897BFA4B" box="[431,606,1424,1450]" pageId="6" pageNumber="52" rank="species">gen. et sp. nov.</taxonomicNameLabel>
Another planthopper species,
<taxonomicName id="0AF34D4E446FFFA22DA522DA8909FA2D" authority="Swierczewski, Malenovsky &amp; Stroinski, 2017" authorityName="Swierczewski, Malenovsky &amp; Stroinski" authorityYear="2017" class="Insecta" family="Flatidae" genus="Haloflata" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="arthrocnemi">
<emphasis id="FF87EADF446FFFA22DA522DA8FE0FA48" box="[959,1221,1423,1449]" italics="true" pageId="6" pageNumber="52">Haloflata arthrocnemi</emphasis>
<bibRefCitation id="A9624B3C446FFFA22AD622C58909FA2D" author="Swierczewski D. &amp; Malenovsky I. &amp; Stroinski A." pageId="6" pageNumber="52" pagination="261 - 278" refId="ref8325" refString="Swierczewski D., Malenovsky I. &amp; Stroinski A. 2017. Haloflata, a new genus from salt marshes in Socotra Island (Hemiptera: Fulgoromorpha: Flatidae). Annales Zoologici 67 (2): 261 - 278. https: // doi. org / 10.3161 / 00034541 ANZ 2017.67.2.007" type="journal article" year="2017">Świerczewski, Malenovský &amp; Stroiński, 2017</bibRefCitation>
</taxonomicName>
from the family of
<taxonomicName id="0AF34D4E446FFFA22D0122E78858FA2D" authorityName="Spinola" authorityYear="1839" box="[795,893,1458,1484]" class="Insecta" family="Flatidae" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="52" phylum="Arthropoda" rank="family">Flatidae</taxonomicName>
, co-occurred at the site (see
<bibRefCitation id="A9624B3C446FFFA22ACE22E68A1FFA11" author="Swierczewski D. &amp; Malenovsky I. &amp; Stroinski A." pageId="6" pageNumber="52" pagination="261 - 278" refId="ref8325" refString="Swierczewski D., Malenovsky I. &amp; Stroinski A. 2017. Haloflata, a new genus from salt marshes in Socotra Island (Hemiptera: Fulgoromorpha: Flatidae). Annales Zoologici 67 (2): 261 - 278. https: // doi. org / 10.3161 / 00034541 ANZ 2017.67.2.007" type="journal article" year="2017">
Świerczewski
<emphasis id="FF87EADF446FFFA22EA722838BDDFA0E" box="[189,248,1493,1519]" italics="true" pageId="6" pageNumber="52">et al.</emphasis>
2017
</bibRefCitation>
for details).
</paragraph>
</subSubSection>
</treatment>
</document>
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